215 lines
18 KiB
XML
215 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e86666" ID-Pensoft-Pub="1864-8312-80-603" ID-Pensoft-UUID="A8EB0928930E59EFB70FAE839C7D88B7" ID-ZooBank="5B8D112EAAC74F2EAC98A057F6094793" ModsDocID="1864-8312-80-603" checkinTime="1669182032150" checkinUser="pensoft" docAuthor="Barbosa, Diego N., Hermes, Marcel Gustavo & Lepeco, Anderson" docDate="2022" docId="F6FF02388F955E21B8917BADFDA8B727" docLanguage="en" docName="ArthropodSystPhyl 80: 603-625" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-11-22" docSource="http://dx.doi.org/10.3897/asp.80.e86666" docTitle="Brachymesitius Barbosa 2022, gen. nov." docType="treatment" docVersion="1" id="A8EB0928930E59EFB70FAE839C7D88B7" lastPageNumber="603" masterDocId="A8EB0928930E59EFB70FAE839C7D88B7" masterDocTitle="Phylogeny of Mesitiinae (Hymenoptera: Bethylidae): assessing their classification, character evolution and diversification" masterLastPageNumber="625" masterPageNumber="603" pageNumber="603" updateTime="1669182032150" updateUser="pensoft">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Phylogeny of Mesitiinae (Hymenoptera: Bethylidae): assessing their classification, character evolution and diversification</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Barbosa, Diego N.</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0667-8598</mods:nameIdentifier>
|
|
<mods:affiliation>Universidade Federal do Parana, Campus III - Centro Politecnico, Av. Coronel Francisco Heraclito dos Santos, s / n, Jardim das Americas, 81.531 - 980, Curitiba, PR, Brazil</mods:affiliation>
|
|
<mods:nameIdentifier type="email">barbosa.laelius@gmail.com</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Hermes, Marcel Gustavo</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9322-4518</mods:nameIdentifier>
|
|
<mods:affiliation>Universidade Federal de Lavras, Campus Universitario, Departamento de Biologia, Av. Central s / n, 37.200 - 900, Lavras, MG, Brazil</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Lepeco, Anderson</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7467-5244</mods:nameIdentifier>
|
|
<mods:affiliation>Laboratorio de Biologia Comparada e Abelhas, Departamento de Biologia, Faculdade de Filosofia, Ciencias e Letras de Ribeirao Preto, Universidade de Sao Paulo, Avenida Bandeirantes, 3900. 14040 - 901, Ribeirao Preto, SP, Brazil</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2022</mods:date>
|
|
<mods:detail type="pubDate">
|
|
<mods:number>2022-11-22</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="volume">
|
|
<mods:number>80</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>603</mods:start>
|
|
<mods:end>625</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/asp.80.e86666</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e86666</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1864-8312-80-603</mods:identifier>
|
|
<mods:identifier type="ZooBank">5B8D112EAAC74F2EAC98A057F6094793</mods:identifier>
|
|
<mods:identifier type="Pensoft-UUID">A8EB0928930E59EFB70FAE839C7D88B7</mods:identifier>
|
|
</mods:mods>
|
|
<treatment LSID="urn:lsid:zoobank.org:act:gEnus/CBF4DC2A-6F41-4527-8168-8440C6044FA7" httpUri="http://treatment.plazi.org/id/F6FF02388F955E21B8917BADFDA8B727" lastPageNumber="603" pageId="0" pageNumber="603">
|
|
<subSubSection pageId="0" pageNumber="603" type="nomenclature">
|
|
<paragraph pageId="0" pageNumber="603">
|
|
<taxonomicName LSID="http://zoobank.org/genus/CBF4DC2A-6F41-4527-8168-8440C6044FA7" authority="Barbosa" authorityName="Barbosa" authorityYear="2022" family="Bethylidae" genus="Brachymesitius" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Brachymesitius" order="Hymenoptera" pageId="0" pageNumber="603" rank="genus" status="gen. nov.">Brachymesitius Barbosa</taxonomicName>
|
|
<taxonomicNameLabel pageId="0" pageNumber="603">gen. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="603" type="type species">
|
|
<paragraph pageId="0" pageNumber="603">Type species.</paragraph>
|
|
<paragraph pageId="0" pageNumber="603">
|
|
<taxonomicName class="Insecta" family="Bethylidae" genus="Incertosulcus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Incertosulcus krombeini" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="species" species="krombeini">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Incertosulcus krombeini</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Móczár">Moczar</normalizedToken>
|
|
, 1970 by original designation.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="603" type="diagnosis">
|
|
<paragraph pageId="0" pageNumber="603">Diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="603">
|
|
The malar space convergent (#7:0), apex of median clypeal carina [in profile] inclined (#11:2), eye very small (#21:2), pubescence of eye absent (#22:0), posterior propodeal projection wide (#64:0), number of distal hamuli of hind wing four (#81:1), hypopygium wider than long (#94:2), anterolateral hypopygial apodeme present (#100:1) were found to be autapomorphies for
|
|
<taxonomicName authorityName="Barbosa" authorityYear="2022" family="Bethylidae" genus="Brachymesitius" kingdom="Animalia" lsidName="Brachymesitius" order="Hymenoptera" pageId="0" pageNumber="603" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Brachymesitius</emphasis>
|
|
</taxonomicName>
|
|
. The type species was first described as a species of
|
|
<taxonomicName authorityName="Moczar" authorityYear="1970" class="Insecta" family="Bethylidae" genus="Incertosulcus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Incertosulcus" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Incertosulcus</emphasis>
|
|
</taxonomicName>
|
|
. The genus was characterized by the presence or absence of a long posterior propodeal projection, making the identification dubious.
|
|
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4489.1.1" author="Azevedo, CO" journalOrPublisher="Zootaxa" pageId="0" pageNumber="603" pagination="1 - 294" refId="B3" refString="Azevedo, CO, Alencar, IDCCA, Ramos, MS, Barbosa, DN, Colombo, WD, Vargas, JM, Lim, J, 2018. Global guide of the flat wasps (Hymenoptera, Bethylidae). Zootaxa 4489: 1 - 294, DOI: https://doi.org/10.11646/zootaxa.4489.1.1" title="Global guide of the flat wasps (Hymenoptera, Bethylidae)." url="https://doi.org/10.11646/zootaxa.4489.1.1" volume="4489" year="2018">Azevedo et al. (2018)</bibRefCitation>
|
|
proposed a new interpretation for the diagnostic characteristics for
|
|
<taxonomicName authorityName="Moczar" authorityYear="1970" class="Insecta" family="Bethylidae" genus="Incertosulcus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Incertosulcus" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Incertosulcus</emphasis>
|
|
</taxonomicName>
|
|
, making
|
|
<taxonomicName class="Insecta" family="Bethylidae" genus="Incertosulcus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Incertosulcus krombeini" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="species" species="krombeini">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Incertosulcus krombeini</emphasis>
|
|
</taxonomicName>
|
|
, as junior synonym of
|
|
<taxonomicName class="Insecta" family="Bethylidae" genus="Parvoculus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parvoculus indicus" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="species" species="indicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Parvoculus indicus</emphasis>
|
|
</taxonomicName>
|
|
Kieffer, 1905.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="603">
|
|
This genus shares similarities with
|
|
<taxonomicName authorityName="Moczar" authorityYear="1970" class="Insecta" family="Bethylidae" genus="Anaylax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anaylax" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Anaylax</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName authorityName="Nagy" authorityYear="1972" class="Insecta" family="Bethylidae" genus="Clytrovorus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clytrovorus" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Clytrovorus</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName authorityName="Barbosa" authorityYear="2022" family="Bethylidae" genus="Hadesmesitius" kingdom="Animalia" lsidName="Hadesmesitius" order="Hymenoptera" pageId="0" pageNumber="603" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Hadesmesitius</emphasis>
|
|
</taxonomicName>
|
|
which have the head, dorsal pronotal area and mesoscutum coriaceous, and the median pronotal line and median mesonotal sulcus absent. Other characteristics distinguish
|
|
<taxonomicName authorityName="Barbosa" authorityYear="2022" family="Bethylidae" genus="Brachymesitius" kingdom="Animalia" lsidName="Brachymesitius" order="Hymenoptera" pageId="0" pageNumber="603" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Brachymesitius</emphasis>
|
|
</taxonomicName>
|
|
from
|
|
<taxonomicName authorityName="Moczar" authorityYear="1970" class="Insecta" family="Bethylidae" genus="Anaylax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anaylax" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Anaylax</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName authorityName="Nagy" authorityYear="1972" class="Insecta" family="Bethylidae" genus="Clytrovorus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Clytrovorus" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Clytrovorus</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName authorityName="Barbosa" authorityYear="2022" family="Bethylidae" genus="Hadesmesitius" kingdom="Animalia" lsidName="Hadesmesitius" order="Hymenoptera" pageId="0" pageNumber="603" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Hadesmesitius</emphasis>
|
|
</taxonomicName>
|
|
, including eyes very small, similar to
|
|
<taxonomicName class="Insecta" genus="Bradepyris" kingdom="Animalia" lsidName="Bradepyris" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Bradepyris</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName authorityName="Barbosa & Azevedo" authorityYear="2014" class="Insecta" family="Bethylidae" genus="Moczariella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Moczariella" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Moczariella</emphasis>
|
|
</taxonomicName>
|
|
; the anterolateral hypopygial apodeme similar to
|
|
<taxonomicName class="Insecta" family="Bethylidae" genus="Mesitius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mesitius" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Mesitius</emphasis>
|
|
</taxonomicName>
|
|
; and the hind wing with four distal hamuli is shared with
|
|
<taxonomicName authorityName="Argaman" authorityYear="2003" class="Insecta" family="Bethylidae" genus="Zimankos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zimankos" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Zimankos</emphasis>
|
|
</taxonomicName>
|
|
; the presence of frontal carina and the propodeal declivity and lateral surface of metapectal-propodeal complex areolate are exclusive for
|
|
<taxonomicName authorityName="Barbosa" authorityYear="2022" family="Bethylidae" genus="Brachymesitius" kingdom="Animalia" lsidName="Brachymesitius" order="Hymenoptera" pageId="0" pageNumber="603" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Brachymesitius</emphasis>
|
|
</taxonomicName>
|
|
as diagnostic characteristics.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="603" type="description">
|
|
<paragraph pageId="0" pageNumber="603">Description.</paragraph>
|
|
<paragraph pageId="0" pageNumber="603">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="603">Wings</emphasis>
|
|
: hyaline.
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="603">Head</emphasis>
|
|
: As long as wide; malar space as long as VOL, convergent; clypeus with median lobe rounded, median clypeal carina inclined; antenna with pubescence sparse and short; pedicel cylindrical, first flagellomere as long as pedicel, flagellomeres short; eye very small, without pubescence; frons foveolate, with frontal carina; ocelli very small; anterior ocellus crossing supra-ocular line; dorsal and ventral half of occipital carina low.
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="603">Pronotum</emphasis>
|
|
: Dorsal pronotal area shorter than wide, foveolate, with humeral angle rounded, side straight, anterior margin outcurved, posterior margin incurved, median pronotal line absent; mesoscutum coriaceous, median mesonotal sulcus absent, notaulus present and narrow; mesoscutellum not touching the metapectal-propodeal disc; metapectal-propodeal disc as long as its half width, metapostnotal median carina complete, with longitudinal ridge between metapostnotal median carina and metapostnotal-propodeal carina, posterior propodeal projection very short and thick; spiracle shape elliptical; propodeal declivity areolate, with median and lateral carinae; lateral surface of metapectal-propodeal complex areolate, without carinae.
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="603">Wings</emphasis>
|
|
: Hind wing with four distal hamuli. Metasoma dorsal and ventral region of terga III-VI polished; hypopygium bilobate, spiculum short, with lobate and short branch, wider than long, lateral margins convergent, with lateral anterior projection.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="603" type="etymology">
|
|
<paragraph pageId="0" pageNumber="603">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="603">
|
|
The name
|
|
<taxonomicName authorityName="Barbosa" authorityYear="2022" family="Bethylidae" genus="Brachymesitius" kingdom="Animalia" lsidName="Brachymesitius" order="Hymenoptera" pageId="0" pageNumber="603" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Brachymesitius</emphasis>
|
|
</taxonomicName>
|
|
, masculine, is a combination of the names
|
|
<normalizedToken originalValue="“brachy”">"brachy"</normalizedToken>
|
|
, from the Greek
|
|
<normalizedToken originalValue="“short”">"short"</normalizedToken>
|
|
, and refers to the reduced size of structures, such as eye size, flagellomeres, ocelli, length of dorsal pronotal area, posterior propodeal projection, and hypopygium, which are diagnostic for the group, and the name "
|
|
<taxonomicName class="Insecta" family="Bethylidae" genus="Mesitius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mesitius" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Mesitius</emphasis>
|
|
</taxonomicName>
|
|
", the type genus of
|
|
<taxonomicName authorityName="Kieffer" authorityYear="1914" lsidName="" pageId="0" pageNumber="603" rank="subFamily" subFamily="Mesitiinae">Mesitiinae</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="603" type="distribution">
|
|
<paragraph pageId="0" pageNumber="603">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="603">Iraq.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="603" type="species included">
|
|
<paragraph pageId="0" pageNumber="603">Species included.</paragraph>
|
|
<paragraph pageId="0" pageNumber="603">
|
|
<taxonomicName class="Insecta" family="Bethylidae" genus="Incertosulcus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Incertosulcus krombeini" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="species" species="krombeini">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Incertosulcus krombeini</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Móczár">Moczar</normalizedToken>
|
|
, 1970, now
|
|
<taxonomicName family="Bethylidae" genus="Brachymesitius" kingdom="Animalia" lsidName="Brachymesitius krombeini" order="Hymenoptera" pageId="0" pageNumber="603" rank="species" species="krombeini">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Brachymesitius krombeini</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<normalizedToken originalValue="Móczár">Moczar</normalizedToken>
|
|
, 1970)
|
|
<emphasis bold="true" pageId="0" pageNumber="603">stat. rev. et comb. nov.</emphasis>
|
|
, removed from the synonym of
|
|
<taxonomicName class="Insecta" family="Bethylidae" genus="Parvoculus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Parvoculus indicus" order="Hymenoptera" pageId="0" pageNumber="603" phylum="Arthropoda" rank="species" species="indicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="603">Parvoculus indicus</emphasis>
|
|
</taxonomicName>
|
|
Kieffer, 1905.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |