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<document id="BA38C6BA7B77E17747A7FB6B5534D4E0" ID-DOI="10.4202/app.00554.2018" ID-ISSN="1732-2421" ID-Zenodo-Dep="10980900" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1713284217429" checkinUser="felipe" docAuthor="Hryniewicz, Krzysztof, Amano, Kazutaka, Bitner, Maria Aleksandra, Hagström, Jonas, Kiel, Steffen, Klompmaker, Adiël A., Mörs, Thomas, Robins, Cristina M. &amp; Kaim, Andrzej" docDate="2019" docId="320C87F44657FFC3FFEFF9AAFBF4DA79" docLanguage="en" docName="ActaPalaeontolPol.64.1.101-141.pdf" docOrigin="Acta Palaeontologica Polonica 64 (1)" docSource="http://dx.doi.org/10.4202/app.00554.2018" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.2:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="2" docTitle="Inoperna plenicostata" docType="treatment" docVersion="3" lastPageNumber="115" masterDocId="CE35FF8C465AFFCDFF8DFFFEFFA8DF58" masterDocTitle="A late Paleocene fauna from shallow-water chemosynthesis-based ecosystems, Spitsbergen, Svalbard" masterLastPageNumber="141" masterPageNumber="101" pageNumber="114" updateTime="1713358443740" updateUser="valdenar" zenodo-license-document="CC-BY-4.0">
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<mods:namePart id="6807219E1828314D09982851078B5432">Amano, Kazutaka</mods:namePart>
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<mods:namePart id="7BF3CDCC4F6AFD564B155BD50ED96412">Bitner, Maria Aleksandra</mods:namePart>
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<mods:namePart id="5CB79DEAF09EAD68521E73E2EC0ABF7A">Hagström, Jonas</mods:namePart>
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<mods:namePart id="190E6F0683AA5E9436EAC81CFB251CCD">Kiel, Steffen</mods:namePart>
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<mods:namePart id="E4A36F2388BD68354462FC05F54C72E3">Klompmaker, Adiël A.</mods:namePart>
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<mods:namePart id="134892D3B226978F9B07D7FED9508BF6">Kaim, Andrzej</mods:namePart>
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<mods:date id="C2F9C9D6F6E7A1568B6C73155E3CCC37">2019</mods:date>
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<subSubSection id="F2BF65694657FFC0FFEFF9AAFDCDD929" box="[98,613,1619,1649]" pageId="13" pageNumber="114" type="nomenclature">
<paragraph id="BA1A36E24657FFC0FFEFF9AAFDCDD929" blockId="13.[98,613,1619,1649]" box="[98,613,1619,1649]" pageId="13" pageNumber="114">
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<taxonomicName id="7DA54D614657FFC0FFEFF9AAFDCDD929" ID-CoL="7RR54" authority="(Anderson, 1970)" baseAuthorityName="Anderson" baseAuthorityYear="1970" box="[98,613,1619,1649]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="plenicostata">
<emphasis id="88D1EAF04657FFC0FFEFF9AAFED3D928" box="[98,379,1619,1648]" italics="true" pageId="13" pageNumber="114">Inoperna plenicostata</emphasis>
(
<bibRefCitation id="DE344B134657FFC0FE01F9ADFDF4D929" author="Anderson, H. - J." box="[396,604,1619,1649]" pageId="13" pageNumber="114" pagination="89 - 101" refId="ref25056" refString="Anderson, H. - J. 1970. Pelecypoda. In: K. Vonderbank (ed.), Geologie und Fauna der tertiaren Ablagerungen Zentral-Spitzbergens. Norsk Polarinstituts Skrifter 153: 89 - 101." type="journal article" year="1970">Anderson, 1970</bibRefCitation>
)
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<subSubSection id="F2BF65694657FFC0FFEFF97EFF02D9CD" box="[98,170,1664,1686]" pageId="13" pageNumber="114" type="description">
<paragraph id="BA1A36E24657FFC0FFEFF97EFF02D9CD" blockId="13.[98,170,1664,1686]" box="[98,170,1664,1686]" pageId="13" pageNumber="114">
<figureCitation id="229E2A674657FFC0FFEFF97EFF0ED9CD" box="[98,166,1664,1686]" captionStart="Fig" captionStartId="14.[151,184,1095,1117]" captionTargetBox="[153,1533,161,1071]" captionTargetId="figure-464@14.[153,1533,161,1071]" captionTargetPageId="14" captionText="Fig.10.Mytilid bivalve Inoperna plenicostata (Anderson, 1970) from the upper Paleocene,Basilika Formation,Zachariassendalen, Spitsbergen,Svalbard. A. NRM-PZ Mo 183949, shell, dorsal view of both valves (A1), right valve (A2), enlarged dorsal view of both valves, showing ornamentation of the early growth stages (A3), oblique anterodorsal view (A4). B. NRM-PZ Mo 183947, partial shell, left valve (B1), ventral view of both valves (B2). C. NRM-PZ Mo 183948, partial shell, left valve (C1), ventral view of partially preserved valves (C2), oblique anterior view of left valve, showing anterior adductor muscle scar (C3)." figureDoi="http://doi.org/10.5281/zenodo.10980922" httpUri="https://zenodo.org/record/10980922/files/figure.png" pageId="13" pageNumber="114">Fig. 10</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F2BF65694657FFC0FFEFF959FD18D981" pageId="13" pageNumber="114" type="reference_group">
<paragraph id="BA1A36E24657FFC0FFEFF959FD18D981" blockId="13.[98,719,1703,1754]" pageId="13" pageNumber="114">
<treatmentCitationGroup id="9AB511CC4657FFC0FFEFF959FD67D9E5" box="[98,719,1703,1725]" pageId="13" pageNumber="114">
1970
<taxonomicName id="7DA54D614657FFC0FF15F959FECDD9E4" ID-CoL="7S8WW" authority="Hryniewicz &amp; Amano &amp; Bitner &amp; Hagström &amp; Kiel &amp; Klompmaker &amp; Mörs &amp; Robins &amp; Kaim, 2019" authorityName="Hryniewicz &amp; Amano &amp; Bitner &amp; Hagström &amp; Kiel &amp; Klompmaker &amp; Mörs &amp; Robins &amp; Kaim" authorityYear="2019" box="[152,357,1703,1724]" class="Bivalvia" family="Mytilidae" genus="Mytilus" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="plenicostatus" status="sp. nov.">
<emphasis id="88D1EAF04657FFC0FF15F959FECDD9E4" box="[152,357,1703,1724]" italics="true" pageId="13" pageNumber="114">Mytilus plenicostatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="93E2578B4657FFC0FEE7F956FE08D9E5" box="[362,416,1704,1725]" pageId="13" pageNumber="114" rank="species">n. sp.</taxonomicNameLabel>
;
<treatmentCitation id="3B0410F34657FFC0FE24F959FDCBD9E4" author="Anderson, H. - J." box="[425,611,1703,1725]" page="93" pageId="13" pageNumber="114" year="1970">
<bibRefCitation id="DE344B134657FFC0FE24F959FDCBD9E4" author="Anderson, H. - J." box="[425,611,1703,1725]" pageId="13" pageNumber="114" pagination="89 - 101" refId="ref25056" refString="Anderson, H. - J. 1970. Pelecypoda. In: K. Vonderbank (ed.), Geologie und Fauna der tertiaren Ablagerungen Zentral-Spitzbergens. Norsk Polarinstituts Skrifter 153: 89 - 101." type="journal article" year="1970">Anderson 1970: 93</bibRefCitation>
</treatmentCitation>
, pl. 9:
<date id="CE1B10224657FFC0FD29F959FF3AD981" pageId="13" pageNumber="114" value="2016-07-06">
67.
<treatmentCitationGroup id="9AB511CC4657FFC0FFEFF93DFD18D981" box="[98,688,1731,1754]" pageId="13" pageNumber="114">
2016
<taxonomicName id="7DA54D614657FFC0FF15F93AFF58D981" ID-CoL="7P5SG" authorityName="Conrad" authorityYear="1875" box="[152,240,1732,1753]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="114" phylum="Mollusca" rank="genus">
<emphasis id="88D1EAF04657FFC0FF15F93AFF58D981" box="[152,240,1732,1753]" italics="true" pageId="13" pageNumber="114">Inoperna</emphasis>
</taxonomicName>
? sp.;
<bibRefCitation id="DE344B134657FFC0FEA5F93AFDAFD981" author="Hryniewicz, K. &amp; Bitner, M. A. &amp; Durska, E. &amp; Hagstrom, J. &amp; Hjalmarsdottir H. R. &amp; Jenkins, R. G. &amp; Little, C. T. S. &amp; Miyajima, Y. &amp; Nakrem, H. A. &amp; Kaim, A." box="[296,519,1731,1754]" pageId="13" pageNumber="114" pagination="41 - 56" refId="ref29314" refString="Hryniewicz, K., Bitner, M. A., Durska, E., Hagstrom, J., Hjalmarsdottir H. R., Jenkins, R. G., Little, C. T. S., Miyajima, Y., Nakrem, H. A., and Kaim, A. 2016. Paleocene methane seep and wood-fall marine environments from Spitsbergen, Svalbard. Palaeogeography, Palaeoclimatology, Palaeoecology 462: 41 - 56." type="journal article" year="2016">Hryniewicz et al. 2016</bibRefCitation>
: table 2, fig. 12E.
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</paragraph>
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<paragraph id="BA1A36E24657FFC0FFEFF915FDE8D8FF" blockId="13.[98,771,1771,1959]" pageId="13" pageNumber="114">
<emphasis id="88D1EAF04657FFC0FFEFF915FF6ED85D" box="[98,198,1771,1797]" italics="true" pageId="13" pageNumber="114">Material</emphasis>
.—
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<specimenCount id="ACA3FD6B4657FFC0FF64F915FE34D85D" box="[233,412,1771,1797]" count="6" pageId="13" pageNumber="114" type="generic">Six specimens</specimenCount>
, all fragmentarily preserved shells, including three (
<specimenCode id="EA039E994657FFC0FEF0F8F2FD90D87E" box="[381,568,1803,1830]" collectionCode="GPIBo" pageId="13" pageNumber="114">GPIBo 155156</specimenCode>
) from the
<location id="BF7A60394657FFC0FD4CF8F2FF7FD81E" LSID="urn:lsid:plazi:treatment:320C87F44657FFC3FFEFF9AAFBF4DA79:BF7A60394657FFC0FD4CF8F2FF7FD81E" country="Svalbard and Jan Mayen" county="Spitsbergen" municipality="Hollendarbukta" name="upper Paleocene" pageId="13" pageNumber="114">upper Paleocene</location>
, locality
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<elevation id="3188D1D14657FFC0FECFF8D2FE24D81E" box="[322,396,1836,1863]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="114" unit="m" value="500.0">500 m</elevation>
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west of
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point 25,
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,
<collectingCounty id="537B4E6E4657FFC0FE92F8B2FE0ED83E" box="[287,422,1868,1894]" pageId="13" pageNumber="114">Spitsbergen</collectingCounty>
,
<collectingCountry id="C2B276724657FFC0FE3DF8B2FDBFD83E" box="[432,535,1868,1894]" name="Svalbard and Jan Mayen" pageId="13" pageNumber="114">Svalbard</collectingCountry>
</materialsCitation>
,
<materialsCitation id="0ACD3CBF4657FFC0FDADF8B2FD93D8FF" collectionCode="NRM-PZ" country="Svalbard and Jan Mayen" county="Spitsbergen" location="upper Paleocene" municipality="Paleocene of Zachariassendalen" pageId="13" pageNumber="114" specimenCount="1">
and three (
<collectionCode id="DCB4AE274657FFC0FD19F8B3FCAAD83F" box="[660,770,1869,1895]" pageId="13" pageNumber="114">NRM-PZ</collectionCode>
Mo 183947183949) specimens from the
<location id="BF7A60394657FFC0FDA0F893FD4ED8DF" LSID="urn:lsid:plazi:treatment:320C87F44657FFC3FFEFF9AAFBF4DA79:BF7A60394657FFC0FDA0F893FD4ED8DF" box="[557,742,1901,1927]" country="Svalbard and Jan Mayen" county="Spitsbergen" municipality="Paleocene of Zachariassendalen" name="upper Paleocene" pageId="13" pageNumber="114">
upper
<collectingMunicipality id="5A7EAC984657FFC0FDF9F893FE9FD8FF" pageId="13" pageNumber="114">Paleocene of Zachariassendalen</collectingMunicipality>
</location>
,
<collectingCounty id="537B4E6E4657FFC0FECFF873FE61D8FF" box="[322,457,1933,1959]" pageId="13" pageNumber="114">Spitsbergen</collectingCounty>
,
<collectingCountry id="C2B276724657FFC0FE59F873FD93D8FF" box="[468,571,1933,1959]" name="Svalbard and Jan Mayen" pageId="13" pageNumber="114">Svalbard</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="BA1A36E24657FFC0FCA5FF61FC25DFA1" blockId="13.[808,1481,158,249]" pageId="13" pageNumber="114">
<emphasis id="88D1EAF04657FFC0FCA5FF61FC64DFE0" box="[808,972,159,184]" italics="true" pageId="13" pageNumber="114">Measurements</emphasis>
.—L, 11.4~
<quantity id="7D5D9B074657FFC0FBD7FF60FB6ADFE1" box="[1114,1218,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="5.9799999999999995" pageId="13" pageNumber="114" unit="mm" value="59.8">59.8 mm</quantity>
; n = 2. Other measurements difficult to estimate due to the poor quality of the material.
</paragraph>
<paragraph id="BA1A36E24657FFC0FCA5FEF5FA3EDE3D" blockId="13.[808,1482,266,942]" pageId="13" pageNumber="114">
<emphasis id="88D1EAF04657FFC0FCA5FEF5FC06DE7C" box="[808,942,267,292]" italics="true" pageId="13" pageNumber="114">Description</emphasis>
.—A description of the species was provided by
<bibRefCitation id="DE344B134657FFC0FCC6FED5FB9BDE1D" author="Anderson, H. - J." box="[843,1075,299,325]" pageId="13" pageNumber="114" pagination="89 - 101" refId="ref25056" refString="Anderson, H. - J. 1970. Pelecypoda. In: K. Vonderbank (ed.), Geologie und Fauna der tertiaren Ablagerungen Zentral-Spitzbergens. Norsk Polarinstituts Skrifter 153: 89 - 101." type="journal article" year="1970">Anderson (1970: 94)</bibRefCitation>
. We hereby provide a description of juvenile specimens, which were not studied previously.
</paragraph>
<paragraph id="BA1A36E24657FFC0FCC3FE92FAF4DCF6" blockId="13.[808,1482,266,942]" pageId="13" pageNumber="114">
Earliest growth stages unknown. Shell shorter than
<quantity id="7D5D9B074657FFC0FCA5FE72FC2DDEFF" box="[808,901,396,423]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="13" pageNumber="114" unit="mm" value="1.75">1.75 mm</quantity>
ornamented with regularly spaced commarginal corrugations with similarly wide depressions separating them. Commarginal growth lines superimposed on fine radial ornament visible in commarginal depressions but diminishing on commarginal corrugations. Possible growth halt visible ~
<quantity id="7D5D9B074657FFC0FCB8FDD0FC32DD11" box="[821,922,558,585]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="13" pageNumber="114" unit="mm" value="0.75">0.75 mm</quantity>
from umbones; second growth halt at ~
<quantity id="7D5D9B074657FFC0FAEDFDD0FA6CDD11" box="[1376,1476,558,585]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="13" pageNumber="114" unit="mm" value="1.75">1.75 mm</quantity>
, roughly coincides with change of ornamentation from regular and strong commarginal ornament superimposed on fine radial ornament to less regular and weaker commarginal currugations and no radial ornament. Latter ornamentation pattern present on remainder of shell; commarginal ornament of largest specimen (
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155; L ~
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) grouped in bundles of 35, whereas commarginal corrugations on shorter specimens show no obvious grouping. Anterior adductor muscle scar minute, rounded, located ventrally to umbo and separated from pallial line. Pallial line visible only in anteroventral part of shell; remainder unknown.
</paragraph>
<paragraph id="BA1A36E24657FFC0FCA5FC41FBA2D83E" blockId="13.[808,1482,959,1960]" pageId="13" pageNumber="114">
<emphasis id="88D1EAF04657FFC0FCA5FC41FC23DC81" box="[808,907,959,985]" italics="true" pageId="13" pageNumber="114">Remarks</emphasis>
.—Some morphological features of this mytilid from Spitsbergen suggest its inclusion in the poorly known genus
<taxonomicName id="7DA54D614657FFC0FCF9FBFFFB2BDB43" authority="Conrad, 1875" authorityName="Conrad" authorityYear="1875" box="[884,1155,1024,1051]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="114" phylum="Mollusca" rank="genus">
<emphasis id="88D1EAF04657FFC0FCF9FBFFFC73DB42" box="[884,987,1025,1050]" italics="true" pageId="13" pageNumber="114">Inoperna</emphasis>
<bibRefCitation id="DE344B134657FFC0FC6BFBFEFB2BDB43" author="Conrad, T. A." box="[998,1155,1024,1051]" pageId="13" pageNumber="114" pagination="1 - 13" refId="ref26694" refString="Conrad, T. A. 1875. Descriptions of new genera and species of fossil shells of North Carolina in the State Cabinet at Raleigh. In: W. C. Kerr (ed.), Appendix A. Report of the Geological Survey of North Carolina 6: 1 - 13." type="journal article" year="1875">Conrad, 1875</bibRefCitation>
</taxonomicName>
. The specimens from locality
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west of Trigonometric point 25 are partial shells with a slightly different shape from the specimens from Zachariassendalen. The
<typeStatus id="651E88404657FFC0FBB6FB9FFB35DB23" box="[1083,1181,1121,1147]" pageId="13" pageNumber="114" type="holotype">holotype</typeStatus>
(
<collectionCode id="DCB4AE274657FFC0FB21FB9CFB51DB24" box="[1196,1273,1122,1148]" pageId="13" pageNumber="114">GPIBo</collectionCode>
155; L ~
<quantity id="7D5D9B074657FFC0FAD3FB9FFA6ADB24" box="[1374,1474,1121,1148]" metricMagnitude="-2" metricUnit="m" metricValue="5.9799999999999995" pageId="13" pageNumber="114" unit="mm" value="59.8">59.8 mm</quantity>
) is an incomplete mytiliform shell with the posterior lobe broken off; therefore, it was originally larger than the measurement given above. The shell has a terminal umbo and no well-defined carina, whereas the
<typeStatus id="651E88404657FFC0FB42FB1AFA99DBA6" box="[1231,1329,1252,1278]" pageId="13" pageNumber="114" type="paratype">paratype</typeStatus>
(
<collectionCode id="DCB4AE274657FFC0FACFFB1DFA27DBA5" box="[1346,1423,1251,1277]" pageId="13" pageNumber="114">GPIBo</collectionCode>
156; L ≥
<quantity id="7D5D9B074657FFC0FCDEFAFAFC1EDA46" box="[851,950,1284,1310]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="13" pageNumber="114" unit="mm" value="11.4">11.4 mm</quantity>
) has a more complete, modioliform shell and a subterminal umbo. Shell shape changes during mytiloid ontogeny (e.g.,
<bibRefCitation id="DE344B134657FFC0FC55FABBFB65DA07" author="Fuller, S. C. &amp; Lutz, R. A." box="[984,1229,1349,1375]" pageId="13" pageNumber="114" pagination="181 - 218" refId="ref28469" refString="Fuller, S. C. and Lutz, R. A. 1989. Shell morphology of larval and postlarval mytilids from the north-western Atlantic. Journal of the Marine Biological Association of the United Kingdom 69: 181 - 218." type="journal article" year="1989">Fuller and Lutz 1989</bibRefCitation>
;
<bibRefCitation id="DE344B134657FFC0FB51FABBFA6DDA07" author="Cosel, R. von &amp; Olu, K." box="[1244,1477,1349,1375]" pageId="13" pageNumber="114" pagination="655 - 663" refId="ref26852" refString="Cosel, R. von and Olu, K. 1998. Gigantism in Mytilidae. A new Bathymodiolus from the cold seep areas of the Barbados accretionary prism. Comptes Rendus de l'Academie des Sciences Serie III 321: 655 - 663." type="journal article" year="1998">Cosel and Olu 1998</bibRefCitation>
;
<bibRefCitation id="DE344B134657FFC0FCA5FA9BFC5CDA27" author="Saether, K. P. &amp; Little, C. T. S. &amp; Campbell, K. A. &amp; Marshall, B. A. &amp; Collins, M. &amp; Alfaro, A. C." box="[808,1012,1381,1408]" pageId="13" pageNumber="114" pagination="1 - 45" refId="ref34890" refString="Saether, K. P., Little, C. T. S., Campbell, K. A., Marshall, B. A., Collins, M., and Alfaro, A. C. 2010. New fossil mussels (Bivalvia: Mytilidae) from Miocene hydrocarbon seep deposits, North Island, New Zealand, with general remarks on vent and seep mussels. Zootaxa 2577: 1 - 45." type="journal article" year="2010">Saether et al. 2010</bibRefCitation>
); therefore, the observed shape differences among specimens of different size are likely an ontogenetic feature. In general, the shell shapes observed in the material from
<collectingCountry id="C2B276724657FFC0FCE8FA39FC60DAB9" box="[869,968,1479,1505]" name="Svalbard and Jan Mayen" pageId="13" pageNumber="114">Svalbard</collectingCountry>
are similar to those known from other species of
<taxonomicName id="7DA54D614657FFC0FCC4FA16FC18D959" authorityName="Conrad" authorityYear="1875" box="[841,944,1512,1537]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="114" phylum="Mollusca" rank="genus">
<emphasis id="88D1EAF04657FFC0FCC4FA16FC18D959" box="[841,944,1512,1537]" italics="true" pageId="13" pageNumber="114">Inoperna</emphasis>
</taxonomicName>
(reviewed by
<bibRefCitation id="DE344B134657FFC0FBD4FA19FA20D95A" author="Stenzel, H. B. &amp; Krause, E. K. &amp; Twining, J. T." box="[1113,1416,1511,1538]" pageId="13" pageNumber="114" pagination="3 - 196" refId="ref36534" refString="Stenzel, H. B., Krause, E. K., and Twining, J. T. 1957. Pelecypoda from the type locality of the Stone City beds (Middle Eocene) of Texas. The University of Texas Publication 5704: 3 - 196." type="journal article" year="1957">Stenzel et al. 1957: 7475</bibRefCitation>
). The curved mytiloid shell of
<taxonomicName id="7DA54D614657FFC0FBBAF9F6FB7CD97A" baseAuthorityName="Anderson" baseAuthorityYear="1970" box="[1079,1236,1544,1570]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="plenicostata">
<emphasis id="88D1EAF04657FFC0FBBAF9F6FB97D979" box="[1079,1087,1544,1569]" italics="true" pageId="13" pageNumber="114">I</emphasis>
.
<emphasis id="88D1EAF04657FFC0FBC1F9F6FB7CD97A" box="[1100,1236,1544,1570]" italics="true" pageId="13" pageNumber="114">plenicostata</emphasis>
</taxonomicName>
is somewhat reminiscent of some specimens of
<taxonomicName id="7DA54D614657FFC0FBD2F9D7FA24D91B" authority="Forbes, 1845" authorityName="Forbes" authorityYear="1845" box="[1119,1420,1576,1603]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="flagellifera">
<emphasis id="88D1EAF04657FFC0FBD2F9D7FBCFD91A" box="[1119,1127,1577,1602]" italics="true" pageId="13" pageNumber="114">I</emphasis>
.
<emphasis id="88D1EAF04657FFC0FBFAF9D6FB5BD91A" box="[1143,1267,1576,1602]" italics="true" pageId="13" pageNumber="114">flagellifera</emphasis>
<bibRefCitation id="DE344B134657FFC0FB71F9D6FA24D91B" author="Forbes, E." box="[1276,1420,1576,1603]" pageId="13" pageNumber="114" pagination="97 - 174" refId="ref28296" refString="Forbes, E. 1845. Report on the fossil Invertebrata from Southern India, collected by Mr. Kaye and Mr. Cunliffe. Transactions of the Geological Society of London. Series 2 7: 97 - 174." type="journal article" year="1845">Forbes, 1845</bibRefCitation>
</taxonomicName>
(e.g.,
<bibRefCitation id="DE344B134657FFC0FCA5F9B7FC18D93B" author="Forbes, E." box="[808,944,1608,1635]" pageId="13" pageNumber="114" pagination="97 - 174" refId="ref28296" refString="Forbes, E. 1845. Report on the fossil Invertebrata from Southern India, collected by Mr. Kaye and Mr. Cunliffe. Transactions of the Geological Society of London. Series 2 7: 97 - 174." type="journal article" year="1845">Forbes 1845</bibRefCitation>
: pl. 16: 9;
<bibRefCitation id="DE344B134657FFC0FB97F9B7FB0FD93B" author="Woods, H." box="[1050,1191,1609,1635]" pageId="13" pageNumber="114" pagination="73 - 112" refId="ref37596" refString="Woods, H. 1900. A monograph of the Cretaceous Lamellibranchia of England. Part. 2. Palaeontographical Society Monographs 54: 73 - 112." type="journal article" year="1900">Woods 1900</bibRefCitation>
: pl. 17: 1), especially of the specimens from the Santonian (Late Cretaceous) Hochmoos Formation of the Gosau Group in
<collectingCountry id="C2B276724657FFC0FB48F974FAB0D9FC" box="[1221,1304,1674,1700]" name="Austria" pageId="13" pageNumber="114">Austria</collectingCountry>
(
<bibRefCitation id="DE344B134657FFC0FAA6F974FA6DD9FC" author="Dhondt, A. V." box="[1323,1477,1674,1700]" pageId="13" pageNumber="114" pagination="41 - 101" refId="ref28042" refString="Dhondt, A. V. 1987. Bivalves from the Hochmoos Formation (Gosau Group, Oberosterreich, Austria). Annalen des Naturhistorischen Museums in Wien A 88: 41 - 101." type="journal article" year="1987">Dhondt 1987</bibRefCitation>
: pl. 1: 13). Most of the species of
<taxonomicName id="7DA54D614657FFC0FB1FF955FB51D99C" authorityName="Conrad" authorityYear="1875" box="[1170,1273,1707,1732]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="114" phylum="Mollusca" rank="genus">
<emphasis id="88D1EAF04657FFC0FB1FF955FB51D99C" box="[1170,1273,1707,1732]" italics="true" pageId="13" pageNumber="114">Inoperna</emphasis>
</taxonomicName>
are known mostly from partially preserved specimens (e.g.,
<bibRefCitation id="DE344B134657FFC0FA8EF934FA6DD9BD" author="Stephenson, L. W." box="[1283,1477,1738,1765]" pageId="13" pageNumber="114" pagination="1 - 408" refId="ref36585" refString="Stephenson, L. W. 1923. The Cretaceous formations of North Carolina. Part 1. Invertebrate fossils of the Upper Cretaceous formations. North Carolina Geological and Economic Survey 5: 1 - 408." type="journal article" year="1923">Stephenson 1923</bibRefCitation>
;
<bibRefCitation id="DE344B134657FFC0FCA5F915FC03D85D" author="Wade, B." box="[808,939,1771,1797]" pageId="13" pageNumber="114" pagination="1 - 272" refId="ref37468" refString="Wade, B. 1926. The Fauna of the Ripley Formation on Coon Creek, Tennessee. United States Geological Survey Professional Paper 137: 1 - 272." type="journal article" year="1926">Wade 1926</bibRefCitation>
;
<bibRefCitation id="DE344B134657FFC0FC37F912FBF4D85D" author="Popenoe, W. P." box="[954,1116,1771,1798]" pageId="13" pageNumber="114" pagination="379 - 402" refId="ref34224" refString="Popenoe, W. P. 1937. Upper Cretaceous Mollusca from Southern California. Journal of Paleontology 11: 379 - 402." type="journal article" year="1937">Popenoe 1937</bibRefCitation>
); therefore, it is difficult to determine their external shapes, and to estimate the degree to which the shell shape of
<taxonomicName id="7DA54D614657FFC0FBB8F8D3FB34D81E" authorityName="Conrad" authorityYear="1875" box="[1077,1180,1837,1862]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="114" phylum="Mollusca" rank="genus">
<emphasis id="88D1EAF04657FFC0FBB8F8D3FB34D81E" box="[1077,1180,1837,1862]" italics="true" pageId="13" pageNumber="114">Inoperna</emphasis>
</taxonomicName>
is taxonomically or phenotypically controlled.
</paragraph>
<paragraph id="BA1A36E24657FFC3FCC3F893FDF1D861" blockId="13.[808,1482,959,1960]" lastBlockId="14.[151,825,1254,1849]" lastPageId="14" lastPageNumber="115" pageId="13" pageNumber="114">
The
<typeStatus id="651E88404657FFC0FC0DF893FC49D8DF" box="[896,993,1901,1927]" pageId="13" pageNumber="114" type="holotype">holotype</typeStatus>
(
<collectionCode id="DCB4AE274657FFC0FC62F893FB94D8DF" box="[1007,1084,1901,1927]" pageId="13" pageNumber="114">GPIBo</collectionCode>
155) of
<taxonomicName id="7DA54D614657FFC0FB1BF890FA22D8DF" baseAuthorityName="Anderson" baseAuthorityYear="1970" box="[1174,1418,1901,1927]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="13" pageNumber="114" phylum="Mollusca" rank="species" species="plenicostata">
<emphasis id="88D1EAF04657FFC0FB1BF890FA22D8DF" box="[1174,1418,1901,1927]" italics="true" pageId="13" pageNumber="114">Inoperna plenicostata</emphasis>
</taxonomicName>
has a characteristic external ornament composed of commarginal corrugations arranged in groups of three to five, separated by broader furrows devoid of any visible ornament. The
<typeStatus id="651E88404654FFC3FD72FAF6FF60DA1C" pageId="14" pageNumber="115" type="paratype">paratype</typeStatus>
(
<collectionCode id="DCB4AE274654FFC3FF58FAD7FE8ADA1B" box="[213,290,1321,1347]" pageId="14" pageNumber="115">GPIBo</collectionCode>
156) has commarginal corrugations not grouped into bundles. We infer that this difference in ornamentation between
<specimenCount id="ACA3FD6B4654FFC3FF76FA93FEF4DADF" box="[251,348,1389,1415]" count="2" pageId="14" pageNumber="115" type="generic">two type</specimenCount>
specimens is likely an ontogenetic feature, as the two are of very different size. An ornamentation composed of commarginal corrugations is typical for
<taxonomicName id="7DA54D614654FFC3FD5FFA4EFC90DA91" authorityName="Conrad" authorityYear="1875" box="[722,824,1456,1481]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="14" pageNumber="115" phylum="Mollusca" rank="genus">
<emphasis id="88D1EAF04654FFC3FD5FFA4EFC90DA91" box="[722,824,1456,1481]" italics="true" pageId="14" pageNumber="115">Inoperna</emphasis>
</taxonomicName>
(
<bibRefCitation id="DE344B134654FFC3FF12FA2EFE86DAB2" author="Woods, H." box="[159,302,1488,1514]" pageId="14" pageNumber="115" pagination="73 - 112" refId="ref37596" refString="Woods, H. 1900. A monograph of the Cretaceous Lamellibranchia of England. Part. 2. Palaeontographical Society Monographs 54: 73 - 112." type="journal article" year="1900">Woods 1900</bibRefCitation>
;
<bibRefCitation id="DE344B134654FFC3FEB7FA2EFE51DAB2" author="Stephenson, L. W." box="[314,505,1488,1514]" pageId="14" pageNumber="115" pagination="1 - 408" refId="ref36585" refString="Stephenson, L. W. 1923. The Cretaceous formations of North Carolina. Part 1. Invertebrate fossils of the Upper Cretaceous formations. North Carolina Geological and Economic Survey 5: 1 - 408." type="journal article" year="1923">Stephenson 1923</bibRefCitation>
;
<bibRefCitation id="DE344B134654FFC3FD89FA2EFD2ADAB2" author="Wade, B." box="[516,642,1488,1514]" pageId="14" pageNumber="115" pagination="1 - 272" refId="ref37468" refString="Wade, B. 1926. The Fauna of the Ripley Formation on Coon Creek, Tennessee. United States Geological Survey Professional Paper 137: 1 - 272." type="journal article" year="1926">Wade 1926</bibRefCitation>
;
<bibRefCitation id="DE344B134654FFC3FD03FA2FFC85DAB2" author="Popenoe, W. P." box="[654,813,1488,1515]" pageId="14" pageNumber="115" pagination="379 - 402" refId="ref34224" refString="Popenoe, W. P. 1937. Upper Cretaceous Mollusca from Southern California. Journal of Paleontology 11: 379 - 402." type="journal article" year="1937">Popenoe 1937</bibRefCitation>
). The ornament of
<taxonomicName id="7DA54D614654FFC3FEE7FA0DFDA6D954" baseAuthorityName="Anderson" baseAuthorityYear="1970" box="[362,526,1522,1548]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="14" pageNumber="115" phylum="Mollusca" rank="species" species="plenicostata">
<emphasis id="88D1EAF04654FFC3FEE7FA0DFEDAD954" box="[362,370,1523,1548]" italics="true" pageId="14" pageNumber="115">I</emphasis>
.
<emphasis id="88D1EAF04654FFC3FE0BFA0CFDA6D954" box="[390,526,1522,1548]" italics="true" pageId="14" pageNumber="115">plenicostata</emphasis>
</taxonomicName>
differs from that known from other species of
<taxonomicName id="7DA54D614654FFC3FE02F9EAFE5DD975" authorityName="Conrad" authorityYear="1875" box="[399,501,1556,1581]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="14" pageNumber="115" phylum="Mollusca" rank="genus">
<emphasis id="88D1EAF04654FFC3FE02F9EAFE5DD975" box="[399,501,1556,1581]" italics="true" pageId="14" pageNumber="115">Inoperna</emphasis>
</taxonomicName>
because the corrugations do not merge close to the dorsal margin to form broader commarginal folds as seen in
<taxonomicName id="7DA54D614654FFC3FE4CF9A9FDC9D928" authorityName="Conrad" authorityYear="1875" box="[449,609,1622,1649]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="14" pageNumber="115" phylum="Mollusca" rank="species" species="carolinensis">
<emphasis id="88D1EAF04654FFC3FE4CF9A9FE61D928" box="[449,457,1623,1648]" italics="true" pageId="14" pageNumber="115">I</emphasis>
.
<emphasis id="88D1EAF04654FFC3FE57F9A8FDC9D928" box="[474,609,1622,1648]" italics="true" pageId="14" pageNumber="115">carolinensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="DE344B134654FFC3FDFFF9A8FC9CD928" author="Stephenson, L. W." box="[626,820,1622,1648]" pageId="14" pageNumber="115" pagination="1 - 408" refId="ref36585" refString="Stephenson, L. W. 1923. The Cretaceous formations of North Carolina. Part 1. Invertebrate fossils of the Upper Cretaceous formations. North Carolina Geological and Economic Survey 5: 1 - 408." type="journal article" year="1923">Stephenson 1923</bibRefCitation>
: pl. 62: 14;
<bibRefCitation id="DE344B134654FFC3FE94F989FE35D9C9" author="Wade, B." box="[281,413,1655,1681]" pageId="14" pageNumber="115" pagination="1 - 272" refId="ref37468" refString="Wade, B. 1926. The Fauna of the Ripley Formation on Coon Creek, Tennessee. United States Geological Survey Professional Paper 137: 1 - 272." type="journal article" year="1926">Wade 1926</bibRefCitation>
: pl. 13: 10) and
<taxonomicName id="7DA54D614654FFC3FDEEF986FD50D9C9" authorityName="Forbes" authorityYear="1845" box="[611,760,1655,1682]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="14" pageNumber="115" phylum="Mollusca" rank="species" species="flagellifera">
<emphasis id="88D1EAF04654FFC3FDEEF986FDC3D9C9" box="[611,619,1656,1681]" italics="true" pageId="14" pageNumber="115">I</emphasis>
.
<emphasis id="88D1EAF04654FFC3FDF0F989FD50D9C9" box="[637,760,1655,1681]" italics="true" pageId="14" pageNumber="115">flagellifera</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation id="DE344B134654FFC3FF1AF967FE8FD9EB" author="Dhondt, A. V." box="[151,295,1689,1715]" pageId="14" pageNumber="115" pagination="41 - 101" refId="ref28042" refString="Dhondt, A. V. 1987. Bivalves from the Hochmoos Formation (Gosau Group, Oberosterreich, Austria). Annalen des Naturhistorischen Museums in Wien A 88: 41 - 101." type="journal article" year="1987">Dhondt 1987</bibRefCitation>
: pl. 1: 1113), and, to some extent,
<taxonomicName id="7DA54D614654FFC3FD12F964FC90D9EB" authorityName="Popenoe" authorityYear="1937" box="[671,824,1689,1715]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="14" pageNumber="115" phylum="Mollusca" rank="species" species="bellarugosa">
<emphasis id="88D1EAF04654FFC3FD12F964FD0FD9EB" box="[671,679,1690,1715]" italics="true" pageId="14" pageNumber="115">I</emphasis>
.
<emphasis id="88D1EAF04654FFC3FD39F967FC90D9EB" box="[692,824,1689,1715]" italics="true" pageId="14" pageNumber="115">bellarugosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="DE344B134654FFC3FF12F944FE94D98C" author="Popenoe, W. P." box="[159,316,1722,1748]" pageId="14" pageNumber="115" pagination="379 - 402" refId="ref34224" refString="Popenoe, W. P. 1937. Upper Cretaceous Mollusca from Southern California. Journal of Paleontology 11: 379 - 402." type="journal article" year="1937">Popenoe 1937</bibRefCitation>
: pl. 45: 7 [not fig. 6]). The commarginal corrugations of most of the species are more regular than those of
<taxonomicName id="7DA54D614654FFC3FF1AF900FE9BD84F" baseAuthorityName="Anderson" baseAuthorityYear="1970" box="[151,307,1789,1816]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="14" pageNumber="115" phylum="Mollusca" rank="species" species="plenicostata">
<emphasis id="88D1EAF04654FFC3FF1AF900FF37D84F" box="[151,159,1790,1815]" italics="true" pageId="14" pageNumber="115">I</emphasis>
.
<emphasis id="88D1EAF04654FFC3FF20F903FE9BD84F" box="[173,307,1789,1815]" italics="true" pageId="14" pageNumber="115">plenicostata</emphasis>
</taxonomicName>
, but we are unable to determine to what extent this feature is of taxonomic significance.
</paragraph>
<caption id="EEDA666A4654FFC3FF1AFBB9FE9BDB95" ID-DOI="http://doi.org/10.5281/zenodo.10980922" ID-Zenodo-Dep="10980922" httpUri="https://zenodo.org/record/10980922/files/figure.png" pageId="14" pageNumber="115" startId="14.[151,184,1095,1117]" targetBox="[153,1533,161,1071]" targetPageId="14" targetType="figure">
<paragraph id="BA1A36E24654FFC3FF1AFBB9FE9BDB95" blockId="14.[151,1536,1095,1232]" pageId="14" pageNumber="115">
Fig. 10. Mytilid bivalve
<taxonomicName id="7DA54D614654FFC3FEE2FBB6FD70DB06" authority="(Anderson, 1970)" baseAuthorityName="Anderson" baseAuthorityYear="1970" box="[367,728,1095,1118]" class="Bivalvia" family="Mytilidae" genus="Inoperna" kingdom="Animalia" order="Mytilida" pageId="14" pageNumber="115" phylum="Mollusca" rank="species" species="plenicostata">
<emphasis id="88D1EAF04654FFC3FEE2FBB6FD9EDB04" box="[367,566,1095,1117]" italics="true" pageId="14" pageNumber="115">Inoperna plenicostata</emphasis>
(
<bibRefCitation id="DE344B134654FFC3FDCDFBB9FD7BDB06" author="Anderson, H. - J." box="[576,723,1095,1118]" pageId="14" pageNumber="115" pagination="89 - 101" refId="ref25056" refString="Anderson, H. - J. 1970. Pelecypoda. In: K. Vonderbank (ed.), Geologie und Fauna der tertiaren Ablagerungen Zentral-Spitzbergens. Norsk Polarinstituts Skrifter 153: 89 - 101." type="journal article" year="1970">Anderson, 1970</bibRefCitation>
)
</taxonomicName>
from the upper Paleocene, Basilika Formation, Zachariassendalen, Spitsbergen, Svalbard.
<emphasis id="88D1EAF04654FFC3FF1AFB9DFF0FDB20" bold="true" box="[151,167,1123,1144]" pageId="14" pageNumber="115">A</emphasis>
. NRM-PZ Mo 183949, shell, dorsal view of both valves (A
<subScript id="262134A74654FFC3FD4CFB90FD61DB24" attach="right" box="[705,713,1134,1148]" fontSize="6" pageId="14" pageNumber="115">1</subScript>
), right valve (A
<subScript id="262134A74654FFC3FCD4FB90FCC9DB24" attach="left" box="[857,865,1134,1148]" fontSize="6" pageId="14" pageNumber="115">2</subScript>
), enlarged dorsal view of both valves, showing ornamentation of the early growth stages (A
<subScript id="262134A74654FFC3FEBFFB74FE92DBC0" attach="left" box="[306,314,1162,1176]" fontSize="6" pageId="14" pageNumber="115">3</subScript>
), oblique anterodorsal view (A
<subScript id="262134A74654FFC3FDDAFB74FDF7DBC0" attach="right" box="[599,607,1162,1176]" fontSize="6" pageId="14" pageNumber="115">4</subScript>
).
<emphasis id="88D1EAF04654FFC3FDFCFB7EFD28DBCD" bold="true" box="[625,640,1152,1173]" pageId="14" pageNumber="115">B</emphasis>
. NRM-PZ Mo 183947, partial shell, left valve (B
<subScript id="262134A74654FFC3FBC9FB74FBE4DBC0" attach="left" box="[1092,1100,1162,1176]" fontSize="6" pageId="14" pageNumber="115">1</subScript>
), ventral view of both valves (B
<subScript id="262134A74654FFC3FAFFFB74FAD2DBC0" attach="left" box="[1394,1402,1162,1176]" fontSize="6" pageId="14" pageNumber="115">2</subScript>
).
<emphasis id="88D1EAF04654FFC3FA00FB81FA35DBCD" bold="true" box="[1421,1437,1151,1173]" pageId="14" pageNumber="115">C</emphasis>
. NRM-PZ Mo 183948, partial shell, left valve (C
<subScript id="262134A74654FFC3FE75FB58FDA8DBEC" attach="left" box="[504,512,1190,1204]" fontSize="6" pageId="14" pageNumber="115">1</subScript>
), ventral view of partially preserved valves (C
<subScript id="262134A74654FFC3FC20FB58FC1DDBEC" attach="left" box="[941,949,1190,1204]" fontSize="6" pageId="14" pageNumber="115">2</subScript>
), oblique anterior view of left valve, showing anterior adductor muscle scar (C
<subScript id="262134A74654FFC3FE93FB3CFE8EDB88" attach="left" box="[286,294,1218,1232]" fontSize="6" pageId="14" pageNumber="115">3</subScript>
).
</paragraph>
</caption>
<paragraph id="BA1A36E24654FFC3FF1AF8B5FC90D8FF" blockId="14.[151,824,1867,1959]" pageId="14" pageNumber="115">
<emphasis id="88D1EAF04654FFC3FF1AF8B5FDE4D83C" box="[151,588,1867,1893]" italics="true" pageId="14" pageNumber="115">Stratigraphic and geographic range</emphasis>
.—Upper Paleocene cold seep carbonate and associated sunken driftwood from the Basilika Formation, Zachariassendalen and locality
</paragraph>
<paragraph id="BA1A36E24654FFC3FCD3FB18FBF4DA79" blockId="14.[862,1534,1254,1313]" pageId="14" pageNumber="115">
<quantity id="7D5D9B074654FFC3FCD3FB18FC0FDA59" box="[862,935,1254,1281]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="115" unit="m" value="500.0">500 m</quantity>
west from Trigonometric point 25, Hollendarbukta, Spitsbergen,
<collectingCountry id="C2B276724654FFC3FC7DFAF9FBFFDA79" box="[1008,1111,1287,1313]" name="Svalbard and Jan Mayen" pageId="14" pageNumber="115">Svalbard</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>