2529 lines
264 KiB
XML
2529 lines
264 KiB
XML
<document ID-DOI="10.3161/15081109ACC2022.24.1.002" ID-GBIF-Dataset="ba996155-bf80-4449-9cc2-73621e4b5500" ID-ISSN="1733-5329" ID-Zenodo-Dep="7738166" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1681319635082" checkinUser="felipe" docAuthor="Curran, Michael, Kopp, Mirjam, Ruedi, Manuel & Bayliss, Julian" docDate="2022" docId="F02387A6FFE4FFB7FC2CFA1AFD91F84F" docLanguage="en" docName="ActaChiropterol.24.1.19-40.pdf" docOrigin="Acta Chiropterologica 24 (1)" docStyle="DocumentStyle:D17EC282BABB8B0847763DCC9F0C3514.1:ActaChiropterol.2016-.journal_article" docStyleId="D17EC282BABB8B0847763DCC9F0C3514" docStyleName="ActaChiropterol.2016-.journal_article" docStyleVersion="1" docTitle="Rhinolophus namuli Curran & Kopp & Ruedi & Bayliss 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="36" masterDocId="0C1AFFDEFFECFFA5FF9FFF83FE49FFFA" masterDocTitle="A new species of horseshoe bat (Chiroptera: Rhinolophidae) from Mount Namuli, Mozambique" masterLastPageNumber="40" masterPageNumber="19" pageNumber="26" updateTime="1682354699395" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>A new species of horseshoe bat (Chiroptera: Rhinolophidae) from Mount Namuli, Mozambique</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Curran, Michael</mods:namePart>
|
||
<mods:affiliation>Formerly: Department of Environmental Sciences, Biogeography Section, University of Basel, 4056 Basel, Switzerland & Present address: Department of Socioeconomic Sciences, Research Institute for Organic Agriculture (FiBL), Ackerstrasse 113, P. O. Box 219, CH- 5070 Frick, Switzerland & Corresponding author: E-mail: michael. curran @ fibl. org</mods:affiliation>
|
||
<mods:nameIdentifier type="email">michael.curran@fibl.org</mods:nameIdentifier>
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||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Kopp, Mirjam</mods:namePart>
|
||
<mods:affiliation>Freelance consultant, 3550 Langnau i. E., Switzerland</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Ruedi, Manuel</mods:namePart>
|
||
<mods:affiliation>Natural History Museum of Geneva, Genève 6, Switzerland</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Bayliss, Julian</mods:namePart>
|
||
<mods:affiliation>Department of Biological and Medical Sciences, Oxford Brookes University, Oxford, OX 3 0 BP, United Kingdom & African Butterfly Research Institute, P. O. Box 14308, 0800 Nairobi, Kenya</mods:affiliation>
|
||
</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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||
<mods:titleInfo>
|
||
<mods:title>Acta Chiropterologica</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2022</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2022-08-18</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>24</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>1</mods:number>
|
||
</mods:detail>
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||
<mods:extent unit="page">
|
||
<mods:start>19</mods:start>
|
||
<mods:end>40</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
|
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<mods:identifier type="DOI">10.3161/15081109ACC2022.24.1.002</mods:identifier>
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<mods:identifier type="GBIF-Dataset">ba996155-bf80-4449-9cc2-73621e4b5500</mods:identifier>
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<mods:identifier type="ISSN">1733-5329</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7738166</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7846158" ID-GBIF-Taxon="209561542" ID-Zenodo-Dep="7846158" LSID="urn:lsid:plazi:treatment:F02387A6FFE4FFB7FC2CFA1AFD91F84F" httpUri="http://treatment.plazi.org/id/F02387A6FFE4FFB7FC2CFA1AFD91F84F" lastPageId="18" lastPageNumber="36" pageId="8" pageNumber="26">
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<subSubSection pageId="8" pageNumber="26" type="nomenclature">
|
||
<paragraph blockId="8.[878,1352,1351,1534]" pageId="8" pageNumber="26">
|
||
<heading box="[947,1284,1433,1459]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="26" reason="2">
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[947,1184,1433,1459]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[947,1184,1433,1459]" italics="true" pageId="8" pageNumber="26">Rhinolophus namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1192,1284,1433,1459]" pageId="8" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
Curran, Kopp, Ruedi, and Bayliss, 2022
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[882,1348,1508,1534]" pageId="8" pageNumber="26" type="vernacular_names">
|
||
<paragraph blockId="8.[878,1352,1351,1534]" box="[882,1348,1508,1534]" pageId="8" pageNumber="26">Common name: Namuli Horseshoe Bat</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="26" type="reference_group">
|
||
<paragraph blockId="8.[809,1422,1555,1973]" pageId="8" pageNumber="26">
|
||
2009
|
||
<taxonomicName box="[870,1033,1555,1576]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis box="[870,991,1555,1576]" italics="true" pageId="8" pageNumber="26">Rhinolophus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation author="TIMBERLAKE, J. & J. BAYLISS & T. ALVES & S. BAENA & J. FRANCISCO & T. HARRIS & C. DA SOUSA & C. BENTO & K. COOK & P. SMITH" box="[1082,1341,1555,1576]" pageId="8" pageNumber="26" refId="ref17507" refString="TIMBERLAKE, J., J. BAYLISS, T. ALVES, S. BAENA, J. FRANCISCO, T. HARRIS, C. DA SOUSA, F. DOWSETT- LEMAIRE, C. BENTO, K. COOK, and P. SMITH. 2008. Mount Namuli, Mozambique: Biodiversity and Conservation. Report Produced under the Darwin Initiative Award 15 / 036." type="journal volume" year="2008">
|
||
Timberlake
|
||
<emphasis box="[1205,1261,1555,1576]" italics="true" pageId="8" pageNumber="26">et al.</emphasis>
|
||
(2008)
|
||
</bibRefCitation>
|
||
: Mount Namuli,
|
||
<collectingCountry box="[944,1079,1584,1605]" name="Mozambique" pageId="8" pageNumber="26">Mozambique</collectingCountry>
|
||
: biodiversity and conservation. Report produced under the Darwin Initiative Award 15/036. Pp. 60 and 110.
|
||
</paragraph>
|
||
<paragraph blockId="8.[152,230,172,1973]" pageId="8" pageNumber="26">= this FA characteristics. in</paragraph>
|
||
<paragraph blockId="8.[152,230,172,1973]" pageId="8" pageNumber="26">species combined comparative related were Key closely which,. 2 genetically specimens</paragraph>
|
||
<paragraph blockId="8.[809,1422,1555,1973]" pageId="8" pageNumber="26">
|
||
2010
|
||
<taxonomicName box="[865,1130,1669,1690]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" isUncertain="true" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[865,986,1669,1690]" italics="true" pageId="8" pageNumber="26">Rhinolophus</emphasis>
|
||
cf.
|
||
<emphasis box="[1026,1130,1669,1690]" italics="true" pageId="8" pageNumber="26">maendeleo</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation author="MONADJEM, A. & P. J. TAYLOR & F. P. D. COTTERILL & M. C. SCHOEMAN" box="[1169,1416,1669,1690]" pageId="8" pageNumber="26" refId="ref16683" refString="MONADJEM, A., P. J. TAYLOR, F. P. D. COTTERILL, and M. C. SCHOEMAN. 2010 b. Bats of Southern and Central Africa: a biogeographic and taxonomic synthesis. Wits University Press, Johannesburg, 596 pp." type="book" year="2010">
|
||
Monadjem
|
||
<emphasis box="[1281,1332,1669,1690]" italics="true" pageId="8" pageNumber="26">et al.</emphasis>
|
||
(2010
|
||
<emphasis box="[1394,1406,1669,1690]" italics="true" pageId="8" pageNumber="26">b</emphasis>
|
||
)
|
||
</bibRefCitation>
|
||
: Act. Chiropterol., 12: 371–391 — partim: Mount Namuli.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="26" type="description">
|
||
<paragraph blockId="8.[809,1422,1555,1973]" pageId="8" pageNumber="26">
|
||
2010
|
||
<taxonomicName box="[862,1093,1725,1746]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[862,1093,1725,1746]" italics="true" pageId="8" pageNumber="26">Rhinolophus maendeleo</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation author="MONADJEM, A. & P. J. TAYLOR & F. P. D. COTTERILL & M. C. SCHOEMAN" box="[1128,1368,1725,1746]" pageId="8" pageNumber="26" refId="ref16683" refString="MONADJEM, A., P. J. TAYLOR, F. P. D. COTTERILL, and M. C. SCHOEMAN. 2010 b. Bats of Southern and Central Africa: a biogeographic and taxonomic synthesis. Wits University Press, Johannesburg, 596 pp." type="book" year="2010">
|
||
Monadjem
|
||
<emphasis box="[1237,1286,1725,1746]" italics="true" pageId="8" pageNumber="26">et al.</emphasis>
|
||
(2010
|
||
<emphasis box="[1346,1358,1725,1746]" italics="true" pageId="8" pageNumber="26">b</emphasis>
|
||
)
|
||
</bibRefCitation>
|
||
: Bats of Southern and Central Africa: a biogeographic and taxonomic synthesis. Wits University Press, Johannesburg: 220–223, 560 — partim: Mount Namuli.
|
||
</paragraph>
|
||
<paragraph blockId="8.[809,1422,1555,1973]" pageId="8" pageNumber="26">
|
||
2019
|
||
<taxonomicName authorityName="Thomas" authorityYear="1904" box="[865,1074,1839,1860]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" isUncertain="true" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="denti">
|
||
<emphasis box="[865,986,1839,1860]" italics="true" pageId="8" pageNumber="26">Rhinolophus</emphasis>
|
||
cf.
|
||
<emphasis box="[1028,1074,1839,1860]" italics="true" pageId="8" pageNumber="26">denti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis box="[1074,1175,1839,1860]" italics="true" pageId="8" pageNumber="26">
|
||
/
|
||
<taxonomicName box="[1085,1175,1839,1860]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="26" phylum="Chordata" rank="species" species="simulator">simulator</taxonomicName>
|
||
</emphasis>
|
||
—
|
||
<bibRefCitation author="DEMOS, T. C. & P. W. WEBALA & S. M. GOODMAN & J. C. KERBIS PETERHANS & M. BARTONJO & B. D. PATTERSON" box="[1216,1418,1839,1860]" pageId="8" pageNumber="26" pagination="166" refId="ref16014" refString="DEMOS, T. C., P. W. WEBALA, S. M. GOODMAN, J. C. KERBIS PETERHANS, M. BARTONJO, and B. D. PATTERSON. 2019. Molecular phylogenetics of the African horseshoe bats (Chiroptera: Rhinolophidae): expanded geographic and taxonomic sampling of the Afrotropics. BMC Evolutionary Biology, 19: 166." type="journal article" year="2019">
|
||
Demos
|
||
<emphasis box="[1292,1344,1839,1860]" italics="true" pageId="8" pageNumber="26">et al.</emphasis>
|
||
(2019)
|
||
</bibRefCitation>
|
||
. Molecular phylogenetics of the African horseshoe bats (
|
||
<taxonomicName box="[853,963,1895,1916]" class="Mammalia" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="26" phylum="Chordata" rank="order">Chiroptera</taxonomicName>
|
||
:
|
||
<taxonomicName authorityName="Gray" authorityYear="1825" box="[981,1130,1895,1916]" class="Mammalia" family="Rhinolophidae" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="26" phylum="Chordata" rank="family">Rhinolophidae</taxonomicName>
|
||
): expanded geographic and taxonomic sampling of the Afrotropics. BMC Evolutionary Biology, 19: 166.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<paragraph blockId="8.[152,230,172,1973]" pageId="8" pageNumber="26">
|
||
<subScript attach="both" box="[156,172,1913,1960]" fontSize="7" pageId="8" pageNumber="26">ABLE</subScript>
|
||
T
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7859656" ID-Zenodo-Dep="7859656" httpUri="https://zenodo.org/record/7859656/files/figure.png" pageId="9" pageNumber="27" startId="9.[151,164,1836,1857]" targetBox="[193,1372,177,1809]" targetPageId="9" targetType="figure">
|
||
<paragraph blockId="9.[151,1421,1836,1971]" pageId="9" pageNumber="27">
|
||
FIG. 3. Maximum likelihood tree based on CYT-B sequences and reconstructed with IQ-tree. Sequences of
|
||
<taxonomicName baseAuthorityName="Bechstein" baseAuthorityYear="1800" box="[1214,1366,1836,1857]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="9" pageNumber="27" phylum="Chordata" rank="species" species="hipposideros">
|
||
<emphasis box="[1214,1366,1836,1857]" italics="true" pageId="9" pageNumber="27">R. hipposideros</emphasis>
|
||
</taxonomicName>
|
||
were used as outgroups. Bootstrap support levels are explained in the box; the scale indicates nucleotide substitutions per site. Newly sequenced taxa are associated with their GenBank accession numbers (see Table 2 and Supplementary Appendix S3 for more details), while the species groups to which they belong appear to the right of the tree. The clade representing the new species (sp. nov.)
|
||
</paragraph>
|
||
<paragraph blockId="9.[151,1421,1836,1971]" box="[683,890,1950,1971]" pageId="9" pageNumber="27">is highlighted in grey</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="10" pageNumber="28" type="materials_examined">
|
||
<paragraph blockId="10.[151,764,178,1973]" pageId="10" pageNumber="28">
|
||
<materialsCitation ID-GBIF-Occurrence="4081226301" collectingDate="2008-11-21" collectingMethod="2 - tiered harp trap" collectionCode="CYT-B, COI" country="Mozambique" county="Male" elevation="1650" latitude="-15.36925" location="Ukalini" longLatPrecision="1" longitude="37.06136" municipality="Mount Namuli" pageId="10" pageNumber="28" specimenCode="MHNG 1971.068" specimenCount="1" stateProvince="Zambezia Province" typeStatus="holotype">
|
||
<emphasis box="[189,330,178,204]" italics="true" pageId="10" pageNumber="28">
|
||
<typeStatus box="[189,296,178,204]" pageId="10" pageNumber="28">Holotype</typeStatus>
|
||
.—
|
||
</emphasis>
|
||
<specimenCount box="[330,457,178,204]" pageId="10" pageNumber="28" type="male">Male adult</specimenCount>
|
||
(alcohol preserved, skull, baculum preserved in glycerol),
|
||
<specimenCode box="[552,763,212,238]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="10" pageNumber="28" type="Museum">MHNG 1971.068</specimenCode>
|
||
(field collection number MZ 120).
|
||
<collectingCountry box="[601,759,246,272]" name="Mozambique" pageId="10" pageNumber="28">Mozambique</collectingCountry>
|
||
,
|
||
<collectingRegion box="[151,389,280,306]" country="Mozambique" name="Zambezia" pageId="10" pageNumber="28">Zambezia Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality box="[408,592,280,306]" pageId="10" pageNumber="28">Mount Namuli</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:F02387A6FFE4FFB7FC2CFA1AFD91F84F:7D55606BFFE6FFAFFDFBFE9BFCF5FEC8" box="[612,700,280,306]" country="Mozambique" county="Male" latitude="-15.36925" longLatPrecision="1" longitude="37.06136" municipality="Mount Namuli" name="Ukalini" pageId="10" pageNumber="28" stateProvince="Zambezia Province">Ukalini</location>
|
||
saddle forest, below mountain peak,
|
||
<geoCoordinate box="[602,759,314,340]" degrees="15.369250" direction="south" orientation="latitude" pageId="10" pageNumber="28" precision="1" value="-15.36925">15.369250°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[151,305,348,374]" degrees="37.061361" direction="east" orientation="longitude" pageId="10" pageNumber="28" precision="1" value="37.06136">37.061361°E</geoCoordinate>
|
||
ca.
|
||
<quantity box="[357,447,348,374]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="10" pageNumber="28" unit="m" value="1650.0">
|
||
<elevation box="[357,447,348,374]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="10" pageNumber="28" unit="m" value="1650.0">1650 m</elevation>
|
||
</quantity>
|
||
a.s.l.,
|
||
<date box="[527,759,348,374]" pageId="10" pageNumber="28" value="2008-11-21">
|
||
<collectingDate box="[527,759,348,374]" pageId="10" pageNumber="28" value="2008-11-21">21 November 2008</collectingDate>
|
||
</date>
|
||
. Captured in
|
||
<collectingMethod box="[314,537,382,408]" pageId="10" pageNumber="28">2-tiered harp trap</collectingMethod>
|
||
set over a small stream leading to large clearing in montane forest, at ca. 18h00. Reference sequences of the mitochondrial CYT-B and
|
||
<emphasis box="[334,385,484,510]" italics="true" pageId="10" pageNumber="28">COI</emphasis>
|
||
genes issued from the holotype are deposited in the GenBank under accession numbers
|
||
<accessionNumber box="[208,340,552,578]" pageId="10" pageNumber="28">MZ936306</accessionNumber>
|
||
and
|
||
<accessionNumber box="[399,533,552,578]" pageId="10" pageNumber="28">MZ936310</accessionNumber>
|
||
, respectively.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="10.[151,764,178,1973]" pageId="10" pageNumber="28">
|
||
<materialsCitation ID-GBIF-Occurrence="4081226302" collectingDate="2008-11-21" collectingMethod="2 - tiered harp trap" collectionCode="MHNG" country="Mozambique" elevation="1650" latitude="-15.36925" location="Ukalini" longLatPrecision="1" longitude="37.06136" municipality="Mount Namuli" pageId="10" pageNumber="28" specimenCode="MHNG 1971.069" specimenCount="1" stateProvince="Zambezia Province" typeStatus="paratype">
|
||
<emphasis box="[189,347,586,612]" italics="true" pageId="10" pageNumber="28">
|
||
<typeStatus box="[189,312,586,612]" pageId="10" pageNumber="28">Paratypes</typeStatus>
|
||
.—
|
||
</emphasis>
|
||
<specimenCount box="[348,493,586,612]" pageId="10" pageNumber="28" type="male">Male adult</specimenCount>
|
||
(alcohol preserved, skull, baculum preserved in glycerol),
|
||
<specimenCode pageId="10" pageNumber="28">MHNG 1971.069</specimenCode>
|
||
(field collection number MZ257).
|
||
<collectingCountry name="Mozambique" pageId="10" pageNumber="28">Mozambique</collectingCountry>
|
||
,
|
||
<collectingRegion box="[295,544,688,714]" country="Mozambique" name="Zambezia" pageId="10" pageNumber="28">Zambezia Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality box="[567,759,688,714]" pageId="10" pageNumber="28">Mount Namuli</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:F02387A6FFE4FFB7FC2CFA1AFD91F84F:7D55606BFFE6FFAFFF08FD51FEBAFD16" box="[151,243,722,748]" country="Mozambique" latitude="-15.36925" longLatPrecision="1" longitude="37.06136" municipality="Mount Namuli" name="Ukalini" pageId="10" pageNumber="28" stateProvince="Zambezia Province">Ukalini</location>
|
||
saddle forest, below mountain peak,
|
||
<geoCoordinate box="[151,306,756,782]" degrees="15.369250" direction="south" orientation="latitude" pageId="10" pageNumber="28" precision="1" value="-15.36925">15.369250°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[325,482,756,782]" degrees="37.061361" direction="east" orientation="longitude" pageId="10" pageNumber="28" precision="1" value="37.06136">37.061361°E</geoCoordinate>
|
||
, ca.
|
||
<quantity box="[549,644,756,782]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="10" pageNumber="28" unit="m" value="1650.0">
|
||
<elevation box="[549,644,756,782]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="10" pageNumber="28" unit="m" value="1650.0">1650 m</elevation>
|
||
</quantity>
|
||
a.s.l.,
|
||
<date pageId="10" pageNumber="28" value="2008-11-21">
|
||
<collectingDate pageId="10" pageNumber="28" value="2008-11-21">21 November 2008</collectingDate>
|
||
</date>
|
||
. Captured in
|
||
<collectingMethod box="[501,704,790,816]" pageId="10" pageNumber="28">2-tiered harp trap</collectingMethod>
|
||
over small stream leading to large clearing in montane forest, ca. 18h00.
|
||
</materialsCitation>
|
||
<materialsCitation ID-GBIF-Occurrence="4081226303" collectingDate="2008-11-21" collectingMethod="2 - tiered harp trap" collectionCode="MHNG" country="Mozambique" elevation="1650" latitude="-15.36925" location="Ukalini" longLatPrecision="1" longitude="37.06136" municipality="Mount Namuli" pageId="10" pageNumber="28" specimenCode="MHNG 1971.067" specimenCount="1" stateProvince="Zambezia Province">
|
||
<specimenCount box="[370,526,858,884]" pageId="10" pageNumber="28" type="female">Female adult</specimenCount>
|
||
(alcohol preserved, skull),
|
||
<specimenCode box="[239,453,892,918]" pageId="10" pageNumber="28">MHNG 1971.067</specimenCode>
|
||
(field collection number MZ261).
|
||
<collectingCountry box="[266,424,926,952]" name="Mozambique" pageId="10" pageNumber="28">Mozambique</collectingCountry>
|
||
,
|
||
<collectingRegion box="[438,671,926,952]" country="Mozambique" name="Zambezia" pageId="10" pageNumber="28">Zambezia Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality pageId="10" pageNumber="28">Mount Namuli</collectingMunicipality>
|
||
, camp on mountain plateau,
|
||
<geoCoordinate box="[603,759,960,986]" degrees="15.405933" direction="south" orientation="latitude" pageId="10" pageNumber="28" precision="1" value="-15.405933">15.405933°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[151,308,994,1020]" degrees="37.067006" direction="east" orientation="longitude" pageId="10" pageNumber="28" precision="1" value="37.067005">37.067006°E</geoCoordinate>
|
||
, ca.
|
||
<quantity box="[360,456,994,1020]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="28" unit="m" value="1200.0">
|
||
<elevation box="[360,456,994,1020]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="28" unit="m" value="1200.0">1,200 m</elevation>
|
||
</quantity>
|
||
a.s.l.,
|
||
<date box="[531,760,994,1020]" pageId="10" pageNumber="28" value="2008-11-20">
|
||
<collectingDate box="[531,760,994,1020]" pageId="10" pageNumber="28" value="2008-11-20">20 November 2008</collectingDate>
|
||
</date>
|
||
.
|
||
<location LSID="urn:lsid:plazi:treatment:F02387A6FFE4FFB7FC2CFA1AFD91F84F:7D55606BFFE6FFAFFF08FB87FE84FBE4" box="[151,205,1028,1054]" country="Mozambique" latitude="-15.405933" longLatPrecision="1" longitude="37.067005" municipality="Mount Namuli" name="Mist" pageId="10" pageNumber="28" stateProvince="Zambezia Province">Mist</location>
|
||
netted over river in fragment of mid-altitude forest on mountain plateau, ca. 18:45 h. Reference sequences of the mitochondrial CYT-B and
|
||
<emphasis box="[713,764,1096,1122]" italics="true" pageId="10" pageNumber="28">COI</emphasis>
|
||
genes issued from these paratypes are deposited in the GenBank under accession numbers
|
||
<accessionNumber box="[617,749,1164,1190]" pageId="10" pageNumber="28">MZ936305</accessionNumber>
|
||
–
|
||
<accessionNumber box="[151,285,1198,1224]" pageId="10" pageNumber="28">MZ936308</accessionNumber>
|
||
and
|
||
<accessionNumber box="[366,502,1198,1224]" pageId="10" pageNumber="28">MZ936309</accessionNumber>
|
||
–
|
||
<accessionNumber box="[517,654,1198,1224]" pageId="10" pageNumber="28">MZ936312</accessionNumber>
|
||
, respectively.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="10.[151,764,178,1973]" lastBlockId="10.[809,1422,177,510]" pageId="10" pageNumber="28">
|
||
<emphasis box="[189,436,1267,1293]" italics="true" pageId="10" pageNumber="28">Referred material.—</emphasis>
|
||
Female adult (alcohol preserved, skull), DM 10839.
|
||
<collectingCountry box="[476,634,1301,1327]" name="Mozambique" pageId="10" pageNumber="28">Mozambique</collectingCountry>
|
||
,
|
||
<collectingRegion country="Mozambique" name="Zambezia" pageId="10" pageNumber="28">Zambezia Province</collectingRegion>
|
||
, Mount Namuli, Manho Plateau Forest,
|
||
<geoCoordinate box="[151,277,1369,1395]" degrees="15.3975" direction="south" orientation="latitude" pageId="10" pageNumber="28" precision="5" value="-15.3975">15.3975°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[295,422,1369,1395]" degrees="37.0372" direction="east" orientation="longitude" pageId="10" pageNumber="28" precision="5" value="37.0372">37.0372°E</geoCoordinate>
|
||
, ca.
|
||
<quantity box="[486,588,1369,1395]" metricMagnitude="3" metricUnit="m" metricValue="1.734" pageId="10" pageNumber="28" unit="m" value="1734.0">1,734 m</quantity>
|
||
a.s.l.,
|
||
<date pageId="10" pageNumber="28" value="2008-11-20">20 November 2008</date>
|
||
. Mist netted over small stream in forest clearing at approximately 18:30 h. Male adult (alcohol preserved, skull, baculum preserved in glycerol), DM 10833 (field collection number MZ099). Mount Namuli, base camp on mountain plateau,
|
||
<geoCoordinate box="[259,414,1573,1599]" degrees="15.405933" direction="south" orientation="latitude" pageId="10" pageNumber="28" precision="1" value="-15.405933">15.405933°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[434,591,1573,1599]" degrees="37.067006" direction="east" orientation="longitude" pageId="10" pageNumber="28" precision="1" value="37.067005">37.067006°E</geoCoordinate>
|
||
, ca.
|
||
<quantity box="[659,764,1573,1599]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="28" unit="m" value="1200.0">1,200 m</quantity>
|
||
a.s.l.,
|
||
<date box="[223,457,1607,1633]" pageId="10" pageNumber="28" value="2008-11-20">20 November 2008</date>
|
||
. Mist netted over river in remnant fragment of mid-altitude forest on mountain plateau, ca. 18:00 h. Other referred material are
|
||
<specimenCount box="[151,346,1709,1735]" pageId="10" pageNumber="28" type="generic">five specimens</specimenCount>
|
||
previously reported as
|
||
<taxonomicName authorityName="Thomas" authorityYear="1904" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" isUncertain="true" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="denti">
|
||
<emphasis box="[686,711,1709,1735]" italics="true" pageId="10" pageNumber="28">R.</emphasis>
|
||
cf.
|
||
<emphasis box="[151,207,1743,1769]" italics="true" pageId="10" pageNumber="28">denti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis box="[207,330,1743,1769]" italics="true" pageId="10" pageNumber="28">
|
||
/
|
||
<taxonomicName box="[221,330,1743,1769]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="simulator">simulator</taxonomicName>
|
||
</emphasis>
|
||
by
|
||
<bibRefCitation author="DEMOS, T. C. & P. W. WEBALA & S. M. GOODMAN & J. C. KERBIS PETERHANS & M. BARTONJO & B. D. PATTERSON" box="[378,621,1743,1769]" pageId="10" pageNumber="28" pagination="166" refId="ref16014" refString="DEMOS, T. C., P. W. WEBALA, S. M. GOODMAN, J. C. KERBIS PETERHANS, M. BARTONJO, and B. D. PATTERSON. 2019. Molecular phylogenetics of the African horseshoe bats (Chiroptera: Rhinolophidae): expanded geographic and taxonomic sampling of the Afrotropics. BMC Evolutionary Biology, 19: 166." type="journal article" year="2019">
|
||
Demos
|
||
<emphasis box="[471,531,1743,1769]" italics="true" pageId="10" pageNumber="28">et al.</emphasis>
|
||
(2019)
|
||
</bibRefCitation>
|
||
, which also proved to belong to
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[385,498,1777,1803]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[385,498,1777,1803]" italics="true" pageId="10" pageNumber="28">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[505,595,1777,1803]" pageId="10" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
based on very similar CYT-B lineages. These specimens were collected approximatively
|
||
<quantity box="[429,489,1845,1871]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="28" unit="km" value="8.0">8 km</quantity>
|
||
north and
|
||
<quantity box="[618,677,1845,1871]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="28" unit="km" value="8.0">8 km</quantity>
|
||
east of Gurue,
|
||
<collectingCountry box="[259,419,1879,1905]" name="Mozambique" pageId="10" pageNumber="28">Mozambique</collectingCountry>
|
||
(FMNH 17708–177110), in the Zovo Chipolo Forest, Nyika National Park,
|
||
<collectingCountry box="[151,241,1947,1973]" name="Malawi" pageId="10" pageNumber="28">Malawi</collectingCountry>
|
||
(FMNH 191585), and in the Ndundulu Forest,
|
||
<quantity box="[903,968,177,203]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="28" unit="km" value="9.0">9 km</quantity>
|
||
east of Udekwa Udzungwa Mts, in
|
||
<collectingCountry box="[809,918,212,238]" name="Tanzania" pageId="10" pageNumber="28">Tanzania</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="28" type="distribution">
|
||
<paragraph blockId="10.[809,1422,177,510]" pageId="10" pageNumber="28">
|
||
<emphasis box="[847,1242,246,272]" italics="true" pageId="10" pageNumber="28">Type locality and distribution.—</emphasis>
|
||
Mount Namuli (
|
||
<geoCoordinate box="[817,975,280,306]" degrees="15.369250" direction="south" orientation="latitude" pageId="10" pageNumber="28" precision="1" value="-15.36925">15.369250°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[988,1144,280,306]" degrees="37.061361" direction="east" orientation="longitude" pageId="10" pageNumber="28" precision="1" value="37.06136">37.061361°E</geoCoordinate>
|
||
, ca.
|
||
<quantity box="[1200,1298,280,306]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="10" pageNumber="28" unit="m" value="1650.0">1,650 m</quantity>
|
||
a.s.l.), ca.
|
||
<quantity box="[809,890,314,340]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="28" unit="km" value="10.0">10 km</quantity>
|
||
north-west of Gurue,
|
||
<collectingRegion box="[1179,1417,314,340]" country="Mozambique" name="Zambezia" pageId="10" pageNumber="28">Zambezia Province</collectingRegion>
|
||
, north-central
|
||
<collectingCountry box="[975,1130,348,374]" name="Mozambique" pageId="10" pageNumber="28">Mozambique</collectingCountry>
|
||
(central
|
||
<collectingCountry box="[1249,1410,348,374]" name="Mozambique" pageId="10" pageNumber="28">Mozambique</collectingCountry>
|
||
). Recorded in montane evergreen forest and nearby forest clearings on the high plateau and below the peak of the mountain at altitudes ranging
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.435" metricValueMax="1.65" metricValueMin="1.22" pageId="10" pageNumber="28" unit="m" value="1435.0" valueMax="1650.0" valueMin="1220.0">1,220 to 1,650 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="28" type="etymology">
|
||
<paragraph blockId="10.[809,934,552,578]" box="[809,934,552,578]" pageId="10" pageNumber="28">
|
||
<heading box="[809,934,552,578]" fontSize="36" level="3" pageId="10" pageNumber="28" reason="6">
|
||
<emphasis box="[809,934,552,578]" italics="true" pageId="10" pageNumber="28">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="10.[809,1422,586,816]" pageId="10" pageNumber="28">
|
||
The species epithet
|
||
<taxonomicName authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[1088,1169,586,612]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="namuli">
|
||
<emphasis box="[1088,1169,586,612]" italics="true" pageId="10" pageNumber="28">namuli</emphasis>
|
||
</taxonomicName>
|
||
is a noun in apposition and is a derivative of the
|
||
<typeStatus box="[1183,1234,620,646]" pageId="10" pageNumber="28">type</typeStatus>
|
||
locality Mount Namuli,
|
||
<collectingRegion box="[912,1143,654,680]" country="Mozambique" name="Zambezia" pageId="10" pageNumber="28">Zambezia Province</collectingRegion>
|
||
,
|
||
<collectingCountry box="[1156,1315,654,680]" name="Mozambique" pageId="10" pageNumber="28">Mozambique</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="28" type="description">
|
||
<paragraph blockId="10.[809,1422,586,816]" pageId="10" pageNumber="28">
|
||
<emphasis box="[847,1149,688,714]" italics="true" pageId="10" pageNumber="28">Nomenclatural statement</emphasis>
|
||
.—A life number was obtained for the new species
|
||
<taxonomicName authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[1172,1412,722,748]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="namuli">
|
||
<emphasis box="[1172,1412,722,748]" italics="true" pageId="10" pageNumber="28">Rhinolophus namuli</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<uri pageId="10" pageNumber="28">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid pageId="10" pageNumber="28">7A9806D9-CCC6-41A2- B898-6DCDEFFFBB51</uuid>
|
||
</uri>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="28" type="diagnosis">
|
||
<paragraph blockId="10.[809,928,858,884]" box="[809,928,858,884]" pageId="10" pageNumber="28">
|
||
<heading box="[809,928,858,884]" fontSize="36" level="3" pageId="10" pageNumber="28" reason="6">
|
||
<emphasis box="[809,928,858,884]" italics="true" pageId="10" pageNumber="28">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="10.[809,1422,892,1803]" pageId="10" pageNumber="28">
|
||
Medium sized bat (forearm length
|
||
<quantity metricMagnitude="-2" metricUnit="m" metricValue="4.835" metricValueMax="4.95" metricValueMin="4.72" pageId="10" pageNumber="28" unit="mm" value="48.35" valueMax="49.5" valueMin="47.2">47.2–49.5 mm</quantity>
|
||
) with medium sized ears and three well-defined mental groves on lower lip (Supplementary Appendix S2
|
||
<figureCitation box="[895,987,994,1020]" captionStart="FIG" captionStartId="2.[151,164,1920,1941]" captionTargetBox="[152,1413,198,1885]" captionTargetId="figure-20@2.[151,1423,173,1904]" captionTargetPageId="2" captionText="FIG. 1. Regional overview (A) and zoom in of the study area on Mount Namuli (B), showing specimen collection sites and extant montane forest and woodland cover, overlaid onto a hillshade DEM (SRTM, 90 m resolution)" figureDoi="http://doi.org/10.5281/zenodo.7859652" httpUri="https://zenodo.org/record/7859652/files/figure.png" pageId="10" pageNumber="28">Fig. A1</figureCitation>
|
||
). Horseshoe and sella wide (
|
||
<figureCitation box="[1333,1406,994,1020]" captionStart="FIG" captionStartId="11.[151,164,1178,1199]" captionTargetBox="[151,763,174,1162]" captionTargetId="figure-171@11.[151,765,174,1163]" captionTargetPageId="11" captionText="FIG. 4. Live photographs of R. namuli sp. nov. in the field (♀, MHNG 1971.067), illustrating pelage colour and features of the (a) head and ears, (b) sella and lips and (c) connecting process" figureDoi="http://doi.org/10.5281/zenodo.7859658" httpUri="https://zenodo.org/record/7859658/files/figure.png" pageId="10" pageNumber="28">Fig. 4</figureCitation>
|
||
), lancet concave with a narrow tip (hastate), connecting process high and rounded, with scattered hairs present along its ridge (
|
||
<figureCitation box="[1098,1172,1096,1122]" captionStart="FIG" captionStartId="12.[151,164,893,914]" captionTargetBox="[361,1218,174,872]" captionTargetId="figure-451@12.[348,1226,173,879]" captionTargetPageId="12" captionText="FIG. 5. Head shape (a and b) and noseleaf (c and d) of holotypes of R. maendeleo SMF 79.643 and R. namuli sp. nov. MHNG 1971.068, respectively; connecting process of R. maendeleo SMF 79.643 (e), SMF 99.760 (f), and of R. namuli sp. nov. MHNG 1971.068 (g)" figureDoi="http://doi.org/10.5281/zenodo.7859660" httpUri="https://zenodo.org/record/7859660/files/figure.png" pageId="10" pageNumber="28">Fig. 5</figureCitation>
|
||
, Supplementary Appendix S2
|
||
<figureCitation box="[937,1026,1130,1156]" captionStart="FIG" captionStartId="7.[151,164,1894,1915]" captionTargetBox="[164,1399,1144,1876]" captionTargetId="figure-494@7.[154,1407,1134,1884]" captionTargetPageId="7" captionText="FIG. 2. PCA biplot for selected (A) craniodental and (B) external measurements. Ellipses represent 95% confidence intervals around each species group (not possible for R. maendeleo due to the low sample size). Large symbols represent the species mean value. Percentage values on each axis are the variances explained. For specimen accession numbers and input data, see Tables 3 and 4" figureDoi="http://doi.org/10.5281/zenodo.7859654" httpUri="https://zenodo.org/record/7859654/files/figure.png" pageId="10" pageNumber="28">Fig. A2</figureCitation>
|
||
and A
|
||
<figureCitation box="[1109,1123,1130,1156]" captionStart="FIG" captionStartId="9.[151,164,1836,1857]" captionTargetBox="[193,1372,177,1809]" captionTargetId="graphics-12@9.[194,1370,177,1776]" captionTargetPageId="9" captionText="FIG. 3. Maximum likelihood tree based on CYT-B sequences and reconstructed with IQ-tree. Sequences of R. hipposideros were used as outgroups. Bootstrap support levels are explained in the box; the scale indicates nucleotide substitutions per site. Newly sequenced taxa are associated with their GenBank accession numbers (see Table 2 and Supplementary Appendix S3 for more details), while the species groups to which they belong appear to the right of the tree. The clade representing the new species (sp. nov.) is highlighted in grey" figureDoi="http://doi.org/10.5281/zenodo.7859656" httpUri="https://zenodo.org/record/7859656/files/figure.png" pageId="10" pageNumber="28">3</figureCitation>
|
||
). Elongated first phalanx of third digit relative to metacarpal (
|
||
<tableCitation box="[1242,1333,1164,1190]" captionStart="TABLE" captionStartId="13.[99,112,574,595]" captionTargetPageId="13" captionText="TABLE 3. External measurements of specimens of R. namuli sp. nov., R. maendeleo and R. simulator. Measurement with an asterisk in the header have non-overlapping ranges between R. namuli sp. nov. and R. maendeleo. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’" httpUri="http://table.plazi.org/id/2CF56638FFE1FFA8FFFCFDBDF895FD76" pageId="10" pageNumber="28" tableUuid="2CF56638FFE1FFA8FFFCFDBDF895FD76">Table 3</tableCitation>
|
||
). Bony bar forming the infraorbital foramen is present and slender, inflated nasal swellings (
|
||
<figureCitation box="[1201,1272,1232,1258]" captionStart="FIG" captionStartId="14.[151,164,1031,1052]" captionTargetBox="[196,1383,174,1015]" captionTargetId="figure-411@14.[196,1383,174,1016]" captionTargetPageId="14" captionText="FIG. 6. Cranium in dorsoventral (top), lateral (middle) and ventrodorsal (bottom; only for MHNG 1971.168) views of holotypes of R. namuli sp. nov. (left) and R. maendeleo (right)" figureDoi="http://doi.org/10.5281/zenodo.7859662" httpUri="https://zenodo.org/record/7859662/files/figure.png" pageId="10" pageNumber="28">Fig. 6</figureCitation>
|
||
and Supplementary Appendix S2
|
||
<figureCitation box="[1092,1190,1267,1293]" captionStart="FIG" captionStartId="11.[151,164,1178,1199]" captionTargetBox="[151,763,174,1162]" captionTargetId="figure-171@11.[151,765,174,1163]" captionTargetPageId="11" captionText="FIG. 4. Live photographs of R. namuli sp. nov. in the field (♀, MHNG 1971.067), illustrating pelage colour and features of the (a) head and ears, (b) sella and lips and (c) connecting process" figureDoi="http://doi.org/10.5281/zenodo.7859658" httpUri="https://zenodo.org/record/7859658/files/figure.png" pageId="10" pageNumber="28">Fig. A4</figureCitation>
|
||
), length of palatal bridge 36% of upper toothrow length (
|
||
<tableCitation box="[1307,1407,1301,1327]" captionStart="TABLE" captionStartId="15.[99,112,575,596]" captionTargetPageId="15" captionText="TABLE 4. Craniodental measurements of specimens of R. namuli sp. nov., R. maendeleo and R. simulator. Measurement with an asterisk in the header have non-overlapping ranges between R. namuli sp. nov. and R. maendeleo. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’" pageId="10" pageNumber="28">Table 4</tableCitation>
|
||
). Minute pm3 either present or absent in the lower toothrow, with pm2 and pm4 not making contact (
|
||
<figureCitation box="[818,890,1403,1429]" captionStart="FIG" captionStartId="16.[151,164,988,1009]" captionTargetBox="[183,736,182,961]" captionTargetId="figure-96@16.[151,764,173,970]" captionTargetPageId="16" captionText="FIG. 7. Mandibles of the holotype of R. namuli sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7859664" httpUri="https://zenodo.org/record/7859664/files/figure.png" pageId="10" pageNumber="28">Fig. 7</figureCitation>
|
||
). Baculum medium-sized, dorsal and ventral parts of the basal cone subequal in size with dorsal and ventral incision shallow (
|
||
<figureCitation box="[1154,1224,1471,1497]" captionStart="FIG" captionStartId="17.[151,164,1598,1619]" captionTargetBox="[212,1360,176,1577]" captionTargetId="figure-126@17.[199,1386,173,1579]" captionTargetPageId="17" captionText="FIG. 8. Baculum of the holotype (MHNG 1971.168) of R. namuli sp. nov. in dorsal-ventral (a), ventral-dorsal (c), lateral (b and d) and proximal-distal (g) views. Baculum of the DM 10833 also shown in dorsal-ventral (e) and lateral (f) view. For the holotype (MHNG 1971.168), total baculum length is ca. 2.85 mm, greatest lateral width of basal cone is 0.78 mm and greatest dorsoventral width of basal cone is 0.87 mm" figureDoi="http://doi.org/10.5281/zenodo.7859666" httpUri="https://zenodo.org/record/7859666/files/figure.png" pageId="10" pageNumber="28">Fig. 8</figureCitation>
|
||
). In lateral view, baculum shaft narrows towards tip, undulating in shape and forms an evenly rounded ventrally bent tip. In dorsoventral view, shaft straight with shallow constriction at inflection point of spatulated tip (Supplementary Appendix S2
|
||
<figureCitation box="[1162,1253,1641,1667]" captionStart="FIG" captionStartId="12.[151,164,893,914]" captionTargetBox="[361,1218,174,872]" captionTargetId="figure-451@12.[348,1226,173,879]" captionTargetPageId="12" captionText="FIG. 5. Head shape (a and b) and noseleaf (c and d) of holotypes of R. maendeleo SMF 79.643 and R. namuli sp. nov. MHNG 1971.068, respectively; connecting process of R. maendeleo SMF 79.643 (e), SMF 99.760 (f), and of R. namuli sp. nov. MHNG 1971.068 (g)" figureDoi="http://doi.org/10.5281/zenodo.7859660" httpUri="https://zenodo.org/record/7859660/files/figure.png" pageId="10" pageNumber="28">Fig. A5</figureCitation>
|
||
).
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis italics="true" pageId="10" pageNumber="28">Rhinolophus namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[905,1003,1675,1701]" pageId="10" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
echolocates at a mean peak frequency of 76.9 kHz, SD = 0.8, range = 75.9–78.0 (
|
||
<figureCitation box="[818,891,1743,1769]" captionStart="FIG" captionStartId="18.[151,164,713,734]" captionTargetBox="[288,1292,174,691]" captionTargetId="figure-537@18.[373,1247,171,639]" captionTargetPageId="18" captionText="FIG. 9. Echolocation call of R. namuli sp. nov. (X, MHNG 1971.067; recorded at Mount Namuli, Zambesia Province, Mozambique, -15.405933 / 37.067006, ca. 1,200 m a.s.l.)" figureDoi="http://doi.org/10.5281/zenodo.7859668" httpUri="https://zenodo.org/record/7859668/files/figure.png" pageId="10" pageNumber="28">Fig. 9</figureCitation>
|
||
and
|
||
<tableCitation box="[953,1045,1743,1769]" captionStart="TABLE" captionStartId="16.[151,164,1459,1480]" captionTargetPageId="16" captionText="TABLE 5. Echolocation call parameters of R. namuli sp. nov. and species with similar echolocation peak frequency from nearby localities. Table highlights mean peak frequency (PF), standard deviation (SD), range and number of individuals recorded (n). Data collected by M. Curran and M. Kopp (unpublished material). MZ = Mozambique, MW = Malawi" httpUri="http://table.plazi.org/id/2CF56638FFFCFFB5FF08FA30FA0FF9FA" pageId="10" pageNumber="28" tableUuid="2CF56638FFFCFFB5FF08FA30FA0FF9FA">Table 5</tableCitation>
|
||
). Phylogenetically, this species belongs to the
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[983,1084,1777,1803]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="28" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[983,1084,1777,1803]" italics="true" pageId="10" pageNumber="28">capensis</emphasis>
|
||
</taxonomicName>
|
||
-group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="12" lastPageNumber="30" pageId="10" pageNumber="28" type="description">
|
||
<paragraph blockId="10.[809,948,1845,1871]" box="[809,948,1845,1871]" pageId="10" pageNumber="28">
|
||
<heading box="[809,948,1845,1871]" fontSize="36" level="3" pageId="10" pageNumber="28" reason="6">
|
||
<emphasis box="[809,948,1845,1871]" italics="true" pageId="10" pageNumber="28">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="10.[809,1422,1879,1973]" lastBlockId="11.[151,764,1342,1972]" lastPageId="11" lastPageNumber="29" pageId="10" pageNumber="28">
|
||
Dorsal pelage near uniform beige brown, slightly darker towards head, turning near rusty on both sides of the muzzle. Ventral fur paler than dorsal. Hair length ca.
|
||
<quantity box="[271,340,1342,1368]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="29" unit="mm" value="7.0">7 mm</quantity>
|
||
, with dark base becoming lighter towards tips causing slight shimmering effect. At time of capture, both male and female live animals exhibited musky and somewhat unpleasant smell.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7859658" ID-Zenodo-Dep="7859658" httpUri="https://zenodo.org/record/7859658/files/figure.png" pageId="11" pageNumber="29" startId="11.[151,164,1178,1199]" targetBox="[151,763,174,1162]" targetPageId="11" targetType="figure">
|
||
<paragraph blockId="11.[151,764,1178,1284]" pageId="11" pageNumber="29">
|
||
FIG. 4. Live photographs of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[448,547,1178,1199]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[448,547,1178,1199]" italics="true" pageId="11" pageNumber="29">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[558,638,1178,1199]" pageId="11" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in the field (♀, MHNG 1971.067), illustrating pelage colour and features of the (a) head and ears, (b) sella and lips and (c) connecting
|
||
</paragraph>
|
||
<paragraph blockId="11.[151,764,1178,1284]" box="[421,493,1263,1284]" pageId="11" pageNumber="29">process</paragraph>
|
||
</caption>
|
||
<paragraph blockId="11.[151,764,1342,1972]" lastBlockId="11.[809,1422,175,1973]" pageId="11" pageNumber="29">
|
||
<emphasis box="[189,253,1476,1502]" italics="true" pageId="11" pageNumber="29">Head</emphasis>
|
||
.—Ears medium sized, just reaching tip of muzzle when laid forward, tips pointed in
|
||
<typeStatus box="[661,764,1510,1536]" pageId="11" pageNumber="29">holotype</typeStatus>
|
||
but bluntly rounded in both
|
||
<typeStatus box="[477,592,1543,1569]" pageId="11" pageNumber="29">paratypes</typeStatus>
|
||
, 8–9 ear folds. Three well-developed mental grooves on the lower lip, both lips devoid of hair (
|
||
<figureCitation box="[526,602,1610,1636]" captionStart="FIG" captionStartId="11.[151,164,1178,1199]" captionTargetBox="[151,763,174,1162]" captionTargetId="figure-171@11.[151,765,174,1163]" captionTargetPageId="11" captionText="FIG. 4. Live photographs of R. namuli sp. nov. in the field (♀, MHNG 1971.067), illustrating pelage colour and features of the (a) head and ears, (b) sella and lips and (c) connecting process" figureDoi="http://doi.org/10.5281/zenodo.7859658" httpUri="https://zenodo.org/record/7859658/files/figure.png" pageId="11" pageNumber="29">Fig. 4</figureCitation>
|
||
and Supplementary Appendix S2
|
||
<figureCitation box="[416,507,1644,1670]" captionStart="FIG" captionStartId="2.[151,164,1920,1941]" captionTargetBox="[152,1413,198,1885]" captionTargetId="figure-20@2.[151,1423,173,1904]" captionTargetPageId="2" captionText="FIG. 1. Regional overview (A) and zoom in of the study area on Mount Namuli (B), showing specimen collection sites and extant montane forest and woodland cover, overlaid onto a hillshade DEM (SRTM, 90 m resolution)" figureDoi="http://doi.org/10.5281/zenodo.7859652" httpUri="https://zenodo.org/record/7859652/files/figure.png" pageId="11" pageNumber="29">Fig. A1</figureCitation>
|
||
). The mental grooves were diagnosed according to
|
||
<bibRefCitation author="CSORBA, G. & P. UJHELYI & N. THOMAS" box="[515,760,1678,1704]" pageId="11" pageNumber="29" refId="ref15877" refString="CSORBA, G., P. UJHELYI, and N. THOMAS. 2003. Horseshoe bats of the World (Chiroptera: Rhinolophidae), 1 st edition. Alana Books, Shropshire, UK, 160 pp." type="book" year="2003">
|
||
Csorba
|
||
<emphasis box="[611,675,1678,1704]" italics="true" pageId="11" pageNumber="29">et al.</emphasis>
|
||
(2003
|
||
</bibRefCitation>
|
||
: Fig. vi, p. xxviii) and are shown in close-up in Supplementary Appendix S2 (
|
||
<figureCitation box="[529,629,1745,1771]" captionStart="FIG" captionStartId="2.[151,164,1920,1941]" captionTargetBox="[152,1413,198,1885]" captionTargetId="figure-20@2.[151,1423,173,1904]" captionTargetPageId="2" captionText="FIG. 1. Regional overview (A) and zoom in of the study area on Mount Namuli (B), showing specimen collection sites and extant montane forest and woodland cover, overlaid onto a hillshade DEM (SRTM, 90 m resolution)" figureDoi="http://doi.org/10.5281/zenodo.7859652" httpUri="https://zenodo.org/record/7859652/files/figure.png" pageId="11" pageNumber="29">Fig. A1</figureCitation>
|
||
). Noseleaf completely covers muzzle, horseshoe wide and secondary noseleaf absent. Sella naked, wide at base, slightly constricted in the middle but otherwise almost parallel sided with rounded tip (
|
||
<figureCitation box="[631,707,1879,1905]" captionStart="FIG" captionStartId="12.[151,164,893,914]" captionTargetBox="[361,1218,174,872]" captionTargetId="figure-451@12.[348,1226,173,879]" captionTargetPageId="12" captionText="FIG. 5. Head shape (a and b) and noseleaf (c and d) of holotypes of R. maendeleo SMF 79.643 and R. namuli sp. nov. MHNG 1971.068, respectively; connecting process of R. maendeleo SMF 79.643 (e), SMF 99.760 (f), and of R. namuli sp. nov. MHNG 1971.068 (g)" figureDoi="http://doi.org/10.5281/zenodo.7859660" httpUri="https://zenodo.org/record/7859660/files/figure.png" pageId="11" pageNumber="29">Fig. 5</figureCitation>
|
||
and Supplementary Appendix S2
|
||
<figureCitation box="[516,615,1913,1939]" captionStart="FIG" captionStartId="7.[151,164,1894,1915]" captionTargetBox="[164,1399,1144,1876]" captionTargetId="figure-494@7.[154,1407,1134,1884]" captionTargetPageId="7" captionText="FIG. 2. PCA biplot for selected (A) craniodental and (B) external measurements. Ellipses represent 95% confidence intervals around each species group (not possible for R. maendeleo due to the low sample size). Large symbols represent the species mean value. Percentage values on each axis are the variances explained. For specimen accession numbers and input data, see Tables 3 and 4" figureDoi="http://doi.org/10.5281/zenodo.7859654" httpUri="https://zenodo.org/record/7859654/files/figure.png" pageId="11" pageNumber="29">Fig. A2</figureCitation>
|
||
). Internarial cup rather well developed. Internarial septum is not markedly expanded. Connecting process somewhat hairy, high and rounded, basal part rises near parallel to sella. Tip of the lancet is hastate, with concave sides and a pointed, tapering tip. The hastate lancet differentiates
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[971,1086,311,337]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[971,1086,311,337]" italics="true" pageId="11" pageNumber="29">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1095,1187,311,337]" pageId="11" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from
|
||
<taxonomicName box="[1260,1421,311,337]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[1260,1421,311,337]" italics="true" pageId="11" pageNumber="29">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
and other members of the
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[1127,1200,345,371]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[1127,1200,345,371]" italics="true" pageId="11" pageNumber="29">adami</emphasis>
|
||
</taxonomicName>
|
||
-group (
|
||
<figureCitation box="[1297,1369,345,371]" captionStart="FIG" captionStartId="12.[151,164,893,914]" captionTargetBox="[361,1218,174,872]" captionTargetId="figure-451@12.[348,1226,173,879]" captionTargetPageId="12" captionText="FIG. 5. Head shape (a and b) and noseleaf (c and d) of holotypes of R. maendeleo SMF 79.643 and R. namuli sp. nov. MHNG 1971.068, respectively; connecting process of R. maendeleo SMF 79.643 (e), SMF 99.760 (f), and of R. namuli sp. nov. MHNG 1971.068 (g)" figureDoi="http://doi.org/10.5281/zenodo.7859660" httpUri="https://zenodo.org/record/7859660/files/figure.png" pageId="11" pageNumber="29">Fig. 5</figureCitation>
|
||
and Supplementary Appendix S2
|
||
<figureCitation box="[1156,1248,379,405]" captionStart="FIG" captionStartId="7.[151,164,1894,1915]" captionTargetBox="[164,1399,1144,1876]" captionTargetId="figure-494@7.[154,1407,1134,1884]" captionTargetPageId="7" captionText="FIG. 2. PCA biplot for selected (A) craniodental and (B) external measurements. Ellipses represent 95% confidence intervals around each species group (not possible for R. maendeleo due to the low sample size). Large symbols represent the species mean value. Percentage values on each axis are the variances explained. For specimen accession numbers and input data, see Tables 3 and 4" figureDoi="http://doi.org/10.5281/zenodo.7859654" httpUri="https://zenodo.org/record/7859654/files/figure.png" pageId="11" pageNumber="29">Fig. A2</figureCitation>
|
||
), more closely resembling members of the
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[1177,1278,413,439]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[1177,1278,413,439]" italics="true" pageId="11" pageNumber="29">capensis</emphasis>
|
||
</taxonomicName>
|
||
-group (see Supplementary Appendix S2
|
||
<figureCitation box="[1155,1244,447,473]" captionStart="FIG" captionStartId="9.[151,164,1836,1857]" captionTargetBox="[193,1372,177,1809]" captionTargetId="graphics-12@9.[194,1370,177,1776]" captionTargetPageId="9" captionText="FIG. 3. Maximum likelihood tree based on CYT-B sequences and reconstructed with IQ-tree. Sequences of R. hipposideros were used as outgroups. Bootstrap support levels are explained in the box; the scale indicates nucleotide substitutions per site. Newly sequenced taxa are associated with their GenBank accession numbers (see Table 2 and Supplementary Appendix S3 for more details), while the species groups to which they belong appear to the right of the tree. The clade representing the new species (sp. nov.) is highlighted in grey" figureDoi="http://doi.org/10.5281/zenodo.7859656" httpUri="https://zenodo.org/record/7859656/files/figure.png" pageId="11" pageNumber="29">Fig. A3</figureCitation>
|
||
for a comparison of lancet shape with
|
||
<taxonomicName box="[1103,1249,482,508]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="29" phylum="Chordata" rank="species" species="simulator">
|
||
<emphasis box="[1103,1249,482,508]" italics="true" pageId="11" pageNumber="29">R. simulator</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="11.[809,1422,175,1973]" pageId="11" pageNumber="29">
|
||
<emphasis box="[847,981,516,542]" italics="true" pageId="11" pageNumber="29">Wing bones</emphasis>
|
||
.—Metacarpal of the 4th and 5th digit subequal in length, always longer than the 3rd digit (
|
||
<tableCitation box="[817,908,584,610]" captionStart="TABLE" captionStartId="13.[99,112,574,595]" captionTargetPageId="13" captionText="TABLE 3. External measurements of specimens of R. namuli sp. nov., R. maendeleo and R. simulator. Measurement with an asterisk in the header have non-overlapping ranges between R. namuli sp. nov. and R. maendeleo. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’" httpUri="http://table.plazi.org/id/2CF56638FFE1FFA8FFFCFDBDF895FD76" pageId="11" pageNumber="29" tableUuid="2CF56638FFE1FFA8FFFCFDBDF895FD76">Table 3</tableCitation>
|
||
). Length of the 1st phalanx greatest in 3rd digit, followed by 5th, then 4th digit. Length of the 2nd phalanx greatest in 3rd digit, followed by 4th, then 5th digit. 2nd phalanx of 4th digit greater than twice the length of 1st phalanx.
|
||
</paragraph>
|
||
<paragraph blockId="11.[809,1422,175,1973]" pageId="11" pageNumber="29">
|
||
<emphasis box="[847,951,754,780]" italics="true" pageId="11" pageNumber="29">Cranium</emphasis>
|
||
.—Skull is narrow (mastoid width exceeds zygomatic width) in MHNG 1971.068 (
|
||
<typeStatus pageId="11" pageNumber="29">holotype</typeStatus>
|
||
, male), MHNG 1971.069 (
|
||
<typeStatus box="[1201,1307,822,848]" pageId="11" pageNumber="29">paratype</typeStatus>
|
||
, female), and DM 108333 (referred material, male), but robust (zygomatic width exceeds mastoid width) in MHNG 1971.067 (
|
||
<specimenCount box="[936,959,925,951]" pageId="11" pageNumber="29" type="male">♂</specimenCount>
|
||
) and DM 10839 (referred material,
|
||
<specimenCount box="[1382,1403,925,951]" pageId="11" pageNumber="29" type="female">♀</specimenCount>
|
||
). Differences are slight, indicating within-species variation in skull shape (
|
||
<tableCitation box="[1094,1185,993,1019]" captionStart="TABLE" captionStartId="15.[99,112,575,596]" captionTargetPageId="15" captionText="TABLE 4. Craniodental measurements of specimens of R. namuli sp. nov., R. maendeleo and R. simulator. Measurement with an asterisk in the header have non-overlapping ranges between R. namuli sp. nov. and R. maendeleo. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’" pageId="11" pageNumber="29">Table 4</tableCitation>
|
||
). Weakly developed sagittal crest extends roughly three quarters along the length of the braincase from well-developed supraorbital crest (sagittal crest equally developed in both male and female). Posterior-most point of skull (meeting point of lambdoid and sagittal crest) positioned markedly above the lateral plane formed by the zygomatic arches. Anterior nasal swelling inflated, moderately wide and high, angled forward and protruding anteriorly in lateral view (
|
||
<figureCitation box="[1300,1371,1300,1326]" captionStart="FIG" captionStartId="14.[151,164,1031,1052]" captionTargetBox="[196,1383,174,1015]" captionTargetId="figure-411@14.[196,1383,174,1016]" captionTargetPageId="14" captionText="FIG. 6. Cranium in dorsoventral (top), lateral (middle) and ventrodorsal (bottom; only for MHNG 1971.168) views of holotypes of R. namuli sp. nov. (left) and R. maendeleo (right)" figureDoi="http://doi.org/10.5281/zenodo.7859662" httpUri="https://zenodo.org/record/7859662/files/figure.png" pageId="11" pageNumber="29">Fig. 6</figureCitation>
|
||
and Supplementary Appendix S2
|
||
<figureCitation box="[1156,1249,1334,1360]" captionStart="FIG" captionStartId="11.[151,164,1178,1199]" captionTargetBox="[151,763,174,1162]" captionTargetId="figure-171@11.[151,765,174,1163]" captionTargetPageId="11" captionText="FIG. 4. Live photographs of R. namuli sp. nov. in the field (♀, MHNG 1971.067), illustrating pelage colour and features of the (a) head and ears, (b) sella and lips and (c) connecting process" figureDoi="http://doi.org/10.5281/zenodo.7859658" httpUri="https://zenodo.org/record/7859658/files/figure.png" pageId="11" pageNumber="29">Fig. A4</figureCitation>
|
||
). Anterior median and lateral nasal swellings well-developed but posterior median swellings deflated. Moderate rostral depression between supraorbital ridges creates a concave rostral profile. Posterior-most point of rostral depression reaches, but does not extend beyond, constriction point between the orbits (
|
||
<figureCitation box="[1333,1406,1538,1564]" captionStart="FIG" captionStartId="14.[151,164,1031,1052]" captionTargetBox="[196,1383,174,1015]" captionTargetId="figure-411@14.[196,1383,174,1016]" captionTargetPageId="14" captionText="FIG. 6. Cranium in dorsoventral (top), lateral (middle) and ventrodorsal (bottom; only for MHNG 1971.168) views of holotypes of R. namuli sp. nov. (left) and R. maendeleo (right)" figureDoi="http://doi.org/10.5281/zenodo.7859662" httpUri="https://zenodo.org/record/7859662/files/figure.png" pageId="11" pageNumber="29">Fig. 6</figureCitation>
|
||
). The infraorbital foramena are enclosed by a bony bar on both sides in all specimens collected, which is illustrated in close-up in Supplementary Appendix S2 (
|
||
<figureCitation box="[856,950,1675,1701]" captionStart="FIG" captionStartId="11.[151,164,1178,1199]" captionTargetBox="[151,763,174,1162]" captionTargetId="figure-171@11.[151,765,174,1163]" captionTargetPageId="11" captionText="FIG. 4. Live photographs of R. namuli sp. nov. in the field (♀, MHNG 1971.067), illustrating pelage colour and features of the (a) head and ears, (b) sella and lips and (c) connecting process" figureDoi="http://doi.org/10.5281/zenodo.7859658" httpUri="https://zenodo.org/record/7859658/files/figure.png" pageId="11" pageNumber="29">Fig. A4</figureCitation>
|
||
). Note that in specimen DM 10833, initial inspection concluded the bar was absent on the left side (as reported by
|
||
<bibRefCitation author="MONADJEM, A. & M. C. SCHOEMAN & A. RESIDE & D. V. PIO & S. STOFFBERG & J. BAYLISS & F. P. D. COTTERILL & M. CURRAN & M. KOPP & P. J. TAYLOR" box="[1112,1404,1743,1769]" pageId="11" pageNumber="29" pagination="371 - 391" refId="ref16601" refString="MONADJEM, A., M. C. SCHOEMAN, A. RESIDE, D. V. PIO, S. STOFFBERG, J. BAYLISS, F. P. D. COTTERILL, M. CURRAN, M. KOPP, and P. J. TAYLOR. 2010 a. A recent inventory of the bats of Mozambique with documentation of seven new species for the country. Acta Chiropterologica, 12: 371 - 391." type="journal article" year="2010">
|
||
Monadjem
|
||
<emphasis box="[1250,1312,1743,1769]" italics="true" pageId="11" pageNumber="29">et al.</emphasis>
|
||
, 2010
|
||
<emphasis box="[1389,1404,1743,1769]" italics="true" pageId="11" pageNumber="29">a</emphasis>
|
||
</bibRefCitation>
|
||
), but subsequent investigation confirms it presence on both sides (L. Richards, personal communication). Length of the palatal bridge moderate, about one third CM3 length (34.1–37.3%).
|
||
</paragraph>
|
||
<paragraph blockId="11.[809,1422,175,1973]" lastBlockId="12.[151,764,1058,1973]" lastPageId="12" lastPageNumber="30" pageId="11" pageNumber="29">
|
||
<emphasis box="[847,909,1913,1939]" italics="true" pageId="11" pageNumber="29">Teeth</emphasis>
|
||
.
|
||
<emphasis box="[916,942,1913,1939]" italics="true" pageId="11" pageNumber="29">—</emphasis>
|
||
Dental formula I 1/2, C 1/1, PM 2/2, M 3/3. Upper canines moderately well developed, extending about twice the length of PM4 (
|
||
<figureCitation box="[673,749,1058,1084]" captionStart="FIG" captionStartId="14.[151,164,1031,1052]" captionTargetBox="[196,1383,174,1015]" captionTargetId="figure-411@14.[196,1383,174,1016]" captionTargetPageId="14" captionText="FIG. 6. Cranium in dorsoventral (top), lateral (middle) and ventrodorsal (bottom; only for MHNG 1971.168) views of holotypes of R. namuli sp. nov. (left) and R. maendeleo (right)" figureDoi="http://doi.org/10.5281/zenodo.7859662" httpUri="https://zenodo.org/record/7859662/files/figure.png" pageId="12" pageNumber="30">Fig. 6</figureCitation>
|
||
). PM2 small, slightly extruded, separating C1 and PM4 and present on both sides. Minute pm3 present on left side and partly hidden by crown of pm2, lacking on right side, with pm2 and pm4 separate on both sides (
|
||
<figureCitation box="[287,359,1229,1255]" captionStart="FIG" captionStartId="16.[151,164,988,1009]" captionTargetBox="[183,736,182,961]" captionTargetId="figure-96@16.[151,764,173,970]" captionTargetPageId="16" captionText="FIG. 7. Mandibles of the holotype of R. namuli sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7859664" httpUri="https://zenodo.org/record/7859664/files/figure.png" pageId="12" pageNumber="30">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7859660" ID-Zenodo-Dep="7859660" httpUri="https://zenodo.org/record/7859660/files/figure.png" pageId="12" pageNumber="30" startId="12.[151,164,893,914]" targetBox="[361,1218,174,872]" targetPageId="12" targetType="figure">
|
||
<paragraph blockId="12.[151,1421,893,971]" pageId="12" pageNumber="30">
|
||
FIG. 5. Head shape (a and b) and noseleaf (c and d) of holotypes of
|
||
<taxonomicName authority="SMF" authorityName="SMF" box="[843,1033,893,914]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[843,976,893,914]" italics="true" pageId="12" pageNumber="30">R. maendeleo</emphasis>
|
||
SMF
|
||
</taxonomicName>
|
||
79.643 and
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[1159,1255,893,914]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[1159,1255,893,914]" italics="true" pageId="12" pageNumber="30">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1263,1340,893,914]" pageId="12" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
MHNG 1971.068, respectively; connecting process of
|
||
<taxonomicName authority="SMF" authorityName="SMF" box="[608,797,921,942]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[608,741,921,942]" italics="true" pageId="12" pageNumber="30">R. maendeleo</emphasis>
|
||
SMF
|
||
</taxonomicName>
|
||
79.643 (e), SMF 99.760 (f), and of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[1160,1256,921,942]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[1160,1256,921,942]" italics="true" pageId="12" pageNumber="30">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1264,1341,921,942]" pageId="12" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
MHNG
|
||
</paragraph>
|
||
<paragraph blockId="12.[151,1421,893,971]" box="[724,849,950,971]" pageId="12" pageNumber="30">1971.068 (g)</paragraph>
|
||
</caption>
|
||
<paragraph blockId="12.[151,764,1058,1973]" lastBlockId="12.[809,1421,1058,1255]" pageId="12" pageNumber="30">
|
||
<emphasis box="[189,293,1263,1289]" italics="true" pageId="12" pageNumber="30">Baculum</emphasis>
|
||
.—The baculum is medium-sized with a total baculum length of
|
||
<quantity box="[466,574,1297,1323]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" pageId="12" pageNumber="30" unit="mm" value="2.85">2.85 mm</quantity>
|
||
(range ca.
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.875" metricValueMax="2.95" metricValueMin="2.8" pageId="12" pageNumber="30" unit="mm" value="2.875" valueMax="2.95" valueMin="2.8">2.8– 2.95 mm</quantity>
|
||
), greatest lateral width of basal cone of
|
||
<quantity box="[151,260,1366,1392]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="12" pageNumber="30" unit="mm" value="0.78">0.78 mm</quantity>
|
||
and greatest dorsoventral width of basal cone of
|
||
<quantity box="[246,350,1400,1426]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="12" pageNumber="30" unit="mm" value="0.87">0.87 mm</quantity>
|
||
(
|
||
<figureCitation box="[366,436,1400,1426]" captionStart="FIG" captionStartId="17.[151,164,1598,1619]" captionTargetBox="[212,1360,176,1577]" captionTargetId="figure-126@17.[199,1386,173,1579]" captionTargetPageId="17" captionText="FIG. 8. Baculum of the holotype (MHNG 1971.168) of R. namuli sp. nov. in dorsal-ventral (a), ventral-dorsal (c), lateral (b and d) and proximal-distal (g) views. Baculum of the DM 10833 also shown in dorsal-ventral (e) and lateral (f) view. For the holotype (MHNG 1971.168), total baculum length is ca. 2.85 mm, greatest lateral width of basal cone is 0.78 mm and greatest dorsoventral width of basal cone is 0.87 mm" figureDoi="http://doi.org/10.5281/zenodo.7859666" httpUri="https://zenodo.org/record/7859666/files/figure.png" pageId="12" pageNumber="30">Fig. 8</figureCitation>
|
||
and Supplementary Appendix S2
|
||
<figureCitation box="[234,326,1434,1460]" captionStart="FIG" captionStartId="12.[151,164,893,914]" captionTargetBox="[361,1218,174,872]" captionTargetId="figure-451@12.[348,1226,173,879]" captionTargetPageId="12" captionText="FIG. 5. Head shape (a and b) and noseleaf (c and d) of holotypes of R. maendeleo SMF 79.643 and R. namuli sp. nov. MHNG 1971.068, respectively; connecting process of R. maendeleo SMF 79.643 (e), SMF 99.760 (f), and of R. namuli sp. nov. MHNG 1971.068 (g)" figureDoi="http://doi.org/10.5281/zenodo.7859660" httpUri="https://zenodo.org/record/7859660/files/figure.png" pageId="12" pageNumber="30">Fig. A5</figureCitation>
|
||
). The basal cone occupies about 20% of the total length, with dorsal and ventral parts subequal in size, neither exhibiting a relative proximally projection. Both ventral and dorsal incisions of the basal cone are shallow. In lateral view, the baculum shaft constricts gradually, before reaching a constant width at about half the length to the tip. The shaft is not straight, but rather undulating. The tip is evenly rounded and distinctively curves ventrally at an angle of about 30° from the proximaldistal axis. In dorsoventral view, the shaft extends from the basal cone largely straight, unflattened and of equal width almost along its entire length. At the inflection point where the tip begins to curve ventrally (about 4/5th along the total baculum length), there is a shallow constriction in the shaft when viewed from a dorsoventral perspective. The tip thus appears spatulated and rounded (
|
||
<figureCitation box="[1199,1270,1092,1118]" captionStart="FIG" captionStartId="17.[151,164,1598,1619]" captionTargetBox="[212,1360,176,1577]" captionTargetId="figure-126@17.[199,1386,173,1579]" captionTargetPageId="17" captionText="FIG. 8. Baculum of the holotype (MHNG 1971.168) of R. namuli sp. nov. in dorsal-ventral (a), ventral-dorsal (c), lateral (b and d) and proximal-distal (g) views. Baculum of the DM 10833 also shown in dorsal-ventral (e) and lateral (f) view. For the holotype (MHNG 1971.168), total baculum length is ca. 2.85 mm, greatest lateral width of basal cone is 0.78 mm and greatest dorsoventral width of basal cone is 0.87 mm" figureDoi="http://doi.org/10.5281/zenodo.7859666" httpUri="https://zenodo.org/record/7859666/files/figure.png" pageId="12" pageNumber="30">Fig. 8</figureCitation>
|
||
and Supplementary Appendix S2
|
||
<figureCitation box="[1074,1167,1126,1152]" captionStart="FIG" captionStartId="12.[151,164,893,914]" captionTargetBox="[361,1218,174,872]" captionTargetId="figure-451@12.[348,1226,173,879]" captionTargetPageId="12" captionText="FIG. 5. Head shape (a and b) and noseleaf (c and d) of holotypes of R. maendeleo SMF 79.643 and R. namuli sp. nov. MHNG 1971.068, respectively; connecting process of R. maendeleo SMF 79.643 (e), SMF 99.760 (f), and of R. namuli sp. nov. MHNG 1971.068 (g)" figureDoi="http://doi.org/10.5281/zenodo.7859660" httpUri="https://zenodo.org/record/7859660/files/figure.png" pageId="12" pageNumber="30">Fig. A5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="12.[809,1421,1058,1255]" pageId="12" pageNumber="30">
|
||
<emphasis box="[847,1003,1161,1187]" italics="true" pageId="12" pageNumber="30">Echolocation</emphasis>
|
||
.—Calls recorded from seven individuals had a mean peak frequency of 76.9 kHz and a range of 75.9–78.0 kHz (
|
||
<figureCitation box="[1127,1197,1229,1255]" captionStart="FIG" captionStartId="18.[151,164,713,734]" captionTargetBox="[288,1292,174,691]" captionTargetId="figure-537@18.[373,1247,171,639]" captionTargetPageId="18" captionText="FIG. 9. Echolocation call of R. namuli sp. nov. (X, MHNG 1971.067; recorded at Mount Namuli, Zambesia Province, Mozambique, -15.405933 / 37.067006, ca. 1,200 m a.s.l.)" figureDoi="http://doi.org/10.5281/zenodo.7859668" httpUri="https://zenodo.org/record/7859668/files/figure.png" pageId="12" pageNumber="30">Fig. 9</figureCitation>
|
||
and
|
||
<tableCitation box="[1255,1345,1229,1255]" captionStart="TABLE" captionStartId="16.[151,164,1459,1480]" captionTargetPageId="16" captionText="TABLE 5. Echolocation call parameters of R. namuli sp. nov. and species with similar echolocation peak frequency from nearby localities. Table highlights mean peak frequency (PF), standard deviation (SD), range and number of individuals recorded (n). Data collected by M. Curran and M. Kopp (unpublished material). MZ = Mozambique, MW = Malawi" httpUri="http://table.plazi.org/id/2CF56638FFFCFFB5FF08FA30FA0FF9FA" pageId="12" pageNumber="30" tableUuid="2CF56638FFFCFFB5FF08FA30FA0FF9FA">Table 5</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="18" lastPageNumber="36" pageId="12" pageNumber="30" type="discussion">
|
||
<paragraph blockId="12.[809,965,1297,1323]" box="[809,965,1297,1323]" pageId="12" pageNumber="30">
|
||
<heading box="[809,965,1297,1323]" fontSize="36" level="3" pageId="12" pageNumber="30" reason="6">
|
||
<emphasis box="[809,965,1297,1323]" italics="true" pageId="12" pageNumber="30">Comparisons</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="12.[809,1422,1332,1973]" lastBlockId="14.[809,1422,1159,1973]" lastPageId="14" lastPageNumber="32" pageId="12" pageNumber="30">
|
||
<emphasis box="[847,1078,1332,1358]" italics="true" pageId="12" pageNumber="30">Taxonomic position</emphasis>
|
||
.—The connecting process is well-developed, high and rounded, but is neither continuously arched (
|
||
<emphasis box="[1107,1214,1400,1426]" italics="true" pageId="12" pageNumber="30">pearsoni</emphasis>
|
||
-group,
|
||
<emphasis italics="true" pageId="12" pageNumber="30">ferrumequinum</emphasis>
|
||
-group or
|
||
<taxonomicName box="[1066,1186,1434,1460]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="fumigatus">
|
||
<emphasis box="[1066,1186,1434,1460]" italics="true" pageId="12" pageNumber="30">fumigatus</emphasis>
|
||
</taxonomicName>
|
||
-group), low and rounded (
|
||
<taxonomicName baseAuthorityName="Bechstein" baseAuthorityYear="1800" box="[927,1077,1468,1494]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="hipposideros">
|
||
<emphasis box="[927,1077,1468,1494]" italics="true" pageId="12" pageNumber="30">hipposideros</emphasis>
|
||
</taxonomicName>
|
||
-group), low and poorly developed (
|
||
<emphasis box="[930,1039,1503,1529]" italics="true" pageId="12" pageNumber="30">maclaudi</emphasis>
|
||
-group) nor sharply triangular (
|
||
<taxonomicName authorityName="Martin" authorityYear="1832" box="[819,904,1537,1563]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="landeri">
|
||
<emphasis box="[819,904,1537,1563]" italics="true" pageId="12" pageNumber="30">landeri</emphasis>
|
||
</taxonomicName>
|
||
-group) (
|
||
<bibRefCitation author="CSORBA, G. & P. UJHELYI & N. THOMAS" box="[1011,1255,1537,1563]" pageId="12" pageNumber="30" refId="ref15877" refString="CSORBA, G., P. UJHELYI, and N. THOMAS. 2003. Horseshoe bats of the World (Chiroptera: Rhinolophidae), 1 st edition. Alana Books, Shropshire, UK, 160 pp." type="book" year="2003">
|
||
Csorba
|
||
<emphasis box="[1108,1172,1537,1563]" italics="true" pageId="12" pageNumber="30">et al.</emphasis>
|
||
, 2003
|
||
</bibRefCitation>
|
||
). The sella is naked and constricted in the middle rather than hairy (
|
||
<taxonomicName box="[819,935,1605,1631]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="fumigatus">
|
||
<emphasis box="[819,935,1605,1631]" italics="true" pageId="12" pageNumber="30">fumigatus</emphasis>
|
||
</taxonomicName>
|
||
-group) and/or parallel-sided (
|
||
<emphasis box="[1323,1411,1605,1631]" italics="true" pageId="12" pageNumber="30">euryale</emphasis>
|
||
- group). Three mental groves are well developed (Supplementary Appendix S2
|
||
<figureCitation box="[1179,1275,1674,1700]" captionStart="FIG" captionStartId="2.[151,164,1920,1941]" captionTargetBox="[152,1413,198,1885]" captionTargetId="figure-20@2.[151,1423,173,1904]" captionTargetPageId="2" captionText="FIG. 1. Regional overview (A) and zoom in of the study area on Mount Namuli (B), showing specimen collection sites and extant montane forest and woodland cover, overlaid onto a hillshade DEM (SRTM, 90 m resolution)" figureDoi="http://doi.org/10.5281/zenodo.7859652" httpUri="https://zenodo.org/record/7859652/files/figure.png" pageId="12" pageNumber="30">Fig. A1</figureCitation>
|
||
), not partly obliterated like in members of
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[1231,1334,1708,1734]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[1231,1334,1708,1734]" italics="true" pageId="12" pageNumber="30">capensis</emphasis>
|
||
</taxonomicName>
|
||
-group. These features, in addition to the characters described above, indicate
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[1109,1231,1776,1802]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[1109,1231,1776,1802]" italics="true" pageId="12" pageNumber="30">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1246,1345,1776,1802]" pageId="12" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
bears most resemblance morphologically to
|
||
<taxonomicName box="[1255,1414,1810,1836]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[1255,1414,1810,1836]" italics="true" pageId="12" pageNumber="30">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
, which was moved, with
|
||
<emphasis box="[1104,1219,1845,1871]" italics="true" pageId="12" pageNumber="30">
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[1104,1215,1845,1871]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="adami">R. adami</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
from the
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis italics="true" pageId="12" pageNumber="30">capensis</emphasis>
|
||
</taxonomicName>
|
||
-group into a newly created
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[1164,1237,1879,1905]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[1164,1237,1879,1905]" italics="true" pageId="12" pageNumber="30">adami</emphasis>
|
||
</taxonomicName>
|
||
-group by
|
||
<bibRefCitation author="KOCK, D. & G. CSORBA & K. M. HOWELL" pageId="12" pageNumber="30" pagination="233 - 239" refId="ref16521" refString="KOCK, D., G. CSORBA, and K. M. HOWELL. 2000. Rhinolophus maendeleo n. sp. from Tanzania, a horseshoe bat noteworthy for its systematics and biogeography. Senckenbergiana Biologica, 80: 233 - 239." type="journal article" year="2000">
|
||
Kock
|
||
<emphasis box="[809,864,1913,1939]" italics="true" pageId="12" pageNumber="30">et al</emphasis>
|
||
. (2000)
|
||
</bibRefCitation>
|
||
. Regarding the defining characters of the
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[851,924,1947,1973]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[851,924,1947,1973]" italics="true" pageId="12" pageNumber="30">adami</emphasis>
|
||
</taxonomicName>
|
||
-group,
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[1016,1129,1947,1973]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="12" pageNumber="30" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[1016,1129,1947,1973]" italics="true" pageId="12" pageNumber="30">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1135,1225,1947,1973]" pageId="12" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a large sella, straight-sided or rounded lancet, three well-defined mental grooves, small second premolar in the toothrow and more bulbous narial swellings. The nasal swellings are pronounced and clearly align to species of the
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[339,412,1296,1322]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[339,412,1296,1322]" italics="true" pageId="14" pageNumber="32">adami</emphasis>
|
||
</taxonomicName>
|
||
- rather than
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[584,685,1296,1322]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[584,685,1296,1322]" italics="true" pageId="14" pageNumber="32">capensis</emphasis>
|
||
</taxonomicName>
|
||
-group when
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[225,341,1330,1356]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[225,341,1330,1356]" italics="true" pageId="14" pageNumber="32">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[352,447,1330,1356]" pageId="14" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[462,598,1330,1356]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[462,598,1330,1356]" italics="true" pageId="14" pageNumber="32">R. capensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName box="[616,763,1330,1356]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="simulator">
|
||
<emphasis box="[616,763,1330,1356]" italics="true" pageId="14" pageNumber="32">R. simulator</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[213,333,1364,1390]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[213,333,1364,1390]" italics="true" pageId="14" pageNumber="32">R. adami</emphasis>
|
||
</taxonomicName>
|
||
are compared side-by-side (see Supplementary Appendix S2
|
||
<figureCitation box="[495,586,1399,1425]" captionStart="FIG" captionStartId="14.[151,164,1031,1052]" captionTargetBox="[196,1383,174,1015]" captionTargetId="figure-411@14.[196,1383,174,1016]" captionTargetPageId="14" captionText="FIG. 6. Cranium in dorsoventral (top), lateral (middle) and ventrodorsal (bottom; only for MHNG 1971.168) views of holotypes of R. namuli sp. nov. (left) and R. maendeleo (right)" figureDoi="http://doi.org/10.5281/zenodo.7859662" httpUri="https://zenodo.org/record/7859662/files/figure.png" pageId="14" pageNumber="32">Fig. A6</figureCitation>
|
||
). However, the ear length of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[320,438,1433,1459]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[320,438,1433,1459]" italics="true" pageId="14" pageNumber="32">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[450,545,1433,1459]" pageId="14" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is medium rather than large, only three of the
|
||
<specimenCount box="[479,654,1467,1493]" pageId="14" pageNumber="32" type="generic">five specimens</specimenCount>
|
||
of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis italics="true" pageId="14" pageNumber="32">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[218,316,1501,1527]" pageId="14" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
have a narrow skull, and all have a moderate palate (36% of CM
|
||
<quantity box="[526,571,1536,1562]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="14" pageNumber="32" unit="in" value="3.0">3 in</quantity>
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[580,661,1536,1562]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[580,661,1536,1562]" italics="true" pageId="14" pageNumber="32">namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[671,764,1536,1562]" pageId="14" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
against> 40% in
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[361,434,1570,1596]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[361,434,1570,1596]" italics="true" pageId="14" pageNumber="32">adami</emphasis>
|
||
</taxonomicName>
|
||
-group). Furthermore, when viewed in profile,
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[380,461,1604,1630]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[380,461,1604,1630]" italics="true" pageId="14" pageNumber="32">namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[474,569,1604,1630]" pageId="14" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a high and rounded connecting process, which is lower and regularly arched in the
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[416,489,1673,1699]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[416,489,1673,1699]" italics="true" pageId="14" pageNumber="32">adami</emphasis>
|
||
</taxonomicName>
|
||
-group (
|
||
<bibRefCitation author="CSORBA, G. & P. UJHELYI & N. THOMAS" pageId="14" pageNumber="32" refId="ref15877" refString="CSORBA, G., P. UJHELYI, and N. THOMAS. 2003. Horseshoe bats of the World (Chiroptera: Rhinolophidae), 1 st edition. Alana Books, Shropshire, UK, 160 pp." type="book" year="2003">
|
||
Csorba
|
||
<emphasis box="[690,756,1673,1699]" italics="true" pageId="14" pageNumber="32">et al.</emphasis>
|
||
, 2003
|
||
</bibRefCitation>
|
||
). Morphologically,
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[454,565,1707,1733]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[454,565,1707,1733]" italics="true" pageId="14" pageNumber="32">R namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[578,673,1707,1733]" pageId="14" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
should therefore be retained within the
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[524,625,1741,1767]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[524,625,1741,1767]" italics="true" pageId="14" pageNumber="32">capensis</emphasis>
|
||
</taxonomicName>
|
||
-group. Our genetic analysis confirms this, with a robust positioning of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[280,389,1810,1836]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[280,389,1810,1836]" italics="true" pageId="14" pageNumber="32">R namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[400,494,1810,1836]" pageId="14" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in the
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[584,685,1810,1836]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[584,685,1810,1836]" italics="true" pageId="14" pageNumber="32">capensis</emphasis>
|
||
</taxonomicName>
|
||
-group alongside
|
||
<taxonomicName box="[275,423,1844,1870]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="simulator">
|
||
<emphasis box="[275,423,1844,1870]" italics="true" pageId="14" pageNumber="32">R. simulator</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Gough" authorityYear="1908" box="[442,569,1844,1870]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="swinnyi">
|
||
<emphasis box="[442,569,1844,1870]" italics="true" pageId="14" pageNumber="32">R. swinnyi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Thomas" authorityYear="1904" box="[633,728,1844,1870]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="denti">
|
||
<emphasis box="[633,728,1844,1870]" italics="true" pageId="14" pageNumber="32">R. denti</emphasis>
|
||
</taxonomicName>
|
||
as closest relatives (
|
||
<figureCitation box="[363,439,1879,1905]" captionStart="FIG" captionStartId="9.[151,164,1836,1857]" captionTargetBox="[193,1372,177,1809]" captionTargetId="graphics-12@9.[194,1370,177,1776]" captionTargetPageId="9" captionText="FIG. 3. Maximum likelihood tree based on CYT-B sequences and reconstructed with IQ-tree. Sequences of R. hipposideros were used as outgroups. Bootstrap support levels are explained in the box; the scale indicates nucleotide substitutions per site. Newly sequenced taxa are associated with their GenBank accession numbers (see Table 2 and Supplementary Appendix S3 for more details), while the species groups to which they belong appear to the right of the tree. The clade representing the new species (sp. nov.) is highlighted in grey" figureDoi="http://doi.org/10.5281/zenodo.7859656" httpUri="https://zenodo.org/record/7859656/files/figure.png" pageId="14" pageNumber="32">Fig. 3</figureCitation>
|
||
). Unfortunately, as no genetic sequences for
|
||
<taxonomicName box="[395,559,1913,1939]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[395,559,1913,1939]" italics="true" pageId="14" pageNumber="32">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[606,716,1913,1939]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[606,716,1913,1939]" italics="true" pageId="14" pageNumber="32">R. adami</emphasis>
|
||
</taxonomicName>
|
||
are currently available, we cannot assess formally their phylogenetic position within the
|
||
<taxonomicName authorityName="Gray" authorityYear="1825" box="[1206,1379,1159,1185]" class="Mammalia" family="Rhinolophidae" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="family">Rhinolophidae</taxonomicName>
|
||
radiation. We therefore leave open the question of the validity of the
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[980,1053,1227,1253]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[980,1053,1227,1253]" italics="true" pageId="14" pageNumber="32">adami</emphasis>
|
||
</taxonomicName>
|
||
-group and whether the striking morphological similarities shared between
|
||
<taxonomicName class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis italics="true" pageId="14" pageNumber="32">maendeleo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[929,1010,1296,1322]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[929,1010,1296,1322]" italics="true" pageId="14" pageNumber="32">namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1018,1109,1296,1322]" pageId="14" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
represent convergences or true synapomorphies.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="2CF56638FFE1FFA8FFFCFDBDF895FD76" httpUri="http://table.plazi.org/id/2CF56638FFE1FFA8FFFCFDBDF895FD76" pageId="13" pageNumber="31" startId="13.[99,112,574,595]" targetBox="[99,1857,684,1570]" targetIsTable="true" targetPageId="13" targetType="table">
|
||
<paragraph blockId="13.[99,1899,574,652]" pageId="13" pageNumber="31">
|
||
TABLE 3. External measurements of specimens of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[588,684,574,595]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="31" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[588,684,574,595]" italics="true" pageId="13" pageNumber="31">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[691,769,574,595]" pageId="13" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName box="[780,912,574,595]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="31" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[780,912,574,595]" italics="true" pageId="13" pageNumber="31">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[962,1082,574,595]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="31" phylum="Chordata" rank="species" species="simulator">
|
||
<emphasis box="[962,1082,574,595]" italics="true" pageId="13" pageNumber="31">R. simulator</emphasis>
|
||
</taxonomicName>
|
||
. Measurement with an asterisk in the header have non-overlapping ranges between
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[99,196,603,624]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="31" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[99,196,603,624]" italics="true" pageId="13" pageNumber="31">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[205,283,603,624]" pageId="13" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName box="[335,469,603,624]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="31" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[335,469,603,624]" italics="true" pageId="13" pageNumber="31">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="13" pageNumber="31">
|
||
<table box="[99,1857,684,1570]" gridcols="19" gridrows="31" pageId="13" pageNumber="31" rowsContinueIn="15.[99,1878,684,1570]">
|
||
<tr box="[99,1857,684,705]" gridrow="0" pageId="13" pageNumber="31">
|
||
<th box="[99,384,684,705]" colspan="3" colspanRight="2" gridcol="0" gridrow="0" pageId="13" pageNumber="31">Collection no. Sex BM</th>
|
||
<th box="[427,481,684,705]" gridcol="3" gridrow="0" pageId="13" pageNumber="31">HB</th>
|
||
<th box="[521,575,684,705]" gridcol="4" gridrow="0" pageId="13" pageNumber="31">Tail*</th>
|
||
<th box="[614,657,684,705]" gridcol="5" gridrow="0" pageId="13" pageNumber="31">HS</th>
|
||
<th box="[695,749,684,705]" gridcol="6" gridrow="0" pageId="13" pageNumber="31">Ear*</th>
|
||
<th box="[790,843,684,705]" gridcol="7" gridrow="0" pageId="13" pageNumber="31">FA</th>
|
||
<th box="[884,938,684,705]" gridcol="8" gridrow="0" pageId="13" pageNumber="31">Tib</th>
|
||
<th box="[977,1019,684,705]" gridcol="9" gridrow="0" pageId="13" pageNumber="31">HF</th>
|
||
<th box="[1054,1117,684,705]" gridcol="10" gridrow="0" pageId="13" pageNumber="31">3Met*</th>
|
||
<th box="[1152,1207,684,705]" gridcol="11" gridrow="0" pageId="13" pageNumber="31">3Ph1</th>
|
||
<th box="[1243,1305,684,705]" gridcol="12" gridrow="0" pageId="13" pageNumber="31">3Ph2*</th>
|
||
<th box="[1341,1396,684,705]" gridcol="13" gridrow="0" pageId="13" pageNumber="31">4Met</th>
|
||
<th box="[1425,1486,684,705]" gridcol="14" gridrow="0" pageId="13" pageNumber="31">4Ph1*</th>
|
||
<th box="[1512,1573,684,705]" gridcol="15" gridrow="0" pageId="13" pageNumber="31">4Ph2*</th>
|
||
<th box="[1608,1663,684,705]" gridcol="16" gridrow="0" pageId="13" pageNumber="31">5Met</th>
|
||
<th box="[1704,1758,684,705]" gridcol="17" gridrow="0" pageId="13" pageNumber="31">5Ph1</th>
|
||
<th box="[1803,1857,684,705]" gridcol="18" gridrow="0" pageId="13" pageNumber="31">5Ph2</th>
|
||
</tr>
|
||
<tr box="[99,1857,720,741]" gridrow="1" pageId="13" pageNumber="31">
|
||
<th box="[99,1857,720,741]" colspan="19" colspanRight="18" gridcol="0" gridrow="1" pageId="13" pageNumber="31">
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[99,193,720,741]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="13" pageNumber="31" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[99,193,720,741]" italics="true" pageId="13" pageNumber="31">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[199,275,720,741]" pageId="13" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
</tr>
|
||
<tr box="[99,1857,748,770]" gridrow="2" pageId="13" pageNumber="31">
|
||
<th box="[99,282,748,770]" gridcol="0" gridrow="2" pageId="13" pageNumber="31">MHNG 1971.067</th>
|
||
<td box="[300,320,748,770]" gridcol="1" gridrow="2" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,748,770]" pageId="13" pageNumber="31">X</emphasis>
|
||
</td>
|
||
<td box="[342,384,748,770]" gridcol="2" gridrow="2" pageId="13" pageNumber="31">10.0</td>
|
||
<td box="[427,481,748,770]" gridcol="3" gridrow="2" pageId="13" pageNumber="31">49.2</td>
|
||
<td box="[521,575,748,770]" gridcol="4" gridrow="2" pageId="13" pageNumber="31">19.8</td>
|
||
<td box="[614,657,748,770]" gridcol="5" gridrow="2" pageId="13" pageNumber="31">8.9</td>
|
||
<td box="[695,749,748,770]" gridcol="6" gridrow="2" pageId="13" pageNumber="31">19.6</td>
|
||
<td box="[790,843,748,770]" gridcol="7" gridrow="2" pageId="13" pageNumber="31">48.3</td>
|
||
<td box="[884,938,748,770]" gridcol="8" gridrow="2" pageId="13" pageNumber="31">19.0</td>
|
||
<td box="[977,1019,748,770]" gridcol="9" gridrow="2" pageId="13" pageNumber="31">8.9</td>
|
||
<td box="[1054,1117,748,770]" gridcol="10" gridrow="2" pageId="13" pageNumber="31">30.7</td>
|
||
<td box="[1152,1207,748,770]" gridcol="11" gridrow="2" pageId="13" pageNumber="31">15.1</td>
|
||
<td box="[1243,1305,748,770]" gridcol="12" gridrow="2" pageId="13" pageNumber="31">26.3</td>
|
||
<td box="[1341,1396,748,770]" gridcol="13" gridrow="2" pageId="13" pageNumber="31">34.4</td>
|
||
<td box="[1425,1486,748,770]" gridcol="14" gridrow="2" pageId="13" pageNumber="31">7.5</td>
|
||
<td box="[1512,1573,748,770]" gridcol="15" gridrow="2" pageId="13" pageNumber="31">16.5</td>
|
||
<td box="[1608,1663,748,770]" gridcol="16" gridrow="2" pageId="13" pageNumber="31">33.7</td>
|
||
<td box="[1704,1758,748,770]" gridcol="17" gridrow="2" pageId="13" pageNumber="31">11.3</td>
|
||
<td box="[1803,1857,748,770]" gridcol="18" gridrow="2" pageId="13" pageNumber="31">13.3</td>
|
||
</tr>
|
||
<tr box="[99,1857,776,797]" gridrow="3" pageId="13" pageNumber="31" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[99,282,776,797]" gridcol="0" gridrow="3" pageId="13" pageNumber="31">(Paratype)</th>
|
||
</tr>
|
||
<tr box="[99,1857,804,826]" gridrow="4" pageId="13" pageNumber="31">
|
||
<th box="[99,282,804,826]" gridcol="0" gridrow="4" pageId="13" pageNumber="31">MHNG 1971.068</th>
|
||
<td box="[300,320,804,826]" gridcol="1" gridrow="4" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,804,826]" pageId="13" pageNumber="31">Y</emphasis>
|
||
</td>
|
||
<td box="[342,384,804,826]" gridcol="2" gridrow="4" pageId="13" pageNumber="31">9.5</td>
|
||
<td box="[427,481,804,826]" gridcol="3" gridrow="4" pageId="13" pageNumber="31">48.2</td>
|
||
<td box="[521,575,804,826]" gridcol="4" gridrow="4" pageId="13" pageNumber="31">23.6</td>
|
||
<td box="[614,657,804,826]" gridcol="5" gridrow="4" pageId="13" pageNumber="31">9.3</td>
|
||
<td box="[695,749,804,826]" gridcol="6" gridrow="4" pageId="13" pageNumber="31">19.5</td>
|
||
<td box="[790,843,804,826]" gridcol="7" gridrow="4" pageId="13" pageNumber="31">49.5</td>
|
||
<td box="[884,938,804,826]" gridcol="8" gridrow="4" pageId="13" pageNumber="31">19.3</td>
|
||
<td box="[977,1019,804,826]" gridcol="9" gridrow="4" pageId="13" pageNumber="31">8.1</td>
|
||
<td box="[1054,1117,804,826]" gridcol="10" gridrow="4" pageId="13" pageNumber="31">31.5</td>
|
||
<td box="[1152,1207,804,826]" gridcol="11" gridrow="4" pageId="13" pageNumber="31">15.6</td>
|
||
<td box="[1243,1305,804,826]" gridcol="12" gridrow="4" pageId="13" pageNumber="31">26.3</td>
|
||
<td box="[1341,1396,804,826]" gridcol="13" gridrow="4" pageId="13" pageNumber="31">35.4</td>
|
||
<td box="[1425,1486,804,826]" gridcol="14" gridrow="4" pageId="13" pageNumber="31">8.0</td>
|
||
<td box="[1512,1573,804,826]" gridcol="15" gridrow="4" pageId="13" pageNumber="31">16.3</td>
|
||
<td box="[1608,1663,804,826]" gridcol="16" gridrow="4" pageId="13" pageNumber="31">34.7</td>
|
||
<td box="[1704,1758,804,826]" gridcol="17" gridrow="4" pageId="13" pageNumber="31">11.3</td>
|
||
<td box="[1803,1857,804,826]" gridcol="18" gridrow="4" pageId="13" pageNumber="31">12.5</td>
|
||
</tr>
|
||
<tr box="[99,1857,833,854]" gridrow="5" pageId="13" pageNumber="31">
|
||
<th box="[99,1857,833,854]" colspan="19" colspanRight="18" gridcol="0" gridrow="5" pageId="13" pageNumber="31">(Holotype)</th>
|
||
</tr>
|
||
<tr box="[99,1857,861,883]" gridrow="6" pageId="13" pageNumber="31">
|
||
<th box="[99,282,861,883]" gridcol="0" gridrow="6" pageId="13" pageNumber="31">MHNG 1971.069</th>
|
||
<td box="[300,320,861,883]" gridcol="1" gridrow="6" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,861,883]" pageId="13" pageNumber="31">Y</emphasis>
|
||
</td>
|
||
<td box="[342,384,861,883]" gridcol="2" gridrow="6" pageId="13" pageNumber="31">9.0</td>
|
||
<td box="[427,481,861,883]" gridcol="3" gridrow="6" pageId="13" pageNumber="31">49.1</td>
|
||
<td box="[521,575,861,883]" gridcol="4" gridrow="6" pageId="13" pageNumber="31">19.0</td>
|
||
<td box="[614,657,861,883]" gridcol="5" gridrow="6" pageId="13" pageNumber="31">9.3</td>
|
||
<td box="[695,749,861,883]" gridcol="6" gridrow="6" pageId="13" pageNumber="31">20.1</td>
|
||
<td box="[790,843,861,883]" gridcol="7" gridrow="6" pageId="13" pageNumber="31">47.3</td>
|
||
<td box="[884,938,861,883]" gridcol="8" gridrow="6" pageId="13" pageNumber="31">18.9</td>
|
||
<td box="[977,1019,861,883]" gridcol="9" gridrow="6" pageId="13" pageNumber="31">8.4</td>
|
||
<td box="[1054,1117,861,883]" gridcol="10" gridrow="6" pageId="13" pageNumber="31">29.8</td>
|
||
<td box="[1152,1207,861,883]" gridcol="11" gridrow="6" pageId="13" pageNumber="31">14.9</td>
|
||
<td box="[1243,1305,861,883]" gridcol="12" gridrow="6" pageId="13" pageNumber="31">26.3</td>
|
||
<td box="[1341,1396,861,883]" gridcol="13" gridrow="6" pageId="13" pageNumber="31">33.6</td>
|
||
<td box="[1425,1486,861,883]" gridcol="14" gridrow="6" pageId="13" pageNumber="31">7.3</td>
|
||
<td box="[1512,1573,861,883]" gridcol="15" gridrow="6" pageId="13" pageNumber="31">15.4</td>
|
||
<td box="[1608,1663,861,883]" gridcol="16" gridrow="6" pageId="13" pageNumber="31">31.9</td>
|
||
<td box="[1704,1758,861,883]" gridcol="17" gridrow="6" pageId="13" pageNumber="31">10.5</td>
|
||
<td box="[1803,1857,861,883]" gridcol="18" gridrow="6" pageId="13" pageNumber="31">12.3</td>
|
||
</tr>
|
||
<tr box="[99,1857,890,911]" gridrow="7" pageId="13" pageNumber="31" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[99,282,890,911]" gridcol="0" gridrow="7" pageId="13" pageNumber="31">(Paratype)</th>
|
||
</tr>
|
||
<tr box="[99,1857,917,939]" gridrow="8" pageId="13" pageNumber="31">
|
||
<th box="[99,282,917,939]" gridcol="0" gridrow="8" pageId="13" pageNumber="31">DM 10833</th>
|
||
<td box="[300,320,917,939]" gridcol="1" gridrow="8" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,917,939]" pageId="13" pageNumber="31">Y</emphasis>
|
||
</td>
|
||
<td box="[342,384,917,939]" gridcol="2" gridrow="8" pageId="13" pageNumber="31">10.0</td>
|
||
<td box="[427,481,917,939]" gridcol="3" gridrow="8" pageId="13" pageNumber="31">46.3</td>
|
||
<td box="[521,575,917,939]" gridcol="4" gridrow="8" pageId="13" pageNumber="31">18.9</td>
|
||
<td box="[614,657,917,939]" gridcol="5" gridrow="8" pageId="13" pageNumber="31">8.7</td>
|
||
<td box="[695,749,917,939]" gridcol="6" gridrow="8" pageId="13" pageNumber="31">19.7</td>
|
||
<td box="[790,843,917,939]" gridcol="7" gridrow="8" pageId="13" pageNumber="31">47.5</td>
|
||
<td box="[884,938,917,939]" gridcol="8" gridrow="8" pageId="13" pageNumber="31">19.2</td>
|
||
<td box="[977,1019,917,939]" gridcol="9" gridrow="8" pageId="13" pageNumber="31">–</td>
|
||
<td box="[1054,1117,917,939]" gridcol="10" gridrow="8" pageId="13" pageNumber="31">–</td>
|
||
<td box="[1152,1207,917,939]" gridcol="11" gridrow="8" pageId="13" pageNumber="31">–</td>
|
||
<td box="[1243,1305,917,939]" gridcol="12" gridrow="8" pageId="13" pageNumber="31">–</td>
|
||
<td box="[1341,1396,917,939]" gridcol="13" gridrow="8" pageId="13" pageNumber="31">–</td>
|
||
<td box="[1425,1486,917,939]" gridcol="14" gridrow="8" pageId="13" pageNumber="31">–</td>
|
||
<td box="[1512,1573,917,939]" gridcol="15" gridrow="8" pageId="13" pageNumber="31">–</td>
|
||
<td box="[1608,1663,917,939]" gridcol="16" gridrow="8" pageId="13" pageNumber="31">–</td>
|
||
<td box="[1704,1758,917,939]" gridcol="17" gridrow="8" pageId="13" pageNumber="31">–</td>
|
||
<td box="[1803,1857,917,939]" gridcol="18" gridrow="8" pageId="13" pageNumber="31">–</td>
|
||
</tr>
|
||
<tr box="[99,1857,946,968]" gridrow="9" pageId="13" pageNumber="31">
|
||
<th box="[99,282,946,968]" gridcol="0" gridrow="9" pageId="13" pageNumber="31">DM 10839</th>
|
||
<td box="[300,320,946,968]" gridcol="1" gridrow="9" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,946,968]" pageId="13" pageNumber="31">X</emphasis>
|
||
</td>
|
||
<td box="[342,384,946,968]" gridcol="2" gridrow="9" pageId="13" pageNumber="31">–</td>
|
||
<td box="[427,481,946,968]" gridcol="3" gridrow="9" pageId="13" pageNumber="31">–</td>
|
||
<td box="[521,575,946,968]" gridcol="4" gridrow="9" pageId="13" pageNumber="31">–</td>
|
||
<td box="[614,657,946,968]" gridcol="5" gridrow="9" pageId="13" pageNumber="31">8.4</td>
|
||
<td box="[695,749,946,968]" gridcol="6" gridrow="9" pageId="13" pageNumber="31">19.6</td>
|
||
<td box="[790,843,946,968]" gridcol="7" gridrow="9" pageId="13" pageNumber="31">48.9</td>
|
||
<td box="[884,938,946,968]" gridcol="8" gridrow="9" pageId="13" pageNumber="31">19.8</td>
|
||
<td box="[977,1019,946,968]" gridcol="9" gridrow="9" pageId="13" pageNumber="31">9.0</td>
|
||
<td box="[1054,1117,946,968]" gridcol="10" gridrow="9" pageId="13" pageNumber="31">31.0</td>
|
||
<td box="[1152,1207,946,968]" gridcol="11" gridrow="9" pageId="13" pageNumber="31">15.3</td>
|
||
<td box="[1243,1305,946,968]" gridcol="12" gridrow="9" pageId="13" pageNumber="31">26.5</td>
|
||
<td box="[1341,1396,946,968]" gridcol="13" gridrow="9" pageId="13" pageNumber="31">34.8</td>
|
||
<td box="[1425,1486,946,968]" gridcol="14" gridrow="9" pageId="13" pageNumber="31">7.6</td>
|
||
<td box="[1512,1573,946,968]" gridcol="15" gridrow="9" pageId="13" pageNumber="31">16.9</td>
|
||
<td box="[1608,1663,946,968]" gridcol="16" gridrow="9" pageId="13" pageNumber="31">35.5</td>
|
||
<td box="[1704,1758,946,968]" gridcol="17" gridrow="9" pageId="13" pageNumber="31">11.5</td>
|
||
<td box="[1803,1857,946,968]" gridcol="18" gridrow="9" pageId="13" pageNumber="31">13.3</td>
|
||
</tr>
|
||
<tr box="[99,1857,974,995]" gridrow="10" pageId="13" pageNumber="31">
|
||
<th box="[99,282,974,995]" gridcol="0" gridrow="10" pageId="13" pageNumber="31">Mean</th>
|
||
<td box="[300,320,974,995]" gridcol="1" gridrow="10" pageId="13" pageNumber="31">–</td>
|
||
<td box="[342,384,974,995]" gridcol="2" gridrow="10" pageId="13" pageNumber="31">9.6</td>
|
||
<td box="[427,481,974,995]" gridcol="3" gridrow="10" pageId="13" pageNumber="31">48.2</td>
|
||
<td box="[521,575,974,995]" gridcol="4" gridrow="10" pageId="13" pageNumber="31">20.3</td>
|
||
<td box="[614,657,974,995]" gridcol="5" gridrow="10" pageId="13" pageNumber="31">8.9</td>
|
||
<td box="[695,749,974,995]" gridcol="6" gridrow="10" pageId="13" pageNumber="31">19.7</td>
|
||
<td box="[790,843,974,995]" gridcol="7" gridrow="10" pageId="13" pageNumber="31">48.3</td>
|
||
<td box="[884,938,974,995]" gridcol="8" gridrow="10" pageId="13" pageNumber="31">19.2</td>
|
||
<td box="[977,1019,974,995]" gridcol="9" gridrow="10" pageId="13" pageNumber="31">8.6</td>
|
||
<td box="[1054,1117,974,995]" gridcol="10" gridrow="10" pageId="13" pageNumber="31">30.7</td>
|
||
<td box="[1152,1207,974,995]" gridcol="11" gridrow="10" pageId="13" pageNumber="31">15.2</td>
|
||
<td box="[1243,1305,974,995]" gridcol="12" gridrow="10" pageId="13" pageNumber="31">26.3</td>
|
||
<td box="[1341,1396,974,995]" gridcol="13" gridrow="10" pageId="13" pageNumber="31">34.5</td>
|
||
<td box="[1425,1486,974,995]" gridcol="14" gridrow="10" pageId="13" pageNumber="31">7.6</td>
|
||
<td box="[1512,1573,974,995]" gridcol="15" gridrow="10" pageId="13" pageNumber="31">16.3</td>
|
||
<td box="[1608,1663,974,995]" gridcol="16" gridrow="10" pageId="13" pageNumber="31">33.9</td>
|
||
<td box="[1704,1758,974,995]" gridcol="17" gridrow="10" pageId="13" pageNumber="31">11.1</td>
|
||
<td box="[1803,1857,974,995]" gridcol="18" gridrow="10" pageId="13" pageNumber="31">12.9</td>
|
||
</tr>
|
||
<tr box="[99,1857,1003,1024]" gridrow="11" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1003,1024]" gridcol="0" gridrow="11" pageId="13" pageNumber="31">Range</th>
|
||
<td box="[300,320,1003,1024]" gridcol="1" gridrow="11" pageId="13" pageNumber="31">–</td>
|
||
<td box="[342,384,1003,1024]" gridcol="2" gridrow="11" pageId="13" pageNumber="31">9.0–</td>
|
||
<td box="[427,481,1003,1024]" gridcol="3" gridrow="11" pageId="13" pageNumber="31">46.3–</td>
|
||
<td box="[521,575,1003,1024]" gridcol="4" gridrow="11" pageId="13" pageNumber="31">18.9–</td>
|
||
<td box="[614,657,1003,1024]" gridcol="5" gridrow="11" pageId="13" pageNumber="31">8.4–</td>
|
||
<td box="[695,749,1003,1024]" gridcol="6" gridrow="11" pageId="13" pageNumber="31">19.5–</td>
|
||
<td box="[790,843,1003,1024]" gridcol="7" gridrow="11" pageId="13" pageNumber="31">47.3–</td>
|
||
<td box="[884,938,1003,1024]" gridcol="8" gridrow="11" pageId="13" pageNumber="31">18.9–</td>
|
||
<td box="[977,1019,1003,1024]" gridcol="9" gridrow="11" pageId="13" pageNumber="31">8.1–</td>
|
||
<td box="[1054,1117,1003,1024]" gridcol="10" gridrow="11" pageId="13" pageNumber="31">29.8–</td>
|
||
<td box="[1152,1207,1003,1024]" gridcol="11" gridrow="11" pageId="13" pageNumber="31">14.9–</td>
|
||
<td box="[1243,1305,1003,1024]" gridcol="12" gridrow="11" pageId="13" pageNumber="31">26.3–</td>
|
||
<td box="[1341,1396,1003,1024]" gridcol="13" gridrow="11" pageId="13" pageNumber="31">34.4–</td>
|
||
<td box="[1425,1486,1003,1024]" gridcol="14" gridrow="11" pageId="13" pageNumber="31">7.3–</td>
|
||
<td box="[1512,1573,1003,1024]" gridcol="15" gridrow="11" pageId="13" pageNumber="31">16.3–</td>
|
||
<td box="[1608,1663,1003,1024]" gridcol="16" gridrow="11" pageId="13" pageNumber="31">31.9–</td>
|
||
<td box="[1704,1758,1003,1024]" gridcol="17" gridrow="11" pageId="13" pageNumber="31">10.5–</td>
|
||
<td box="[1803,1857,1003,1024]" gridcol="18" gridrow="11" pageId="13" pageNumber="31">12.3–</td>
|
||
</tr>
|
||
<tr box="[99,1857,1031,1052]" gridrow="12" pageId="13" pageNumber="31" rowspan-0="1" rowspan-1="1">
|
||
<td box="[342,384,1031,1052]" gridcol="2" gridrow="12" pageId="13" pageNumber="31">10.0</td>
|
||
<td box="[427,481,1031,1052]" gridcol="3" gridrow="12" pageId="13" pageNumber="31">49.2</td>
|
||
<td box="[521,575,1031,1052]" gridcol="4" gridrow="12" pageId="13" pageNumber="31">23.6</td>
|
||
<td box="[614,657,1031,1052]" gridcol="5" gridrow="12" pageId="13" pageNumber="31">9.3</td>
|
||
<td box="[695,749,1031,1052]" gridcol="6" gridrow="12" pageId="13" pageNumber="31">20.1</td>
|
||
<td box="[790,843,1031,1052]" gridcol="7" gridrow="12" pageId="13" pageNumber="31">49.5</td>
|
||
<td box="[884,938,1031,1052]" gridcol="8" gridrow="12" pageId="13" pageNumber="31">19.8</td>
|
||
<td box="[977,1019,1031,1052]" gridcol="9" gridrow="12" pageId="13" pageNumber="31">9.0</td>
|
||
<td box="[1054,1117,1031,1052]" gridcol="10" gridrow="12" pageId="13" pageNumber="31">31.5</td>
|
||
<td box="[1152,1207,1031,1052]" gridcol="11" gridrow="12" pageId="13" pageNumber="31">15.6</td>
|
||
<td box="[1243,1305,1031,1052]" gridcol="12" gridrow="12" pageId="13" pageNumber="31">26.5</td>
|
||
<td box="[1341,1396,1031,1052]" gridcol="13" gridrow="12" pageId="13" pageNumber="31">35.4</td>
|
||
<td box="[1425,1486,1031,1052]" gridcol="14" gridrow="12" pageId="13" pageNumber="31">8.0</td>
|
||
<td box="[1512,1573,1031,1052]" gridcol="15" gridrow="12" pageId="13" pageNumber="31">16.9</td>
|
||
<td box="[1608,1663,1031,1052]" gridcol="16" gridrow="12" pageId="13" pageNumber="31">35.5</td>
|
||
<td box="[1704,1758,1031,1052]" gridcol="17" gridrow="12" pageId="13" pageNumber="31">11.5</td>
|
||
<td box="[1803,1857,1031,1052]" gridcol="18" gridrow="12" pageId="13" pageNumber="31">13.3</td>
|
||
</tr>
|
||
<tr box="[99,1857,1064,1085]" gridrow="13" pageId="13" pageNumber="31" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[99,282,1064,1085]" gridcol="0" gridrow="13" pageId="13" pageNumber="31">
|
||
<emphasis box="[99,230,1064,1085]" italics="true" pageId="13" pageNumber="31">R. maendeleo</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[99,1857,1091,1113]" gridrow="14" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1091,1113]" gridcol="0" gridrow="14" pageId="13" pageNumber="31">SMF 79.643</th>
|
||
<td box="[300,320,1091,1113]" gridcol="1" gridrow="14" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,1091,1113]" pageId="13" pageNumber="31">Y</emphasis>
|
||
</td>
|
||
<td box="[342,384,1091,1113]" gridcol="2" gridrow="14" pageId="13" pageNumber="31">–</td>
|
||
<td box="[427,481,1091,1113]" gridcol="3" gridrow="14" pageId="13" pageNumber="31">–</td>
|
||
<td box="[521,575,1091,1113]" gridcol="4" gridrow="14" pageId="13" pageNumber="31">26.2</td>
|
||
<td box="[614,657,1091,1113]" gridcol="5" gridrow="14" pageId="13" pageNumber="31">8.8</td>
|
||
<td box="[695,749,1091,1113]" gridcol="6" gridrow="14" pageId="13" pageNumber="31">24.2</td>
|
||
<td box="[790,843,1091,1113]" gridcol="7" gridrow="14" pageId="13" pageNumber="31">47.6</td>
|
||
<td box="[884,938,1091,1113]" gridcol="8" gridrow="14" pageId="13" pageNumber="31">19.6</td>
|
||
<td box="[977,1019,1091,1113]" gridcol="9" gridrow="14" pageId="13" pageNumber="31">8.4</td>
|
||
<td box="[1054,1117,1091,1113]" gridcol="10" gridrow="14" pageId="13" pageNumber="31">32.5</td>
|
||
<td box="[1152,1207,1091,1113]" gridcol="11" gridrow="14" pageId="13" pageNumber="31">14.6</td>
|
||
<td box="[1243,1305,1091,1113]" gridcol="12" gridrow="14" pageId="13" pageNumber="31">24.2</td>
|
||
<td box="[1341,1396,1091,1113]" gridcol="13" gridrow="14" pageId="13" pageNumber="31">35.1</td>
|
||
<td box="[1425,1486,1091,1113]" gridcol="14" gridrow="14" pageId="13" pageNumber="31">8.1</td>
|
||
<td box="[1512,1573,1091,1113]" gridcol="15" gridrow="14" pageId="13" pageNumber="31">14.5</td>
|
||
<td box="[1608,1663,1091,1113]" gridcol="16" gridrow="14" pageId="13" pageNumber="31">34.5</td>
|
||
<td box="[1704,1758,1091,1113]" gridcol="17" gridrow="14" pageId="13" pageNumber="31">10.4</td>
|
||
<td box="[1803,1857,1091,1113]" gridcol="18" gridrow="14" pageId="13" pageNumber="31">13.0</td>
|
||
</tr>
|
||
<tr box="[99,1857,1120,1141]" gridrow="15" pageId="13" pageNumber="31">
|
||
<th box="[99,1857,1120,1141]" colspan="19" colspanRight="18" gridcol="0" gridrow="15" pageId="13" pageNumber="31">(Holotype)</th>
|
||
</tr>
|
||
<tr box="[99,1857,1148,1170]" gridrow="16" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1148,1170]" gridcol="0" gridrow="16" pageId="13" pageNumber="31">SMF 66.960</th>
|
||
<td box="[300,320,1148,1170]" gridcol="1" gridrow="16" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,1148,1170]" pageId="13" pageNumber="31">X</emphasis>
|
||
</td>
|
||
<td box="[342,384,1148,1170]" gridcol="2" gridrow="16" pageId="13" pageNumber="31">–</td>
|
||
<td box="[427,481,1148,1170]" gridcol="3" gridrow="16" pageId="13" pageNumber="31">–</td>
|
||
<td box="[521,575,1148,1170]" gridcol="4" gridrow="16" pageId="13" pageNumber="31">25.8</td>
|
||
<td box="[614,657,1148,1170]" gridcol="5" gridrow="16" pageId="13" pageNumber="31">8.5</td>
|
||
<td box="[695,749,1148,1170]" gridcol="6" gridrow="16" pageId="13" pageNumber="31">23.5</td>
|
||
<td box="[790,843,1148,1170]" gridcol="7" gridrow="16" pageId="13" pageNumber="31">49.1</td>
|
||
<td box="[884,938,1148,1170]" gridcol="8" gridrow="16" pageId="13" pageNumber="31">19.1</td>
|
||
<td box="[977,1019,1148,1170]" gridcol="9" gridrow="16" pageId="13" pageNumber="31">8.8</td>
|
||
<td box="[1054,1117,1148,1170]" gridcol="10" gridrow="16" pageId="13" pageNumber="31">33.0</td>
|
||
<td box="[1152,1207,1148,1170]" gridcol="11" gridrow="16" pageId="13" pageNumber="31">14.5</td>
|
||
<td box="[1243,1305,1148,1170]" gridcol="12" gridrow="16" pageId="13" pageNumber="31">25.2</td>
|
||
<td box="[1341,1396,1148,1170]" gridcol="13" gridrow="16" pageId="13" pageNumber="31">36.3</td>
|
||
<td box="[1425,1486,1148,1170]" gridcol="14" gridrow="16" pageId="13" pageNumber="31">8.2</td>
|
||
<td box="[1512,1573,1148,1170]" gridcol="15" gridrow="16" pageId="13" pageNumber="31">14.2</td>
|
||
<td box="[1608,1663,1148,1170]" gridcol="16" gridrow="16" pageId="13" pageNumber="31">36.9</td>
|
||
<td box="[1704,1758,1148,1170]" gridcol="17" gridrow="16" pageId="13" pageNumber="31">10.3</td>
|
||
<td box="[1803,1857,1148,1170]" gridcol="18" gridrow="16" pageId="13" pageNumber="31">13.3</td>
|
||
</tr>
|
||
<tr box="[99,1857,1177,1198]" gridrow="17" pageId="13" pageNumber="31" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[99,282,1177,1198]" gridcol="0" gridrow="17" pageId="13" pageNumber="31">(Paratype)</th>
|
||
</tr>
|
||
<tr box="[99,1857,1204,1226]" gridrow="18" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1204,1226]" gridcol="0" gridrow="18" pageId="13" pageNumber="31">SMF 93.354</th>
|
||
<td box="[300,320,1204,1226]" gridcol="1" gridrow="18" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,1204,1226]" pageId="13" pageNumber="31">Y</emphasis>
|
||
</td>
|
||
<td box="[342,384,1204,1226]" gridcol="2" gridrow="18" pageId="13" pageNumber="31">–</td>
|
||
<td box="[427,481,1204,1226]" gridcol="3" gridrow="18" pageId="13" pageNumber="31">–</td>
|
||
<td box="[521,575,1204,1226]" gridcol="4" gridrow="18" pageId="13" pageNumber="31">25.7</td>
|
||
<td box="[614,657,1204,1226]" gridcol="5" gridrow="18" pageId="13" pageNumber="31">8.8</td>
|
||
<td box="[695,749,1204,1226]" gridcol="6" gridrow="18" pageId="13" pageNumber="31">22.9</td>
|
||
<td box="[790,843,1204,1226]" gridcol="7" gridrow="18" pageId="13" pageNumber="31">48.8</td>
|
||
<td box="[884,938,1204,1226]" gridcol="8" gridrow="18" pageId="13" pageNumber="31">19.5</td>
|
||
<td box="[977,1019,1204,1226]" gridcol="9" gridrow="18" pageId="13" pageNumber="31">9.4</td>
|
||
<td box="[1054,1117,1204,1226]" gridcol="10" gridrow="18" pageId="13" pageNumber="31">32.1</td>
|
||
<td box="[1152,1207,1204,1226]" gridcol="11" gridrow="18" pageId="13" pageNumber="31">14.9</td>
|
||
<td box="[1243,1305,1204,1226]" gridcol="12" gridrow="18" pageId="13" pageNumber="31">24.5</td>
|
||
<td box="[1341,1396,1204,1226]" gridcol="13" gridrow="18" pageId="13" pageNumber="31">36.0</td>
|
||
<td box="[1425,1486,1204,1226]" gridcol="14" gridrow="18" pageId="13" pageNumber="31">8.1</td>
|
||
<td box="[1512,1573,1204,1226]" gridcol="15" gridrow="18" pageId="13" pageNumber="31">14.1</td>
|
||
<td box="[1608,1663,1204,1226]" gridcol="16" gridrow="18" pageId="13" pageNumber="31">35.1</td>
|
||
<td box="[1704,1758,1204,1226]" gridcol="17" gridrow="18" pageId="13" pageNumber="31">11.0</td>
|
||
<td box="[1803,1857,1204,1226]" gridcol="18" gridrow="18" pageId="13" pageNumber="31">13.0</td>
|
||
</tr>
|
||
<tr box="[99,1857,1233,1254]" gridrow="19" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1233,1254]" gridcol="0" gridrow="19" pageId="13" pageNumber="31">Mean</th>
|
||
<td box="[300,320,1233,1254]" gridcol="1" gridrow="19" pageId="13" pageNumber="31">–</td>
|
||
<td box="[342,384,1233,1254]" gridcol="2" gridrow="19" pageId="13" pageNumber="31">–</td>
|
||
<td box="[427,481,1233,1254]" gridcol="3" gridrow="19" pageId="13" pageNumber="31">–</td>
|
||
<td box="[521,575,1233,1254]" gridcol="4" gridrow="19" pageId="13" pageNumber="31">25.9</td>
|
||
<td box="[614,657,1233,1254]" gridcol="5" gridrow="19" pageId="13" pageNumber="31">8.7</td>
|
||
<td box="[695,749,1233,1254]" gridcol="6" gridrow="19" pageId="13" pageNumber="31">23.5</td>
|
||
<td box="[790,843,1233,1254]" gridcol="7" gridrow="19" pageId="13" pageNumber="31">48.5</td>
|
||
<td box="[884,938,1233,1254]" gridcol="8" gridrow="19" pageId="13" pageNumber="31">19.4</td>
|
||
<td box="[977,1019,1233,1254]" gridcol="9" gridrow="19" pageId="13" pageNumber="31">8.9</td>
|
||
<td box="[1054,1117,1233,1254]" gridcol="10" gridrow="19" pageId="13" pageNumber="31">32.5</td>
|
||
<td box="[1152,1207,1233,1254]" gridcol="11" gridrow="19" pageId="13" pageNumber="31">14.7</td>
|
||
<td box="[1243,1305,1233,1254]" gridcol="12" gridrow="19" pageId="13" pageNumber="31">24.6</td>
|
||
<td box="[1341,1396,1233,1254]" gridcol="13" gridrow="19" pageId="13" pageNumber="31">35.8</td>
|
||
<td box="[1425,1486,1233,1254]" gridcol="14" gridrow="19" pageId="13" pageNumber="31">8.1</td>
|
||
<td box="[1512,1573,1233,1254]" gridcol="15" gridrow="19" pageId="13" pageNumber="31">14.3</td>
|
||
<td box="[1608,1663,1233,1254]" gridcol="16" gridrow="19" pageId="13" pageNumber="31">35.5</td>
|
||
<td box="[1704,1758,1233,1254]" gridcol="17" gridrow="19" pageId="13" pageNumber="31">10.6</td>
|
||
<td box="[1803,1857,1233,1254]" gridcol="18" gridrow="19" pageId="13" pageNumber="31">13.1</td>
|
||
</tr>
|
||
<tr box="[99,1857,1262,1283]" gridrow="20" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1262,1283]" gridcol="0" gridrow="20" pageId="13" pageNumber="31">Range</th>
|
||
<td box="[300,320,1262,1283]" gridcol="1" gridrow="20" pageId="13" pageNumber="31">–</td>
|
||
<td box="[342,384,1262,1283]" gridcol="2" gridrow="20" pageId="13" pageNumber="31">–</td>
|
||
<td box="[427,481,1262,1283]" gridcol="3" gridrow="20" pageId="13" pageNumber="31">–</td>
|
||
<td box="[521,575,1262,1283]" gridcol="4" gridrow="20" pageId="13" pageNumber="31">25.7–</td>
|
||
<td box="[614,657,1262,1283]" gridcol="5" gridrow="20" pageId="13" pageNumber="31">8.5–</td>
|
||
<td box="[695,749,1262,1283]" gridcol="6" gridrow="20" pageId="13" pageNumber="31">22.9–</td>
|
||
<td box="[790,843,1262,1283]" gridcol="7" gridrow="20" pageId="13" pageNumber="31">47.6–</td>
|
||
<td box="[884,938,1262,1283]" gridcol="8" gridrow="20" pageId="13" pageNumber="31">19.1–</td>
|
||
<td box="[977,1019,1262,1283]" gridcol="9" gridrow="20" pageId="13" pageNumber="31">8.4–</td>
|
||
<td box="[1054,1117,1262,1283]" gridcol="10" gridrow="20" pageId="13" pageNumber="31">32.1–</td>
|
||
<td box="[1152,1207,1262,1283]" gridcol="11" gridrow="20" pageId="13" pageNumber="31">14.5–</td>
|
||
<td box="[1243,1305,1262,1283]" gridcol="12" gridrow="20" pageId="13" pageNumber="31">24.2–</td>
|
||
<td box="[1341,1396,1262,1283]" gridcol="13" gridrow="20" pageId="13" pageNumber="31">35.1–</td>
|
||
<td box="[1425,1486,1262,1283]" gridcol="14" gridrow="20" pageId="13" pageNumber="31">8.1–</td>
|
||
<td box="[1512,1573,1262,1283]" gridcol="15" gridrow="20" pageId="13" pageNumber="31">14.1–</td>
|
||
<td box="[1608,1663,1262,1283]" gridcol="16" gridrow="20" pageId="13" pageNumber="31">34.5–</td>
|
||
<td box="[1704,1758,1262,1283]" gridcol="17" gridrow="20" pageId="13" pageNumber="31">10.3–</td>
|
||
<td box="[1803,1857,1262,1283]" gridcol="18" gridrow="20" pageId="13" pageNumber="31">13.0–</td>
|
||
</tr>
|
||
<tr box="[99,1857,1290,1311]" gridrow="21" pageId="13" pageNumber="31" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<td box="[521,575,1290,1311]" gridcol="4" gridrow="21" pageId="13" pageNumber="31">26.2</td>
|
||
<td box="[614,657,1290,1311]" gridcol="5" gridrow="21" pageId="13" pageNumber="31">8.8</td>
|
||
<td box="[695,749,1290,1311]" gridcol="6" gridrow="21" pageId="13" pageNumber="31">24.2</td>
|
||
<td box="[790,843,1290,1311]" gridcol="7" gridrow="21" pageId="13" pageNumber="31">49.1</td>
|
||
<td box="[884,938,1290,1311]" gridcol="8" gridrow="21" pageId="13" pageNumber="31">19.6</td>
|
||
<td box="[977,1019,1290,1311]" gridcol="9" gridrow="21" pageId="13" pageNumber="31">9.4</td>
|
||
<td box="[1054,1117,1290,1311]" gridcol="10" gridrow="21" pageId="13" pageNumber="31">33.0</td>
|
||
<td box="[1152,1207,1290,1311]" gridcol="11" gridrow="21" pageId="13" pageNumber="31">14.9</td>
|
||
<td box="[1243,1305,1290,1311]" gridcol="12" gridrow="21" pageId="13" pageNumber="31">25.2</td>
|
||
<td box="[1341,1396,1290,1311]" gridcol="13" gridrow="21" pageId="13" pageNumber="31">36.3</td>
|
||
<td box="[1425,1486,1290,1311]" gridcol="14" gridrow="21" pageId="13" pageNumber="31">8.2</td>
|
||
<td box="[1512,1573,1290,1311]" gridcol="15" gridrow="21" pageId="13" pageNumber="31">14.5</td>
|
||
<td box="[1608,1663,1290,1311]" gridcol="16" gridrow="21" pageId="13" pageNumber="31">36.9</td>
|
||
<td box="[1704,1758,1290,1311]" gridcol="17" gridrow="21" pageId="13" pageNumber="31">11.0</td>
|
||
<td box="[1803,1857,1290,1311]" gridcol="18" gridrow="21" pageId="13" pageNumber="31">13.3</td>
|
||
</tr>
|
||
<tr box="[99,1857,1323,1344]" gridrow="22" pageId="13" pageNumber="31" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-18="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[99,282,1323,1344]" gridcol="0" gridrow="22" pageId="13" pageNumber="31">
|
||
<emphasis box="[99,218,1323,1344]" italics="true" pageId="13" pageNumber="31">R. simulator</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[99,1857,1350,1372]" gridrow="23" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1350,1372]" gridcol="0" gridrow="23" pageId="13" pageNumber="31">MHNG 1971.064</th>
|
||
<td box="[300,320,1350,1372]" gridcol="1" gridrow="23" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,1350,1372]" pageId="13" pageNumber="31">Y</emphasis>
|
||
</td>
|
||
<td box="[342,384,1350,1372]" gridcol="2" gridrow="23" pageId="13" pageNumber="31">8.0</td>
|
||
<td box="[427,481,1350,1372]" gridcol="3" gridrow="23" pageId="13" pageNumber="31">38.5</td>
|
||
<td box="[521,575,1350,1372]" gridcol="4" gridrow="23" pageId="13" pageNumber="31">25.5</td>
|
||
<td box="[614,657,1350,1372]" gridcol="5" gridrow="23" pageId="13" pageNumber="31">8.0</td>
|
||
<td box="[695,749,1350,1372]" gridcol="6" gridrow="23" pageId="13" pageNumber="31">19.8</td>
|
||
<td box="[790,843,1350,1372]" gridcol="7" gridrow="23" pageId="13" pageNumber="31">43.1</td>
|
||
<td box="[884,938,1350,1372]" gridcol="8" gridrow="23" pageId="13" pageNumber="31">17.3</td>
|
||
<td box="[977,1019,1350,1372]" gridcol="9" gridrow="23" pageId="13" pageNumber="31">8.4</td>
|
||
<td box="[1054,1117,1350,1372]" gridcol="10" gridrow="23" pageId="13" pageNumber="31">28.6</td>
|
||
<td box="[1152,1207,1350,1372]" gridcol="11" gridrow="23" pageId="13" pageNumber="31">13.4</td>
|
||
<td box="[1243,1305,1350,1372]" gridcol="12" gridrow="23" pageId="13" pageNumber="31">21.8</td>
|
||
<td box="[1341,1396,1350,1372]" gridcol="13" gridrow="23" pageId="13" pageNumber="31">31.9</td>
|
||
<td box="[1425,1486,1350,1372]" gridcol="14" gridrow="23" pageId="13" pageNumber="31">7.9</td>
|
||
<td box="[1512,1573,1350,1372]" gridcol="15" gridrow="23" pageId="13" pageNumber="31">12.7</td>
|
||
<td box="[1608,1663,1350,1372]" gridcol="16" gridrow="23" pageId="13" pageNumber="31">31.1</td>
|
||
<td box="[1704,1758,1350,1372]" gridcol="17" gridrow="23" pageId="13" pageNumber="31">9.0</td>
|
||
<td box="[1803,1857,1350,1372]" gridcol="18" gridrow="23" pageId="13" pageNumber="31">11.4</td>
|
||
</tr>
|
||
<tr box="[99,1857,1379,1401]" gridrow="24" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1379,1401]" gridcol="0" gridrow="24" pageId="13" pageNumber="31">MoM MW179°</th>
|
||
<td box="[300,320,1379,1401]" gridcol="1" gridrow="24" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,1379,1401]" pageId="13" pageNumber="31">X</emphasis>
|
||
</td>
|
||
<td box="[342,384,1379,1401]" gridcol="2" gridrow="24" pageId="13" pageNumber="31">8.0</td>
|
||
<td box="[427,481,1379,1401]" gridcol="3" gridrow="24" pageId="13" pageNumber="31">45.9</td>
|
||
<td box="[521,575,1379,1401]" gridcol="4" gridrow="24" pageId="13" pageNumber="31">25.1</td>
|
||
<td box="[614,657,1379,1401]" gridcol="5" gridrow="24" pageId="13" pageNumber="31">8.4</td>
|
||
<td box="[695,749,1379,1401]" gridcol="6" gridrow="24" pageId="13" pageNumber="31">19.7</td>
|
||
<td box="[790,843,1379,1401]" gridcol="7" gridrow="24" pageId="13" pageNumber="31">44.5</td>
|
||
<td box="[884,938,1379,1401]" gridcol="8" gridrow="24" pageId="13" pageNumber="31">17.8</td>
|
||
<td box="[977,1019,1379,1401]" gridcol="9" gridrow="24" pageId="13" pageNumber="31">8.0</td>
|
||
<td box="[1054,1117,1379,1401]" gridcol="10" gridrow="24" pageId="13" pageNumber="31">30.5</td>
|
||
<td box="[1152,1207,1379,1401]" gridcol="11" gridrow="24" pageId="13" pageNumber="31">13.7</td>
|
||
<td box="[1243,1305,1379,1401]" gridcol="12" gridrow="24" pageId="13" pageNumber="31">22.9</td>
|
||
<td box="[1341,1396,1379,1401]" gridcol="13" gridrow="24" pageId="13" pageNumber="31">32.8</td>
|
||
<td box="[1425,1486,1379,1401]" gridcol="14" gridrow="24" pageId="13" pageNumber="31">7.4</td>
|
||
<td box="[1512,1573,1379,1401]" gridcol="15" gridrow="24" pageId="13" pageNumber="31">14.0</td>
|
||
<td box="[1608,1663,1379,1401]" gridcol="16" gridrow="24" pageId="13" pageNumber="31">31.7</td>
|
||
<td box="[1704,1758,1379,1401]" gridcol="17" gridrow="24" pageId="13" pageNumber="31">9.5</td>
|
||
<td box="[1803,1857,1379,1401]" gridcol="18" gridrow="24" pageId="13" pageNumber="31">12.7</td>
|
||
</tr>
|
||
<tr box="[99,1857,1407,1429]" gridrow="25" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1407,1429]" gridcol="0" gridrow="25" pageId="13" pageNumber="31">MHNG 1971.066</th>
|
||
<td box="[300,320,1407,1429]" gridcol="1" gridrow="25" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,1407,1429]" pageId="13" pageNumber="31">Y</emphasis>
|
||
</td>
|
||
<td box="[342,384,1407,1429]" gridcol="2" gridrow="25" pageId="13" pageNumber="31">7.5</td>
|
||
<td box="[427,481,1407,1429]" gridcol="3" gridrow="25" pageId="13" pageNumber="31">42.9</td>
|
||
<td box="[521,575,1407,1429]" gridcol="4" gridrow="25" pageId="13" pageNumber="31">20.9</td>
|
||
<td box="[614,657,1407,1429]" gridcol="5" gridrow="25" pageId="13" pageNumber="31">8.2</td>
|
||
<td box="[695,749,1407,1429]" gridcol="6" gridrow="25" pageId="13" pageNumber="31">20.4</td>
|
||
<td box="[790,843,1407,1429]" gridcol="7" gridrow="25" pageId="13" pageNumber="31">42.0</td>
|
||
<td box="[884,938,1407,1429]" gridcol="8" gridrow="25" pageId="13" pageNumber="31">17.8</td>
|
||
<td box="[977,1019,1407,1429]" gridcol="9" gridrow="25" pageId="13" pageNumber="31">8.2</td>
|
||
<td box="[1054,1117,1407,1429]" gridcol="10" gridrow="25" pageId="13" pageNumber="31">27.9</td>
|
||
<td box="[1152,1207,1407,1429]" gridcol="11" gridrow="25" pageId="13" pageNumber="31">13.5</td>
|
||
<td box="[1243,1305,1407,1429]" gridcol="12" gridrow="25" pageId="13" pageNumber="31">22.6</td>
|
||
<td box="[1341,1396,1407,1429]" gridcol="13" gridrow="25" pageId="13" pageNumber="31">30.7</td>
|
||
<td box="[1425,1486,1407,1429]" gridcol="14" gridrow="25" pageId="13" pageNumber="31">7.3</td>
|
||
<td box="[1512,1573,1407,1429]" gridcol="15" gridrow="25" pageId="13" pageNumber="31">13.3</td>
|
||
<td box="[1608,1663,1407,1429]" gridcol="16" gridrow="25" pageId="13" pageNumber="31">31.1</td>
|
||
<td box="[1704,1758,1407,1429]" gridcol="17" gridrow="25" pageId="13" pageNumber="31">9.5</td>
|
||
<td box="[1803,1857,1407,1429]" gridcol="18" gridrow="25" pageId="13" pageNumber="31">11.9</td>
|
||
</tr>
|
||
<tr box="[99,1857,1435,1457]" gridrow="26" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1435,1457]" gridcol="0" gridrow="26" pageId="13" pageNumber="31">MHNG 1971.065</th>
|
||
<td box="[300,320,1435,1457]" gridcol="1" gridrow="26" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,1435,1457]" pageId="13" pageNumber="31">X</emphasis>
|
||
</td>
|
||
<td box="[342,384,1435,1457]" gridcol="2" gridrow="26" pageId="13" pageNumber="31">7.5</td>
|
||
<td box="[427,481,1435,1457]" gridcol="3" gridrow="26" pageId="13" pageNumber="31">39.8</td>
|
||
<td box="[521,575,1435,1457]" gridcol="4" gridrow="26" pageId="13" pageNumber="31">22.8</td>
|
||
<td box="[614,657,1435,1457]" gridcol="5" gridrow="26" pageId="13" pageNumber="31">8.0</td>
|
||
<td box="[695,749,1435,1457]" gridcol="6" gridrow="26" pageId="13" pageNumber="31">20.0</td>
|
||
<td box="[790,843,1435,1457]" gridcol="7" gridrow="26" pageId="13" pageNumber="31">45.3</td>
|
||
<td box="[884,938,1435,1457]" gridcol="8" gridrow="26" pageId="13" pageNumber="31">17.0</td>
|
||
<td box="[977,1019,1435,1457]" gridcol="9" gridrow="26" pageId="13" pageNumber="31">8.6</td>
|
||
<td box="[1054,1117,1435,1457]" gridcol="10" gridrow="26" pageId="13" pageNumber="31">29.0</td>
|
||
<td box="[1152,1207,1435,1457]" gridcol="11" gridrow="26" pageId="13" pageNumber="31">13.8</td>
|
||
<td box="[1243,1305,1435,1457]" gridcol="12" gridrow="26" pageId="13" pageNumber="31">21.5</td>
|
||
<td box="[1341,1396,1435,1457]" gridcol="13" gridrow="26" pageId="13" pageNumber="31">32.6</td>
|
||
<td box="[1425,1486,1435,1457]" gridcol="14" gridrow="26" pageId="13" pageNumber="31">6.9</td>
|
||
<td box="[1512,1573,1435,1457]" gridcol="15" gridrow="26" pageId="13" pageNumber="31">13.6</td>
|
||
<td box="[1608,1663,1435,1457]" gridcol="16" gridrow="26" pageId="13" pageNumber="31">32.4</td>
|
||
<td box="[1704,1758,1435,1457]" gridcol="17" gridrow="26" pageId="13" pageNumber="31">9.4</td>
|
||
<td box="[1803,1857,1435,1457]" gridcol="18" gridrow="26" pageId="13" pageNumber="31">12.0</td>
|
||
</tr>
|
||
<tr box="[99,1857,1463,1485]" gridrow="27" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1463,1485]" gridcol="0" gridrow="27" pageId="13" pageNumber="31">MoM MW273°</th>
|
||
<td box="[300,320,1463,1485]" gridcol="1" gridrow="27" pageId="13" pageNumber="31">
|
||
<emphasis bold="true" box="[300,320,1463,1485]" pageId="13" pageNumber="31">X</emphasis>
|
||
</td>
|
||
<td box="[342,384,1463,1485]" gridcol="2" gridrow="27" pageId="13" pageNumber="31">8.5</td>
|
||
<td box="[427,481,1463,1485]" gridcol="3" gridrow="27" pageId="13" pageNumber="31">42.2</td>
|
||
<td box="[521,575,1463,1485]" gridcol="4" gridrow="27" pageId="13" pageNumber="31">27.3</td>
|
||
<td box="[614,657,1463,1485]" gridcol="5" gridrow="27" pageId="13" pageNumber="31">8.5</td>
|
||
<td box="[695,749,1463,1485]" gridcol="6" gridrow="27" pageId="13" pageNumber="31">19.8</td>
|
||
<td box="[790,843,1463,1485]" gridcol="7" gridrow="27" pageId="13" pageNumber="31">44.2</td>
|
||
<td box="[884,938,1463,1485]" gridcol="8" gridrow="27" pageId="13" pageNumber="31">17.6</td>
|
||
<td box="[977,1019,1463,1485]" gridcol="9" gridrow="27" pageId="13" pageNumber="31">8.2</td>
|
||
<td box="[1054,1117,1463,1485]" gridcol="10" gridrow="27" pageId="13" pageNumber="31">29.5</td>
|
||
<td box="[1152,1207,1463,1485]" gridcol="11" gridrow="27" pageId="13" pageNumber="31">14.0</td>
|
||
<td box="[1243,1305,1463,1485]" gridcol="12" gridrow="27" pageId="13" pageNumber="31">23.2</td>
|
||
<td box="[1341,1396,1463,1485]" gridcol="13" gridrow="27" pageId="13" pageNumber="31">32.8</td>
|
||
<td box="[1425,1486,1463,1485]" gridcol="14" gridrow="27" pageId="13" pageNumber="31">7.8</td>
|
||
<td box="[1512,1573,1463,1485]" gridcol="15" gridrow="27" pageId="13" pageNumber="31">14.4</td>
|
||
<td box="[1608,1663,1463,1485]" gridcol="16" gridrow="27" pageId="13" pageNumber="31">32.2</td>
|
||
<td box="[1704,1758,1463,1485]" gridcol="17" gridrow="27" pageId="13" pageNumber="31">9.6</td>
|
||
<td box="[1803,1857,1463,1485]" gridcol="18" gridrow="27" pageId="13" pageNumber="31">11.4</td>
|
||
</tr>
|
||
<tr box="[99,1857,1492,1513]" gridrow="28" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1492,1513]" gridcol="0" gridrow="28" pageId="13" pageNumber="31">Mean</th>
|
||
<td box="[300,320,1492,1513]" gridcol="1" gridrow="28" pageId="13" pageNumber="31">–</td>
|
||
<td box="[342,384,1492,1513]" gridcol="2" gridrow="28" pageId="13" pageNumber="31">7.9</td>
|
||
<td box="[427,481,1492,1513]" gridcol="3" gridrow="28" pageId="13" pageNumber="31">41.9</td>
|
||
<td box="[521,575,1492,1513]" gridcol="4" gridrow="28" pageId="13" pageNumber="31">24.3</td>
|
||
<td box="[614,657,1492,1513]" gridcol="5" gridrow="28" pageId="13" pageNumber="31">8.2</td>
|
||
<td box="[695,749,1492,1513]" gridcol="6" gridrow="28" pageId="13" pageNumber="31">19.9</td>
|
||
<td box="[790,843,1492,1513]" gridcol="7" gridrow="28" pageId="13" pageNumber="31">43.8</td>
|
||
<td box="[884,938,1492,1513]" gridcol="8" gridrow="28" pageId="13" pageNumber="31">17.5</td>
|
||
<td box="[977,1019,1492,1513]" gridcol="9" gridrow="28" pageId="13" pageNumber="31">8.3</td>
|
||
<td box="[1054,1117,1492,1513]" gridcol="10" gridrow="28" pageId="13" pageNumber="31">29.1</td>
|
||
<td box="[1152,1207,1492,1513]" gridcol="11" gridrow="28" pageId="13" pageNumber="31">13.7</td>
|
||
<td box="[1243,1305,1492,1513]" gridcol="12" gridrow="28" pageId="13" pageNumber="31">22.4</td>
|
||
<td box="[1341,1396,1492,1513]" gridcol="13" gridrow="28" pageId="13" pageNumber="31">32.1</td>
|
||
<td box="[1425,1486,1492,1513]" gridcol="14" gridrow="28" pageId="13" pageNumber="31">7.4</td>
|
||
<td box="[1512,1573,1492,1513]" gridcol="15" gridrow="28" pageId="13" pageNumber="31">13.6</td>
|
||
<td box="[1608,1663,1492,1513]" gridcol="16" gridrow="28" pageId="13" pageNumber="31">31.7</td>
|
||
<td box="[1704,1758,1492,1513]" gridcol="17" gridrow="28" pageId="13" pageNumber="31">9.4</td>
|
||
<td box="[1803,1857,1492,1513]" gridcol="18" gridrow="28" pageId="13" pageNumber="31">11.9</td>
|
||
</tr>
|
||
<tr box="[99,1857,1520,1541]" gridrow="29" pageId="13" pageNumber="31">
|
||
<th box="[99,282,1520,1541]" gridcol="0" gridrow="29" pageId="13" pageNumber="31">Range</th>
|
||
<td box="[300,320,1520,1541]" gridcol="1" gridrow="29" pageId="13" pageNumber="31">–</td>
|
||
<td box="[342,384,1520,1541]" gridcol="2" gridrow="29" pageId="13" pageNumber="31">7.5–</td>
|
||
<td box="[427,481,1520,1541]" gridcol="3" gridrow="29" pageId="13" pageNumber="31">38.5–</td>
|
||
<td box="[521,575,1520,1541]" gridcol="4" gridrow="29" pageId="13" pageNumber="31">20.9–</td>
|
||
<td box="[614,657,1520,1541]" gridcol="5" gridrow="29" pageId="13" pageNumber="31">8.0–</td>
|
||
<td box="[695,749,1520,1541]" gridcol="6" gridrow="29" pageId="13" pageNumber="31">19.7–</td>
|
||
<td box="[790,843,1520,1541]" gridcol="7" gridrow="29" pageId="13" pageNumber="31">42.0–</td>
|
||
<td box="[884,938,1520,1541]" gridcol="8" gridrow="29" pageId="13" pageNumber="31">17.0–</td>
|
||
<td box="[977,1019,1520,1541]" gridcol="9" gridrow="29" pageId="13" pageNumber="31">8.0–</td>
|
||
<td box="[1054,1117,1520,1541]" gridcol="10" gridrow="29" pageId="13" pageNumber="31">27.9–</td>
|
||
<td box="[1152,1207,1520,1541]" gridcol="11" gridrow="29" pageId="13" pageNumber="31">13.4–</td>
|
||
<td box="[1243,1305,1520,1541]" gridcol="12" gridrow="29" pageId="13" pageNumber="31">21.5–</td>
|
||
<td box="[1341,1396,1520,1541]" gridcol="13" gridrow="29" pageId="13" pageNumber="31">30.7–</td>
|
||
<td box="[1425,1486,1520,1541]" gridcol="14" gridrow="29" pageId="13" pageNumber="31">6.9–</td>
|
||
<td box="[1512,1573,1520,1541]" gridcol="15" gridrow="29" pageId="13" pageNumber="31">12.7–</td>
|
||
<td box="[1608,1663,1520,1541]" gridcol="16" gridrow="29" pageId="13" pageNumber="31">31.1–</td>
|
||
<td box="[1704,1758,1520,1541]" gridcol="17" gridrow="29" pageId="13" pageNumber="31">9.0–</td>
|
||
<td box="[1803,1857,1520,1541]" gridcol="18" gridrow="29" pageId="13" pageNumber="31">11.4–</td>
|
||
</tr>
|
||
<tr box="[99,1857,1549,1570]" gridrow="30" pageId="13" pageNumber="31" rowspan-0="1" rowspan-1="1">
|
||
<td box="[342,384,1549,1570]" gridcol="2" gridrow="30" pageId="13" pageNumber="31">9.0</td>
|
||
<td box="[427,481,1549,1570]" gridcol="3" gridrow="30" pageId="13" pageNumber="31">45.9</td>
|
||
<td box="[521,575,1549,1570]" gridcol="4" gridrow="30" pageId="13" pageNumber="31">27.3</td>
|
||
<td box="[614,657,1549,1570]" gridcol="5" gridrow="30" pageId="13" pageNumber="31">8.5</td>
|
||
<td box="[695,749,1549,1570]" gridcol="6" gridrow="30" pageId="13" pageNumber="31">20.4</td>
|
||
<td box="[790,843,1549,1570]" gridcol="7" gridrow="30" pageId="13" pageNumber="31">45.3</td>
|
||
<td box="[884,938,1549,1570]" gridcol="8" gridrow="30" pageId="13" pageNumber="31">17.8</td>
|
||
<td box="[977,1019,1549,1570]" gridcol="9" gridrow="30" pageId="13" pageNumber="31">8.6</td>
|
||
<td box="[1054,1117,1549,1570]" gridcol="10" gridrow="30" pageId="13" pageNumber="31">30.5</td>
|
||
<td box="[1152,1207,1549,1570]" gridcol="11" gridrow="30" pageId="13" pageNumber="31">14.0</td>
|
||
<td box="[1243,1305,1549,1570]" gridcol="12" gridrow="30" pageId="13" pageNumber="31">23.2</td>
|
||
<td box="[1341,1396,1549,1570]" gridcol="13" gridrow="30" pageId="13" pageNumber="31">32.8</td>
|
||
<td box="[1425,1486,1549,1570]" gridcol="14" gridrow="30" pageId="13" pageNumber="31">7.9</td>
|
||
<td box="[1512,1573,1549,1570]" gridcol="15" gridrow="30" pageId="13" pageNumber="31">14.4</td>
|
||
<td box="[1608,1663,1549,1570]" gridcol="16" gridrow="30" pageId="13" pageNumber="31">32.4</td>
|
||
<td box="[1704,1758,1549,1570]" gridcol="17" gridrow="30" pageId="13" pageNumber="31">9.6</td>
|
||
<td box="[1803,1857,1549,1570]" gridcol="18" gridrow="30" pageId="13" pageNumber="31">12.7</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7859662" ID-Zenodo-Dep="7859662" httpUri="https://zenodo.org/record/7859662/files/figure.png" pageId="14" pageNumber="32" startId="14.[151,164,1031,1052]" targetBox="[196,1383,174,1015]" targetPageId="14" targetType="figure">
|
||
<paragraph blockId="14.[151,1421,1031,1081]" pageId="14" pageNumber="32">
|
||
FIG. 6. Cranium in dorsoventral (top), lateral (middle) and ventrodorsal (bottom; only for MHNG 1971.168) views of holotypes of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[548,642,1060,1081]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[548,642,1060,1081]" italics="true" pageId="14" pageNumber="32">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[648,724,1060,1081]" pageId="14" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(left) and
|
||
<taxonomicName box="[825,957,1060,1081]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[825,957,1060,1081]" italics="true" pageId="14" pageNumber="32">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
(right)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="14.[809,1422,1159,1973]" lastBlockId="16.[809,1422,175,1391]" lastPageId="16" lastPageNumber="34" pageId="14" pageNumber="32">
|
||
<emphasis box="[847,1023,1364,1390]" italics="true" pageId="14" pageNumber="32">Specific status</emphasis>
|
||
.—
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[1061,1306,1364,1390]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[1061,1306,1364,1390]" italics="true" pageId="14" pageNumber="32">Rhinolophus namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1322,1422,1364,1390]" pageId="14" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
appears morphologically most similar to
|
||
<taxonomicName class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis italics="true" pageId="14" pageNumber="32">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
, although these two species are currently placed in two separate groups (
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[1189,1290,1467,1493]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[1189,1290,1467,1493]" italics="true" pageId="14" pageNumber="32">capensis</emphasis>
|
||
</taxonomicName>
|
||
-group and
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[809,882,1501,1527]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[809,882,1501,1527]" italics="true" pageId="14" pageNumber="32">adami</emphasis>
|
||
</taxonomicName>
|
||
-group, respectively). Due to this mismatch,
|
||
<taxonomicName box="[809,970,1536,1562]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[809,970,1536,1562]" italics="true" pageId="14" pageNumber="32">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
is used as the primary comparison to assign specific status, while other members of the
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[809,882,1604,1630]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[809,882,1604,1630]" italics="true" pageId="14" pageNumber="32">adami</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[953,1054,1604,1630]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[953,1054,1604,1630]" italics="true" pageId="14" pageNumber="32">capensis</emphasis>
|
||
</taxonomicName>
|
||
groups are considered under ‘Similar species’ below. In comparison to
|
||
<taxonomicName class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis italics="true" pageId="14" pageNumber="32">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[889,1005,1673,1699]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[889,1005,1673,1699]" italics="true" pageId="14" pageNumber="32">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1015,1108,1673,1699]" pageId="14" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
shares several similar external and craniodental measurements, including overlapping forearm length, horseshoe width, metacarpal length of 4th and 5th digit, length of 1st phalanx of 3rd and 5th digit, length of 2nd phalanx of 5th digit, Tib, HF, MW, ZW, BcH, CC, CM3, IOB, ML, cm3, AMSW, PL (
|
||
<tableCitation box="[1188,1286,1879,1905]" captionStart="TABLE" captionStartId="13.[99,112,574,595]" captionTargetPageId="13" captionText="TABLE 3. External measurements of specimens of R. namuli sp. nov., R. maendeleo and R. simulator. Measurement with an asterisk in the header have non-overlapping ranges between R. namuli sp. nov. and R. maendeleo. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’" httpUri="http://table.plazi.org/id/2CF56638FFE1FFA8FFFCFDBDF895FD76" pageId="14" pageNumber="32" tableUuid="2CF56638FFE1FFA8FFFCFDBDF895FD76">Table 3</tableCitation>
|
||
). The two species also exhibit a similar overall cranial shape and size (
|
||
<figureCitation box="[931,1008,1947,1973]" captionStart="FIG" captionStartId="14.[151,164,1031,1052]" captionTargetBox="[196,1383,174,1015]" captionTargetId="figure-411@14.[196,1383,174,1016]" captionTargetPageId="14" captionText="FIG. 6. Cranium in dorsoventral (top), lateral (middle) and ventrodorsal (bottom; only for MHNG 1971.168) views of holotypes of R. namuli sp. nov. (left) and R. maendeleo (right)" figureDoi="http://doi.org/10.5281/zenodo.7859662" httpUri="https://zenodo.org/record/7859662/files/figure.png" pageId="14" pageNumber="32">Fig. 6</figureCitation>
|
||
). However,
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[1161,1280,1947,1973]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="32" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[1161,1280,1947,1973]" italics="true" pageId="14" pageNumber="32">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1293,1389,1947,1973]" pageId="14" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is distinguished from
|
||
<taxonomicName box="[390,556,1092,1118]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[390,556,1092,1118]" italics="true" pageId="16" pageNumber="34">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
by a number of key characters. The characteristic feature of
|
||
<emphasis italics="true" pageId="16" pageNumber="34">
|
||
<taxonomicName class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="maendeleo">R. maendeleo</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
an absence of the bony bars forming the infraorbital foramena (
|
||
<bibRefCitation author="KOCK, D. & G. CSORBA & K. M. HOWELL" box="[392,598,1195,1221]" pageId="16" pageNumber="34" pagination="233 - 239" refId="ref16521" refString="KOCK, D., G. CSORBA, and K. M. HOWELL. 2000. Rhinolophus maendeleo n. sp. from Tanzania, a horseshoe bat noteworthy for its systematics and biogeography. Senckenbergiana Biologica, 80: 233 - 239." type="journal article" year="2000">
|
||
Kock
|
||
<emphasis box="[464,521,1195,1221]" italics="true" pageId="16" pageNumber="34">et al.</emphasis>
|
||
, 2000
|
||
</bibRefCitation>
|
||
), is not shared in
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[184,300,1229,1255]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="namuli" status="sp. nov">
|
||
<emphasis box="[184,300,1229,1255]" italics="true" pageId="16" pageNumber="34">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[311,399,1229,1255]" pageId="16" pageNumber="34" rank="species">sp. nov</taxonomicNameLabel>
|
||
, where the bony bars forming the infraorbital foramena are present in all collected specimens (Supplementary Appendix S2
|
||
<figureCitation box="[654,751,1297,1323]" captionStart="FIG" captionStartId="11.[151,164,1178,1199]" captionTargetBox="[151,763,174,1162]" captionTargetId="figure-171@11.[151,765,174,1163]" captionTargetPageId="11" captionText="FIG. 4. Live photographs of R. namuli sp. nov. in the field (♀, MHNG 1971.067), illustrating pelage colour and features of the (a) head and ears, (b) sella and lips and (c) connecting process" figureDoi="http://doi.org/10.5281/zenodo.7859658" httpUri="https://zenodo.org/record/7859658/files/figure.png" pageId="16" pageNumber="34">Fig. A4</figureCitation>
|
||
).
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[151,269,1331,1357]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[151,269,1331,1357]" italics="true" pageId="16" pageNumber="34">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[281,376,1331,1357]" pageId="16" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can also be distinguished from
|
||
<taxonomicName box="[151,322,1365,1391]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[151,322,1365,1391]" italics="true" pageId="16" pageNumber="34">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
by a number of non-overlapping craniodental measurements that indicate a slightly shorter overall skull length (SL —
|
||
<tableCitation box="[1302,1405,209,235]" captionStart="TABLE" captionStartId="15.[99,112,575,596]" captionTargetPageId="15" captionText="TABLE 4. Craniodental measurements of specimens of R. namuli sp. nov., R. maendeleo and R. simulator. Measurement with an asterisk in the header have non-overlapping ranges between R. namuli sp. nov. and R. maendeleo. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’" pageId="16" pageNumber="34">Table 4</tableCitation>
|
||
), a slightly wider and shorter rostrum (M3M3 and PB —
|
||
<tableCitation box="[845,933,277,303]" captionStart="TABLE" captionStartId="15.[99,112,575,596]" captionTargetPageId="15" captionText="TABLE 4. Craniodental measurements of specimens of R. namuli sp. nov., R. maendeleo and R. simulator. Measurement with an asterisk in the header have non-overlapping ranges between R. namuli sp. nov. and R. maendeleo. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’" pageId="16" pageNumber="34">Table 4</tableCitation>
|
||
) and subtle differences in rostral and narial inflation shape (
|
||
<figureCitation box="[1038,1111,311,337]" captionStart="FIG" captionStartId="14.[151,164,1031,1052]" captionTargetBox="[196,1383,174,1015]" captionTargetId="figure-411@14.[196,1383,174,1016]" captionTargetPageId="14" captionText="FIG. 6. Cranium in dorsoventral (top), lateral (middle) and ventrodorsal (bottom; only for MHNG 1971.168) views of holotypes of R. namuli sp. nov. (left) and R. maendeleo (right)" figureDoi="http://doi.org/10.5281/zenodo.7859662" httpUri="https://zenodo.org/record/7859662/files/figure.png" pageId="16" pageNumber="34">Fig. 6</figureCitation>
|
||
).
|
||
<taxonomicName box="[1134,1295,311,337]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[1134,1295,311,337]" italics="true" pageId="16" pageNumber="34">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
also bears a deeper and longer rostral depression whose posterior most point extends beyond the constriction point between the orbits. In
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[1154,1270,413,439]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[1154,1270,413,439]" italics="true" pageId="16" pageNumber="34">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1281,1375,413,439]" pageId="16" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
the depression is shallower, and does not extend as far back between the orbits (
|
||
<figureCitation box="[1113,1187,481,507]" captionStart="FIG" captionStartId="14.[151,164,1031,1052]" captionTargetBox="[196,1383,174,1015]" captionTargetId="figure-411@14.[196,1383,174,1016]" captionTargetPageId="14" captionText="FIG. 6. Cranium in dorsoventral (top), lateral (middle) and ventrodorsal (bottom; only for MHNG 1971.168) views of holotypes of R. namuli sp. nov. (left) and R. maendeleo (right)" figureDoi="http://doi.org/10.5281/zenodo.7859662" httpUri="https://zenodo.org/record/7859662/files/figure.png" pageId="16" pageNumber="34">Fig. 6</figureCitation>
|
||
). These differences in rostrum shape, inflation angle and depression are captured in the two angle measurements (Ang1, Ang2), of which the first is non-overlapping between the species (
|
||
<tableCitation box="[1031,1124,617,643]" captionStart="TABLE" captionStartId="15.[99,112,575,596]" captionTargetPageId="15" captionText="TABLE 4. Craniodental measurements of specimens of R. namuli sp. nov., R. maendeleo and R. simulator. Measurement with an asterisk in the header have non-overlapping ranges between R. namuli sp. nov. and R. maendeleo. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’" pageId="16" pageNumber="34">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption pageId="15" pageNumber="33" startId="15.[99,112,575,596]" targetBox="[99,1878,684,1570]" targetIsTable="true" targetPageId="15" targetType="table">
|
||
<paragraph blockId="15.[99,1899,575,652]" pageId="15" pageNumber="33">
|
||
TABLE 4. Craniodental measurements of specimens of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[619,713,575,596]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="33" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[619,713,575,596]" italics="true" pageId="15" pageNumber="33">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[719,795,575,596]" pageId="15" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName box="[804,935,575,596]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="33" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[804,935,575,596]" italics="true" pageId="15" pageNumber="33">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[981,1100,575,596]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="33" phylum="Chordata" rank="species" species="simulator">
|
||
<emphasis box="[981,1100,575,596]" italics="true" pageId="15" pageNumber="33">R. simulator</emphasis>
|
||
</taxonomicName>
|
||
. Measurement with an asterisk in the header have non-overlapping ranges between
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[99,199,603,624]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="33" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[99,199,603,624]" italics="true" pageId="15" pageNumber="33">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[211,292,603,624]" pageId="15" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName box="[351,488,603,624]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="33" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[351,488,603,624]" italics="true" pageId="15" pageNumber="33">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="15" pageNumber="33">
|
||
<table box="[99,1878,684,1570]" gridcols="18" gridrows="31" pageId="15" pageNumber="33" rowsContinueFrom="13.[99,1857,684,1570]">
|
||
<tr box="[99,1878,684,705]" gridrow="0" pageId="15" pageNumber="33">
|
||
<th box="[99,328,684,705]" colspan="2" colspanRight="1" gridcol="0" gridrow="0" pageId="15" pageNumber="33">Collection no. Sex</th>
|
||
<th box="[345,411,684,705]" gridcol="2" gridrow="0" pageId="15" pageNumber="33">SL*</th>
|
||
<th box="[461,515,684,705]" gridcol="3" gridrow="0" pageId="15" pageNumber="33">MW</th>
|
||
<th box="[555,610,684,705]" gridcol="4" gridrow="0" pageId="15" pageNumber="33">ZW</th>
|
||
<th box="[646,700,684,705]" gridcol="5" gridrow="0" pageId="15" pageNumber="33">BcH</th>
|
||
<th box="[741,794,684,705]" gridcol="6" gridrow="0" pageId="15" pageNumber="33">CC</th>
|
||
<th box="[823,901,684,705]" gridcol="7" gridrow="0" pageId="15" pageNumber="33">M3M3*</th>
|
||
<th box="[929,984,684,705]" gridcol="8" gridrow="0" pageId="15" pageNumber="33">CM3</th>
|
||
<th box="[1024,1078,684,705]" gridcol="9" gridrow="0" pageId="15" pageNumber="33">IOB</th>
|
||
<th box="[1120,1186,684,705]" gridcol="10" gridrow="0" pageId="15" pageNumber="33">ML</th>
|
||
<th box="[1228,1283,684,705]" gridcol="11" gridrow="0" pageId="15" pageNumber="33">cm3</th>
|
||
<th box="[1310,1389,684,705]" gridcol="12" gridrow="0" pageId="15" pageNumber="33">ALSW*</th>
|
||
<th box="[1407,1482,684,705]" gridcol="13" gridrow="0" pageId="15" pageNumber="33">AMSW</th>
|
||
<th box="[1512,1565,684,705]" gridcol="14" gridrow="0" pageId="15" pageNumber="33">PL</th>
|
||
<th box="[1602,1656,684,705]" gridcol="15" gridrow="0" pageId="15" pageNumber="33">PB*</th>
|
||
<th box="[1698,1765,684,705]" gridcol="16" gridrow="0" pageId="15" pageNumber="33">Ang1*</th>
|
||
<th box="[1812,1878,684,705]" gridcol="17" gridrow="0" pageId="15" pageNumber="33">Ang2</th>
|
||
</tr>
|
||
<tr box="[99,1878,720,741]" gridrow="1" pageId="15" pageNumber="33">
|
||
<th box="[99,1878,720,741]" colspan="18" colspanRight="17" gridcol="0" gridrow="1" pageId="15" pageNumber="33">
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[99,193,720,741]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="33" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[99,193,720,741]" italics="true" pageId="15" pageNumber="33">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[199,275,720,741]" pageId="15" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
</tr>
|
||
<tr box="[99,1878,748,770]" gridrow="2" pageId="15" pageNumber="33">
|
||
<th box="[99,282,748,770]" gridcol="0" gridrow="2" pageId="15" pageNumber="33">MHNG 1971.067</th>
|
||
<td box="[300,328,748,770]" gridcol="1" gridrow="2" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,748,770]" pageId="15" pageNumber="33">X</emphasis>
|
||
</td>
|
||
<td box="[345,411,748,770]" gridcol="2" gridrow="2" pageId="15" pageNumber="33">19.09</td>
|
||
<td box="[461,515,748,770]" gridcol="3" gridrow="2" pageId="15" pageNumber="33">9.30</td>
|
||
<td box="[555,610,748,770]" gridcol="4" gridrow="2" pageId="15" pageNumber="33">9.29</td>
|
||
<td box="[646,700,748,770]" gridcol="5" gridrow="2" pageId="15" pageNumber="33">5.79</td>
|
||
<td box="[741,794,748,770]" gridcol="6" gridrow="2" pageId="15" pageNumber="33">4.78</td>
|
||
<td box="[823,901,748,770]" gridcol="7" gridrow="2" pageId="15" pageNumber="33">6.90</td>
|
||
<td box="[929,984,748,770]" gridcol="8" gridrow="2" pageId="15" pageNumber="33">7.08</td>
|
||
<td box="[1024,1078,748,770]" gridcol="9" gridrow="2" pageId="15" pageNumber="33">2.51</td>
|
||
<td box="[1120,1186,748,770]" gridcol="10" gridrow="2" pageId="15" pageNumber="33">12.67</td>
|
||
<td box="[1228,1283,748,770]" gridcol="11" gridrow="2" pageId="15" pageNumber="33">7.43</td>
|
||
<td box="[1310,1389,748,770]" gridcol="12" gridrow="2" pageId="15" pageNumber="33">3.34</td>
|
||
<td box="[1407,1482,748,770]" gridcol="13" gridrow="2" pageId="15" pageNumber="33">4.97</td>
|
||
<td box="[1512,1565,748,770]" gridcol="14" gridrow="2" pageId="15" pageNumber="33">2.64</td>
|
||
<td box="[1602,1656,748,770]" gridcol="15" gridrow="2" pageId="15" pageNumber="33">3.01</td>
|
||
<td box="[1698,1765,748,770]" gridcol="16" gridrow="2" pageId="15" pageNumber="33">16.11</td>
|
||
<td box="[1812,1878,748,770]" gridcol="17" gridrow="2" pageId="15" pageNumber="33">19.46</td>
|
||
</tr>
|
||
<tr box="[99,1878,777,798]" gridrow="3" pageId="15" pageNumber="33" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[99,282,777,798]" gridcol="0" gridrow="3" pageId="15" pageNumber="33">(Paratype)</th>
|
||
</tr>
|
||
<tr box="[99,1878,804,826]" gridrow="4" pageId="15" pageNumber="33">
|
||
<th box="[99,282,804,826]" gridcol="0" gridrow="4" pageId="15" pageNumber="33">MHNG 1971.068</th>
|
||
<td box="[300,328,804,826]" gridcol="1" gridrow="4" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,804,826]" pageId="15" pageNumber="33">Y</emphasis>
|
||
</td>
|
||
<td box="[345,411,804,826]" gridcol="2" gridrow="4" pageId="15" pageNumber="33">19.60</td>
|
||
<td box="[461,515,804,826]" gridcol="3" gridrow="4" pageId="15" pageNumber="33">9.56</td>
|
||
<td box="[555,610,804,826]" gridcol="4" gridrow="4" pageId="15" pageNumber="33">9.64</td>
|
||
<td box="[646,700,804,826]" gridcol="5" gridrow="4" pageId="15" pageNumber="33">6.01</td>
|
||
<td box="[741,794,804,826]" gridcol="6" gridrow="4" pageId="15" pageNumber="33">5.10</td>
|
||
<td box="[823,901,804,826]" gridcol="7" gridrow="4" pageId="15" pageNumber="33">7.07</td>
|
||
<td box="[929,984,804,826]" gridcol="8" gridrow="4" pageId="15" pageNumber="33">7.19</td>
|
||
<td box="[1024,1078,804,826]" gridcol="9" gridrow="4" pageId="15" pageNumber="33">2.51</td>
|
||
<td box="[1120,1186,804,826]" gridcol="10" gridrow="4" pageId="15" pageNumber="33">12.95</td>
|
||
<td box="[1228,1283,804,826]" gridcol="11" gridrow="4" pageId="15" pageNumber="33">7.56</td>
|
||
<td box="[1310,1389,804,826]" gridcol="12" gridrow="4" pageId="15" pageNumber="33">3.31</td>
|
||
<td box="[1407,1482,804,826]" gridcol="13" gridrow="4" pageId="15" pageNumber="33">5.17</td>
|
||
<td box="[1512,1565,804,826]" gridcol="14" gridrow="4" pageId="15" pageNumber="33">2.59</td>
|
||
<td box="[1602,1656,804,826]" gridcol="15" gridrow="4" pageId="15" pageNumber="33">3.25</td>
|
||
<td box="[1698,1765,804,826]" gridcol="16" gridrow="4" pageId="15" pageNumber="33">15.97</td>
|
||
<td box="[1812,1878,804,826]" gridcol="17" gridrow="4" pageId="15" pageNumber="33">17.95</td>
|
||
</tr>
|
||
<tr box="[99,1878,833,854]" gridrow="5" pageId="15" pageNumber="33">
|
||
<th box="[99,1878,833,854]" colspan="18" colspanRight="17" gridcol="0" gridrow="5" pageId="15" pageNumber="33">(Holotype)</th>
|
||
</tr>
|
||
<tr box="[99,1878,861,883]" gridrow="6" pageId="15" pageNumber="33">
|
||
<th box="[99,282,861,883]" gridcol="0" gridrow="6" pageId="15" pageNumber="33">MHNG 1971.069</th>
|
||
<td box="[300,328,861,883]" gridcol="1" gridrow="6" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,861,883]" pageId="15" pageNumber="33">Y</emphasis>
|
||
</td>
|
||
<td box="[345,411,861,883]" gridcol="2" gridrow="6" pageId="15" pageNumber="33">19.27</td>
|
||
<td box="[461,515,861,883]" gridcol="3" gridrow="6" pageId="15" pageNumber="33">9.25</td>
|
||
<td box="[555,610,861,883]" gridcol="4" gridrow="6" pageId="15" pageNumber="33">9.18</td>
|
||
<td box="[646,700,861,883]" gridcol="5" gridrow="6" pageId="15" pageNumber="33">5.82</td>
|
||
<td box="[741,794,861,883]" gridcol="6" gridrow="6" pageId="15" pageNumber="33">4.84</td>
|
||
<td box="[823,901,861,883]" gridcol="7" gridrow="6" pageId="15" pageNumber="33">6.72</td>
|
||
<td box="[929,984,861,883]" gridcol="8" gridrow="6" pageId="15" pageNumber="33">7.04</td>
|
||
<td box="[1024,1078,861,883]" gridcol="9" gridrow="6" pageId="15" pageNumber="33">2.33</td>
|
||
<td box="[1120,1186,861,883]" gridcol="10" gridrow="6" pageId="15" pageNumber="33">12.75</td>
|
||
<td box="[1228,1283,861,883]" gridcol="11" gridrow="6" pageId="15" pageNumber="33">7.33</td>
|
||
<td box="[1310,1389,861,883]" gridcol="12" gridrow="6" pageId="15" pageNumber="33">3.21</td>
|
||
<td box="[1407,1482,861,883]" gridcol="13" gridrow="6" pageId="15" pageNumber="33">4.85</td>
|
||
<td box="[1512,1565,861,883]" gridcol="14" gridrow="6" pageId="15" pageNumber="33">2.46</td>
|
||
<td box="[1602,1656,861,883]" gridcol="15" gridrow="6" pageId="15" pageNumber="33">3.13</td>
|
||
<td box="[1698,1765,861,883]" gridcol="16" gridrow="6" pageId="15" pageNumber="33">15.70</td>
|
||
<td box="[1812,1878,861,883]" gridcol="17" gridrow="6" pageId="15" pageNumber="33">17.66</td>
|
||
</tr>
|
||
<tr box="[99,1878,890,911]" gridrow="7" pageId="15" pageNumber="33">
|
||
<th box="[99,1878,890,911]" colspan="18" colspanRight="17" gridcol="0" gridrow="7" pageId="15" pageNumber="33">(Paratype)</th>
|
||
</tr>
|
||
<tr box="[99,1878,918,940]" gridrow="8" pageId="15" pageNumber="33">
|
||
<th box="[99,282,918,940]" gridcol="0" gridrow="8" pageId="15" pageNumber="33">DM 10833</th>
|
||
<td box="[300,328,918,940]" gridcol="1" gridrow="8" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,918,940]" pageId="15" pageNumber="33">Y</emphasis>
|
||
</td>
|
||
<td box="[345,411,918,940]" gridcol="2" gridrow="8" pageId="15" pageNumber="33">19.39</td>
|
||
<td box="[461,515,918,940]" gridcol="3" gridrow="8" pageId="15" pageNumber="33">9.65</td>
|
||
<td box="[555,610,918,940]" gridcol="4" gridrow="8" pageId="15" pageNumber="33">9.41</td>
|
||
<td box="[646,700,918,940]" gridcol="5" gridrow="8" pageId="15" pageNumber="33">5.64</td>
|
||
<td box="[741,794,918,940]" gridcol="6" gridrow="8" pageId="15" pageNumber="33">4.89</td>
|
||
<td box="[823,901,918,940]" gridcol="7" gridrow="8" pageId="15" pageNumber="33">6.77</td>
|
||
<td box="[929,984,918,940]" gridcol="8" gridrow="8" pageId="15" pageNumber="33">7.08</td>
|
||
<td box="[1024,1078,918,940]" gridcol="9" gridrow="8" pageId="15" pageNumber="33">2.64</td>
|
||
<td box="[1120,1186,918,940]" gridcol="10" gridrow="8" pageId="15" pageNumber="33">12.84</td>
|
||
<td box="[1228,1283,918,940]" gridcol="11" gridrow="8" pageId="15" pageNumber="33">7.44</td>
|
||
<td box="[1310,1389,918,940]" gridcol="12" gridrow="8" pageId="15" pageNumber="33">3.27</td>
|
||
<td box="[1407,1482,918,940]" gridcol="13" gridrow="8" pageId="15" pageNumber="33">5.03</td>
|
||
<td box="[1512,1565,918,940]" gridcol="14" gridrow="8" pageId="15" pageNumber="33">2.60</td>
|
||
<td box="[1602,1656,918,940]" gridcol="15" gridrow="8" pageId="15" pageNumber="33">3.27</td>
|
||
<td box="[1698,1765,918,940]" gridcol="16" gridrow="8" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1725,1737,918,939]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
<td box="[1812,1878,918,940]" gridcol="17" gridrow="8" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1839,1851,918,939]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr box="[99,1878,946,968]" gridrow="9" pageId="15" pageNumber="33">
|
||
<th box="[99,282,946,968]" gridcol="0" gridrow="9" pageId="15" pageNumber="33">DM 10839</th>
|
||
<td box="[300,328,946,968]" gridcol="1" gridrow="9" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,946,968]" pageId="15" pageNumber="33">X</emphasis>
|
||
</td>
|
||
<td box="[345,411,946,968]" gridcol="2" gridrow="9" pageId="15" pageNumber="33">19.48</td>
|
||
<td box="[461,515,946,968]" gridcol="3" gridrow="9" pageId="15" pageNumber="33">9.40</td>
|
||
<td box="[555,610,946,968]" gridcol="4" gridrow="9" pageId="15" pageNumber="33">9.53</td>
|
||
<td box="[646,700,946,968]" gridcol="5" gridrow="9" pageId="15" pageNumber="33">5.7</td>
|
||
<td box="[741,794,946,968]" gridcol="6" gridrow="9" pageId="15" pageNumber="33">4.70</td>
|
||
<td box="[823,901,946,968]" gridcol="7" gridrow="9" pageId="15" pageNumber="33">6.96</td>
|
||
<td box="[929,984,946,968]" gridcol="8" gridrow="9" pageId="15" pageNumber="33">7.03</td>
|
||
<td box="[1024,1078,946,968]" gridcol="9" gridrow="9" pageId="15" pageNumber="33">2.51</td>
|
||
<td box="[1120,1186,946,968]" gridcol="10" gridrow="9" pageId="15" pageNumber="33">12.74</td>
|
||
<td box="[1228,1283,946,968]" gridcol="11" gridrow="9" pageId="15" pageNumber="33">7.35</td>
|
||
<td box="[1310,1389,946,968]" gridcol="12" gridrow="9" pageId="15" pageNumber="33">3.29</td>
|
||
<td box="[1407,1482,946,968]" gridcol="13" gridrow="9" pageId="15" pageNumber="33">4.88</td>
|
||
<td box="[1512,1565,946,968]" gridcol="14" gridrow="9" pageId="15" pageNumber="33">2.48</td>
|
||
<td box="[1602,1656,946,968]" gridcol="15" gridrow="9" pageId="15" pageNumber="33">2.98</td>
|
||
<td box="[1698,1765,946,968]" gridcol="16" gridrow="9" pageId="15" pageNumber="33">16.38</td>
|
||
<td box="[1812,1878,946,968]" gridcol="17" gridrow="9" pageId="15" pageNumber="33">18.45</td>
|
||
</tr>
|
||
<tr box="[99,1878,975,996]" gridrow="10" pageId="15" pageNumber="33">
|
||
<th box="[99,282,975,996]" gridcol="0" gridrow="10" pageId="15" pageNumber="33">Mean</th>
|
||
<td box="[300,328,975,996]" gridcol="1" gridrow="10" pageId="15" pageNumber="33">–</td>
|
||
<td box="[345,411,975,996]" gridcol="2" gridrow="10" pageId="15" pageNumber="33">19.37</td>
|
||
<td box="[461,515,975,996]" gridcol="3" gridrow="10" pageId="15" pageNumber="33">9.43</td>
|
||
<td box="[555,610,975,996]" gridcol="4" gridrow="10" pageId="15" pageNumber="33">9.41</td>
|
||
<td box="[646,700,975,996]" gridcol="5" gridrow="10" pageId="15" pageNumber="33">5.79</td>
|
||
<td box="[741,794,975,996]" gridcol="6" gridrow="10" pageId="15" pageNumber="33">4.86</td>
|
||
<td box="[823,901,975,996]" gridcol="7" gridrow="10" pageId="15" pageNumber="33">6.88</td>
|
||
<td box="[929,984,975,996]" gridcol="8" gridrow="10" pageId="15" pageNumber="33">7.08</td>
|
||
<td box="[1024,1078,975,996]" gridcol="9" gridrow="10" pageId="15" pageNumber="33">2.50</td>
|
||
<td box="[1120,1186,975,996]" gridcol="10" gridrow="10" pageId="15" pageNumber="33">12.79</td>
|
||
<td box="[1228,1283,975,996]" gridcol="11" gridrow="10" pageId="15" pageNumber="33">7.42</td>
|
||
<td box="[1310,1389,975,996]" gridcol="12" gridrow="10" pageId="15" pageNumber="33">3.28</td>
|
||
<td box="[1407,1482,975,996]" gridcol="13" gridrow="10" pageId="15" pageNumber="33">4.98</td>
|
||
<td box="[1512,1565,975,996]" gridcol="14" gridrow="10" pageId="15" pageNumber="33">2.55</td>
|
||
<td box="[1602,1656,975,996]" gridcol="15" gridrow="10" pageId="15" pageNumber="33">3.13</td>
|
||
<td box="[1698,1765,975,996]" gridcol="16" gridrow="10" pageId="15" pageNumber="33">16.04</td>
|
||
<td box="[1812,1878,975,996]" gridcol="17" gridrow="10" pageId="15" pageNumber="33">18.38</td>
|
||
</tr>
|
||
<tr box="[99,1878,1003,1024]" gridrow="11" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1003,1024]" gridcol="0" gridrow="11" pageId="15" pageNumber="33">Range</th>
|
||
<td box="[300,328,1003,1024]" gridcol="1" gridrow="11" pageId="15" pageNumber="33">–</td>
|
||
<td box="[345,411,1003,1024]" gridcol="2" gridrow="11" pageId="15" pageNumber="33">19.09–</td>
|
||
<td box="[461,515,1003,1024]" gridcol="3" gridrow="11" pageId="15" pageNumber="33">9.25–</td>
|
||
<td box="[555,610,1003,1024]" gridcol="4" gridrow="11" pageId="15" pageNumber="33">9.18–</td>
|
||
<td box="[646,700,1003,1024]" gridcol="5" gridrow="11" pageId="15" pageNumber="33">5.64–</td>
|
||
<td box="[741,794,1003,1024]" gridcol="6" gridrow="11" pageId="15" pageNumber="33">4.70–</td>
|
||
<td box="[823,901,1003,1024]" gridcol="7" gridrow="11" pageId="15" pageNumber="33">6.72–</td>
|
||
<td box="[929,984,1003,1024]" gridcol="8" gridrow="11" pageId="15" pageNumber="33">7.03–</td>
|
||
<td box="[1024,1078,1003,1024]" gridcol="9" gridrow="11" pageId="15" pageNumber="33">2.33–</td>
|
||
<td box="[1120,1186,1003,1024]" gridcol="10" gridrow="11" pageId="15" pageNumber="33">12.67–</td>
|
||
<td box="[1228,1283,1003,1024]" gridcol="11" gridrow="11" pageId="15" pageNumber="33">7.33–</td>
|
||
<td box="[1310,1389,1003,1024]" gridcol="12" gridrow="11" pageId="15" pageNumber="33">3.21–</td>
|
||
<td box="[1407,1482,1003,1024]" gridcol="13" gridrow="11" pageId="15" pageNumber="33">4.85–</td>
|
||
<td box="[1512,1565,1003,1024]" gridcol="14" gridrow="11" pageId="15" pageNumber="33">2.46–</td>
|
||
<td box="[1602,1656,1003,1024]" gridcol="15" gridrow="11" pageId="15" pageNumber="33">2.98–</td>
|
||
<td box="[1698,1765,1003,1024]" gridcol="16" gridrow="11" pageId="15" pageNumber="33">15.7–</td>
|
||
<td box="[1812,1878,1003,1024]" gridcol="17" gridrow="11" pageId="15" pageNumber="33">17.66–</td>
|
||
</tr>
|
||
<tr box="[99,1878,1031,1052]" gridrow="12" pageId="15" pageNumber="33" rowspan-0="1" rowspan-1="1">
|
||
<td box="[345,411,1031,1052]" gridcol="2" gridrow="12" pageId="15" pageNumber="33">19.60</td>
|
||
<td box="[461,515,1031,1052]" gridcol="3" gridrow="12" pageId="15" pageNumber="33">9.65</td>
|
||
<td box="[555,610,1031,1052]" gridcol="4" gridrow="12" pageId="15" pageNumber="33">9.64</td>
|
||
<td box="[646,700,1031,1052]" gridcol="5" gridrow="12" pageId="15" pageNumber="33">6.01</td>
|
||
<td box="[741,794,1031,1052]" gridcol="6" gridrow="12" pageId="15" pageNumber="33">5.10</td>
|
||
<td box="[823,901,1031,1052]" gridcol="7" gridrow="12" pageId="15" pageNumber="33">7.07</td>
|
||
<td box="[929,984,1031,1052]" gridcol="8" gridrow="12" pageId="15" pageNumber="33">7.19</td>
|
||
<td box="[1024,1078,1031,1052]" gridcol="9" gridrow="12" pageId="15" pageNumber="33">2.61</td>
|
||
<td box="[1120,1186,1031,1052]" gridcol="10" gridrow="12" pageId="15" pageNumber="33">12.95</td>
|
||
<td box="[1228,1283,1031,1052]" gridcol="11" gridrow="12" pageId="15" pageNumber="33">7.56</td>
|
||
<td box="[1310,1389,1031,1052]" gridcol="12" gridrow="12" pageId="15" pageNumber="33">3.34</td>
|
||
<td box="[1407,1482,1031,1052]" gridcol="13" gridrow="12" pageId="15" pageNumber="33">5.17</td>
|
||
<td box="[1512,1565,1031,1052]" gridcol="14" gridrow="12" pageId="15" pageNumber="33">2.6</td>
|
||
<td box="[1602,1656,1031,1052]" gridcol="15" gridrow="12" pageId="15" pageNumber="33">3.27</td>
|
||
<td box="[1698,1765,1031,1052]" gridcol="16" gridrow="12" pageId="15" pageNumber="33">16.38</td>
|
||
<td box="[1812,1878,1031,1052]" gridcol="17" gridrow="12" pageId="15" pageNumber="33">19.46</td>
|
||
</tr>
|
||
<tr box="[99,1878,1064,1085]" gridrow="13" pageId="15" pageNumber="33">
|
||
<th box="[99,1878,1064,1085]" colspan="18" colspanRight="17" gridcol="0" gridrow="13" pageId="15" pageNumber="33">
|
||
<emphasis box="[99,230,1064,1085]" italics="true" pageId="15" pageNumber="33">R. maendeleo</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[99,1878,1092,1114]" gridrow="14" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1092,1114]" gridcol="0" gridrow="14" pageId="15" pageNumber="33">SMF 79.643</th>
|
||
<td box="[300,328,1092,1114]" gridcol="1" gridrow="14" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,1092,1114]" pageId="15" pageNumber="33">Y</emphasis>
|
||
</td>
|
||
<td box="[345,411,1092,1114]" gridcol="2" gridrow="14" pageId="15" pageNumber="33">19.96</td>
|
||
<td box="[461,515,1092,1114]" gridcol="3" gridrow="14" pageId="15" pageNumber="33">9.39</td>
|
||
<td box="[555,610,1092,1114]" gridcol="4" gridrow="14" pageId="15" pageNumber="33">9.39</td>
|
||
<td box="[646,700,1092,1114]" gridcol="5" gridrow="14" pageId="15" pageNumber="33">6.06</td>
|
||
<td box="[741,794,1092,1114]" gridcol="6" gridrow="14" pageId="15" pageNumber="33">4.74</td>
|
||
<td box="[823,901,1092,1114]" gridcol="7" gridrow="14" pageId="15" pageNumber="33">6.60</td>
|
||
<td box="[929,984,1092,1114]" gridcol="8" gridrow="14" pageId="15" pageNumber="33">7.25</td>
|
||
<td box="[1024,1078,1092,1114]" gridcol="9" gridrow="14" pageId="15" pageNumber="33">2.54</td>
|
||
<td box="[1120,1186,1092,1114]" gridcol="10" gridrow="14" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1147,1159,1092,1113]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
<td box="[1228,1283,1092,1114]" gridcol="11" gridrow="14" pageId="15" pageNumber="33">7.61</td>
|
||
<td box="[1310,1389,1092,1114]" gridcol="12" gridrow="14" pageId="15" pageNumber="33">3.51</td>
|
||
<td box="[1407,1482,1092,1114]" gridcol="13" gridrow="14" pageId="15" pageNumber="33">5.03</td>
|
||
<td box="[1512,1565,1092,1114]" gridcol="14" gridrow="14" pageId="15" pageNumber="33">2.73</td>
|
||
<td box="[1602,1656,1092,1114]" gridcol="15" gridrow="14" pageId="15" pageNumber="33">2.87</td>
|
||
<td box="[1698,1765,1092,1114]" gridcol="16" gridrow="14" pageId="15" pageNumber="33">17.56</td>
|
||
<td box="[1812,1878,1092,1114]" gridcol="17" gridrow="14" pageId="15" pageNumber="33">18.4</td>
|
||
</tr>
|
||
<tr box="[99,1878,1120,1141]" gridrow="15" pageId="15" pageNumber="33" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-16="1" rowspan-17="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th box="[99,282,1120,1141]" gridcol="0" gridrow="15" pageId="15" pageNumber="33">(Holotype)</th>
|
||
</tr>
|
||
<tr box="[99,1878,1148,1170]" gridrow="16" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1148,1170]" gridcol="0" gridrow="16" pageId="15" pageNumber="33">SMF 66.960</th>
|
||
<td box="[300,328,1148,1170]" gridcol="1" gridrow="16" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,1148,1170]" pageId="15" pageNumber="33">X</emphasis>
|
||
</td>
|
||
<td box="[345,411,1148,1170]" gridcol="2" gridrow="16" pageId="15" pageNumber="33">19.99</td>
|
||
<td box="[461,515,1148,1170]" gridcol="3" gridrow="16" pageId="15" pageNumber="33">9.43</td>
|
||
<td box="[555,610,1148,1170]" gridcol="4" gridrow="16" pageId="15" pageNumber="33">9.38</td>
|
||
<td box="[646,700,1148,1170]" gridcol="5" gridrow="16" pageId="15" pageNumber="33">5.78</td>
|
||
<td box="[741,794,1148,1170]" gridcol="6" gridrow="16" pageId="15" pageNumber="33">4.76</td>
|
||
<td box="[823,901,1148,1170]" gridcol="7" gridrow="16" pageId="15" pageNumber="33">6.55</td>
|
||
<td box="[929,984,1148,1170]" gridcol="8" gridrow="16" pageId="15" pageNumber="33">7.20</td>
|
||
<td box="[1024,1078,1148,1170]" gridcol="9" gridrow="16" pageId="15" pageNumber="33">2.43</td>
|
||
<td box="[1120,1186,1148,1170]" gridcol="10" gridrow="16" pageId="15" pageNumber="33">13.16</td>
|
||
<td box="[1228,1283,1148,1170]" gridcol="11" gridrow="16" pageId="15" pageNumber="33">7.41</td>
|
||
<td box="[1310,1389,1148,1170]" gridcol="12" gridrow="16" pageId="15" pageNumber="33">3.65</td>
|
||
<td box="[1407,1482,1148,1170]" gridcol="13" gridrow="16" pageId="15" pageNumber="33">4.99</td>
|
||
<td box="[1512,1565,1148,1170]" gridcol="14" gridrow="16" pageId="15" pageNumber="33">2.54</td>
|
||
<td box="[1602,1656,1148,1170]" gridcol="15" gridrow="16" pageId="15" pageNumber="33">2.79</td>
|
||
<td box="[1698,1765,1148,1170]" gridcol="16" gridrow="16" pageId="15" pageNumber="33">16.72</td>
|
||
<td box="[1812,1878,1148,1170]" gridcol="17" gridrow="16" pageId="15" pageNumber="33">17.88</td>
|
||
</tr>
|
||
<tr box="[99,1878,1177,1198]" gridrow="17" pageId="15" pageNumber="33">
|
||
<th box="[99,1878,1177,1198]" colspan="18" colspanRight="17" gridcol="0" gridrow="17" pageId="15" pageNumber="33">(Paratype)</th>
|
||
</tr>
|
||
<tr box="[99,1878,1205,1227]" gridrow="18" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1205,1227]" gridcol="0" gridrow="18" pageId="15" pageNumber="33">SMF 93.354</th>
|
||
<td box="[300,328,1205,1227]" gridcol="1" gridrow="18" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,1205,1227]" pageId="15" pageNumber="33">Y</emphasis>
|
||
</td>
|
||
<td box="[345,411,1205,1227]" gridcol="2" gridrow="18" pageId="15" pageNumber="33">19.91</td>
|
||
<td box="[461,515,1205,1227]" gridcol="3" gridrow="18" pageId="15" pageNumber="33">9.42</td>
|
||
<td box="[555,610,1205,1227]" gridcol="4" gridrow="18" pageId="15" pageNumber="33">9.21</td>
|
||
<td box="[646,700,1205,1227]" gridcol="5" gridrow="18" pageId="15" pageNumber="33">6.02</td>
|
||
<td box="[741,794,1205,1227]" gridcol="6" gridrow="18" pageId="15" pageNumber="33">4.59</td>
|
||
<td box="[823,901,1205,1227]" gridcol="7" gridrow="18" pageId="15" pageNumber="33">6.46</td>
|
||
<td box="[929,984,1205,1227]" gridcol="8" gridrow="18" pageId="15" pageNumber="33">7.10</td>
|
||
<td box="[1024,1078,1205,1227]" gridcol="9" gridrow="18" pageId="15" pageNumber="33">2.49</td>
|
||
<td box="[1120,1186,1205,1227]" gridcol="10" gridrow="18" pageId="15" pageNumber="33">12.89</td>
|
||
<td box="[1228,1283,1205,1227]" gridcol="11" gridrow="18" pageId="15" pageNumber="33">7.48</td>
|
||
<td box="[1310,1389,1205,1227]" gridcol="12" gridrow="18" pageId="15" pageNumber="33">3.48</td>
|
||
<td box="[1407,1482,1205,1227]" gridcol="13" gridrow="18" pageId="15" pageNumber="33">5.05</td>
|
||
<td box="[1512,1565,1205,1227]" gridcol="14" gridrow="18" pageId="15" pageNumber="33">2.64</td>
|
||
<td box="[1602,1656,1205,1227]" gridcol="15" gridrow="18" pageId="15" pageNumber="33">2.68</td>
|
||
<td box="[1698,1765,1205,1227]" gridcol="16" gridrow="18" pageId="15" pageNumber="33">17.69</td>
|
||
<td box="[1812,1878,1205,1227]" gridcol="17" gridrow="18" pageId="15" pageNumber="33">19.26</td>
|
||
</tr>
|
||
<tr box="[99,1878,1234,1255]" gridrow="19" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1234,1255]" gridcol="0" gridrow="19" pageId="15" pageNumber="33">Mean</th>
|
||
<td box="[300,328,1234,1255]" gridcol="1" gridrow="19" pageId="15" pageNumber="33">–</td>
|
||
<td box="[345,411,1234,1255]" gridcol="2" gridrow="19" pageId="15" pageNumber="33">19.95</td>
|
||
<td box="[461,515,1234,1255]" gridcol="3" gridrow="19" pageId="15" pageNumber="33">9.41</td>
|
||
<td box="[555,610,1234,1255]" gridcol="4" gridrow="19" pageId="15" pageNumber="33">9.33</td>
|
||
<td box="[646,700,1234,1255]" gridcol="5" gridrow="19" pageId="15" pageNumber="33">5.95</td>
|
||
<td box="[741,794,1234,1255]" gridcol="6" gridrow="19" pageId="15" pageNumber="33">4.70</td>
|
||
<td box="[823,901,1234,1255]" gridcol="7" gridrow="19" pageId="15" pageNumber="33">6.54</td>
|
||
<td box="[929,984,1234,1255]" gridcol="8" gridrow="19" pageId="15" pageNumber="33">7.18</td>
|
||
<td box="[1024,1078,1234,1255]" gridcol="9" gridrow="19" pageId="15" pageNumber="33">2.49</td>
|
||
<td box="[1120,1186,1234,1255]" gridcol="10" gridrow="19" pageId="15" pageNumber="33">13.03</td>
|
||
<td box="[1228,1283,1234,1255]" gridcol="11" gridrow="19" pageId="15" pageNumber="33">7.50</td>
|
||
<td box="[1310,1389,1234,1255]" gridcol="12" gridrow="19" pageId="15" pageNumber="33">3.55</td>
|
||
<td box="[1407,1482,1234,1255]" gridcol="13" gridrow="19" pageId="15" pageNumber="33">5.02</td>
|
||
<td box="[1512,1565,1234,1255]" gridcol="14" gridrow="19" pageId="15" pageNumber="33">2.64</td>
|
||
<td box="[1602,1656,1234,1255]" gridcol="15" gridrow="19" pageId="15" pageNumber="33">2.78</td>
|
||
<td box="[1698,1765,1234,1255]" gridcol="16" gridrow="19" pageId="15" pageNumber="33">17.32</td>
|
||
<td box="[1812,1878,1234,1255]" gridcol="17" gridrow="19" pageId="15" pageNumber="33">18.51</td>
|
||
</tr>
|
||
<tr box="[99,1878,1262,1283]" gridrow="20" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1262,1283]" gridcol="0" gridrow="20" pageId="15" pageNumber="33">Range</th>
|
||
<td box="[300,328,1262,1283]" gridcol="1" gridrow="20" pageId="15" pageNumber="33">–</td>
|
||
<td box="[345,411,1262,1283]" gridcol="2" gridrow="20" pageId="15" pageNumber="33">19.91–</td>
|
||
<td box="[461,515,1262,1283]" gridcol="3" gridrow="20" pageId="15" pageNumber="33">9.39–</td>
|
||
<td box="[555,610,1262,1283]" gridcol="4" gridrow="20" pageId="15" pageNumber="33">9.21–</td>
|
||
<td box="[646,700,1262,1283]" gridcol="5" gridrow="20" pageId="15" pageNumber="33">5.78–</td>
|
||
<td box="[741,794,1262,1283]" gridcol="6" gridrow="20" pageId="15" pageNumber="33">4.59–</td>
|
||
<td box="[823,901,1262,1283]" gridcol="7" gridrow="20" pageId="15" pageNumber="33">6.46–</td>
|
||
<td box="[929,984,1262,1283]" gridcol="8" gridrow="20" pageId="15" pageNumber="33">7.10–</td>
|
||
<td box="[1024,1078,1262,1283]" gridcol="9" gridrow="20" pageId="15" pageNumber="33">2.43–</td>
|
||
<td box="[1120,1186,1262,1283]" gridcol="10" gridrow="20" pageId="15" pageNumber="33">12.89–</td>
|
||
<td box="[1228,1283,1262,1283]" gridcol="11" gridrow="20" pageId="15" pageNumber="33">7.41–</td>
|
||
<td box="[1310,1389,1262,1283]" gridcol="12" gridrow="20" pageId="15" pageNumber="33">3.48–</td>
|
||
<td box="[1407,1482,1262,1283]" gridcol="13" gridrow="20" pageId="15" pageNumber="33">4.99–</td>
|
||
<td box="[1512,1565,1262,1283]" gridcol="14" gridrow="20" pageId="15" pageNumber="33">2.54–</td>
|
||
<td box="[1602,1656,1262,1283]" gridcol="15" gridrow="20" pageId="15" pageNumber="33">2.68–</td>
|
||
<td box="[1698,1765,1262,1283]" gridcol="16" gridrow="20" pageId="15" pageNumber="33">16.72–</td>
|
||
<td box="[1812,1878,1262,1283]" gridcol="17" gridrow="20" pageId="15" pageNumber="33">17.88–</td>
|
||
</tr>
|
||
<tr box="[99,1878,1290,1311]" gridrow="21" pageId="15" pageNumber="33" rowspan-0="1" rowspan-1="1">
|
||
<td box="[345,411,1290,1311]" gridcol="2" gridrow="21" pageId="15" pageNumber="33">19.99</td>
|
||
<td box="[461,515,1290,1311]" gridcol="3" gridrow="21" pageId="15" pageNumber="33">9.43</td>
|
||
<td box="[555,610,1290,1311]" gridcol="4" gridrow="21" pageId="15" pageNumber="33">9.39</td>
|
||
<td box="[646,700,1290,1311]" gridcol="5" gridrow="21" pageId="15" pageNumber="33">6.06</td>
|
||
<td box="[741,794,1290,1311]" gridcol="6" gridrow="21" pageId="15" pageNumber="33">4.76</td>
|
||
<td box="[823,901,1290,1311]" gridcol="7" gridrow="21" pageId="15" pageNumber="33">6.60</td>
|
||
<td box="[929,984,1290,1311]" gridcol="8" gridrow="21" pageId="15" pageNumber="33">7.25</td>
|
||
<td box="[1024,1078,1290,1311]" gridcol="9" gridrow="21" pageId="15" pageNumber="33">2.54</td>
|
||
<td box="[1120,1186,1290,1311]" gridcol="10" gridrow="21" pageId="15" pageNumber="33">13.16</td>
|
||
<td box="[1228,1283,1290,1311]" gridcol="11" gridrow="21" pageId="15" pageNumber="33">7.61</td>
|
||
<td box="[1310,1389,1290,1311]" gridcol="12" gridrow="21" pageId="15" pageNumber="33">3.65</td>
|
||
<td box="[1407,1482,1290,1311]" gridcol="13" gridrow="21" pageId="15" pageNumber="33">5.05</td>
|
||
<td box="[1512,1565,1290,1311]" gridcol="14" gridrow="21" pageId="15" pageNumber="33">2.73</td>
|
||
<td box="[1602,1656,1290,1311]" gridcol="15" gridrow="21" pageId="15" pageNumber="33">2.87</td>
|
||
<td box="[1698,1765,1290,1311]" gridcol="16" gridrow="21" pageId="15" pageNumber="33">17.69</td>
|
||
<td box="[1812,1878,1290,1311]" gridcol="17" gridrow="21" pageId="15" pageNumber="33">19.26</td>
|
||
</tr>
|
||
<tr box="[99,1878,1323,1344]" gridrow="22" pageId="15" pageNumber="33">
|
||
<th box="[99,1878,1323,1344]" colspan="18" colspanRight="17" gridcol="0" gridrow="22" pageId="15" pageNumber="33">
|
||
<emphasis box="[99,218,1323,1344]" italics="true" pageId="15" pageNumber="33">R. simulator</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[99,1878,1351,1373]" gridrow="23" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1351,1373]" gridcol="0" gridrow="23" pageId="15" pageNumber="33">MHNG 1971.064</th>
|
||
<td box="[300,328,1351,1373]" gridcol="1" gridrow="23" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,1351,1373]" pageId="15" pageNumber="33">Y</emphasis>
|
||
</td>
|
||
<td box="[345,411,1351,1373]" gridcol="2" gridrow="23" pageId="15" pageNumber="33">18.08</td>
|
||
<td box="[461,515,1351,1373]" gridcol="3" gridrow="23" pageId="15" pageNumber="33">8.85</td>
|
||
<td box="[555,610,1351,1373]" gridcol="4" gridrow="23" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[576,588,1351,1372]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
<td box="[646,700,1351,1373]" gridcol="5" gridrow="23" pageId="15" pageNumber="33">5.71</td>
|
||
<td box="[741,794,1351,1373]" gridcol="6" gridrow="23" pageId="15" pageNumber="33">4.70</td>
|
||
<td box="[823,901,1351,1373]" gridcol="7" gridrow="23" pageId="15" pageNumber="33">6.25</td>
|
||
<td box="[929,984,1351,1373]" gridcol="8" gridrow="23" pageId="15" pageNumber="33">6.71</td>
|
||
<td box="[1024,1078,1351,1373]" gridcol="9" gridrow="23" pageId="15" pageNumber="33">2.27</td>
|
||
<td box="[1120,1186,1351,1373]" gridcol="10" gridrow="23" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1147,1159,1351,1372]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
<td box="[1228,1283,1351,1373]" gridcol="11" gridrow="23" pageId="15" pageNumber="33">6.88</td>
|
||
<td box="[1310,1389,1351,1373]" gridcol="12" gridrow="23" pageId="15" pageNumber="33">3.04</td>
|
||
<td box="[1407,1482,1351,1373]" gridcol="13" gridrow="23" pageId="15" pageNumber="33">4.63</td>
|
||
<td box="[1512,1565,1351,1373]" gridcol="14" gridrow="23" pageId="15" pageNumber="33">2.25</td>
|
||
<td box="[1602,1656,1351,1373]" gridcol="15" gridrow="23" pageId="15" pageNumber="33">2.67</td>
|
||
<td box="[1698,1765,1351,1373]" gridcol="16" gridrow="23" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1725,1737,1351,1372]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
<td box="[1812,1878,1351,1373]" gridcol="17" gridrow="23" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1839,1851,1351,1372]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr box="[99,1878,1379,1401]" gridrow="24" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1379,1401]" gridcol="0" gridrow="24" pageId="15" pageNumber="33">MoM MW179°</th>
|
||
<td box="[300,328,1379,1401]" gridcol="1" gridrow="24" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,1379,1401]" pageId="15" pageNumber="33">X</emphasis>
|
||
</td>
|
||
<td box="[345,411,1379,1401]" gridcol="2" gridrow="24" pageId="15" pageNumber="33">18.59</td>
|
||
<td box="[461,515,1379,1401]" gridcol="3" gridrow="24" pageId="15" pageNumber="33">9.07</td>
|
||
<td box="[555,610,1379,1401]" gridcol="4" gridrow="24" pageId="15" pageNumber="33">9.12</td>
|
||
<td box="[646,700,1379,1401]" gridcol="5" gridrow="24" pageId="15" pageNumber="33">5.72</td>
|
||
<td box="[741,794,1379,1401]" gridcol="6" gridrow="24" pageId="15" pageNumber="33">4.56</td>
|
||
<td box="[823,901,1379,1401]" gridcol="7" gridrow="24" pageId="15" pageNumber="33">6.56</td>
|
||
<td box="[929,984,1379,1401]" gridcol="8" gridrow="24" pageId="15" pageNumber="33">6.73</td>
|
||
<td box="[1024,1078,1379,1401]" gridcol="9" gridrow="24" pageId="15" pageNumber="33">2.51</td>
|
||
<td box="[1120,1186,1379,1401]" gridcol="10" gridrow="24" pageId="15" pageNumber="33">11.94</td>
|
||
<td box="[1228,1283,1379,1401]" gridcol="11" gridrow="24" pageId="15" pageNumber="33">7.00</td>
|
||
<td box="[1310,1389,1379,1401]" gridcol="12" gridrow="24" pageId="15" pageNumber="33">3.38</td>
|
||
<td box="[1407,1482,1379,1401]" gridcol="13" gridrow="24" pageId="15" pageNumber="33">4.66</td>
|
||
<td box="[1512,1565,1379,1401]" gridcol="14" gridrow="24" pageId="15" pageNumber="33">2.55</td>
|
||
<td box="[1602,1656,1379,1401]" gridcol="15" gridrow="24" pageId="15" pageNumber="33">2.76</td>
|
||
<td box="[1698,1765,1379,1401]" gridcol="16" gridrow="24" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1725,1737,1379,1400]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
<td box="[1812,1878,1379,1401]" gridcol="17" gridrow="24" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1839,1851,1379,1400]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr box="[99,1878,1407,1429]" gridrow="25" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1407,1429]" gridcol="0" gridrow="25" pageId="15" pageNumber="33">MHNG 1971.066</th>
|
||
<td box="[300,328,1407,1429]" gridcol="1" gridrow="25" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,1407,1429]" pageId="15" pageNumber="33">Y</emphasis>
|
||
</td>
|
||
<td box="[345,411,1407,1429]" gridcol="2" gridrow="25" pageId="15" pageNumber="33">18.41</td>
|
||
<td box="[461,515,1407,1429]" gridcol="3" gridrow="25" pageId="15" pageNumber="33">8.80</td>
|
||
<td box="[555,610,1407,1429]" gridcol="4" gridrow="25" pageId="15" pageNumber="33">8.90</td>
|
||
<td box="[646,700,1407,1429]" gridcol="5" gridrow="25" pageId="15" pageNumber="33">5.87</td>
|
||
<td box="[741,794,1407,1429]" gridcol="6" gridrow="25" pageId="15" pageNumber="33">4.87</td>
|
||
<td box="[823,901,1407,1429]" gridcol="7" gridrow="25" pageId="15" pageNumber="33">6.38</td>
|
||
<td box="[929,984,1407,1429]" gridcol="8" gridrow="25" pageId="15" pageNumber="33">6.76</td>
|
||
<td box="[1024,1078,1407,1429]" gridcol="9" gridrow="25" pageId="15" pageNumber="33">2.16</td>
|
||
<td box="[1120,1186,1407,1429]" gridcol="10" gridrow="25" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1147,1159,1408,1429]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
<td box="[1228,1283,1407,1429]" gridcol="11" gridrow="25" pageId="15" pageNumber="33">7.06</td>
|
||
<td box="[1310,1389,1407,1429]" gridcol="12" gridrow="25" pageId="15" pageNumber="33">3.40</td>
|
||
<td box="[1407,1482,1407,1429]" gridcol="13" gridrow="25" pageId="15" pageNumber="33">4.74</td>
|
||
<td box="[1512,1565,1407,1429]" gridcol="14" gridrow="25" pageId="15" pageNumber="33">2.22</td>
|
||
<td box="[1602,1656,1407,1429]" gridcol="15" gridrow="25" pageId="15" pageNumber="33">2.81</td>
|
||
<td box="[1698,1765,1407,1429]" gridcol="16" gridrow="25" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1725,1737,1408,1429]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
<td box="[1812,1878,1407,1429]" gridcol="17" gridrow="25" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1839,1851,1408,1429]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr box="[99,1878,1435,1457]" gridrow="26" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1435,1457]" gridcol="0" gridrow="26" pageId="15" pageNumber="33">MHNG 1971.065</th>
|
||
<td box="[300,328,1435,1457]" gridcol="1" gridrow="26" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,1435,1457]" pageId="15" pageNumber="33">X</emphasis>
|
||
</td>
|
||
<td box="[345,411,1435,1457]" gridcol="2" gridrow="26" pageId="15" pageNumber="33">18.09</td>
|
||
<td box="[461,515,1435,1457]" gridcol="3" gridrow="26" pageId="15" pageNumber="33">9.00</td>
|
||
<td box="[555,610,1435,1457]" gridcol="4" gridrow="26" pageId="15" pageNumber="33">8.79</td>
|
||
<td box="[646,700,1435,1457]" gridcol="5" gridrow="26" pageId="15" pageNumber="33">6.04</td>
|
||
<td box="[741,794,1435,1457]" gridcol="6" gridrow="26" pageId="15" pageNumber="33">4.35</td>
|
||
<td box="[823,901,1435,1457]" gridcol="7" gridrow="26" pageId="15" pageNumber="33">6.35</td>
|
||
<td box="[929,984,1435,1457]" gridcol="8" gridrow="26" pageId="15" pageNumber="33">6.63</td>
|
||
<td box="[1024,1078,1435,1457]" gridcol="9" gridrow="26" pageId="15" pageNumber="33">2.54</td>
|
||
<td box="[1120,1186,1435,1457]" gridcol="10" gridrow="26" pageId="15" pageNumber="33">11.29</td>
|
||
<td box="[1228,1283,1435,1457]" gridcol="11" gridrow="26" pageId="15" pageNumber="33">6.90</td>
|
||
<td box="[1310,1389,1435,1457]" gridcol="12" gridrow="26" pageId="15" pageNumber="33">3.26</td>
|
||
<td box="[1407,1482,1435,1457]" gridcol="13" gridrow="26" pageId="15" pageNumber="33">4.59</td>
|
||
<td box="[1512,1565,1435,1457]" gridcol="14" gridrow="26" pageId="15" pageNumber="33">2.07</td>
|
||
<td box="[1602,1656,1435,1457]" gridcol="15" gridrow="26" pageId="15" pageNumber="33">2.72</td>
|
||
<td box="[1698,1765,1435,1457]" gridcol="16" gridrow="26" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1725,1737,1436,1457]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
<td box="[1812,1878,1435,1457]" gridcol="17" gridrow="26" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1839,1851,1436,1457]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr box="[99,1878,1464,1486]" gridrow="27" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1464,1486]" gridcol="0" gridrow="27" pageId="15" pageNumber="33">MoM MW273°</th>
|
||
<td box="[300,328,1464,1486]" gridcol="1" gridrow="27" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[300,320,1464,1486]" pageId="15" pageNumber="33">X</emphasis>
|
||
</td>
|
||
<td box="[345,411,1464,1486]" gridcol="2" gridrow="27" pageId="15" pageNumber="33">18.17</td>
|
||
<td box="[461,515,1464,1486]" gridcol="3" gridrow="27" pageId="15" pageNumber="33">8.89</td>
|
||
<td box="[555,610,1464,1486]" gridcol="4" gridrow="27" pageId="15" pageNumber="33">8.52</td>
|
||
<td box="[646,700,1464,1486]" gridcol="5" gridrow="27" pageId="15" pageNumber="33">5.79</td>
|
||
<td box="[741,794,1464,1486]" gridcol="6" gridrow="27" pageId="15" pageNumber="33">4.35</td>
|
||
<td box="[823,901,1464,1486]" gridcol="7" gridrow="27" pageId="15" pageNumber="33">6.26</td>
|
||
<td box="[929,984,1464,1486]" gridcol="8" gridrow="27" pageId="15" pageNumber="33">6.36</td>
|
||
<td box="[1024,1078,1464,1486]" gridcol="9" gridrow="27" pageId="15" pageNumber="33">2.19</td>
|
||
<td box="[1120,1186,1464,1486]" gridcol="10" gridrow="27" pageId="15" pageNumber="33">11.10</td>
|
||
<td box="[1228,1283,1464,1486]" gridcol="11" gridrow="27" pageId="15" pageNumber="33">6.69</td>
|
||
<td box="[1310,1389,1464,1486]" gridcol="12" gridrow="27" pageId="15" pageNumber="33">3.08</td>
|
||
<td box="[1407,1482,1464,1486]" gridcol="13" gridrow="27" pageId="15" pageNumber="33">4.38</td>
|
||
<td box="[1512,1565,1464,1486]" gridcol="14" gridrow="27" pageId="15" pageNumber="33">2.34</td>
|
||
<td box="[1602,1656,1464,1486]" gridcol="15" gridrow="27" pageId="15" pageNumber="33">2.60</td>
|
||
<td box="[1698,1765,1464,1486]" gridcol="16" gridrow="27" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1725,1737,1464,1485]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
<td box="[1812,1878,1464,1486]" gridcol="17" gridrow="27" pageId="15" pageNumber="33">
|
||
<emphasis bold="true" box="[1839,1851,1464,1485]" pageId="15" pageNumber="33">–</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr box="[99,1878,1492,1513]" gridrow="28" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1492,1513]" gridcol="0" gridrow="28" pageId="15" pageNumber="33">Mean</th>
|
||
<td box="[300,328,1492,1513]" gridcol="1" gridrow="28" pageId="15" pageNumber="33">–</td>
|
||
<td box="[345,411,1492,1513]" gridcol="2" gridrow="28" pageId="15" pageNumber="33">18.27</td>
|
||
<td box="[461,515,1492,1513]" gridcol="3" gridrow="28" pageId="15" pageNumber="33">8.92</td>
|
||
<td box="[555,610,1492,1513]" gridcol="4" gridrow="28" pageId="15" pageNumber="33">8.83</td>
|
||
<td box="[646,700,1492,1513]" gridcol="5" gridrow="28" pageId="15" pageNumber="33">5.83</td>
|
||
<td box="[741,794,1492,1513]" gridcol="6" gridrow="28" pageId="15" pageNumber="33">4.57</td>
|
||
<td box="[823,901,1492,1513]" gridcol="7" gridrow="28" pageId="15" pageNumber="33">6.36</td>
|
||
<td box="[929,984,1492,1513]" gridcol="8" gridrow="28" pageId="15" pageNumber="33">6.64</td>
|
||
<td box="[1024,1078,1492,1513]" gridcol="9" gridrow="28" pageId="15" pageNumber="33">2.33</td>
|
||
<td box="[1120,1186,1492,1513]" gridcol="10" gridrow="28" pageId="15" pageNumber="33">11.44</td>
|
||
<td box="[1228,1283,1492,1513]" gridcol="11" gridrow="28" pageId="15" pageNumber="33">6.91</td>
|
||
<td box="[1310,1389,1492,1513]" gridcol="12" gridrow="28" pageId="15" pageNumber="33">3.23</td>
|
||
<td box="[1407,1482,1492,1513]" gridcol="13" gridrow="28" pageId="15" pageNumber="33">4.60</td>
|
||
<td box="[1512,1565,1492,1513]" gridcol="14" gridrow="28" pageId="15" pageNumber="33">2.29</td>
|
||
<td box="[1602,1656,1492,1513]" gridcol="15" gridrow="28" pageId="15" pageNumber="33">2.71</td>
|
||
<td box="[1698,1765,1492,1513]" gridcol="16" gridrow="28" pageId="15" pageNumber="33">–</td>
|
||
<td box="[1812,1878,1492,1513]" gridcol="17" gridrow="28" pageId="15" pageNumber="33">–</td>
|
||
</tr>
|
||
<tr box="[99,1878,1521,1542]" gridrow="29" pageId="15" pageNumber="33">
|
||
<th box="[99,282,1521,1542]" gridcol="0" gridrow="29" pageId="15" pageNumber="33">Range</th>
|
||
<td box="[300,328,1521,1542]" gridcol="1" gridrow="29" pageId="15" pageNumber="33">–</td>
|
||
<td box="[345,411,1521,1542]" gridcol="2" gridrow="29" pageId="15" pageNumber="33">18.08–</td>
|
||
<td box="[461,515,1521,1542]" gridcol="3" gridrow="29" pageId="15" pageNumber="33">8.80–</td>
|
||
<td box="[555,610,1521,1542]" gridcol="4" gridrow="29" pageId="15" pageNumber="33">8.52–</td>
|
||
<td box="[646,700,1521,1542]" gridcol="5" gridrow="29" pageId="15" pageNumber="33">5.71–</td>
|
||
<td box="[741,794,1521,1542]" gridcol="6" gridrow="29" pageId="15" pageNumber="33">4.35–</td>
|
||
<td box="[823,901,1521,1542]" gridcol="7" gridrow="29" pageId="15" pageNumber="33">6.25–</td>
|
||
<td box="[929,984,1521,1542]" gridcol="8" gridrow="29" pageId="15" pageNumber="33">6.36–</td>
|
||
<td box="[1024,1078,1521,1542]" gridcol="9" gridrow="29" pageId="15" pageNumber="33">2.16–</td>
|
||
<td box="[1120,1186,1521,1542]" gridcol="10" gridrow="29" pageId="15" pageNumber="33">11.10–</td>
|
||
<td box="[1228,1283,1521,1542]" gridcol="11" gridrow="29" pageId="15" pageNumber="33">6.69–</td>
|
||
<td box="[1310,1389,1521,1542]" gridcol="12" gridrow="29" pageId="15" pageNumber="33">3.04–</td>
|
||
<td box="[1407,1482,1521,1542]" gridcol="13" gridrow="29" pageId="15" pageNumber="33">4.38–</td>
|
||
<td box="[1512,1565,1521,1542]" gridcol="14" gridrow="29" pageId="15" pageNumber="33">2.07–</td>
|
||
<td box="[1602,1656,1521,1542]" gridcol="15" gridrow="29" pageId="15" pageNumber="33">2.60–</td>
|
||
<td box="[1698,1765,1521,1542]" gridcol="16" gridrow="29" pageId="15" pageNumber="33">–</td>
|
||
<td box="[1812,1878,1521,1542]" gridcol="17" gridrow="29" pageId="15" pageNumber="33">–</td>
|
||
</tr>
|
||
<tr box="[99,1878,1549,1570]" gridrow="30" pageId="15" pageNumber="33" rowspan-0="1" rowspan-1="1" rowspan-16="1" rowspan-17="1">
|
||
<td box="[345,411,1549,1570]" gridcol="2" gridrow="30" pageId="15" pageNumber="33">18.59</td>
|
||
<td box="[461,515,1549,1570]" gridcol="3" gridrow="30" pageId="15" pageNumber="33">9.07</td>
|
||
<td box="[555,610,1549,1570]" gridcol="4" gridrow="30" pageId="15" pageNumber="33">9.12</td>
|
||
<td box="[646,700,1549,1570]" gridcol="5" gridrow="30" pageId="15" pageNumber="33">6.04</td>
|
||
<td box="[741,794,1549,1570]" gridcol="6" gridrow="30" pageId="15" pageNumber="33">4.87</td>
|
||
<td box="[823,901,1549,1570]" gridcol="7" gridrow="30" pageId="15" pageNumber="33">6.56</td>
|
||
<td box="[929,984,1549,1570]" gridcol="8" gridrow="30" pageId="15" pageNumber="33">6.76</td>
|
||
<td box="[1024,1078,1549,1570]" gridcol="9" gridrow="30" pageId="15" pageNumber="33">2.54</td>
|
||
<td box="[1120,1186,1549,1570]" gridcol="10" gridrow="30" pageId="15" pageNumber="33">11.94</td>
|
||
<td box="[1228,1283,1549,1570]" gridcol="11" gridrow="30" pageId="15" pageNumber="33">7.06</td>
|
||
<td box="[1310,1389,1549,1570]" gridcol="12" gridrow="30" pageId="15" pageNumber="33">3.40</td>
|
||
<td box="[1407,1482,1549,1570]" gridcol="13" gridrow="30" pageId="15" pageNumber="33">4.74</td>
|
||
<td box="[1512,1565,1549,1570]" gridcol="14" gridrow="30" pageId="15" pageNumber="33">2.55</td>
|
||
<td box="[1602,1656,1549,1570]" gridcol="15" gridrow="30" pageId="15" pageNumber="33">2.81</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="16.[809,1422,175,1391]" pageId="16" pageNumber="34">
|
||
The two species also differ completely in bacular morphology (
|
||
<figureCitation box="[974,1049,685,711]" captionStart="FIG" captionStartId="17.[151,164,1598,1619]" captionTargetBox="[212,1360,176,1577]" captionTargetId="figure-126@17.[199,1386,173,1579]" captionTargetPageId="17" captionText="FIG. 8. Baculum of the holotype (MHNG 1971.168) of R. namuli sp. nov. in dorsal-ventral (a), ventral-dorsal (c), lateral (b and d) and proximal-distal (g) views. Baculum of the DM 10833 also shown in dorsal-ventral (e) and lateral (f) view. For the holotype (MHNG 1971.168), total baculum length is ca. 2.85 mm, greatest lateral width of basal cone is 0.78 mm and greatest dorsoventral width of basal cone is 0.87 mm" figureDoi="http://doi.org/10.5281/zenodo.7859666" httpUri="https://zenodo.org/record/7859666/files/figure.png" pageId="16" pageNumber="34">Fig. 8</figureCitation>
|
||
and Supplementary Appendix S2
|
||
<figureCitation box="[846,937,719,745]" captionStart="FIG" captionStartId="12.[151,164,893,914]" captionTargetBox="[361,1218,174,872]" captionTargetId="figure-451@12.[348,1226,173,879]" captionTargetPageId="12" captionText="FIG. 5. Head shape (a and b) and noseleaf (c and d) of holotypes of R. maendeleo SMF 79.643 and R. namuli sp. nov. MHNG 1971.068, respectively; connecting process of R. maendeleo SMF 79.643 (e), SMF 99.760 (f), and of R. namuli sp. nov. MHNG 1971.068 (g)" figureDoi="http://doi.org/10.5281/zenodo.7859660" httpUri="https://zenodo.org/record/7859660/files/figure.png" pageId="16" pageNumber="34">Fig. A5</figureCitation>
|
||
). In lateral view, the baculum of
|
||
<taxonomicName class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis italics="true" pageId="16" pageNumber="34">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
narrows gradually from the basal cone to the tip, the shaft being straight and flattened dorsoventrally, especially towards the distal end (
|
||
<bibRefCitation author="KOCK, D. & G. CSORBA & K. M. HOWELL" pageId="16" pageNumber="34" pagination="233 - 239" refId="ref16521" refString="KOCK, D., G. CSORBA, and K. M. HOWELL. 2000. Rhinolophus maendeleo n. sp. from Tanzania, a horseshoe bat noteworthy for its systematics and biogeography. Senckenbergiana Biologica, 80: 233 - 239." type="journal article" year="2000">
|
||
Kock
|
||
<emphasis box="[1355,1414,821,847]" italics="true" pageId="16" pageNumber="34">et al.</emphasis>
|
||
, 2000
|
||
</bibRefCitation>
|
||
). In contrast, the baculum of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[1211,1324,855,881]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[1211,1324,855,881]" italics="true" pageId="16" pageNumber="34">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1331,1422,855,881]" pageId="16" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is undulating, uncompressed, and is distinctively downturned (ventrally angled) at the tip. The inflations forming the basal cone of the baculum of
|
||
<taxonomicName box="[809,970,991,1017]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[809,970,991,1017]" italics="true" pageId="16" pageNumber="34">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
are unequal with the dorsal inflations protruding proximally relative to the ventral. This is in contrast to
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[974,1089,1059,1085]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[974,1089,1059,1085]" italics="true" pageId="16" pageNumber="34">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1098,1193,1059,1085]" pageId="16" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, which bears subequal inflations with no clear protrusion. The species also differ in the extent of the ventral incision in the basal cone, which is deep in
|
||
<taxonomicName box="[1147,1307,1161,1187]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[1147,1307,1161,1187]" italics="true" pageId="16" pageNumber="34">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
and shallow in
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[891,1005,1195,1221]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[891,1005,1195,1221]" italics="true" pageId="16" pageNumber="34">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1013,1105,1195,1221]" pageId="16" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Supplementary Appendix S2
|
||
<figureCitation box="[848,941,1229,1255]" captionStart="FIG" captionStartId="12.[151,164,893,914]" captionTargetBox="[361,1218,174,872]" captionTargetId="figure-451@12.[348,1226,173,879]" captionTargetPageId="12" captionText="FIG. 5. Head shape (a and b) and noseleaf (c and d) of holotypes of R. maendeleo SMF 79.643 and R. namuli sp. nov. MHNG 1971.068, respectively; connecting process of R. maendeleo SMF 79.643 (e), SMF 99.760 (f), and of R. namuli sp. nov. MHNG 1971.068 (g)" figureDoi="http://doi.org/10.5281/zenodo.7859660" httpUri="https://zenodo.org/record/7859660/files/figure.png" pageId="16" pageNumber="34">Fig. A5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7859664" ID-Zenodo-Dep="7859664" box="[151,684,988,1009]" httpUri="https://zenodo.org/record/7859664/files/figure.png" pageId="16" pageNumber="34" startId="16.[151,164,988,1009]" targetBox="[183,736,182,961]" targetPageId="16" targetType="figure">
|
||
<paragraph blockId="16.[151,684,988,1009]" box="[151,684,988,1009]" pageId="16" pageNumber="34">
|
||
FIG. 7. Mandibles of the holotype of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[508,603,988,1009]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[508,603,988,1009]" italics="true" pageId="16" pageNumber="34">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[609,684,988,1009]" pageId="16" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="16.[809,1422,175,1391]" lastBlockId="17.[809,1421,1779,1973]" lastPageId="17" lastPageNumber="35" pageId="16" pageNumber="34">
|
||
Externally, the two species are distinguished by the length of the tail, size of the ears and differences in horseshoe structure, and in the dimensions of the wing bones (
|
||
<tableCitation box="[958,1048,1365,1391]" captionStart="TABLE" captionStartId="13.[99,112,574,595]" captionTargetPageId="13" captionText="TABLE 3. External measurements of specimens of R. namuli sp. nov., R. maendeleo and R. simulator. Measurement with an asterisk in the header have non-overlapping ranges between R. namuli sp. nov. and R. maendeleo. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’" httpUri="http://table.plazi.org/id/2CF56638FFE1FFA8FFFCFDBDF895FD76" pageId="16" pageNumber="34" tableUuid="2CF56638FFE1FFA8FFFCFDBDF895FD76">Table 3</tableCitation>
|
||
). At a mean length of
|
||
<quantity box="[1310,1417,1365,1391]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="16" pageNumber="34" unit="mm" value="25.9">25.9 mm</quantity>
|
||
, the tail of
|
||
<taxonomicName box="[270,429,1779,1805]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="17" pageNumber="35" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[270,429,1779,1805]" italics="true" pageId="17" pageNumber="35">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
is markedly longer than that of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[183,296,1813,1839]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="17" pageNumber="35" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[183,296,1813,1839]" italics="true" pageId="17" pageNumber="35">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[304,395,1813,1839]" pageId="17" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(mean of
|
||
<quantity box="[514,623,1813,1839]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="17" pageNumber="35" unit="mm" value="20.3">20.3 mm</quantity>
|
||
). Ear length follows a similar pattern (means of
|
||
<quantity box="[575,681,1846,1872]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="17" pageNumber="35" unit="mm" value="23.5">23.5 mm</quantity>
|
||
versus
|
||
<quantity box="[151,255,1880,1906]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="17" pageNumber="35" unit="mm" value="19.3">19.3 mm</quantity>
|
||
in
|
||
<taxonomicName box="[291,451,1880,1906]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="17" pageNumber="35" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[291,451,1880,1906]" italics="true" pageId="17" pageNumber="35">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[507,620,1880,1906]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="17" pageNumber="35" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[507,620,1880,1906]" italics="true" pageId="17" pageNumber="35">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[627,720,1880,1906]" pageId="17" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, respectively). The connecting process of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[609,722,1914,1940]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="17" pageNumber="35" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[609,722,1914,1940]" italics="true" pageId="17" pageNumber="35">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="17" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is high and rounded, not continuously arched as in
|
||
<taxonomicName box="[841,1002,1779,1805]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="17" pageNumber="35" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[841,1002,1779,1805]" italics="true" pageId="17" pageNumber="35">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
, and with a more pronounced constriction in the sella (
|
||
<figureCitation box="[1081,1169,1813,1839]" captionStart="FIG" captionStartId="11.[151,164,1178,1199]" captionTargetBox="[151,763,174,1162]" captionTargetId="figure-171@11.[151,765,174,1163]" captionTargetPageId="11" captionText="FIG. 4. Live photographs of R. namuli sp. nov. in the field (♀, MHNG 1971.067), illustrating pelage colour and features of the (a) head and ears, (b) sella and lips and (c) connecting process" figureDoi="http://doi.org/10.5281/zenodo.7859658" httpUri="https://zenodo.org/record/7859658/files/figure.png" pageId="17" pageNumber="35">Figs. 4</figureCitation>
|
||
and
|
||
<figureCitation box="[1238,1254,1813,1839]" captionStart="FIG" captionStartId="12.[151,164,893,914]" captionTargetBox="[361,1218,174,872]" captionTargetId="figure-451@12.[348,1226,173,879]" captionTargetPageId="12" captionText="FIG. 5. Head shape (a and b) and noseleaf (c and d) of holotypes of R. maendeleo SMF 79.643 and R. namuli sp. nov. MHNG 1971.068, respectively; connecting process of R. maendeleo SMF 79.643 (e), SMF 99.760 (f), and of R. namuli sp. nov. MHNG 1971.068 (g)" figureDoi="http://doi.org/10.5281/zenodo.7859660" httpUri="https://zenodo.org/record/7859660/files/figure.png" pageId="17" pageNumber="35">5</figureCitation>
|
||
, and Supplementary Appendix S2
|
||
<figureCitation box="[1092,1189,1846,1872]" captionStart="FIG" captionStartId="7.[151,164,1894,1915]" captionTargetBox="[164,1399,1144,1876]" captionTargetId="figure-494@7.[154,1407,1134,1884]" captionTargetPageId="7" captionText="FIG. 2. PCA biplot for selected (A) craniodental and (B) external measurements. Ellipses represent 95% confidence intervals around each species group (not possible for R. maendeleo due to the low sample size). Large symbols represent the species mean value. Percentage values on each axis are the variances explained. For specimen accession numbers and input data, see Tables 3 and 4" figureDoi="http://doi.org/10.5281/zenodo.7859654" httpUri="https://zenodo.org/record/7859654/files/figure.png" pageId="17" pageNumber="35">Fig. A2</figureCitation>
|
||
). Within the wing bones, measurements of metacarpal and phalanx length illustrate differences between the species, particularly in the 3rd and 4th digits (
|
||
<tableCitation box="[1251,1342,1947,1973]" captionStart="TABLE" captionStartId="13.[99,112,574,595]" captionTargetPageId="13" captionText="TABLE 3. External measurements of specimens of R. namuli sp. nov., R. maendeleo and R. simulator. Measurement with an asterisk in the header have non-overlapping ranges between R. namuli sp. nov. and R. maendeleo. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’" httpUri="http://table.plazi.org/id/2CF56638FFE1FFA8FFFCFDBDF895FD76" pageId="17" pageNumber="35" tableUuid="2CF56638FFE1FFA8FFFCFDBDF895FD76">Table 3</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-Table-UUID="2CF56638FFFCFFB5FF08FA30FA0FF9FA" httpUri="http://table.plazi.org/id/2CF56638FFFCFFB5FF08FA30FA0FF9FA" pageId="16" pageNumber="34" startId="16.[151,164,1459,1480]" targetBox="[151,1342,1563,1965]" targetIsTable="true" targetPageId="16" targetType="table">
|
||
<paragraph blockId="16.[151,1421,1459,1536]" pageId="16" pageNumber="34">
|
||
TABLE 5. Echolocation call parameters of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[570,666,1459,1480]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[570,666,1459,1480]" italics="true" pageId="16" pageNumber="34">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[675,753,1459,1480]" pageId="16" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and species with similar echolocation peak frequency from nearby localities. Table highlights mean peak frequency (PF), standard deviation (SD), range and number of individuals recorded (
|
||
<emphasis box="[1343,1355,1487,1508]" italics="true" pageId="16" pageNumber="34">n</emphasis>
|
||
). Data collected by M. Curran and M. Kopp (unpublished material). MZ = Mozambique, MW = Malawi
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="16" pageNumber="34">
|
||
<table box="[151,1342,1563,1965]" gridcols="6" gridrows="13" pageId="16" pageNumber="34">
|
||
<tr box="[151,1342,1563,1618]" gridrow="0" pageId="16" pageNumber="34">
|
||
<th box="[151,327,1563,1618]" gridcol="0" gridrow="0" pageId="16" pageNumber="34">Species</th>
|
||
<th box="[378,585,1563,1618]" gridcol="1" gridrow="0" pageId="16" pageNumber="34">Location</th>
|
||
<th box="[729,784,1563,1618]" gridcol="2" gridrow="0" pageId="16" pageNumber="34">Mean</th>
|
||
<th box="[900,990,1563,1618]" gridcol="3" gridrow="0" pageId="16" pageNumber="34">PF (kHz) SD</th>
|
||
<th box="[1086,1182,1563,1618]" gridcol="4" gridrow="0" pageId="16" pageNumber="34">Range</th>
|
||
<th box="[1318,1342,1563,1618]" gridcol="5" gridrow="0" pageId="16" pageNumber="34">
|
||
<emphasis box="[1324,1336,1581,1602]" italics="true" pageId="16" pageNumber="34">n</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[151,1342,1633,1654]" gridrow="1" pageId="16" pageNumber="34" rowspan-3="1" rowspan-4="1">
|
||
<th box="[151,327,1633,1654]" gridcol="0" gridrow="1" pageId="16" pageNumber="34">
|
||
<emphasis box="[151,247,1633,1654]" italics="true" pageId="16" pageNumber="34">R. landeri</emphasis>
|
||
</th>
|
||
<td box="[378,585,1633,1654]" gridcol="1" gridrow="1" pageId="16" pageNumber="34">Mount Mabu, MZ</td>
|
||
<td box="[729,784,1633,1654]" gridcol="2" gridrow="1" pageId="16" pageNumber="34">102.2</td>
|
||
<td box="[1318,1342,1633,1654]" gridcol="5" gridrow="1" pageId="16" pageNumber="34">1</td>
|
||
</tr>
|
||
<tr box="[151,1342,1661,1682]" gridrow="2" pageId="16" pageNumber="34">
|
||
<th box="[151,327,1661,1682]" gridcol="0" gridrow="2" pageId="16" pageNumber="34">
|
||
<emphasis box="[151,231,1661,1682]" italics="true" pageId="16" pageNumber="34">R. blasii</emphasis>
|
||
</th>
|
||
<td box="[378,585,1661,1682]" gridcol="1" gridrow="2" pageId="16" pageNumber="34">Mount Namuli, MZ</td>
|
||
<td box="[729,784,1661,1682]" gridcol="2" gridrow="2" pageId="16" pageNumber="34">94.0</td>
|
||
<td box="[900,990,1661,1682]" gridcol="3" gridrow="2" pageId="16" pageNumber="34">1.1</td>
|
||
<td box="[1086,1182,1661,1682]" gridcol="4" gridrow="2" pageId="16" pageNumber="34">93.2–94.7</td>
|
||
<td box="[1318,1342,1661,1682]" gridcol="5" gridrow="2" pageId="16" pageNumber="34">2</td>
|
||
</tr>
|
||
<tr box="[151,1342,1690,1711]" gridrow="3" pageId="16" pageNumber="34" rowspan-0="1">
|
||
<td box="[378,585,1690,1711]" gridcol="1" gridrow="3" pageId="16" pageNumber="34">Mount Mabu, MZ</td>
|
||
<td box="[729,784,1690,1711]" gridcol="2" gridrow="3" pageId="16" pageNumber="34">94.9</td>
|
||
<td box="[900,990,1690,1711]" gridcol="3" gridrow="3" pageId="16" pageNumber="34">0.8</td>
|
||
<td box="[1086,1182,1690,1711]" gridcol="4" gridrow="3" pageId="16" pageNumber="34">93.8–95.5</td>
|
||
<td box="[1318,1342,1690,1711]" gridcol="5" gridrow="3" pageId="16" pageNumber="34">4</td>
|
||
</tr>
|
||
<tr box="[151,1342,1718,1739]" gridrow="4" pageId="16" pageNumber="34" rowspan-0="1">
|
||
<td box="[378,585,1718,1739]" gridcol="1" gridrow="4" pageId="16" pageNumber="34">Mount Mulanje, MW</td>
|
||
<td box="[729,784,1718,1739]" gridcol="2" gridrow="4" pageId="16" pageNumber="34">93.9</td>
|
||
<td box="[900,990,1718,1739]" gridcol="3" gridrow="4" pageId="16" pageNumber="34">1.0</td>
|
||
<td box="[1086,1182,1718,1739]" gridcol="4" gridrow="4" pageId="16" pageNumber="34">92.2–96.0</td>
|
||
<td box="[1318,1342,1718,1739]" gridcol="5" gridrow="4" pageId="16" pageNumber="34">17</td>
|
||
</tr>
|
||
<tr box="[151,1342,1746,1767]" gridrow="5" pageId="16" pageNumber="34">
|
||
<th box="[151,327,1746,1767]" gridcol="0" gridrow="5" pageId="16" pageNumber="34">
|
||
<emphasis box="[151,270,1746,1767]" italics="true" pageId="16" pageNumber="34">R. simulator</emphasis>
|
||
</th>
|
||
<td box="[378,585,1746,1767]" gridcol="1" gridrow="5" pageId="16" pageNumber="34">Mount Mulanje, MW</td>
|
||
<td box="[729,784,1746,1767]" gridcol="2" gridrow="5" pageId="16" pageNumber="34">86.0</td>
|
||
<td box="[900,990,1746,1767]" gridcol="3" gridrow="5" pageId="16" pageNumber="34">0.8</td>
|
||
<td box="[1086,1182,1746,1767]" gridcol="4" gridrow="5" pageId="16" pageNumber="34">84.0–86.7</td>
|
||
<td box="[1318,1342,1746,1767]" gridcol="5" gridrow="5" pageId="16" pageNumber="34">13</td>
|
||
</tr>
|
||
<tr box="[151,1342,1775,1796]" gridrow="6" pageId="16" pageNumber="34">
|
||
<th box="[151,327,1775,1796]" gridcol="0" gridrow="6" pageId="16" pageNumber="34">
|
||
<emphasis box="[151,255,1775,1796]" italics="true" pageId="16" pageNumber="34">R. clivosus</emphasis>
|
||
</th>
|
||
<td box="[378,585,1775,1796]" gridcol="1" gridrow="6" pageId="16" pageNumber="34">Mount Namuli, MZ</td>
|
||
<td box="[729,784,1775,1796]" gridcol="2" gridrow="6" pageId="16" pageNumber="34">81.2</td>
|
||
<td box="[900,990,1775,1796]" gridcol="3" gridrow="6" pageId="16" pageNumber="34">0.3</td>
|
||
<td box="[1086,1182,1775,1796]" gridcol="4" gridrow="6" pageId="16" pageNumber="34">81.0–81.4</td>
|
||
<td box="[1318,1342,1775,1796]" gridcol="5" gridrow="6" pageId="16" pageNumber="34">2</td>
|
||
</tr>
|
||
<tr box="[151,1342,1803,1824]" gridrow="7" pageId="16" pageNumber="34" rowspan-0="1">
|
||
<td box="[378,585,1803,1824]" gridcol="1" gridrow="7" pageId="16" pageNumber="34">Mount Mabu, MZ</td>
|
||
<td box="[729,784,1803,1824]" gridcol="2" gridrow="7" pageId="16" pageNumber="34">80.5</td>
|
||
<td box="[900,990,1803,1824]" gridcol="3" gridrow="7" pageId="16" pageNumber="34">0.5</td>
|
||
<td box="[1086,1182,1803,1824]" gridcol="4" gridrow="7" pageId="16" pageNumber="34">79.8–81.0</td>
|
||
<td box="[1318,1342,1803,1824]" gridcol="5" gridrow="7" pageId="16" pageNumber="34">4</td>
|
||
</tr>
|
||
<tr box="[151,1342,1831,1852]" gridrow="8" pageId="16" pageNumber="34" rowspan-0="1">
|
||
<td box="[378,585,1831,1852]" gridcol="1" gridrow="8" pageId="16" pageNumber="34">Mount Mulanje, MW</td>
|
||
<td box="[729,784,1831,1852]" gridcol="2" gridrow="8" pageId="16" pageNumber="34">81.3</td>
|
||
<td box="[900,990,1831,1852]" gridcol="3" gridrow="8" pageId="16" pageNumber="34">0.8</td>
|
||
<td box="[1086,1182,1831,1852]" gridcol="4" gridrow="8" pageId="16" pageNumber="34">79.9–82.6</td>
|
||
<td box="[1318,1342,1831,1852]" gridcol="5" gridrow="8" pageId="16" pageNumber="34">15</td>
|
||
</tr>
|
||
<tr box="[151,1342,1859,1880]" gridrow="9" pageId="16" pageNumber="34">
|
||
<th box="[151,327,1859,1880]" gridcol="0" gridrow="9" pageId="16" pageNumber="34">
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[151,245,1859,1880]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="34" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[151,245,1859,1880]" italics="true" pageId="16" pageNumber="34">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[252,327,1859,1880]" pageId="16" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<td box="[378,585,1859,1880]" gridcol="1" gridrow="9" pageId="16" pageNumber="34">Mount Namuli, MZ</td>
|
||
<td box="[729,784,1859,1880]" gridcol="2" gridrow="9" pageId="16" pageNumber="34">76.9</td>
|
||
<td box="[900,990,1859,1880]" gridcol="3" gridrow="9" pageId="16" pageNumber="34">0.8</td>
|
||
<td box="[1086,1182,1859,1880]" gridcol="4" gridrow="9" pageId="16" pageNumber="34">75.9–78.0</td>
|
||
<td box="[1318,1342,1859,1880]" gridcol="5" gridrow="9" pageId="16" pageNumber="34">7</td>
|
||
</tr>
|
||
<tr box="[151,1342,1888,1909]" gridrow="10" pageId="16" pageNumber="34">
|
||
<th box="[151,327,1888,1909]" gridcol="0" gridrow="10" pageId="16" pageNumber="34">
|
||
<emphasis box="[151,273,1888,1909]" italics="true" pageId="16" pageNumber="34">R. fumigatus</emphasis>
|
||
</th>
|
||
<td box="[378,585,1888,1909]" gridcol="1" gridrow="10" pageId="16" pageNumber="34">Mount Mulanje, MW</td>
|
||
<td box="[729,784,1888,1909]" gridcol="2" gridrow="10" pageId="16" pageNumber="34">58.4</td>
|
||
<td box="[900,990,1888,1909]" gridcol="3" gridrow="10" pageId="16" pageNumber="34">0.4</td>
|
||
<td box="[1086,1182,1888,1909]" gridcol="4" gridrow="10" pageId="16" pageNumber="34">58.1–59.0</td>
|
||
<td box="[1318,1342,1888,1909]" gridcol="5" gridrow="10" pageId="16" pageNumber="34">4</td>
|
||
</tr>
|
||
<tr box="[151,1342,1916,1937]" gridrow="11" pageId="16" pageNumber="34">
|
||
<th box="[151,327,1916,1937]" gridcol="0" gridrow="11" pageId="16" pageNumber="34">
|
||
<emphasis box="[151,279,1916,1937]" italics="true" pageId="16" pageNumber="34">R. mabuensis</emphasis>
|
||
</th>
|
||
<td box="[378,585,1916,1937]" gridcol="1" gridrow="11" pageId="16" pageNumber="34">Mount Mabu, MZ</td>
|
||
<td box="[729,784,1916,1937]" gridcol="2" gridrow="11" pageId="16" pageNumber="34">38.1</td>
|
||
<td box="[900,990,1916,1937]" gridcol="3" gridrow="11" pageId="16" pageNumber="34">0.3</td>
|
||
<td box="[1086,1182,1916,1937]" gridcol="4" gridrow="11" pageId="16" pageNumber="34">37.5–37.9</td>
|
||
<td box="[1318,1342,1916,1937]" gridcol="5" gridrow="11" pageId="16" pageNumber="34">2</td>
|
||
</tr>
|
||
<tr box="[151,1342,1944,1965]" gridrow="12" pageId="16" pageNumber="34" rowspan-0="1">
|
||
<td box="[378,585,1944,1965]" gridcol="1" gridrow="12" pageId="16" pageNumber="34">Mount Mulanje, MW</td>
|
||
<td box="[729,784,1944,1965]" gridcol="2" gridrow="12" pageId="16" pageNumber="34">37.7</td>
|
||
<td box="[900,990,1944,1965]" gridcol="3" gridrow="12" pageId="16" pageNumber="34">0.3</td>
|
||
<td box="[1086,1182,1944,1965]" gridcol="4" gridrow="12" pageId="16" pageNumber="34">37.9–38.3</td>
|
||
<td box="[1318,1342,1944,1965]" gridcol="5" gridrow="12" pageId="16" pageNumber="34">2</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7859666" ID-Zenodo-Dep="7859666" httpUri="https://zenodo.org/record/7859666/files/figure.png" pageId="17" pageNumber="35" startId="17.[151,164,1598,1619]" targetBox="[212,1360,176,1577]" targetPageId="17" targetType="figure">
|
||
<paragraph blockId="17.[151,1421,1598,1704]" pageId="17" pageNumber="35">
|
||
FIG. 8. Baculum of the holotype (MHNG 1971.168) of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[693,788,1598,1619]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="17" pageNumber="35" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[693,788,1598,1619]" italics="true" pageId="17" pageNumber="35">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[795,871,1598,1619]" pageId="17" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in dorsal-ventral (a), ventral-dorsal (c), lateral (b and d) and proximal-distal (g) views. Baculum of the DM 10833 also shown in dorsal-ventral (e) and lateral (f) view. For the holotype (MHNG 1971.168), total baculum length is ca. 2.85 mm, greatest lateral width of basal cone is 0.78 mm and greatest dorsoventral
|
||
</paragraph>
|
||
<paragraph blockId="17.[151,1421,1598,1704]" box="[634,938,1683,1704]" pageId="17" pageNumber="35">width of basal cone is 0.87 mm</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7859668" ID-Zenodo-Dep="7859668" httpUri="https://zenodo.org/record/7859668/files/figure.png" pageId="18" pageNumber="36" startId="18.[151,164,713,734]" targetBox="[288,1292,174,691]" targetPageId="18" targetType="figure">
|
||
<paragraph blockId="18.[151,1422,713,763]" pageId="18" pageNumber="36">
|
||
FIG. 9. Echolocation call of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[420,514,713,734]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[420,514,713,734]" italics="true" pageId="18" pageNumber="36">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[520,595,713,734]" pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<emphasis bold="true" box="[608,628,713,735]" pageId="18" pageNumber="36">X</emphasis>
|
||
, MHNG 1971.067; recorded at Mount Namuli, Zambesia Province, Mozambique, -15.405933 / 37.067006, ca. 1,200 m a.s.l.)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="18.[151,764,843,1973]" pageId="18" pageNumber="36">
|
||
<emphasis box="[189,382,843,869]" italics="true" pageId="18" pageNumber="36">Similar species.</emphasis>
|
||
—
|
||
<bibRefCitation author="KOCK, D. & G. CSORBA & K. M. HOWELL" box="[412,648,843,869]" pageId="18" pageNumber="36" pagination="233 - 239" refId="ref16521" refString="KOCK, D., G. CSORBA, and K. M. HOWELL. 2000. Rhinolophus maendeleo n. sp. from Tanzania, a horseshoe bat noteworthy for its systematics and biogeography. Senckenbergiana Biologica, 80: 233 - 239." type="journal article" year="2000">
|
||
Kock
|
||
<emphasis box="[489,555,843,869]" italics="true" pageId="18" pageNumber="36">et al.</emphasis>
|
||
(2000)
|
||
</bibRefCitation>
|
||
compare
|
||
<taxonomicName box="[151,330,878,904]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[151,330,878,904]" italics="true" pageId="18" pageNumber="36">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
to the morphologically similar
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[151,260,912,938]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[151,260,912,938]" italics="true" pageId="18" pageNumber="36">R. adami</emphasis>
|
||
</taxonomicName>
|
||
, illustrating a set of characters that differ between the species. We confirmed that
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[646,763,947,973]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[646,763,947,973]" italics="true" pageId="18" pageNumber="36">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[151,244,981,1007]" pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
possesses many of the same distinguishing characters. Specifically, both
|
||
<taxonomicName box="[531,702,1016,1042]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[531,702,1016,1042]" italics="true" pageId="18" pageNumber="36">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[151,264,1050,1076]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[151,264,1050,1076]" italics="true" pageId="18" pageNumber="36">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[272,362,1050,1076]" pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis italics="true" pageId="18" pageNumber="36">R. adami</emphasis>
|
||
</taxonomicName>
|
||
by a smaller and concave-sided lancet, a less arched connecting process, a near-parallel sided interpterygoid groove, differing bacular morphologies, external and craniodental measurements, including palate length and inflation breadth (
|
||
<bibRefCitation author="KOCK, D. & G. CSORBA & K. M. HOWELL" pageId="18" pageNumber="36" pagination="233 - 239" refId="ref16521" refString="KOCK, D., G. CSORBA, and K. M. HOWELL. 2000. Rhinolophus maendeleo n. sp. from Tanzania, a horseshoe bat noteworthy for its systematics and biogeography. Senckenbergiana Biologica, 80: 233 - 239." type="journal article" year="2000">
|
||
Kock
|
||
<emphasis box="[689,756,1223,1249]" italics="true" pageId="18" pageNumber="36">et al.</emphasis>
|
||
, 2000
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="18.[151,764,843,1973]" pageId="18" pageNumber="36">
|
||
When compared to members of the
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[653,754,1292,1318]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[653,754,1292,1318]" italics="true" pageId="18" pageNumber="36">capensis</emphasis>
|
||
</taxonomicName>
|
||
- group,
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[234,347,1326,1352]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[234,347,1326,1352]" italics="true" pageId="18" pageNumber="36">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[355,445,1326,1352]" pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is larger than
|
||
<taxonomicName box="[611,756,1326,1352]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="simulator">
|
||
<emphasis box="[611,756,1326,1352]" italics="true" pageId="18" pageNumber="36">R. simulator</emphasis>
|
||
</taxonomicName>
|
||
, with non-overlapping forearm, tibia and wing bone lengths (
|
||
<tableCitation box="[252,342,1395,1421]" captionStart="TABLE" captionStartId="13.[99,112,574,595]" captionTargetPageId="13" captionText="TABLE 3. External measurements of specimens of R. namuli sp. nov., R. maendeleo and R. simulator. Measurement with an asterisk in the header have non-overlapping ranges between R. namuli sp. nov. and R. maendeleo. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’" httpUri="http://table.plazi.org/id/2CF56638FFE1FFA8FFFCFDBDF895FD76" pageId="18" pageNumber="36" tableUuid="2CF56638FFE1FFA8FFFCFDBDF895FD76">Table 3</tableCitation>
|
||
) and several non-overlapping craniodental measurements, such as greatest skull length, mastoid width, zygomatic width and lower toothrow length (
|
||
<tableCitation box="[351,442,1499,1525]" captionStart="TABLE" captionStartId="15.[99,112,575,596]" captionTargetPageId="15" captionText="TABLE 4. Craniodental measurements of specimens of R. namuli sp. nov., R. maendeleo and R. simulator. Measurement with an asterisk in the header have non-overlapping ranges between R. namuli sp. nov. and R. maendeleo. DM = Durban Natural Science Museum; MHNG = Muséum d’Histoire Naturelle de Genève; MoM = National Museums of Malawi; SMF = Senckenberg Museum Frankfurt. * = accession numbers unknown for MoM, field codes used. Abbreviations are explained in the methods under ‘Character measurements’" pageId="18" pageNumber="36">Table 4</tableCitation>
|
||
). Based on a comparison of published measurements of
|
||
<emphasis box="[504,757,1533,1559]" italics="true" pageId="18" pageNumber="36">
|
||
<taxonomicName authorityName="Thomas" authorityYear="1904" box="[504,605,1533,1559]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="denti">R. denti</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Gough" authorityYear="1908" box="[626,757,1533,1559]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="swinnyi">R. swinnyi</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName box="[151,323,1568,1594]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="gorongosae">
|
||
<emphasis box="[151,323,1568,1594]" italics="true" pageId="18" pageNumber="36">R. gorongosae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Roberts" authorityYear="1946" box="[385,535,1568,1594]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="rhodesiae">
|
||
<emphasis box="[385,535,1568,1594]" italics="true" pageId="18" pageNumber="36">R. rhodesiae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[552,661,1568,1594]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[552,661,1568,1594]" italics="true" pageId="18" pageNumber="36">R namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[671,764,1568,1594]" pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is much larger and more robust as highlighted by non-overlapping forearm length and greatest skull length. It also echolocates at a much lower frequency that any of these species (Table 2).
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[650,763,1706,1732]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[650,763,1706,1732]" italics="true" pageId="18" pageNumber="36">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[151,246,1740,1766]" pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has more inflated anterior nasal swellings than members of the
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[420,521,1775,1801]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[420,521,1775,1801]" italics="true" pageId="18" pageNumber="36">capensis</emphasis>
|
||
</taxonomicName>
|
||
-group (Supplementary Appendix S2
|
||
<figureCitation box="[363,454,1809,1835]" captionStart="FIG" captionStartId="14.[151,164,1031,1052]" captionTargetBox="[196,1383,174,1015]" captionTargetId="figure-411@14.[196,1383,174,1016]" captionTargetPageId="14" captionText="FIG. 6. Cranium in dorsoventral (top), lateral (middle) and ventrodorsal (bottom; only for MHNG 1971.168) views of holotypes of R. namuli sp. nov. (left) and R. maendeleo (right)" figureDoi="http://doi.org/10.5281/zenodo.7859662" httpUri="https://zenodo.org/record/7859662/files/figure.png" pageId="18" pageNumber="36">Fig. A6</figureCitation>
|
||
). Finally, the bacular morphology of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[289,404,1844,1870]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[289,404,1844,1870]" italics="true" pageId="18" pageNumber="36">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[414,506,1844,1870]" pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
sets the species apart from all potentially similar species from the
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[690,763,1878,1904]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[690,763,1878,1904]" italics="true" pageId="18" pageNumber="36">adami</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[203,304,1913,1939]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[203,304,1913,1939]" italics="true" pageId="18" pageNumber="36">capensis</emphasis>
|
||
</taxonomicName>
|
||
groups to which it bears closest morphological affinity.
|
||
</paragraph>
|
||
<paragraph blockId="18.[809,1422,843,1973]" pageId="18" pageNumber="36">
|
||
Summarizing the differences highlighted above, the PCA results comparing
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[1155,1275,878,904]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[1155,1275,878,904]" italics="true" pageId="18" pageNumber="36">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1289,1385,878,904]" pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
to both
|
||
<taxonomicName box="[871,1033,912,938]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[871,1033,912,938]" italics="true" pageId="18" pageNumber="36">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[1096,1244,912,938]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="simulator">
|
||
<emphasis box="[1096,1244,912,938]" italics="true" pageId="18" pageNumber="36">R. simulator</emphasis>
|
||
</taxonomicName>
|
||
(the two most similar species from the
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[1096,1169,947,973]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[1096,1169,947,973]" italics="true" pageId="18" pageNumber="36">adami</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[1226,1327,947,973]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[1226,1327,947,973]" italics="true" pageId="18" pageNumber="36">capensis</emphasis>
|
||
</taxonomicName>
|
||
groups, respectively) illustrate the distinct position of the three species (
|
||
<figureCitation box="[979,1053,1016,1042]" captionStart="FIG" captionStartId="7.[151,164,1894,1915]" captionTargetBox="[164,1399,1144,1876]" captionTargetId="figure-494@7.[154,1407,1134,1884]" captionTargetPageId="7" captionText="FIG. 2. PCA biplot for selected (A) craniodental and (B) external measurements. Ellipses represent 95% confidence intervals around each species group (not possible for R. maendeleo due to the low sample size). Large symbols represent the species mean value. Percentage values on each axis are the variances explained. For specimen accession numbers and input data, see Tables 3 and 4" figureDoi="http://doi.org/10.5281/zenodo.7859654" httpUri="https://zenodo.org/record/7859654/files/figure.png" pageId="18" pageNumber="36">Fig. 2</figureCitation>
|
||
). While 95% confidence intervals (CIs) for the first two PCs overlap slightly between
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[887,1001,1085,1111]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[887,1001,1085,1111]" italics="true" pageId="18" pageNumber="36">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1010,1101,1085,1111]" pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName box="[1159,1304,1085,1111]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="simulator">
|
||
<emphasis box="[1159,1304,1085,1111]" italics="true" pageId="18" pageNumber="36">R. simulator</emphasis>
|
||
</taxonomicName>
|
||
, such uncertainty is to be expected given the small sample size. For
|
||
<taxonomicName box="[930,1097,1154,1180]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="maendeleo">
|
||
<emphasis box="[930,1097,1154,1180]" italics="true" pageId="18" pageNumber="36">R. maendeleo</emphasis>
|
||
</taxonomicName>
|
||
, the sample size was too small to estimate CIs. The first two PCs explained a large amount of variance in the data, with a cumulative 76% for craniodental measurements and 82.2% for external measurements.
|
||
</paragraph>
|
||
<paragraph blockId="18.[809,1422,843,1973]" pageId="18" pageNumber="36">
|
||
Phylogenetic reconstructions based on the CYT-B gene (
|
||
<figureCitation box="[974,1045,1361,1387]" captionStart="FIG" captionStartId="9.[151,164,1836,1857]" captionTargetBox="[193,1372,177,1809]" captionTargetId="graphics-12@9.[194,1370,177,1776]" captionTargetPageId="9" captionText="FIG. 3. Maximum likelihood tree based on CYT-B sequences and reconstructed with IQ-tree. Sequences of R. hipposideros were used as outgroups. Bootstrap support levels are explained in the box; the scale indicates nucleotide substitutions per site. Newly sequenced taxa are associated with their GenBank accession numbers (see Table 2 and Supplementary Appendix S3 for more details), while the species groups to which they belong appear to the right of the tree. The clade representing the new species (sp. nov.) is highlighted in grey" figureDoi="http://doi.org/10.5281/zenodo.7859656" httpUri="https://zenodo.org/record/7859656/files/figure.png" pageId="18" pageNumber="36">Fig. 3</figureCitation>
|
||
) demonstrate that
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[1265,1379,1361,1387]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[1265,1379,1361,1387]" italics="true" pageId="18" pageNumber="36">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
belong to the
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[1036,1137,1395,1421]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[1036,1137,1395,1421]" italics="true" pageId="18" pageNumber="36">capensis</emphasis>
|
||
</taxonomicName>
|
||
-group with strong support. Compared to the closely related and sympatric
|
||
<emphasis box="[809,1090,1464,1490]" italics="true" pageId="18" pageNumber="36">
|
||
<taxonomicName box="[809,957,1464,1490]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="simulator">R. simulator</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[973,1090,1464,1490]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">R. namuli</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[1102,1196,1464,1490]" pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is genetically distinct, confirming the morphological differences presented above. In the future, additional studies on the phylogenetic relationships between the species of the
|
||
<taxonomicName authorityName="Aellen & Brosset" authorityYear="1968" box="[862,935,1602,1628]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="adami">
|
||
<emphasis box="[862,935,1602,1628]" italics="true" pageId="18" pageNumber="36">adami</emphasis>
|
||
</taxonomicName>
|
||
- and
|
||
<taxonomicName authorityName="Lichtenstein" authorityYear="1823" box="[1022,1123,1602,1628]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis box="[1022,1123,1602,1628]" italics="true" pageId="18" pageNumber="36">capensis</emphasis>
|
||
</taxonomicName>
|
||
-group would be desirable to ascertain relationships or test their taxonomic validity.
|
||
</paragraph>
|
||
<paragraph blockId="18.[809,1422,843,1973]" pageId="18" pageNumber="36">
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[847,1083,1706,1732]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[847,1083,1706,1732]" italics="true" pageId="18" pageNumber="36">Rhinolophus namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1090,1180,1706,1732]" pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
echolocates at a frequency that is not shared by other species in the region (
|
||
<tableCitation box="[875,965,1775,1801]" captionStart="TABLE" captionStartId="16.[151,164,1459,1480]" captionTargetPageId="16" captionText="TABLE 5. Echolocation call parameters of R. namuli sp. nov. and species with similar echolocation peak frequency from nearby localities. Table highlights mean peak frequency (PF), standard deviation (SD), range and number of individuals recorded (n). Data collected by M. Curran and M. Kopp (unpublished material). MZ = Mozambique, MW = Malawi" httpUri="http://table.plazi.org/id/2CF56638FFFCFFB5FF08FA30FA0FF9FA" pageId="18" pageNumber="36" tableUuid="2CF56638FFFCFFB5FF08FA30FA0FF9FA">Table 5</tableCitation>
|
||
). At a mean frequency of 76.9 kHz, the calls of
|
||
<taxonomicName authority="Curran & Kopp & Ruedi & Bayliss, 2022" authorityName="Curran & Kopp & Ruedi & Bayliss" authorityYear="2022" box="[912,1031,1809,1835]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="namuli" status="sp. nov.">
|
||
<emphasis box="[912,1031,1809,1835]" italics="true" pageId="18" pageNumber="36">R. namuli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1044,1140,1809,1835]" pageId="18" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs markedly from
|
||
<taxonomicName box="[809,955,1844,1870]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="simulator">
|
||
<emphasis box="[809,955,1844,1870]" italics="true" pageId="18" pageNumber="36">R. simulator</emphasis>
|
||
</taxonomicName>
|
||
(86 kHz),
|
||
<taxonomicName box="[1088,1218,1844,1870]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="clivosus">
|
||
<emphasis box="[1088,1218,1844,1870]" italics="true" pageId="18" pageNumber="36">R. clivosus</emphasis>
|
||
</taxonomicName>
|
||
(80.5–81.3 kHz) and
|
||
<taxonomicName box="[862,1015,1878,1904]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="18" pageNumber="36" phylum="Chordata" rank="species" species="fumigatus">
|
||
<emphasis box="[862,1015,1878,1904]" italics="true" pageId="18" pageNumber="36">R. fumigatus</emphasis>
|
||
</taxonomicName>
|
||
(58.4 kHz), the three most similarly echolocating species in our dataset for the region (
|
||
<tableCitation box="[877,969,1947,1973]" captionStart="TABLE" captionStartId="16.[151,164,1459,1480]" captionTargetPageId="16" captionText="TABLE 5. Echolocation call parameters of R. namuli sp. nov. and species with similar echolocation peak frequency from nearby localities. Table highlights mean peak frequency (PF), standard deviation (SD), range and number of individuals recorded (n). Data collected by M. Curran and M. Kopp (unpublished material). MZ = Mozambique, MW = Malawi" httpUri="http://table.plazi.org/id/2CF56638FFFCFFB5FF08FA30FA0FF9FA" pageId="18" pageNumber="36" tableUuid="2CF56638FFFCFFB5FF08FA30FA0FF9FA">Table 5</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |