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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.137.1764" ID-GBIF-Dataset="5e467153-180c-4d2a-a1ed-691204811d36" ID-PMC="PMC3260888" ID-Pensoft-Pub="1313-2970-137-41" ID-PubMed="22259304" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-137-41" ModsDocOrigin="ZooKeys 137" ModsDocTitle="Six new species of Acomoptera from North America (Diptera, Mycetophilidae)" checkinTime="1451249779108" checkinUser="pensoft" docAuthor="Kerr, Peter H." docDate="2011" docId="C37E8835124AC225272FF287537BDE19" docLanguage="en" docName="ZooKeys 137: 41-76" docOrigin="ZooKeys 137" docSource="http://dx.doi.org/10.3897/zookeys.137.1764" docTitle="Acomoptera vockerothi Kerr, 2011, sp. n." docType="treatment" docVersion="4" lastPageNumber="67" masterDocId="217BCD19C84AFFAF7F58FFEDFFE37B3C" masterDocTitle="Six new species of Acomoptera from North America (Diptera, Mycetophilidae)" masterLastPageNumber="76" masterPageNumber="41" pageNumber="65" updateTime="1668152407328" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Six new species of Acomoptera from North America (Diptera, Mycetophilidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kerr, Peter H.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>137</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>41</mods:start>
<mods:end>76</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.137.1764</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.137.1764</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-137-41</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152031468" LSID="urn:lsid:zoobank.org:act:A2DAD9AC-96F0-4478-B172-A5C4BB21EF14" httpUri="http://treatment.plazi.org/id/C37E8835124AC225272FF287537BDE19" lastPageId="26" lastPageNumber="67" pageId="24" pageNumber="65">
<subSubSection pageId="24" pageNumber="65" type="nomenclature">
<paragraph pageId="24" pageNumber="65">
<taxonomicName LSID="urn:lsid:zoobank.org:act:A2DAD9AC-96F0-4478-B172-A5C4BB21EF14" class="Insecta" family="Mycetophilidae" genus="Acomoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acomoptera vockerothi" order="Diptera" pageId="24" pageNumber="65" phylum="Arthropoda" rank="species" species="vockerothi">Acomoptera vockerothi</taxonomicName>
<taxonomicNameLabel pageId="24" pageNumber="65">sp. n.</taxonomicNameLabel>
Figs 2426
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="65" type="type material">
<paragraph pageId="24" pageNumber="65">Type Material.</paragraph>
<paragraph pageId="24" pageNumber="65">
Holotype: ♂, &quot;Can: Manitoba, 2mi., ne. Treesbank, along, Souris R. 11.viii.1993,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="49.666668">49°40'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-99.6">99°36'W</geoCoordinate>
, B. Gallaway MT&quot; / &quot;HOLOTYPE 11G627 ♂
<taxonomicName class="Insecta" family="Mycetophilidae" genus="Acomoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acomoptera vockerothi" order="Diptera" pageId="24" pageNumber="65" phylum="Arthropoda" rank="species" species="vockerothi">Acomoptera vockerothi</taxonomicName>
Kerr 2011&quot; [red label]. Deposited in CNC, complete specimen in excellent condition, glued directly to the pin.
</paragraph>
<paragraph pageId="24" pageNumber="65">
Paratypes: CANADA: 1 ♂, same data as holotype [CNC]; 4 ♂♂, &quot;Ont., Iroquois Falls, 30.vi.1987, J.R. Vockeroth&quot; [48.768°, -80.673°; CNC]; 1 ♂, &quot;Ont., Iroquois Falls, 7.vii.1987, J.R. Vockeroth / 11G495&quot; [48.768°, -80.673°; CNC]; 1 ♂, &quot;N.Burgess Twp., Lanark Co., ONT., 7.ix.1970, D.M. Wood&quot; [45.010°, -76.359°; CNC]; 1 ♂, &quot;Thwartway Island, St. Lawrence Is. National Park / A. Carter, Aug 27, 1976, Malaise Trap, Code 4529-G&quot; [44.294°, -76.150°; CNC]; 1 ♂, &quot;King Mt., Old Chelsea, QUE., June 16-1960, J.G. Chillcott&quot; [45.489°, -75.864°; CNC]; 1 ♂, &quot;Duncan Lake, Nr. Rupert, Que., 1.IX.1971, J.F.McAlpine&quot; [54.690°, -75.989°; CNC]; 1 ♂, &quot;N.S., CBHNt.Pk., Mackenzie Mt., 300m PG645851, 29.VIII.1983 / Picea Betula woods&quot; [46.738°, -60.650°; CNC]; 1 ♂, &quot;N.S., CBHNt.Pk., Lone Shieling, 300m PG731861, 21.VIII.1983 / Maple forest with fern undergrowth, J.R. Vockeroth&quot; [46.738°, -60.650°; CNC]; 1 ♂, &quot;N.S., S. Harbour, Bch. PG962943, 12.VIII.1983, J.R. Vockeroth&quot; [46.866°, -60.468°; CNC]; USA: 1 ♂, &quot;Laurel, MD., 25 May 65, Malaise Trap / 11G482&quot; [39.099°, -76.359°; CNC]; 3 ♂♂, &quot;Laurel, MD., 26 May 65, Malaise Trap&quot; [39.099°, -76.359°; CNC]; 1 ♂, &quot;Highlands, Macon Co., N.C. 3850',
<geoCoordinate direction="north" orientation="latitude" precision="92" value="35.053333">35°3.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.18833">83°11.3'W</geoCoordinate>
, June 21, 1958, Jean L. Laffoon&quot; [ISUI]; 1 ♂, &quot;Highlands, Macon Co., N.C. 3850',
<geoCoordinate direction="north" orientation="latitude" precision="92" value="35.053333">35°3.2'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.18833">83°11.3'W</geoCoordinate>
, at light, VII-5-1958, Jean L. Laffoon&quot; [ISUI]; 1 ♂, &quot;Clear Creek, 1 mile so. Highlands, Macon Co., No. Car. 3000',
<geoCoordinate direction="north" orientation="latitude" precision="92" value="35.025">35°1.5'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.191666">83°11.5'W</geoCoordinate>
, VII-1-1958, J. Laffoon&quot; [ISUI]; 2 ♂♂, &quot;Robin Branch (near Wayah Bald), 4000', Macon Co., No. Car.,
<geoCoordinate direction="north" orientation="latitude" precision="92" value="35.168335">35°10.1'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-83.585">83°35.1'W</geoCoordinate>
, VII-3-1958, J. Laffoon&quot; [ISUI].
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="65" type="diagnosis">
<paragraph pageId="24" pageNumber="65">Diagnosis.</paragraph>
<paragraph pageId="24" pageNumber="65">
This species is most similar to
<taxonomicName class="Insecta" family="Mycetophilidae" genus="Acomoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acomoptera crispa" order="Diptera" pageId="24" pageNumber="65" phylum="Arthropoda" rank="species" species="crispa">Acomoptera crispa</taxonomicName>
sp. n. in having similarly-shaped gonostyli that feature a line of long, curved setae near the dorsal margin, as mentioned above. The gonocoxites of
<taxonomicName class="Insecta" family="Mycetophilidae" genus="Acomoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acomoptera vockerothi" order="Diptera" pageId="24" pageNumber="65" phylum="Arthropoda" rank="species" species="vockerothi">Acomoptera vockerothi</taxonomicName>
bears a posteromedial denticulate ridge ventrally, however (Figs 25B, 26B). It is also distinguished by having a short, unforked ventromedial gonostylus lobe (Fig. 26E) and the outer dorsal lobe is broadly attached dorsally (Figs 25, 26
<normalizedToken originalValue="AB">A-B</normalizedToken>
, 26
<normalizedToken originalValue="DE">D-E</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="26" lastPageNumber="67" pageId="24" pageNumber="65" type="description">
<paragraph pageId="24" pageNumber="65">Description.</paragraph>
<paragraph pageId="24" pageNumber="65">Male. Body length (n=10): 4.6-6.3 mm (avg = 5.6 mm). Wing length: 4.5-5.3 mm (avg = 5.0 mm).</paragraph>
<paragraph pageId="24" pageNumber="65">Coloration (Fig. 24). Head brown; palpomeres yellow to yellowish brown. Antennal scape, pedicel, and flagellomeres yellow to yellowish brown, flagellomeres darker than scape and pedicel. Thorax cream-colored to yellowish or light brown; area of scutum bearing acrostichal and dorsocentral setae defined by darker coloration, scutum setae gold- or golden brown-colored. Legs becoming increasingly brown towards tarsi, coxae yellowish or cream-colored; femur yellowish, tibia yellowish brown, tarsi brown. Wing hyaline without markings, wing veins brown; haltere stem and knob yellow to light yellowish brown. Abdominal segments concolorous, yellowish brown to brown, with scattered yellow or golden brown setae. Terminalia yellowish brown.</paragraph>
<paragraph lastPageId="25" lastPageNumber="66" pageId="24" pageNumber="65">
Head. Ocelli slightly raised; middle ocellus approx. same size as lateral ocelli; lateral ocellus located approx. width of ocellus or less from eye margin, separated from median ocellus by approx. twice its own diameter. Eyes with sparse, inconspicuous microsetae, which are approximately as long as width of facet. Face with brown setae, longest of
<pageBreakToken pageId="25" pageNumber="66" start="start">which</pageBreakToken>
approx. same length as width of face. Antennal length approx. 0.8
<normalizedToken originalValue="×">x</normalizedToken>
length of abdomen. Palpus approx. 1.0
<normalizedToken originalValue="1.25×">-1.25x</normalizedToken>
width of head (anterior view); length of palpomeres 2 and 3 subequal; palpomere 4 approx. 4.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; palpomere 5 approx. 10
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, subequal to or longer than combined length of palpomeres 3 and 4.
</paragraph>
<paragraph pageId="25" pageNumber="66">
Thorax. Antepronotum bearing setae; remaining thoracic sclerites bare. Wing venation similar to others in the genus (
<bibRefCitation author="Vockeroth, JR" journalOrPublisher="Canadian Entomologist" pageId="28" pageNumber="69" pagination="529 - 544" title="New genera and species of Mycetophilidae (Diptera) from the Holarctic region, with notes on other species." volume="112" year="1980">Vockeroth 1980</bibRefCitation>
: fig. 3); costal vein extends beyond R5, approx. 0.33
<normalizedToken originalValue="×">x</normalizedToken>
distance between R5 and M1; R1, R5, M1, and M2 with at least some setae on lower surface.
</paragraph>
<paragraph pageId="25" pageNumber="66">
Male Genitalia (Figs 25-26). Epandrium approx. 3
<normalizedToken originalValue="×">x</normalizedToken>
wider than long. Gonocoxites with dorsomedial comb (Figs 26A). Gonostylus complex, as shown in Figs 25, 26
<normalizedToken originalValue="AB">A-B</normalizedToken>
, 25
<normalizedToken originalValue="DE">D-E</normalizedToken>
.
</paragraph>
<paragraph pageId="26" pageNumber="67">
<pageBreakToken pageId="26" pageNumber="67" start="start">Female</pageBreakToken>
unknown.
</paragraph>
<caption pageId="26" pageNumber="67">
<paragraph pageId="26" pageNumber="67">
Figure 24.
<taxonomicName class="Insecta" family="Mycetophilidae" genus="Acomoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acomoptera vockerothi" order="Diptera" pageId="26" pageNumber="67" phylum="Arthropoda" rank="species" species="vockerothi">Acomoptera vockerothi</taxonomicName>
sp. n., habitus, lateral view [691250]. Scale line = 1 mm.
</paragraph>
</caption>
<caption pageId="26" pageNumber="67">
<paragraph pageId="26" pageNumber="67">
Figure 25.
<taxonomicName class="Insecta" family="Mycetophilidae" genus="Acomoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acomoptera vockerothi" order="Diptera" pageId="26" pageNumber="67" phylum="Arthropoda" rank="species" species="vockerothi">Acomoptera vockerothi</taxonomicName>
sp. n., male genitalia, images: A dorsal view [691246] B ventral view [691247] C lateral view [691248] D posterior view [691249]. Scale line = 0.1 mm.
</paragraph>
</caption>
<caption pageId="26" pageNumber="67">
<paragraph pageId="26" pageNumber="67">
Figure 26.
<taxonomicName class="Insecta" family="Mycetophilidae" genus="Acomoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acomoptera vockerothi" order="Diptera" pageId="26" pageNumber="67" phylum="Arthropoda" rank="species" species="vockerothi">Acomoptera vockerothi</taxonomicName>
sp. n., male genitalia, illustrated: A gonopods, dorsal view [691252] B gonopods, ventral view [691253] C gonocoxites, with gonostyli removed, posterior view D gonostyli, posterior view [691255] E gonopods, lateral view [691254] F epandrium, dorsal view [691251]. Scale line = 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="26" pageNumber="67" type="etymology">
<paragraph pageId="26" pageNumber="67">Etymology.</paragraph>
<paragraph pageId="26" pageNumber="67">The species is named after J.R. Vockeroth, a remarkably friendly and engaging person, legendary figure in the history of Dipterology, author of the genus, and frequent collector of this species.</paragraph>
</subSubSection>
</treatment>
</document>