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<document ID-DOI="http://doi.org/10.5281/zenodo.3812747" ID-GBIF-Dataset="363f3d76-f265-4279-bfc9-266bd2616a33" ID-GBIF-Taxon="163309675" ID-Zenodo-Dep="3812747" ID-ZooBank="urn:lsid:zoobank.org:pub:63D950B1-10B3-4EC1-B4E6-8558F5618DF6" LSID="urn:lsid:zoobank.org:act:C82E3A75-96FF-4D2A-9B52-3ABF4B58BC2B" checkinTime="1585401867630" checkinUser="plazi" docAuthor="Branch, William R., Conradie, Werner, Pinto, Pedro Vaz &amp; Tolley, Krystal A." docDate="2019" docId="A153460A1C13F365848AF8B2FC2E6327" docLanguage="en" docName="ARC_13_2_[Special_Section]_82-95_e199.pdf" docOrigin="Amphibian &amp; Reptile Conservation 13 (2)" docStyle="DocumentStyle{}" docTitle="Nucras broadleyi Branch &amp; Conradie &amp; Pinto &amp; Tolley 2019, sp. nov." docType="treatment" docUuid="C82E3A75-96FF-4D2A-9B52-3ABF4B58BC2B" docUuidSource="ZooBank" docVersion="6" lastPageId="9" lastPageNumber="91" masterDocId="5D6A3E721C17F36C8407FF81FF906471" masterDocTitle="Another Angolan Namib endemic species: a new Nucras Gray, 1838 (Squamata: Lacertidae) from south-western Angola" masterLastPageNumber="95" masterPageNumber="82" pageId="4" pageNumber="86" updateTime="1643473473503" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Another Angolan Namib endemic species: a new Nucras Gray, 1838 (Squamata: Lacertidae) from south-western Angola</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Branch, William R.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Conradie, Werner</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pinto, Pedro Vaz</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tolley, Krystal A.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Amphibian &amp; Reptile Conservation</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-11-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>13</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
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<mods:start>82</mods:start>
<mods:end>95</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3731977</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3812747" ID-GBIF-Taxon="163309675" ID-Zenodo-Dep="3812747" LSID="urn:lsid:zoobank.org:act:C82E3A75-96FF-4D2A-9B52-3ABF4B58BC2B" httpUri="http://treatment.plazi.org/id/A153460A1C13F365848AF8B2FC2E6327" lastPageId="9" lastPageNumber="91" pageId="4" pageNumber="86">
<subSubSection box="[141,415,1842,1866]" pageId="4" pageNumber="86" type="nomenclature">
<paragraph blockId="4.[141,415,1842,1898]" box="[141,415,1842,1866]" pageId="4" pageNumber="86">
<heading bold="true" box="[141,415,1842,1866]" fontSize="10" level="1" pageId="4" pageNumber="86" reason="6">
<emphasis bold="true" box="[141,415,1842,1866]" pageId="4" pageNumber="86">
<taxonomicName authority="Branch &amp; Conradie &amp; Pinto &amp; Tolley, 2019" authorityName="Branch &amp; Conradie &amp; Pinto &amp; Tolley" authorityYear="2019" box="[141,325,1842,1866]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis bold="true" box="[141,325,1842,1866]" italics="true" pageId="4" pageNumber="86">Nucras broadleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[331,415,1842,1866]" pageId="4" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[141,408,1874,1898]" pageId="4" pageNumber="86" type="vernacular_name">
<paragraph blockId="4.[141,415,1842,1898]" box="[141,408,1874,1898]" pageId="4" pageNumber="86">
<vernacularName box="[141,408,1874,1898]" pageId="4" pageNumber="86">Angolan Sandveld Lizard</vernacularName>
</paragraph>
</subSubSection>
<subSubSection box="[141,745,1941,1960]" pageId="4" pageNumber="86" type="nomenclature">
<paragraph blockId="4.[141,745,1941,1960]" box="[141,745,1941,1960]" pageId="4" pageNumber="86">
<emphasis bold="true" box="[141,745,1941,1960]" pageId="4" pageNumber="86">
<uri box="[141,745,1941,1960]" pageId="4" pageNumber="86">
urn:lsid:zoobank.org:act:
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</uri>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection box="[791,931,175,199]" pageId="4" pageNumber="86" type="description">
<paragraph blockId="4.[791,931,175,199]" box="[791,931,175,199]" pageId="4" pageNumber="86">
(
<figureCitation box="[798,924,175,199]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="9.[141,176,1075,1097]" captionStartId-1="10.[141,176,1072,1094]" captionStartId-2="11.[139,174,843,865]" captionTargetBox-0="[141,1394,141,1063]" captionTargetBox-1="[141,1394,141,1060]" captionTargetBox-2="[146,1391,140,834]" captionTargetId-0="figure@9.[141,1394,141,1066]" captionTargetId-1="figure@10.[141,1394,139,1064]" captionTargetId-2="figure@11.[139,1392,133,837]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Fig. 4. Nucras broadleyi sp. nov. A holotype, adult male, PEM R24005 (AG 18) in life; B general habitat photo of type locality, 10 km west of Lola, edge of Bentiaba River valley, Namibe Province, Angola; C lateral close-up of head of holotype; D dorsal close-up of head of holotype; E ventral close-up of head of holotype (Photos: Bill Branch)." captionText-1="Fig. 5. Nucras broadleyi sp. nov. A paratype, adult female, PEM R24157 (AG 166) dorsal view; B ventral view; C dorsal close-up of head of paratype; D ventral close-up of head of paratype; E lateral close-up of head of paratype; F general habitat photo of type locality, 8.8 km southwest of Farm Mucongo, Namibe Province, Angola (Photos: Bill Branch)." captionText-2="Fig. 6. Variation of Nucras broadleyi nov. sp. dorsal color pattern.A TM 40392 from “34 km S of Moçâmedes to Porto Alexandre;” B BM 1970.6.29.10 from Ponang Kuma (=Donquena); C MHNC 91.0524 from Capelongo; D MD 1967 from “km 34 de la route de Moçâmedes à Sa da Bandeira” (Photos: A,B Bill Branch, C, D Luis Ceríaco)." figureDoi-0="http://doi.org/10.5281/zenodo.3731985" figureDoi-1="http://doi.org/10.5281/zenodo.3731987" figureDoi-2="http://doi.org/10.5281/zenodo.3731989" httpUri-0="https://zenodo.org/record/3731985/files/figure.png" httpUri-1="https://zenodo.org/record/3731987/files/figure.png" httpUri-2="https://zenodo.org/record/3731989/files/figure.png" pageId="4" pageNumber="86">Figures 46</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="86" type="reference_group">
<paragraph blockId="4.[791,1396,239,520]" pageId="4" pageNumber="86">
<emphasis bold="true" box="[791,937,239,263]" pageId="4" pageNumber="86">Chersonymy.</emphasis>
<taxonomicName authority="(Bocage 1895: 30)" baseAuthorityName="Bocage" baseAuthorityPageNumber="30" baseAuthorityYear="1895" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="variety" species="tessellata" variety="taeniolata">
<emphasis box="[948,1133,239,263]" italics="true" pageId="4" pageNumber="86">Nucras tessellata</emphasis>
var.
<emphasis box="[1192,1297,239,263]" italics="true" pageId="4" pageNumber="86">taeniolata</emphasis>
(
<bibRefCitation author="Bocage JVB" bookContentInfo="203 p." journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="4" pageNumber="86" refId="ref8226" refString="Bocage JVB. 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal. 203 p., 20 pls." title="Herpetologie d'Angola et du Congo" type="book" year="1895">Bocage 1895: 30</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName authority="(Boulenger 1910: 474)" baseAuthorityName="Boulenger" baseAuthorityPageNumber="474" baseAuthorityYear="1910" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="variety" species="tessellata" variety="taeniolata">
<emphasis box="[915,1100,271,295]" italics="true" pageId="4" pageNumber="86">Nucras tessellata</emphasis>
var.
<emphasis box="[1161,1266,271,295]" italics="true" pageId="4" pageNumber="86">taeniolata</emphasis>
(
<bibRefCitation author="Boulenger GA" journalOrPublisher="Annals of the South African Museum" pageId="4" pageNumber="86" pagination="455 - 538" part="5" refId="ref8255" refString="Boulenger GA. 1910. A revised list of South African reptiles and batrachians, with synoptic tables, special reference of specimens in the South African Museum, and descriptions of new species. Annals of the South African Museum 5: 455 - 538." title="A revised list of South African reptiles and batrachians, with synoptic tables, special reference of specimens in the South African Museum, and descriptions of new species" type="journal article" year="1910">Boulenger 1910: 474</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName authority="(Boulenger 1917: 210)" baseAuthorityName="Boulenger" baseAuthorityPageNumber="210" baseAuthorityYear="1917" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="variety" species="tessellata" variety="holubi">
<emphasis box="[915,1093,303,327]" italics="true" pageId="4" pageNumber="86">Nucras tessellata</emphasis>
var.
<emphasis box="[1141,1207,303,327]" italics="true" pageId="4" pageNumber="86">holubi</emphasis>
(
<bibRefCitation author="Boulenger GA" journalOrPublisher="Annals of the South African Museum" pageId="4" pageNumber="86" pagination="195 - 216" part="13" refId="ref8302" refString="Boulenger GA. 1917. A revision of the lizards of the genus Nucras, Gray. Annals of the South African Museum 13: 195 - 216." title="A revision of the lizards of the genus Nucras, Gray" type="journal article" year="1917">Boulenger 1917: 210</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName authority="(Boulenger 1920: 20)" baseAuthorityName="Boulenger" baseAuthorityPageNumber="20" baseAuthorityYear="1920" box="[852,1390,335,360]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="variety" species="intertexta" variety="holubi">
<emphasis box="[852,1034,336,359]" italics="true" pageId="4" pageNumber="86">Nucras intertexta</emphasis>
var.
<emphasis box="[1088,1154,335,359]" italics="true" pageId="4" pageNumber="86">holubi</emphasis>
(
<bibRefCitation author="Boulenger GA" bookContentInfo="352 p." box="[1169,1382,335,360]" journalOrPublisher="Trustees of the British Museum (Natural History), London, United Kingdom" pageId="4" pageNumber="86" refId="ref8331" refString="Boulenger GA. 1920. Monograph of the Lacertidae. Volume 1. Trustees of the British Museum (Natural History), London, United Kingdom. x + 352 p." title="Monograph of the Lacertidae. Volume 1" type="book" year="1920">Boulenger 1920: 20</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[791,973,367,391]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="species" species="tessellata">
<emphasis box="[791,973,367,391]" italics="true" pageId="4" pageNumber="86">Nucras tessellata</emphasis>
</taxonomicName>
(
<bibRefCitation author="Monard A." box="[989,1181,367,392]" journalOrPublisher="Arquivos do Museu Bocage" pageId="4" pageNumber="86" pagination="19 - 154" part="8" refId="ref9569" refString="Monard A. 1937. Contribution a l'herpetologie d'Angola. Arquivos do Museu Bocage 8: 19 - 154." title="Contribution a l'herpetologie d'Angola" type="journal article" year="1937">Monard 1937: 73</bibRefCitation>
;
<bibRefCitation author="Laurent RF" box="[1195,1383,367,391]" journalOrPublisher="Publicacoes Culturais Com- panhia de Diamantes de Angola" pageId="4" pageNumber="86" pagination="1 - 165" part="67" refId="ref9224" refString="Laurent RF. 1964. Reptiles et amphibiens de l'Angola (troiseme contribution). Publicacoes Culturais Com- panhia de Diamantes de Angola 67: 1 - 165." title="Reptiles et amphibiens de l'Angola (troiseme contribution)" type="journal article" year="1964">Laurent 1964: 56</bibRefCitation>
),
<taxonomicName authority="(Broadley 1965: 23)" baseAuthorityName="Broadley" baseAuthorityPageNumber="23" baseAuthorityYear="1965" box="[791,1190,399,424]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="species" species="ornata">
<emphasis box="[791,949,400,423]" italics="true" pageId="4" pageNumber="86">Nucras ornata</emphasis>
(
<bibRefCitation author="Broadley DG" box="[970,1181,399,424]" journalOrPublisher="Journal of the Herpetological Association of Africa" pageId="4" pageNumber="86" pagination="18 - 23" part="1" refId="ref8624" refString="Broadley DG. 1965. Some problems presented by sand lizards of the Nucras tessellata group. Journal of the Herpetological Association of Africa 1: 18 - 23." title="Some problems presented by sand lizards of the Nucras tessellata group" type="journal article" year="1965">Broadley 1965: 23</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[1208,1395,399,423]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="species" species="tessellata">
<emphasis box="[1208,1395,399,423]" italics="true" pageId="4" pageNumber="86">Nucras tessellata</emphasis>
</taxonomicName>
(
<bibRefCitation author="Broadley DG" box="[798,996,431,456]" journalOrPublisher="Arnoldia" pageId="4" pageNumber="86" pagination="1 - 35" part="20" refId="ref8654" refString="Broadley DG. 1972. A review of the Nucras tessellata group (Sauria: Lacertidae). Arnoldia 20: 1 - 35." title="A review of the Nucras tessellata group (Sauria: Lacertidae)" type="journal article" year="1972">Broadley 1972: 30</bibRefCitation>
;
<bibRefCitation author="Ceriaco LMP &amp; de Sa SC &amp; Bandeira S &amp; Valerio H &amp; Stanley EL &amp; Kuhn AL &amp; Marques M &amp; Vindum JV &amp; Blackburn DC &amp; Bauer AM" box="[1007,1246,431,455]" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="4" pageNumber="86" pagination="15 - 61" part="63" refId="ref8814" refString="Ceriaco LMP, de Sa SC, Bandeira S, Valerio H, Stanley EL, Kuhn AL, Marques M, Vindum JV, Blackburn DC, Bauer AM. 2016. Herpetological survey of Iona National Park and Namibe Regional Natural Park, with a synoptic list of the amphibians and reptiles of Namibe Province, Southwestern Angola. Proceedings of the California Academy of Sciences 63 (2): 15 - 61." title="Herpetological survey of Iona National Park and Namibe Regional Natural Park, with a synoptic list of the amphibians and reptiles of Namibe Province, Southwestern Angola" type="journal article" year="2016">Ceríaco et al. 2016: 56</bibRefCitation>
;
<bibRefCitation author="Burger M." bookContentInfo="485 p." editor="Bates MF &amp; Branch WR &amp; Bauer AM &amp; Burger M &amp; Marais J &amp; Alexander GJ &amp; de Villiers MS" journalOrPublisher="Suricata 1. South African National Biodiversity Institute, Pretoria, South Africa" pageId="4" pageNumber="86" pagination="171 - 172" refId="ref8679" refString="Burger M. 2014. Nucras tessellata (A. Smith, 1838). Pp. 171 - 172 In: Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Editors, Bates MF, Branch WR, Bauer AM, Burger M, Marais J, Alexander GJ, de Villiers MS. Suricata 1. South African National Biodiversity Institute, Pretoria, South Africa. 485 p." title="Nucras tessellata (A. Smith, 1838). Pp" type="book chapter" volumeTitle="Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland" year="2014">Burger 2014: 171</bibRefCitation>
),
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[856,1088,463,487]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="species" species="tessellata">
<emphasis box="[856,931,464,487]" italics="true" pageId="4" pageNumber="86">Nucras</emphasis>
aff.
<emphasis box="[990,1088,463,487]" italics="true" pageId="4" pageNumber="86">tessellata</emphasis>
</taxonomicName>
(
<bibRefCitation author="Marques MP &amp; Ceriaco LMP &amp; Blackburn DC &amp; Bauer AM" box="[1106,1390,463,488]" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="4" pageNumber="86" pagination="1 - 501" part="65" refId="ref9357" refString="Marques MP, Ceriaco LMP, Blackburn DC, Bauer AM 2018. Diversity and distribution of the amphibians and terrestrial reptiles of Angola: atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences Series 4, 65 (Supplement II): 1 - 501." title="Diversity and distribution of the amphibians and terrestrial reptiles of Angola: atlas of historical and bibliographic records (1840 - 2017)" type="journal article" year="2018">Marques et al. 2018: 221</bibRefCitation>
;
<bibRefCitation author="Branch WR &amp; Baptista N &amp; Vaz Pinto P &amp; Conradie W." bookContentInfo="552 p." box="[791,1037,495,520]" editor="Huntley BJ &amp; Ferrand N &amp; Russo V &amp; Lages F." journalOrPublisher="Springer Open, Cham, Switzerland" pageId="4" pageNumber="86" pagination="283 - 326" refId="ref8545" refString="Branch WR, Baptista N, Vaz Pinto P, Conradie W. 2019. The reptiles of Angola - History, updated checklists, endemism, hot spots, and future directions for research. Pp. 283 - 326 In: Biodiversity of Angola. Science and Conservation: A Modern Synthesis. Editors, Huntley BJ, Ferrand N, Russo V, Lages F. Springer Open, Cham, Switzerland. 552 p." title="The reptiles of Angola - History, updated checklists, endemism, hot spots, and future directions for research. Pp" type="book chapter" volumeTitle="Biodiversity of Angola. Science and Conservation: A Modern Synthesis" year="2019">Branch et al. 2019: 317</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="86" type="materials_examined">
<paragraph blockId="4.[791,1395,559,647]" pageId="4" pageNumber="86">
<emphasis bold="true" box="[791,948,559,584]" pageId="4" pageNumber="86">Type material</emphasis>
. The
<typeStatus box="[1010,1055,560,584]" pageId="4" pageNumber="86">type</typeStatus>
series is comprised of the three most recently collected specimens, which are housed in
<collectionCode box="[791,845,624,647]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="4" pageNumber="86" type="Museum">PEM</collectionCode>
and
<collectionCode box="[894,936,624,647]" country="Germany" name="Teylers Museum, Paleontologische" pageId="4" pageNumber="86">TM</collectionCode>
.
</paragraph>
<paragraph blockId="4.[791,1395,687,840]" pageId="4" pageNumber="86">
<materialsCitation ID-GBIF-Occurrence="2592348495" collectingDate="2015-11-02" collectionCode="PEM, R" collectorName="Branch, P. &amp; J. S. de Almeida" country="Angola" elevation="84802" latitude="-14.29028" location="Lola" longLatPrecision="1" longitude="13.53056" municipality="Bentiaba River valley" pageId="4" pageNumber="86" specimenCode="AG 018" specimenCount="1" stateProvince="Namibe" typeStatus="holotype">
<emphasis bold="true" box="[791,899,688,712]" pageId="4" pageNumber="86">
<typeStatus box="[791,894,688,712]" pageId="4" pageNumber="86">Holotype</typeStatus>
.
</emphasis>
A subadult male (
<collectionCode box="[1097,1152,688,712]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="4" pageNumber="86" type="Museum">PEM</collectionCode>
<accessionNumber box="[1161,1246,687,711]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R24005" pageId="4" pageNumber="86">
<collectionCode box="[1161,1179,687,711]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="86">R</collectionCode>
24005
</accessionNumber>
,
<specimenCode box="[1258,1347,688,712]" collectionCode="AG" pageId="4" pageNumber="86">AG 018</specimenCode>
),
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="86" unit="km" value="10.0">10 km</quantity>
west of
<location LSID="urn:lsid:plazi:treatment:A153460A1C13F365848AF8B2FC2E6327:2C25A1C71C13F3688790FD4EFC596696" box="[919,969,719,743]" country="Angola" latitude="-14.29028" longLatPrecision="1" longitude="13.53056" municipality="Bentiaba River valley" name="Lola" pageId="4" pageNumber="86" stateProvince="Namibe">Lola</location>
, edge of
<collectingMunicipality box="[1069,1300,719,743]" pageId="4" pageNumber="86">Bentiaba River valley</collectingMunicipality>
,
<collectingRegion country="Angola" name="Namibe" pageId="4" pageNumber="86">Namibe Province</collectingRegion>
,
<collectingCountry box="[896,973,751,775]" name="Angola" pageId="4" pageNumber="86">Angola</collectingCountry>
(
<geoCoordinate box="[988,1098,752,776]" degrees="14.29028" direction="south" orientation="latitude" pageId="4" pageNumber="86" precision="1" value="-14.29028">-14.29028</geoCoordinate>
,
<geoCoordinate box="[1109,1209,751,775]" degrees="13.53056" direction="east" orientation="longitude" pageId="4" pageNumber="86" precision="1" value="13.53056">13.53056</geoCoordinate>
, WGS
<elevation metricMagnitude="4" metricUnit="m" metricValue="8.4802" pageId="4" pageNumber="86" unit="m" value="84802.0">
<quantity box="[1287,1395,752,776]" metricMagnitude="4" metricUnit="m" metricValue="8.4802" pageId="4" pageNumber="86" unit="m" value="84802.0">84, 802 m</quantity>
asl
</elevation>
). Collected by W.
<collectionCode box="[1015,1034,784,807]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="86">R</collectionCode>
.
<collectorName box="[1045,1154,783,807]" pageId="4" pageNumber="86">Branch, P.</collectorName>
Vaz Pinto, and
<collectorName pageId="4" pageNumber="86">J.S. de Almeida</collectorName>
on
<date box="[919,1110,815,840]" pageId="4" pageNumber="86" value="2015-11-02">
<collectingDate box="[919,1110,815,840]" pageId="4" pageNumber="86" value="2015-11-02">2 November 2015</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph blockId="4.[791,1396,879,1160]" pageId="4" pageNumber="86">
<materialsCitation ID-GBIF-Occurrence="2592348493" collectingDate="2015-11-08" collectionCode="PEM, R" collectorName="Branch, P. &amp; J. S. de Almeida" country="Angola" elevation="84385" latitude="-14.80167" location="Farm Mucungo" longLatPrecision="1" longitude="12.41917" pageId="4" pageNumber="86" specimenCode="AG 166" specimenCount="1" stateProvince="Namibe" typeStatus="paratype">
<emphasis bold="true" box="[791,954,879,904]" pageId="4" pageNumber="86">
<typeStatus box="[791,902,880,904]" pageId="4" pageNumber="86">Paratypes</typeStatus>
(2).
</emphasis>
a) A subadult female (
<collectionCode box="[1234,1289,880,904]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="4" pageNumber="86" type="Museum">PEM</collectionCode>
<accessionNumber box="[1305,1390,879,903]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R24157" pageId="4" pageNumber="86">
<collectionCode box="[1305,1323,879,903]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="86">R</collectionCode>
24157
</accessionNumber>
,
<specimenCode box="[791,880,911,936]" collectionCode="AG" pageId="4" pageNumber="86">AG 166</specimenCode>
),
<quantity box="[902,977,911,936]" metricMagnitude="3" metricUnit="m" metricValue="8.8" pageId="4" pageNumber="86" unit="km" value="8.8">8.8 km</quantity>
southwest of
<location LSID="urn:lsid:plazi:treatment:A153460A1C13F365848AF8B2FC2E6327:2C25A1C71C13F368806CFC11FA8267D9" box="[1131,1298,912,936]" country="Angola" latitude="-14.80167" longLatPrecision="1" longitude="12.41917" name="Farm Mucungo" pageId="4" pageNumber="86" stateProvince="Namibe">Farm Mucungo</location>
,
<collectingRegion country="Angola" name="Namibe" pageId="4" pageNumber="86">Namibe Province</collectingRegion>
,
<collectingCountry box="[896,973,943,967]" name="Angola" pageId="4" pageNumber="86">Angola</collectingCountry>
(
<geoCoordinate box="[988,1098,943,967]" degrees="14.80167" direction="south" orientation="latitude" pageId="4" pageNumber="86" precision="1" value="-14.80167">-14.80167</geoCoordinate>
,
<geoCoordinate box="[1109,1209,944,968]" degrees="12.41917" direction="east" orientation="longitude" pageId="4" pageNumber="86" precision="1" value="12.41917">12.41917</geoCoordinate>
,
<potCollectionCode box="[1220,1278,944,968]" pageId="4" pageNumber="86">WGS</potCollectionCode>
<elevation metricMagnitude="4" metricUnit="m" metricValue="8.4385" pageId="4" pageNumber="86" unit="m" value="84385.0">
<quantity box="[1287,1395,943,968]" metricMagnitude="4" metricUnit="m" metricValue="8.4385" pageId="4" pageNumber="86" unit="m" value="84385.0">84, 385 m</quantity>
asl
</elevation>
). Collected by
<collectionCode box="[984,1011,976,999]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="4" pageNumber="86" type="Herbarium">W</collectionCode>
.
<collectionCode box="[1015,1034,976,999]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="86">R</collectionCode>
.
<collectorName box="[1045,1154,975,999]" pageId="4" pageNumber="86">
Branch,
<collectionCodeEnum box="[1135,1150,976,999]" pageId="4" pageNumber="86">
<collectionCode box="[1135,1150,976,999]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="86" type="Herbarium">P</collectionCode>
</collectionCodeEnum>
.
</collectorName>
Vaz Pinto, and
<collectorName pageId="4" pageNumber="86">
<collectionCode box="[1324,1337,976,1000]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14647" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="4" pageNumber="86" type="Herbarium">
<collectionCodeEnum box="[1324,1337,976,1000]" pageId="4" pageNumber="86">J</collectionCodeEnum>
</collectionCode>
.
<collectionCodeEnum box="[1341,1357,976,1000]" pageId="4" pageNumber="86">
<collectionCode box="[1341,1357,976,1000]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="4" pageNumber="86" type="Herbarium">S</collectionCode>
</collectionCodeEnum>
. de Almeida
</collectorName>
on
<date box="[922,1115,1007,1032]" pageId="4" pageNumber="86" value="2015-11-08">
<collectingDate box="[922,1115,1007,1032]" pageId="4" pageNumber="86" value="2015-11-08">
<date box="[922,1053,1007,1032]" day="08" month="11" monthName="November" pageId="4" pageNumber="86">
8
<date box="[943,1115,1007,1031]" month="11" monthName="November" pageId="4" pageNumber="86" year="2015">November 2015</date>
</date>
</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2592348492" collectingDate="1971-03-30" collectionCode="TM" collectorName="W. D. Haacke" country="Angola" county="Tombwa" latitude="-15.4822" location="An" longLatPrecision="1" longitude="12.18289" municipality="Ca" pageId="4" pageNumber="86" specimenCode="TM 40392" specimenCount="1" stateProvince="Namibe" typeStatus="paratype">
b)
<location LSID="urn:lsid:plazi:treatment:A153460A1C13F365848AF8B2FC2E6327:2C25A1C71C13F3688085FC71FB326079" box="[1154,1186,1008,1032]" country="Angola" county="Tombwa" latitude="-15.4822" longLatPrecision="1" longitude="12.18289" municipality="Ca" name="An" pageId="4" pageNumber="86" stateProvince="Namibe">An</location>
unsexed adult (
<specimenCode collectionCode="TM" country="Germany" name="Teylers Museum, Paleontologische" pageId="4" pageNumber="86">
<potCollectionCode box="[1354,1395,1008,1032]" pageId="4" pageNumber="86">
<collectionCodeEnum box="[1354,1395,1008,1032]" pageId="4" pageNumber="86">TM</collectionCodeEnum>
</potCollectionCode>
40392
</specimenCode>
), “
<locationDeviation box="[893,1167,1039,1064]" location="Mocamedes" pageId="4" pageNumber="86">
<quantity box="[893,967,1039,1064]" metricMagnitude="4" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="86" unit="km" value="34.0">34 km</quantity>
<collectionCodeEnum box="[980,995,1040,1064]" pageId="4" pageNumber="86">
<collectionCode box="[980,995,1040,1064]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="4" pageNumber="86" type="Herbarium">S</collectionCode>
</collectionCodeEnum>
of Moçâmedes
</locationDeviation>
to
<location LSID="urn:lsid:plazi:treatment:A153460A1C13F365848AF8B2FC2E6327:2C25A1C71C13F36880BBFB91FAFF6056" box="[1212,1391,1039,1063]" country="Angola" county="Tombwa" latitude="-15.4822" longLatPrecision="1" longitude="12.18289" municipality="Ca" name="Porto Alexandre" pageId="4" pageNumber="86" stateProvince="Namibe">
<collectingRegion box="[1212,1268,1040,1063]" country="Portugal" name="Porto" pageId="4" pageNumber="86">Porto</collectingRegion>
Alexandre
</location>
,
<collectingCountry box="[791,870,1071,1095]" name="Angola" pageId="4" pageNumber="86">Angola</collectingCountry>
, 1512
<collectingMunicipality box="[942,972,1071,1095]" pageId="4" pageNumber="86">Ca</collectingMunicipality>
” (=
<quantity box="[1035,1109,1071,1096]" metricMagnitude="4" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="86" unit="km" value="34.0">34 km</quantity>
<collectionCode box="[1125,1140,1072,1096]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15668" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="4" pageNumber="86" type="Herbarium">
<collectionCodeEnum box="[1125,1140,1072,1096]" pageId="4" pageNumber="86">S</collectionCodeEnum>
</collectionCode>
<collectingRegion box="[1154,1237,1071,1095]" country="Angola" name="Namibe" pageId="4" pageNumber="86">Namibe</collectingRegion>
to
<collectingCounty box="[1287,1384,1071,1095]" pageId="4" pageNumber="86">Tômbwa</collectingCounty>
),
<collectingRegion box="[791,975,1103,1127]" country="Angola" name="Namibe" pageId="4" pageNumber="86">Namibe Province</collectingRegion>
,
<collectingCountry box="[985,1062,1103,1127]" name="Angola" pageId="4" pageNumber="86">Angola</collectingCountry>
(approx.
<geoCoordinate box="[1163,1271,1103,1127]" degrees="15.48220" direction="south" orientation="latitude" pageId="4" pageNumber="86" precision="1" value="-15.4822">-15.48220</geoCoordinate>
,
<geoCoordinate box="[1282,1384,1104,1128]" degrees="12.18289" direction="east" orientation="longitude" pageId="4" pageNumber="86" precision="1" value="12.18289">12.18289</geoCoordinate>
). Collected by
<collectorName box="[928,1067,1135,1159]" pageId="4" pageNumber="86">W.D. Haacke</collectorName>
on
<date box="[1105,1267,1135,1160]" pageId="4" pageNumber="86" value="1971-03-30">
<collectingDate box="[1105,1267,1135,1160]" pageId="4" pageNumber="86" value="1971-03-30">30 March 1971</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="86" type="materials_examined">
<paragraph blockId="4.[791,1396,1199,1576]" pageId="4" pageNumber="86">
<emphasis bold="true" box="[791,1122,1199,1223]" pageId="4" pageNumber="86">Additional referred material:</emphasis>
The following additional material was used to expand the description of variation within the species: a)
<materialsCitation ID-GBIF-Occurrence="2592348491" collectingDate="1949-10-24" country="Angola" latitude="-15.03333" location="34 km from Namibe on Lubango road" longLatPrecision="1" longitude="12.41667" pageId="4" pageNumber="86" specimenCount="2" specimenCount-adult="1" specimenCount-male="1">
an
<specimenCount box="[1063,1115,1263,1287]" pageId="4" pageNumber="86" type="adult">adult</specimenCount>
<specimenCount box="[1125,1176,1263,1287]" pageId="4" pageNumber="86" type="male">male</specimenCount>
(MD 1967,
<bibRefCitation author="Laurent RF" journalOrPublisher="Publicacoes Culturais Com- panhia de Diamantes de Angola" pageId="4" pageNumber="86" pagination="1 - 165" part="67" refId="ref9224" refString="Laurent RF. 1964. Reptiles et amphibiens de l'Angola (troiseme contribution). Publicacoes Culturais Com- panhia de Diamantes de Angola 67: 1 - 165." title="Reptiles et amphibiens de l'Angola (troiseme contribution)" type="journal article" year="1964">Laurent 1964</bibRefCitation>
), “km 34 de la route de Moçâmedes à Sa da Bandeira” (=
<location LSID="urn:lsid:plazi:treatment:A153460A1C13F365848AF8B2FC2E6327:2C25A1C71C13F368872AFAB1FB606136" box="[813,1264,1327,1352]" latitude="-15.03333" longLatPrecision="1" longitude="12.41667" name="34 km from Namibe on Lubango road" pageId="4" pageNumber="86">
<quantity box="[813,888,1327,1352]" metricMagnitude="4" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="86" unit="km" value="34.0">34 km</quantity>
from Namibe on Lubango road
</location>
,
<geoCoordinate box="[1282,1391,1327,1351]" degrees="15.03333" direction="south" orientation="latitude" pageId="4" pageNumber="86" precision="1" value="-15.03333">-15.03333</geoCoordinate>
,
<geoCoordinate box="[791,892,1359,1383]" degrees="12.41667" direction="east" orientation="longitude" pageId="4" pageNumber="86" precision="1" value="12.41667">12.41667</geoCoordinate>
), collected
<date box="[1027,1221,1359,1384]" pageId="4" pageNumber="86" value="1949-10-24">
<collectingDate box="[1027,1221,1359,1384]" pageId="4" pageNumber="86" value="1949-10-24">24 October 1949</collectingDate>
</date>
</materialsCitation>
, b)
<materialsCitation ID-GBIF-Occurrence="2592348496" collectionCode="MBL" latitude="-15.01667" longLatPrecision="1" longitude="13.2" pageId="4" pageNumber="86" specimenCode="MBL 646, 647" specimenCount="1">
<specimenCode box="[1278,1391,1359,1383]" collectionCode="MBL" country="Portugal" httpUri="http://grbio.org/cool/pyk6-nt7k" name="Museu Nacional de Historia Natural" pageId="4" pageNumber="86">MBL 646</specimenCode>
,
<specimenCode box="[791,832,1391,1415]" collectionCode="MBL" country="Portugal" httpUri="http://grbio.org/cool/pyk6-nt7k" name="Museu Nacional de Historia Natural" pageId="4" pageNumber="86">647</specimenCode>
a, 647b (
<bibRefCitation author="Bocage JVB" bookContentInfo="203 p." box="[931,1120,1391,1416]" journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="4" pageNumber="86" refId="ref8226" refString="Bocage JVB. 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal. 203 p., 20 pls." title="Herpetologie d'Angola et du Congo" type="book" year="1895">Bocage 1895: 30</bibRefCitation>
) from Maconjo (approx.
<geoCoordinate box="[791,899,1423,1447]" degrees="15.01667" direction="south" orientation="latitude" pageId="4" pageNumber="86" precision="1" value="-15.01667">-15.01667</geoCoordinate>
,
<geoCoordinate box="[908,1009,1424,1448]" degrees="13.20000" direction="east" orientation="longitude" pageId="4" pageNumber="86" precision="1" value="13.2">13.20000</geoCoordinate>
), c
</materialsCitation>
)
<materialsCitation ID-GBIF-Occurrence="2592348490" collectionCode="BM" latitude="-17.01667" longLatPrecision="1" longitude="14.71667" pageId="4" pageNumber="86" specimenCount="1">
<collectionCode box="[1052,1093,1424,1447]" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="86">BM</collectionCode>
1970.6.29.1011 (
<bibRefCitation author="Boulenger GA" journalOrPublisher="Annals of the South African Museum" pageId="4" pageNumber="86" pagination="455 - 538" part="5" refId="ref8255" refString="Boulenger GA. 1910. A revised list of South African reptiles and batrachians, with synoptic tables, special reference of specimens in the South African Museum, and descriptions of new species. Annals of the South African Museum 5: 455 - 538." title="A revised list of South African reptiles and batrachians, with synoptic tables, special reference of specimens in the South African Museum, and descriptions of new species" type="journal article" year="1910">Boulenger 1910: 474</bibRefCitation>
) from Ponang Kuma (= Donguena, approx.
<geoCoordinate box="[791,899,1487,1511]" degrees="17.01667" direction="south" orientation="latitude" pageId="4" pageNumber="86" precision="1" value="-17.01667">-17.01667</geoCoordinate>
,
<geoCoordinate box="[914,1015,1487,1511]" degrees="14.71667" direction="east" orientation="longitude" pageId="4" pageNumber="86" precision="1" value="14.71667">14.71667</geoCoordinate>
), and d
</materialsCitation>
)
<materialsCitation ID-GBIF-Occurrence="2592348494" collectingDate="1933-04" collectionCode="MHNC" latitude="-14.88333" longLatPrecision="1" longitude="15.083333" pageId="4" pageNumber="86" specimenCode="MHNC 91.0524" specimenCount="1">
<specimenCode box="[1118,1292,1487,1512]" collectionCode="MHNC" country="0" httpUri="http://grbio.org/cool/25c9-tsqt" name="Museo de Historia Natural de Concepcion (Chile)" pageId="4" pageNumber="86">MHNC 91.0524</specimenCode>
(
<bibRefCitation author="Monard A." journalOrPublisher="Arquivos do Museu Bocage" pageId="4" pageNumber="86" pagination="19 - 154" part="8" refId="ref9569" refString="Monard A. 1937. Contribution a l'herpetologie d'Angola. Arquivos do Museu Bocage 8: 19 - 154." title="Contribution a l'herpetologie d'Angola" type="journal article" year="1937">Monard 1937</bibRefCitation>
) from Capelongo (approx.
<geoCoordinate box="[1146,1254,1520,1544]" degrees="14.88333" direction="south" orientation="latitude" pageId="4" pageNumber="86" precision="1" value="-14.88333">-14.88333</geoCoordinate>
,
<geoCoordinate box="[1269,1384,1519,1543]" degrees="15.083333" direction="east" orientation="longitude" pageId="4" pageNumber="86" precision="1" value="15.083333">15.083333</geoCoordinate>
), collected
<date box="[889,1005,1551,1576]" pageId="4" pageNumber="86" value="1933-04">
<collectingDate box="[889,1005,1551,1576]" pageId="4" pageNumber="86" value="1933-04">April 1933</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="86" type="etymology">
<paragraph blockId="4.[791,1395,1615,1799]" pageId="4" pageNumber="86">
<emphasis bold="true" box="[791,915,1616,1640]" pageId="4" pageNumber="86">Etymology.</emphasis>
The specific epithet is a patronym in honor of Donald G. Broadley for his numerous contributions to the herpetofauna of Africa. Don (as most of us knew him) was the first to recognize the Angolan population as a separate species (
<bibRefCitation author="Broadley DG" box="[882,1044,1743,1768]" journalOrPublisher="Arnoldia" pageId="4" pageNumber="86" pagination="1 - 35" part="20" refId="ref8654" refString="Broadley DG. 1972. A review of the Nucras tessellata group (Sauria: Lacertidae). Arnoldia 20: 1 - 35." title="A review of the Nucras tessellata group (Sauria: Lacertidae)" type="journal article" year="1972">Broadley 1972</bibRefCitation>
). The name is constructed in the masculine genitive.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="89" pageId="4" pageNumber="86" type="diagnosis">
<paragraph blockId="4.[791,1395,1839,1960]" lastBlockId="5.[141,746,1704,1952]" lastPageId="5" lastPageNumber="87" pageId="4" pageNumber="86">
<emphasis bold="true" box="[791,903,1839,1863]" pageId="4" pageNumber="86">Diagnosis.</emphasis>
Assigned to
<taxonomicName authorityName="Gray" authorityYear="1838" box="[1036,1111,1840,1863]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="genus">
<emphasis box="[1036,1111,1840,1863]" italics="true" pageId="4" pageNumber="86">Nucras</emphasis>
</taxonomicName>
due to a well-defined collar (absent in
<taxonomicName authorityName="W.Peters" authorityYear="1854" box="[897,1015,1871,1895]" class="Reptilia" family="Lacertidae" genus="Ichnotropis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="genus">
<emphasis box="[897,1015,1871,1895]" italics="true" pageId="4" pageNumber="86">Ichnotropis</emphasis>
</taxonomicName>
), toes not serrated or fringed (versus serrated or fringed in
<taxonomicName authorityName="Gray" authorityYear="1838" box="[1012,1096,1903,1927]" class="Reptilia" family="Lacertidae" genus="Meroles" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="genus">
<emphasis box="[1012,1096,1903,1927]" italics="true" pageId="4" pageNumber="86">Meroles</emphasis>
</taxonomicName>
), subdigital lamellae smooth (versus keeled in
<taxonomicName box="[978,1102,1935,1959]" class="Reptilia" family="Lacertidae" genus="Pedioplanis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="genus">
<emphasis box="[978,1102,1935,1959]" italics="true" pageId="4" pageNumber="86">Pedioplanis</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Fitzinger" authorityYear="1843" box="[1156,1270,1935,1959]" class="Reptilia" family="Lacertidae" genus="Heliobolus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="86" phylum="Chordata" rank="genus">
<emphasis box="[1156,1270,1935,1959]" italics="true" pageId="4" pageNumber="86">Heliobolus</emphasis>
</taxonomicName>
), subocular bordering lip, the nostril is pierced between two nasals, nasal well separated from upper labial, and dorsal scales small, smooth, and juxtaposed.
</paragraph>
<caption pageId="5" pageNumber="87">
<paragraph blockId="5.[139,1678,316,397]" pageId="5" pageNumber="87">
<emphasis bold="true" box="[139,220,316,339]" pageId="5" pageNumber="87">Table 1.</emphasis>
Meristic and scalation data for Angolan
<taxonomicName authority="Branch &amp; Conradie &amp; Pinto &amp; Tolley, 2019" authorityName="Branch &amp; Conradie &amp; Pinto &amp; Tolley" authorityYear="2019" box="[615,782,316,339]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="87" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis box="[615,782,316,339]" italics="true" pageId="5" pageNumber="87">Nucras broadleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[788,867,316,339]" pageId="5" pageNumber="87" rank="species">
<emphasis bold="true" box="[788,861,316,339]" pageId="5" pageNumber="87">sp. nov</emphasis>
.
</taxonomicNameLabel>
PEM Port Elizabeth Museum, BM Natural History Museum, London (formerly British Museum), MD Museu Dundo Regional do Dundo, Angola, TM Ditsong Museum of Natural History, South Africa (formerly Transvaal Museum), MBL Museu Bocage Lisboa, Portugal (material destroyed), MHNC Musée d' Histoire Naturelle de La Chaux-de-Fonds, Chaux-de-Fonds, Switzerland.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="87">
<table box="[150,1659,421,1471]" gridcols="11" gridrows="23" pageId="5" pageNumber="87">
<tr box="[150,1659,421,480]" gridrow="0" pageId="5" pageNumber="87">
<th box="[150,348,421,480]" gridcol="0" gridrow="0" pageId="5" pageNumber="87">
<emphasis bold="true" box="[150,327,439,462]" pageId="5" pageNumber="87">Museum number</emphasis>
</th>
<th box="[380,469,421,480]" gridcol="1" gridrow="0" pageId="5" pageNumber="87">
<emphasis bold="true" pageId="5" pageNumber="87">PEM R24005</emphasis>
</th>
<th box="[495,579,421,480]" gridcol="2" gridrow="0" pageId="5" pageNumber="87">
<emphasis bold="true" pageId="5" pageNumber="87">PEM R24157</emphasis>
</th>
<th box="[605,689,421,480]" gridcol="3" gridrow="0" pageId="5" pageNumber="87">
<emphasis bold="true" pageId="5" pageNumber="87">TM 40397</emphasis>
</th>
<th box="[717,820,421,480]" gridcol="4" gridrow="0" pageId="5" pageNumber="87">
<emphasis bold="true" box="[722,816,439,462]" pageId="5" pageNumber="87">MD 1967</emphasis>
</th>
<th box="[852,955,421,480]" gridcol="5" gridrow="0" pageId="5" pageNumber="87">
<emphasis bold="true" box="[856,951,439,462]" pageId="5" pageNumber="87">MBL 646</emphasis>
</th>
<th box="[984,1092,421,480]" gridcol="6" gridrow="0" pageId="5" pageNumber="87">
<emphasis bold="true" box="[984,1092,439,462]" pageId="5" pageNumber="87">MBL 647a</emphasis>
</th>
<th box="[1119,1228,421,480]" gridcol="7" gridrow="0" pageId="5" pageNumber="87">
<emphasis bold="true" box="[1119,1228,439,462]" pageId="5" pageNumber="87">MBL 647b</emphasis>
</th>
<th box="[1254,1380,421,480]" gridcol="8" gridrow="0" pageId="5" pageNumber="87">
<emphasis bold="true" pageId="5" pageNumber="87">BM 1907.6.29.10</emphasis>
</th>
<th box="[1404,1528,421,480]" gridcol="9" gridrow="0" pageId="5" pageNumber="87">
<emphasis bold="true" pageId="5" pageNumber="87">BM 1907.6.29.11</emphasis>
</th>
<th box="[1556,1659,421,480]" gridcol="10" gridrow="0" pageId="5" pageNumber="87">
<emphasis bold="true" pageId="5" pageNumber="87">MHNC 91.05245</emphasis>
</th>
</tr>
<tr box="[150,1659,495,554]" gridrow="1" pageId="5" pageNumber="87">
<th box="[150,348,495,554]" gridcol="0" gridrow="1" pageId="5" pageNumber="87">Status</th>
<td box="[380,469,495,554]" gridcol="1" gridrow="1" pageId="5" pageNumber="87">Holotype</td>
<td box="[495,579,495,554]" gridcol="2" gridrow="1" pageId="5" pageNumber="87">Paratype</td>
<td box="[605,689,495,554]" gridcol="3" gridrow="1" pageId="5" pageNumber="87">Paratype</td>
<td box="[717,820,495,554]" gridcol="4" gridrow="1" pageId="5" pageNumber="87">Additional specimen</td>
<td box="[852,955,495,554]" gridcol="5" gridrow="1" pageId="5" pageNumber="87">Additional specimen</td>
<td box="[984,1092,495,554]" gridcol="6" gridrow="1" pageId="5" pageNumber="87">Additional specimen</td>
<td box="[1119,1228,495,554]" gridcol="7" gridrow="1" pageId="5" pageNumber="87">Additional specimen</td>
<td box="[1254,1380,495,554]" gridcol="8" gridrow="1" pageId="5" pageNumber="87">Additional specimen</td>
<td box="[1404,1528,495,554]" gridcol="9" gridrow="1" pageId="5" pageNumber="87">Additional specimen</td>
<td box="[1556,1659,495,554]" gridcol="10" gridrow="1" pageId="5" pageNumber="87">Additional specimens</td>
</tr>
<tr box="[150,1659,569,592]" gridrow="2" pageId="5" pageNumber="87" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th box="[150,348,569,592]" gridcol="0" gridrow="2" pageId="5" pageNumber="87">Sex</th>
<td box="[380,469,569,592]" gridcol="1" gridrow="2" pageId="5" pageNumber="87">Male</td>
<td box="[495,579,569,592]" gridcol="2" gridrow="2" pageId="5" pageNumber="87">Female</td>
<td box="[605,689,569,592]" gridcol="3" gridrow="2" pageId="5" pageNumber="87">Male</td>
<td box="[717,820,569,592]" gridcol="4" gridrow="2" pageId="5" pageNumber="87">Male</td>
<td box="[1556,1659,569,592]" gridcol="10" gridrow="2" pageId="5" pageNumber="87">Juveniles</td>
</tr>
<tr box="[150,1659,607,630]" gridrow="3" pageId="5" pageNumber="87" rowspan-10="1">
<th box="[150,348,607,630]" gridcol="0" gridrow="3" pageId="5" pageNumber="87">SVL</th>
<td box="[380,469,607,630]" gridcol="1" gridrow="3" pageId="5" pageNumber="87">36</td>
<td box="[495,579,607,630]" gridcol="2" gridrow="3" pageId="5" pageNumber="87">44</td>
<td box="[605,689,607,630]" gridcol="3" gridrow="3" pageId="5" pageNumber="87">58</td>
<td box="[717,820,607,630]" gridcol="4" gridrow="3" pageId="5" pageNumber="87">40 (39)</td>
<td box="[852,955,607,630]" gridcol="5" gridrow="3" pageId="5" pageNumber="87">57</td>
<td box="[984,1092,607,630]" gridcol="6" gridrow="3" pageId="5" pageNumber="87">63</td>
<td box="[1119,1228,607,630]" gridcol="7" gridrow="3" pageId="5" pageNumber="87">52</td>
<td box="[1254,1380,607,630]" gridcol="8" gridrow="3" pageId="5" pageNumber="87">74</td>
<td box="[1404,1528,607,630]" gridcol="9" gridrow="3" pageId="5" pageNumber="87">60</td>
</tr>
<tr box="[150,1659,646,669]" gridrow="4" pageId="5" pageNumber="87" rowspan-10="1">
<th box="[150,348,646,669]" gridcol="0" gridrow="4" pageId="5" pageNumber="87">Tail</th>
<td box="[380,469,646,669]" gridcol="1" gridrow="4" pageId="5" pageNumber="87">t</td>
<td box="[495,579,646,669]" gridcol="2" gridrow="4" pageId="5" pageNumber="87">t</td>
<td box="[605,689,646,669]" gridcol="3" gridrow="4" pageId="5" pageNumber="87">110</td>
<td box="[717,820,646,669]" gridcol="4" gridrow="4" pageId="5" pageNumber="87">90 (91)</td>
<td box="[852,955,646,669]" gridcol="5" gridrow="4" pageId="5" pageNumber="87">120+</td>
<td box="[984,1092,646,669]" gridcol="6" gridrow="4" pageId="5" pageNumber="87">133</td>
<td box="[1119,1228,646,669]" gridcol="7" gridrow="4" pageId="5" pageNumber="87">135</td>
<td box="[1254,1380,646,669]" gridcol="8" gridrow="4" pageId="5" pageNumber="87">144+</td>
<td box="[1404,1528,646,669]" gridcol="9" gridrow="4" pageId="5" pageNumber="87">114</td>
</tr>
<tr box="[150,1659,685,708]" gridrow="5" pageId="5" pageNumber="87" rowspan-10="1">
<th box="[150,348,685,708]" gridcol="0" gridrow="5" pageId="5" pageNumber="87">Total</th>
<td box="[380,469,685,708]" gridcol="1" gridrow="5" pageId="5" pageNumber="87">-</td>
<td box="[495,579,685,708]" gridcol="2" gridrow="5" pageId="5" pageNumber="87">-</td>
<td box="[605,689,685,708]" gridcol="3" gridrow="5" pageId="5" pageNumber="87">168</td>
<td box="[717,820,685,708]" gridcol="4" gridrow="5" pageId="5" pageNumber="87">130</td>
<td box="[852,955,685,708]" gridcol="5" gridrow="5" pageId="5" pageNumber="87">177+</td>
<td box="[984,1092,685,708]" gridcol="6" gridrow="5" pageId="5" pageNumber="87">193</td>
<td box="[1119,1228,685,708]" gridcol="7" gridrow="5" pageId="5" pageNumber="87">187</td>
<td box="[1254,1380,685,708]" gridcol="8" gridrow="5" pageId="5" pageNumber="87">218</td>
<td box="[1404,1528,685,708]" gridcol="9" gridrow="5" pageId="5" pageNumber="87">174</td>
</tr>
<tr box="[150,1659,723,746]" gridrow="6" pageId="5" pageNumber="87" rowspan-10="1" rowspan-8="1" rowspan-9="1">
<th box="[150,348,723,746]" gridcol="0" gridrow="6" pageId="5" pageNumber="87">Head length (HL)</th>
<td box="[380,469,723,746]" gridcol="1" gridrow="6" pageId="5" pageNumber="87">10.5</td>
<td box="[495,579,723,746]" gridcol="2" gridrow="6" pageId="5" pageNumber="87">9.7</td>
<td box="[605,689,723,746]" gridcol="3" gridrow="6" pageId="5" pageNumber="87">12.4</td>
<td box="[717,820,723,746]" gridcol="4" gridrow="6" pageId="5" pageNumber="87">9.5</td>
<td box="[852,955,723,746]" gridcol="5" gridrow="6" pageId="5" pageNumber="87">12.1</td>
<td box="[984,1092,723,746]" gridcol="6" gridrow="6" pageId="5" pageNumber="87">13.8</td>
<td box="[1119,1228,723,746]" gridcol="7" gridrow="6" pageId="5" pageNumber="87">12.8</td>
</tr>
<tr box="[150,1659,762,785]" gridrow="7" pageId="5" pageNumber="87" rowspan-10="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th box="[150,348,762,785]" gridcol="0" gridrow="7" pageId="5" pageNumber="87">Head width (HW)</th>
<td box="[380,469,762,785]" gridcol="1" gridrow="7" pageId="5" pageNumber="87">5.88</td>
<td box="[495,579,762,785]" gridcol="2" gridrow="7" pageId="5" pageNumber="87">6.4</td>
<td box="[605,689,762,785]" gridcol="3" gridrow="7" pageId="5" pageNumber="87">6.2</td>
</tr>
<tr box="[150,1659,801,824]" gridrow="8" pageId="5" pageNumber="87" rowspan-10="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th box="[150,348,801,824]" gridcol="0" gridrow="8" pageId="5" pageNumber="87">HW/HL ratio</th>
<td box="[380,469,801,824]" gridcol="1" gridrow="8" pageId="5" pageNumber="87">0.56</td>
<td box="[495,579,801,824]" gridcol="2" gridrow="8" pageId="5" pageNumber="87">0.7</td>
<td box="[605,689,801,824]" gridcol="3" gridrow="8" pageId="5" pageNumber="87">0.5</td>
</tr>
<tr box="[150,1659,840,863]" gridrow="9" pageId="5" pageNumber="87" rowspan-10="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th box="[150,348,840,863]" gridcol="0" gridrow="9" pageId="5" pageNumber="87">Snout length</th>
<td box="[380,469,840,863]" gridcol="1" gridrow="9" pageId="5" pageNumber="87">3.8</td>
<td box="[495,579,840,863]" gridcol="2" gridrow="9" pageId="5" pageNumber="87">4.1</td>
<td box="[605,689,840,863]" gridcol="3" gridrow="9" pageId="5" pageNumber="87">4.9</td>
</tr>
<tr box="[150,1659,879,902]" gridrow="10" pageId="5" pageNumber="87" rowspan-10="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th box="[150,348,879,902]" gridcol="0" gridrow="10" pageId="5" pageNumber="87">Eye length</th>
<td box="[380,469,879,902]" gridcol="1" gridrow="10" pageId="5" pageNumber="87">2.11</td>
<td box="[495,579,879,902]" gridcol="2" gridrow="10" pageId="5" pageNumber="87">2.1</td>
<td box="[605,689,879,902]" gridcol="3" gridrow="10" pageId="5" pageNumber="87">2.4</td>
</tr>
<tr box="[150,1659,918,941]" gridrow="11" pageId="5" pageNumber="87" rowspan-10="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th box="[150,348,918,941]" gridcol="0" gridrow="11" pageId="5" pageNumber="87">Ear-eye length</th>
<td box="[380,469,918,941]" gridcol="1" gridrow="11" pageId="5" pageNumber="87">3.74</td>
<td box="[495,579,918,941]" gridcol="2" gridrow="11" pageId="5" pageNumber="87">3.5</td>
<td box="[605,689,918,941]" gridcol="3" gridrow="11" pageId="5" pageNumber="87">4.2</td>
</tr>
<tr box="[150,1659,956,979]" gridrow="12" pageId="5" pageNumber="87" rowspan-10="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th box="[150,348,956,979]" gridcol="0" gridrow="12" pageId="5" pageNumber="87">UL *</th>
<td box="[380,469,956,979]" gridcol="1" gridrow="12" pageId="5" pageNumber="87">4 (2)</td>
<td box="[495,579,956,979]" gridcol="2" gridrow="12" pageId="5" pageNumber="87">4 (2)</td>
<td box="[605,689,956,979]" gridcol="3" gridrow="12" pageId="5" pageNumber="87">4 (2)</td>
<td box="[1254,1380,956,979]" gridcol="8" gridrow="12" pageId="5" pageNumber="87">4 (2)</td>
<td box="[1404,1528,956,979]" gridcol="9" gridrow="12" pageId="5" pageNumber="87">4 (2)</td>
</tr>
<tr box="[150,1659,995,1018]" gridrow="13" pageId="5" pageNumber="87" rowspan-10="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th box="[150,348,995,1018]" gridcol="0" gridrow="13" pageId="5" pageNumber="87">LL **</th>
<td box="[380,469,995,1018]" gridcol="1" gridrow="13" pageId="5" pageNumber="87">6 (3)</td>
<td box="[495,579,995,1018]" gridcol="2" gridrow="13" pageId="5" pageNumber="87">6 (3)</td>
<td box="[605,689,995,1018]" gridcol="3" gridrow="13" pageId="5" pageNumber="87">6 (3)</td>
<td box="[1254,1380,995,1018]" gridcol="8" gridrow="13" pageId="5" pageNumber="87">6 (3)</td>
<td box="[1404,1528,995,1018]" gridcol="9" gridrow="13" pageId="5" pageNumber="87">6 (3)</td>
</tr>
<tr box="[150,1659,1034,1057]" gridrow="14" pageId="5" pageNumber="87">
<th box="[150,348,1034,1057]" gridcol="0" gridrow="14" pageId="5" pageNumber="87">MSR</th>
<td box="[380,469,1034,1057]" gridcol="1" gridrow="14" pageId="5" pageNumber="87">42</td>
<td box="[495,579,1034,1057]" gridcol="2" gridrow="14" pageId="5" pageNumber="87">48</td>
<td box="[605,689,1034,1057]" gridcol="3" gridrow="14" pageId="5" pageNumber="87">46</td>
<td box="[717,820,1034,1057]" gridcol="4" gridrow="14" pageId="5" pageNumber="87">48 (50)</td>
<td box="[852,955,1034,1057]" gridcol="5" gridrow="14" pageId="5" pageNumber="87">38</td>
<td box="[984,1092,1034,1057]" gridcol="6" gridrow="14" pageId="5" pageNumber="87">44</td>
<td box="[1119,1228,1034,1057]" gridcol="7" gridrow="14" pageId="5" pageNumber="87">46</td>
<td box="[1254,1380,1034,1057]" gridcol="8" gridrow="14" pageId="5" pageNumber="87">48</td>
<td box="[1404,1528,1034,1057]" gridcol="9" gridrow="14" pageId="5" pageNumber="87">48</td>
<td box="[1556,1659,1034,1057]" gridcol="10" gridrow="14" pageId="5" pageNumber="87">4243</td>
</tr>
<tr box="[150,1659,1072,1095]" gridrow="15" pageId="5" pageNumber="87" rowspan-10="1">
<th box="[150,348,1072,1095]" gridcol="0" gridrow="15" pageId="5" pageNumber="87">Ventrals (transverse)</th>
<td box="[380,469,1072,1095]" gridcol="1" gridrow="15" pageId="5" pageNumber="87">8</td>
<td box="[495,579,1072,1095]" gridcol="2" gridrow="15" pageId="5" pageNumber="87">8</td>
<td box="[605,689,1072,1095]" gridcol="3" gridrow="15" pageId="5" pageNumber="87">8</td>
<td box="[717,820,1072,1095]" gridcol="4" gridrow="15" pageId="5" pageNumber="87">8</td>
<td box="[852,955,1072,1095]" gridcol="5" gridrow="15" pageId="5" pageNumber="87">8</td>
<td box="[984,1092,1072,1095]" gridcol="6" gridrow="15" pageId="5" pageNumber="87">8</td>
<td box="[1119,1228,1072,1095]" gridcol="7" gridrow="15" pageId="5" pageNumber="87">8</td>
<td box="[1254,1380,1072,1095]" gridcol="8" gridrow="15" pageId="5" pageNumber="87">8</td>
<td box="[1404,1528,1072,1095]" gridcol="9" gridrow="15" pageId="5" pageNumber="87">8</td>
</tr>
<tr box="[150,1659,1110,1169]" gridrow="16" pageId="5" pageNumber="87">
<th box="[150,348,1110,1169]" gridcol="0" gridrow="16" pageId="5" pageNumber="87">Ventrals (longitudinal)</th>
<td box="[380,469,1110,1169]" gridcol="1" gridrow="16" pageId="5" pageNumber="87">28</td>
<td box="[495,579,1110,1169]" gridcol="2" gridrow="16" pageId="5" pageNumber="87">29</td>
<td box="[605,689,1110,1169]" gridcol="3" gridrow="16" pageId="5" pageNumber="87">27</td>
<td box="[717,820,1110,1169]" gridcol="4" gridrow="16" pageId="5" pageNumber="87">29</td>
<td box="[852,955,1110,1169]" gridcol="5" gridrow="16" pageId="5" pageNumber="87">33</td>
<td box="[984,1092,1110,1169]" gridcol="6" gridrow="16" pageId="5" pageNumber="87">30</td>
<td box="[1119,1228,1110,1169]" gridcol="7" gridrow="16" pageId="5" pageNumber="87">29</td>
<td box="[1254,1380,1110,1169]" gridcol="8" gridrow="16" pageId="5" pageNumber="87">33</td>
<td box="[1404,1528,1110,1169]" gridcol="9" gridrow="16" pageId="5" pageNumber="87">29</td>
<td box="[1556,1659,1110,1169]" gridcol="10" gridrow="16" pageId="5" pageNumber="87">2729</td>
</tr>
<tr box="[150,1659,1184,1207]" gridrow="17" pageId="5" pageNumber="87">
<th box="[150,348,1184,1207]" gridcol="0" gridrow="17" pageId="5" pageNumber="87">Granules SC-SO</th>
<td box="[380,469,1184,1207]" gridcol="1" gridrow="17" pageId="5" pageNumber="87">1/0</td>
<td box="[495,579,1184,1207]" gridcol="2" gridrow="17" pageId="5" pageNumber="87">3/4</td>
<td box="[605,689,1184,1207]" gridcol="3" gridrow="17" pageId="5" pageNumber="87">2/3</td>
<td box="[717,820,1184,1207]" gridcol="4" gridrow="17" pageId="5" pageNumber="87">3</td>
<td box="[852,955,1184,1207]" gridcol="5" gridrow="17" pageId="5" pageNumber="87">1</td>
<td box="[984,1092,1184,1207]" gridcol="6" gridrow="17" pageId="5" pageNumber="87">2</td>
<td box="[1119,1228,1184,1207]" gridcol="7" gridrow="17" pageId="5" pageNumber="87">6/3</td>
<td box="[1254,1380,1184,1207]" gridcol="8" gridrow="17" pageId="5" pageNumber="87">3/2</td>
<td box="[1404,1528,1184,1207]" gridcol="9" gridrow="17" pageId="5" pageNumber="87">3/3</td>
<td box="[1556,1659,1184,1207]" gridcol="10" gridrow="17" pageId="5" pageNumber="87">02</td>
</tr>
<tr box="[150,1659,1223,1246]" gridrow="18" pageId="5" pageNumber="87" rowspan-10="1">
<th box="[150,348,1223,1246]" gridcol="0" gridrow="18" pageId="5" pageNumber="87">Supraciliaries</th>
<td box="[380,469,1223,1246]" gridcol="1" gridrow="18" pageId="5" pageNumber="87">6/6</td>
<td box="[495,579,1223,1246]" gridcol="2" gridrow="18" pageId="5" pageNumber="87">7/8</td>
<td box="[605,689,1223,1246]" gridcol="3" gridrow="18" pageId="5" pageNumber="87">6/6</td>
<td box="[717,820,1223,1246]" gridcol="4" gridrow="18" pageId="5" pageNumber="87">6</td>
<td box="[852,955,1223,1246]" gridcol="5" gridrow="18" pageId="5" pageNumber="87">5</td>
<td box="[984,1092,1223,1246]" gridcol="6" gridrow="18" pageId="5" pageNumber="87">6</td>
<td box="[1119,1228,1223,1246]" gridcol="7" gridrow="18" pageId="5" pageNumber="87">7</td>
<td box="[1254,1380,1223,1246]" gridcol="8" gridrow="18" pageId="5" pageNumber="87">7/6</td>
<td box="[1404,1528,1223,1246]" gridcol="9" gridrow="18" pageId="5" pageNumber="87">6</td>
</tr>
<tr box="[150,1659,1262,1285]" gridrow="19" pageId="5" pageNumber="87" rowspan-10="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th box="[150,348,1262,1285]" gridcol="0" gridrow="19" pageId="5" pageNumber="87">Interparietal Y/N</th>
<td box="[380,469,1262,1285]" gridcol="1" gridrow="19" pageId="5" pageNumber="87">Y</td>
<td box="[495,579,1262,1285]" gridcol="2" gridrow="19" pageId="5" pageNumber="87">Y</td>
<td box="[605,689,1262,1285]" gridcol="3" gridrow="19" pageId="5" pageNumber="87">Y</td>
<td box="[717,820,1262,1285]" gridcol="4" gridrow="19" pageId="5" pageNumber="87">Y</td>
<td box="[1254,1380,1262,1285]" gridcol="8" gridrow="19" pageId="5" pageNumber="87">Y</td>
<td box="[1404,1528,1262,1285]" gridcol="9" gridrow="19" pageId="5" pageNumber="87">Y</td>
</tr>
<tr box="[150,1659,1300,1359]" gridrow="20" pageId="5" pageNumber="87" rowspan-10="1">
<th box="[150,348,1300,1359]" gridcol="0" gridrow="20" pageId="5" pageNumber="87">IP contact with occipital</th>
<td box="[380,469,1300,1359]" gridcol="1" gridrow="20" pageId="5" pageNumber="87">contact</td>
<td box="[495,579,1300,1359]" gridcol="2" gridrow="20" pageId="5" pageNumber="87">contact</td>
<td box="[605,689,1300,1359]" gridcol="3" gridrow="20" pageId="5" pageNumber="87">contact</td>
<td box="[717,820,1300,1359]" gridcol="4" gridrow="20" pageId="5" pageNumber="87">contact</td>
<td box="[852,955,1300,1359]" gridcol="5" gridrow="20" pageId="5" pageNumber="87">separate</td>
<td box="[984,1092,1300,1359]" gridcol="6" gridrow="20" pageId="5" pageNumber="87">broad contact</td>
<td box="[1119,1228,1300,1359]" gridcol="7" gridrow="20" pageId="5" pageNumber="87">broad contact</td>
<td box="[1254,1380,1300,1359]" gridcol="8" gridrow="20" pageId="5" pageNumber="87">contact</td>
<td box="[1404,1528,1300,1359]" gridcol="9" gridrow="20" pageId="5" pageNumber="87">contact</td>
</tr>
<tr box="[150,1659,1374,1433]" gridrow="21" pageId="5" pageNumber="87">
<th box="[150,348,1374,1433]" gridcol="0" gridrow="21" pageId="5" pageNumber="87">Subdigital lamellae below 4th toe</th>
<td box="[380,469,1374,1433]" gridcol="1" gridrow="21" pageId="5" pageNumber="87">23/25</td>
<td box="[495,579,1374,1433]" gridcol="2" gridrow="21" pageId="5" pageNumber="87">27/27</td>
<td box="[605,689,1374,1433]" gridcol="3" gridrow="21" pageId="5" pageNumber="87">23/25</td>
<td box="[717,820,1374,1433]" gridcol="4" gridrow="21" pageId="5" pageNumber="87">28/29 (27)</td>
<td box="[852,955,1374,1433]" gridcol="5" gridrow="21" pageId="5" pageNumber="87">27/26</td>
<td box="[984,1092,1374,1433]" gridcol="6" gridrow="21" pageId="5" pageNumber="87">28/27</td>
<td box="[1119,1228,1374,1433]" gridcol="7" gridrow="21" pageId="5" pageNumber="87">28/28</td>
<td box="[1254,1380,1374,1433]" gridcol="8" gridrow="21" pageId="5" pageNumber="87">28/28</td>
<td box="[1404,1528,1374,1433]" gridcol="9" gridrow="21" pageId="5" pageNumber="87">26/26</td>
<td box="[1556,1659,1374,1433]" gridcol="10" gridrow="21" pageId="5" pageNumber="87">25</td>
</tr>
<tr box="[150,1659,1448,1471]" gridrow="22" pageId="5" pageNumber="87" rowspan-10="1">
<th box="[150,348,1448,1471]" gridcol="0" gridrow="22" pageId="5" pageNumber="87">Femoral pores</th>
<td box="[380,469,1448,1471]" gridcol="1" gridrow="22" pageId="5" pageNumber="87">13/13</td>
<td box="[495,579,1448,1471]" gridcol="2" gridrow="22" pageId="5" pageNumber="87">15/15</td>
<td box="[605,689,1448,1471]" gridcol="3" gridrow="22" pageId="5" pageNumber="87">14/16</td>
<td box="[717,820,1448,1471]" gridcol="4" gridrow="22" pageId="5" pageNumber="87">13/13</td>
<td box="[852,955,1448,1471]" gridcol="5" gridrow="22" pageId="5" pageNumber="87">14/13</td>
<td box="[984,1092,1448,1471]" gridcol="6" gridrow="22" pageId="5" pageNumber="87">12/12</td>
<td box="[1119,1228,1448,1471]" gridcol="7" gridrow="22" pageId="5" pageNumber="87">15/14</td>
<td box="[1254,1380,1448,1471]" gridcol="8" gridrow="22" pageId="5" pageNumber="87">14/14</td>
<td box="[1404,1528,1448,1471]" gridcol="9" gridrow="22" pageId="5" pageNumber="87">13/13</td>
</tr>
</table>
</paragraph>
<paragraph blockId="5.[139,919,1482,1505]" box="[139,919,1482,1505]" pageId="5" pageNumber="87">
<tableNote box="[139,919,1482,1505]" pageId="5" pageNumber="87" targetBox="[150,1659,421,1471]" targetPageId="5">
<emphasis bold="true" box="[139,151,1482,1505]" pageId="5" pageNumber="87">*</emphasis>
anterior subocular (posterior subocular), ** total (largest number, t = truncated)
</tableNote>
</paragraph>
<paragraph blockId="5.[141,746,1704,1952]" lastBlockId="7.[142,703,141,1965]" lastPageId="7" lastPageNumber="89" pageId="5" pageNumber="87">
The new species can be diagnosed from other
<taxonomicName authorityName="Gray" authorityYear="1838" box="[670,745,1800,1823]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="87" phylum="Chordata" rank="genus">
<emphasis box="[670,745,1800,1823]" italics="true" pageId="5" pageNumber="87">Nucras</emphasis>
</taxonomicName>
species based on a combination of the following characters: series of transversely enlarged plates present under forearm (versus absent or only feebly enlarged in
<taxonomicName baseAuthorityName="Milne-edwards" baseAuthorityYear="1829" box="[568,730,1896,1920]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="87" phylum="Chordata" rank="species" species="lalandii">
<emphasis box="[568,730,1896,1920]" italics="true" pageId="5" pageNumber="87">Nucras lalandii</emphasis>
</taxonomicName>
), a small series (06) of small granules present between supraciliaries and supraoculars (versus mostly absent in
<taxonomicName authorityName="Neumann" authorityYear="1900" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="87" phylum="Chordata" rank="species" species="boulengeri">
<emphasis italics="true" pageId="5" pageNumber="87">N. boulengeri</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Milne-edwards" baseAuthorityYear="1829" box="[957,1069,1736,1760]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="87" phylum="Chordata" rank="species" species="lalandii">
<emphasis box="[957,1069,1736,1760]" italics="true" pageId="5" pageNumber="87">N. lalandii</emphasis>
</taxonomicName>
), 2329 lamellae under 4
<superScript attach="left" box="[1343,1355,1733,1748]" fontSize="6" pageId="5" pageNumber="87">th</superScript>
toe (versus less than
<quantity box="[966,1018,1768,1792]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="5" pageNumber="87" unit="in" value="22.0">22 in</quantity>
<taxonomicName baseAuthorityName="Milne-edwards" baseAuthorityYear="1829" box="[1024,1134,1768,1792]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="87" phylum="Chordata" rank="species" species="lalandii">
<emphasis box="[1024,1134,1768,1792]" italics="true" pageId="5" pageNumber="87">N. lalandii</emphasis>
</taxonomicName>
), dorsum with a series of longitudinal pale stripes (versus dark cross bands present in
<taxonomicName baseAuthorityName="Milne-edwards" baseAuthorityYear="1829" box="[819,931,1832,1856]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="87" phylum="Chordata" rank="species" species="lalandii">
<emphasis box="[819,931,1832,1856]" italics="true" pageId="5" pageNumber="87">N. lalandii</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Laurent" authorityYear="1964" box="[984,1098,1832,1856]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="87" phylum="Chordata" rank="species" species="scalaris">
<emphasis box="[984,1098,1832,1856]" italics="true" pageId="5" pageNumber="87">N. scalaris</emphasis>
</taxonomicName>
or a series of pale vertebral spots, sometimes forming irregular transverse bands in
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="87" phylum="Chordata" rank="species" species="intertexta">
<emphasis italics="true" pageId="5" pageNumber="87">N. intertexta</emphasis>
</taxonomicName>
or lack of any dorsal patterns in
<taxonomicName authorityName="Bauer, Childers, Broeckhoven &amp; Mouton" authorityYear="2019" box="[1235,1380,1896,1920]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="87" phylum="Chordata" rank="species" species="aurantiaca">
<emphasis box="[1235,1380,1896,1920]" italics="true" pageId="5" pageNumber="87">N. aurantiaca</emphasis>
</taxonomicName>
), four pale stripes on nape with outer stripes forming a.
</paragraph>
<caption box="[139,1876,486,509]" pageId="6" pageNumber="88">
<paragraph blockId="6.[139,1876,486,509]" box="[139,1876,486,509]" pageId="6" pageNumber="88">
<emphasis bold="true" box="[139,213,486,509]" pageId="6" pageNumber="88">Table 2</emphasis>
. Samples used in genetic analysis. Museum abbreviations: CAS California Academy of Science, PEM Port Elizabeth Museum, NMB National Museum Bloemfontein.
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="88">
<table box="[150,1818,536,1709]" gridcols="8" gridrows="30" pageId="6" pageNumber="88">
<tr box="[150,1818,536,595]" gridrow="0" pageId="6" pageNumber="88">
<th box="[150,219,536,595]" gridcol="0" gridrow="0" pageId="6" pageNumber="88">
<emphasis bold="true" box="[150,215,554,577]" pageId="6" pageNumber="88">Genus</emphasis>
</th>
<th box="[360,535,536,595]" gridcol="1" gridrow="0" pageId="6" pageNumber="88">
<emphasis bold="true" box="[360,435,554,577]" pageId="6" pageNumber="88">Species</emphasis>
</th>
<th box="[600,753,536,595]" gridcol="2" gridrow="0" pageId="6" pageNumber="88">
<emphasis bold="true" pageId="6" pageNumber="88">Field accession ID</emphasis>
</th>
<th box="[788,970,536,595]" gridcol="3" gridrow="0" pageId="6" pageNumber="88">
<emphasis bold="true" box="[789,970,536,595]" pageId="6" pageNumber="88">Museum accession number</emphasis>
</th>
<th box="[1000,1122,536,595]" gridcol="4" gridrow="0" pageId="6" pageNumber="88">
<emphasis bold="true" box="[1000,1038,554,577]" pageId="6" pageNumber="88">16S</emphasis>
</th>
<th box="[1147,1270,536,595]" gridcol="5" gridrow="0" pageId="6" pageNumber="88">
<emphasis bold="true" box="[1147,1194,554,577]" pageId="6" pageNumber="88">ND4</emphasis>
</th>
<th box="[1295,1417,536,595]" gridcol="6" gridrow="0" pageId="6" pageNumber="88">
<emphasis bold="true" box="[1295,1360,554,577]" pageId="6" pageNumber="88">RAG1</emphasis>
</th>
<th box="[1442,1818,536,595]" gridcol="7" gridrow="0" pageId="6" pageNumber="88">
<emphasis bold="true" box="[1442,1526,554,577]" pageId="6" pageNumber="88">Locality</emphasis>
</th>
</tr>
<tr box="[150,1818,610,634]" gridrow="1" pageId="6" pageNumber="88">
<taxonomicName authority="Branch &amp; Conradie &amp; Pinto &amp; Tolley, 2019" authorityName="Branch &amp; Conradie &amp; Pinto &amp; Tolley" authorityYear="2019" box="[150,451,610,634]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="93" phylum="Chordata" rank="species" species="broadleyi" status="nov. sp.">
<th box="[150,219,610,634]" gridcol="0" gridrow="1" pageId="6" pageNumber="88">
<emphasis box="[150,219,610,633]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,610,634]" gridcol="1" gridrow="1" pageId="6" pageNumber="88">
<emphasis box="[360,451,611,634]" italics="true" pageId="6" pageNumber="88">broadleyi</emphasis>
<taxonomicNameLabel box="[457,535,611,634]" pageId="6" pageNumber="88" rank="species">
<emphasis bold="true" box="[457,529,611,634]" pageId="6" pageNumber="88">sp. nov</emphasis>
.
</taxonomicNameLabel>
</td>
</taxonomicName>
<td box="[600,753,610,634]" gridcol="2" gridrow="1" pageId="6" pageNumber="88">AG18</td>
<td box="[788,970,610,634]" gridcol="3" gridrow="1" pageId="6" pageNumber="88">PEM R24005</td>
<td box="[1000,1122,610,634]" gridcol="4" gridrow="1" pageId="6" pageNumber="88">MN265869y</td>
<td box="[1147,1270,610,634]" gridcol="5" gridrow="1" pageId="6" pageNumber="88"></td>
<td box="[1295,1417,610,634]" gridcol="6" gridrow="1" pageId="6" pageNumber="88">MN265872y</td>
<td box="[1442,1818,610,634]" gridcol="7" gridrow="1" pageId="6" pageNumber="88">Angola, Namibe Province</td>
</tr>
<tr box="[150,1818,649,672]" gridrow="2" pageId="6" pageNumber="88">
<taxonomicName authority="Branch &amp; Conradie &amp; Pinto &amp; Tolley, 2019" authorityName="Branch &amp; Conradie &amp; Pinto &amp; Tolley" authorityYear="2019" box="[150,451,649,672]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="93" phylum="Chordata" rank="species" species="broadleyi" status="nov. sp.">
<th box="[150,219,649,672]" gridcol="0" gridrow="2" pageId="6" pageNumber="88">
<emphasis box="[150,219,649,672]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,649,672]" gridcol="1" gridrow="2" pageId="6" pageNumber="88">
<emphasis box="[360,451,649,672]" italics="true" pageId="6" pageNumber="88">broadleyi</emphasis>
<taxonomicNameLabel box="[457,535,649,672]" pageId="6" pageNumber="88" rank="species">
<emphasis bold="true" box="[457,529,649,672]" pageId="6" pageNumber="88">sp. nov</emphasis>
.
</taxonomicNameLabel>
</td>
</taxonomicName>
<td box="[600,753,649,672]" gridcol="2" gridrow="2" pageId="6" pageNumber="88">AG166</td>
<td box="[788,970,649,672]" gridcol="3" gridrow="2" pageId="6" pageNumber="88">PEM R24157</td>
<td box="[1000,1122,649,672]" gridcol="4" gridrow="2" pageId="6" pageNumber="88">MN265870y</td>
<td box="[1147,1270,649,672]" gridcol="5" gridrow="2" pageId="6" pageNumber="88">MN265871y</td>
<td box="[1295,1417,649,672]" gridcol="6" gridrow="2" pageId="6" pageNumber="88">MN265873y</td>
<td box="[1442,1818,649,672]" gridcol="7" gridrow="2" pageId="6" pageNumber="88">Angola, Namibe Province</td>
</tr>
<tr box="[150,1818,687,711]" gridrow="3" pageId="6" pageNumber="88" rowspan-3="1">
<taxonomicName authorityName="Neumann" authorityYear="1900" box="[150,464,687,710]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="boulengeri">
<th box="[150,219,687,711]" gridcol="0" gridrow="3" pageId="6" pageNumber="88">
<emphasis box="[150,219,687,710]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,687,711]" gridcol="1" gridrow="3" pageId="6" pageNumber="88">
<emphasis box="[360,464,687,710]" italics="true" pageId="6" pageNumber="88">boulengeri</emphasis>
</td>
</taxonomicName>
<td box="[600,753,687,711]" gridcol="2" gridrow="3" pageId="6" pageNumber="88">JM02169</td>
<td box="[1000,1122,687,711]" gridcol="4" gridrow="3" pageId="6" pageNumber="88">HG005184</td>
<td box="[1147,1270,687,711]" gridcol="5" gridrow="3" pageId="6" pageNumber="88">HG005212</td>
<td box="[1295,1417,687,711]" gridcol="6" gridrow="3" pageId="6" pageNumber="88">HG005233</td>
<td box="[1442,1818,687,711]" gridcol="7" gridrow="3" pageId="6" pageNumber="88">Kenya</td>
</tr>
<tr box="[150,1818,726,749]" gridrow="4" pageId="6" pageNumber="88">
<th box="[150,219,726,749]" gridcol="0" gridrow="4" pageId="6" pageNumber="88">
<taxonomicName authorityName="Gray" authorityYear="1838" box="[150,219,726,749]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="genus">
<emphasis box="[150,219,726,749]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</taxonomicName>
</th>
<td box="[360,535,726,749]" gridcol="1" gridrow="4" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1882" box="[360,421,726,749]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="species" species="holubi">
<emphasis box="[360,421,726,749]" italics="true" pageId="6" pageNumber="88">holubi</emphasis>
</taxonomicName>
</td>
<td box="[600,753,726,749]" gridcol="2" gridrow="4" pageId="6" pageNumber="88">MBUR00260</td>
<td box="[788,970,726,749]" gridcol="3" gridrow="4" pageId="6" pageNumber="88">PEM R22814</td>
<td box="[1000,1122,726,749]" gridcol="4" gridrow="4" pageId="6" pageNumber="88">HG005188</td>
<td box="[1147,1270,726,749]" gridcol="5" gridrow="4" pageId="6" pageNumber="88">HG005216</td>
<td box="[1295,1417,726,749]" gridcol="6" gridrow="4" pageId="6" pageNumber="88">HG005237</td>
<td box="[1442,1818,726,749]" gridcol="7" gridrow="4" pageId="6" pageNumber="88">South Africa, Limpopo Province</td>
</tr>
<tr box="[150,1818,764,787]" gridrow="5" pageId="6" pageNumber="88">
<th box="[150,219,764,787]" gridcol="0" gridrow="5" pageId="6" pageNumber="88">
<taxonomicName authorityName="Gray" authorityYear="1838" box="[150,219,764,787]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="genus">
<emphasis box="[150,219,764,787]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</taxonomicName>
</th>
<td box="[360,535,764,787]" gridcol="1" gridrow="5" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1882" box="[360,421,764,787]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="species" species="holubi">
<emphasis box="[360,421,764,787]" italics="true" pageId="6" pageNumber="88">holubi</emphasis>
</taxonomicName>
</td>
<td box="[600,753,764,787]" gridcol="2" gridrow="5" pageId="6" pageNumber="88">MCZ38793</td>
<td box="[788,970,764,787]" gridcol="3" gridrow="5" pageId="6" pageNumber="88">CAS 234138</td>
<td box="[1000,1122,764,787]" gridcol="4" gridrow="5" pageId="6" pageNumber="88">HG005186</td>
<td box="[1147,1270,764,787]" gridcol="5" gridrow="5" pageId="6" pageNumber="88">HG005214</td>
<td box="[1295,1417,764,787]" gridcol="6" gridrow="5" pageId="6" pageNumber="88">HG005235</td>
<td box="[1442,1818,764,787]" gridcol="7" gridrow="5" pageId="6" pageNumber="88">South Africa, Limpopo Province</td>
</tr>
<tr box="[150,1818,802,825]" gridrow="6" pageId="6" pageNumber="88">
<th box="[150,219,802,825]" gridcol="0" gridrow="6" pageId="6" pageNumber="88">
<taxonomicName authorityName="Gray" authorityYear="1838" box="[150,219,802,825]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="genus">
<emphasis box="[150,219,802,825]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</taxonomicName>
</th>
<td box="[360,535,802,825]" gridcol="1" gridrow="6" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1882" box="[360,421,802,825]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="species" species="holubi">
<emphasis box="[360,421,802,825]" italics="true" pageId="6" pageNumber="88">holubi</emphasis>
</taxonomicName>
</td>
<td box="[600,753,802,825]" gridcol="2" gridrow="6" pageId="6" pageNumber="88">RSP420</td>
<td box="[788,970,802,825]" gridcol="3" gridrow="6" pageId="6" pageNumber="88">PEM R18647</td>
<td box="[1000,1122,802,825]" gridcol="4" gridrow="6" pageId="6" pageNumber="88">HG005187</td>
<td box="[1147,1270,802,825]" gridcol="5" gridrow="6" pageId="6" pageNumber="88">HG005215</td>
<td box="[1295,1417,802,825]" gridcol="6" gridrow="6" pageId="6" pageNumber="88">HG005236</td>
<td box="[1442,1818,802,825]" gridcol="7" gridrow="6" pageId="6" pageNumber="88">South Africa, Limpopo Province</td>
</tr>
<tr box="[150,1818,840,863]" gridrow="7" pageId="6" pageNumber="88" rowspan-3="1">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[150,452,840,863]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="86" phylum="Chordata" rank="species" species="intertexta">
<th box="[150,219,840,863]" gridcol="0" gridrow="7" pageId="6" pageNumber="88">
<emphasis box="[150,219,840,863]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,840,863]" gridcol="1" gridrow="7" pageId="6" pageNumber="88">
<emphasis box="[360,452,840,863]" italics="true" pageId="6" pageNumber="88">intertexta</emphasis>
</td>
</taxonomicName>
<td box="[600,753,840,863]" gridcol="2" gridrow="7" pageId="6" pageNumber="88">MB20952</td>
<td box="[1000,1122,840,863]" gridcol="4" gridrow="7" pageId="6" pageNumber="88">HG005193</td>
<td box="[1147,1270,840,863]" gridcol="5" gridrow="7" pageId="6" pageNumber="88">HG005221</td>
<td box="[1295,1417,840,863]" gridcol="6" gridrow="7" pageId="6" pageNumber="88">HG005242</td>
<td box="[1442,1818,840,863]" gridcol="7" gridrow="7" pageId="6" pageNumber="88">South Africa, Northern Cape Province</td>
</tr>
<tr box="[150,1818,879,902]" gridrow="8" pageId="6" pageNumber="88" rowspan-3="1">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[150,452,879,902]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="86" phylum="Chordata" rank="species" species="intertexta">
<th box="[150,219,879,902]" gridcol="0" gridrow="8" pageId="6" pageNumber="88">
<emphasis box="[150,219,879,902]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,879,902]" gridcol="1" gridrow="8" pageId="6" pageNumber="88">
<emphasis box="[360,452,879,902]" italics="true" pageId="6" pageNumber="88">intertexta</emphasis>
</td>
</taxonomicName>
<td box="[600,753,879,902]" gridcol="2" gridrow="8" pageId="6" pageNumber="88">MB21183</td>
<td box="[1000,1122,879,902]" gridcol="4" gridrow="8" pageId="6" pageNumber="88">HG005194</td>
<td box="[1147,1270,879,902]" gridcol="5" gridrow="8" pageId="6" pageNumber="88">HG005222</td>
<td box="[1295,1417,879,902]" gridcol="6" gridrow="8" pageId="6" pageNumber="88"></td>
<td box="[1442,1818,879,902]" gridcol="7" gridrow="8" pageId="6" pageNumber="88">South Africa, Northern Cape Province</td>
</tr>
<tr box="[150,1818,917,940]" gridrow="9" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[150,452,917,940]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="86" phylum="Chordata" rank="species" species="intertexta">
<th box="[150,219,917,940]" gridcol="0" gridrow="9" pageId="6" pageNumber="88">
<emphasis box="[150,219,917,940]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,917,940]" gridcol="1" gridrow="9" pageId="6" pageNumber="88">
<emphasis box="[360,452,917,940]" italics="true" pageId="6" pageNumber="88">intertexta</emphasis>
</td>
</taxonomicName>
<td box="[600,753,917,940]" gridcol="2" gridrow="9" pageId="6" pageNumber="88">MCZ38872</td>
<td box="[788,970,917,940]" gridcol="3" gridrow="9" pageId="6" pageNumber="88">CAS 234212</td>
<td box="[1000,1122,917,940]" gridcol="4" gridrow="9" pageId="6" pageNumber="88">HG005192</td>
<td box="[1147,1270,917,940]" gridcol="5" gridrow="9" pageId="6" pageNumber="88">HG005220</td>
<td box="[1295,1417,917,940]" gridcol="6" gridrow="9" pageId="6" pageNumber="88">HG005241</td>
<td box="[1442,1818,917,940]" gridcol="7" gridrow="9" pageId="6" pageNumber="88">South Africa, Limpopo Province</td>
</tr>
<tr box="[150,1818,955,978]" gridrow="10" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[150,452,955,978]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="86" phylum="Chordata" rank="species" species="intertexta">
<th box="[150,219,955,978]" gridcol="0" gridrow="10" pageId="6" pageNumber="88">
<emphasis box="[150,219,955,978]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,955,978]" gridcol="1" gridrow="10" pageId="6" pageNumber="88">
<emphasis box="[360,452,955,978]" italics="true" pageId="6" pageNumber="88">intertexta</emphasis>
</td>
</taxonomicName>
<td box="[600,753,955,978]" gridcol="2" gridrow="10" pageId="6" pageNumber="88">RSP030</td>
<td box="[788,970,955,978]" gridcol="3" gridrow="10" pageId="6" pageNumber="88">PEM R18257</td>
<td box="[1000,1122,955,978]" gridcol="4" gridrow="10" pageId="6" pageNumber="88">HG005191</td>
<td box="[1147,1270,955,978]" gridcol="5" gridrow="10" pageId="6" pageNumber="88">HG005219</td>
<td box="[1295,1417,955,978]" gridcol="6" gridrow="10" pageId="6" pageNumber="88">HG005240</td>
<td box="[1442,1818,955,978]" gridcol="7" gridrow="10" pageId="6" pageNumber="88">South Africa, Northern Cape Province</td>
</tr>
<tr box="[150,1818,993,1016]" gridrow="11" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[150,452,993,1016]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="86" phylum="Chordata" rank="species" species="intertexta">
<th box="[150,219,993,1016]" gridcol="0" gridrow="11" pageId="6" pageNumber="88">
<emphasis box="[150,219,993,1016]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,993,1016]" gridcol="1" gridrow="11" pageId="6" pageNumber="88">
<emphasis box="[360,452,993,1016]" italics="true" pageId="6" pageNumber="88">intertexta</emphasis>
</td>
</taxonomicName>
<td box="[600,753,993,1016]" gridcol="2" gridrow="11" pageId="6" pageNumber="88">RSP277</td>
<td box="[788,970,993,1016]" gridcol="3" gridrow="11" pageId="6" pageNumber="88">PEM R18661</td>
<td box="[1000,1122,993,1016]" gridcol="4" gridrow="11" pageId="6" pageNumber="88">HG005190</td>
<td box="[1147,1270,993,1016]" gridcol="5" gridrow="11" pageId="6" pageNumber="88">HG005218</td>
<td box="[1295,1417,993,1016]" gridcol="6" gridrow="11" pageId="6" pageNumber="88">HG005239</td>
<td box="[1442,1818,993,1016]" gridcol="7" gridrow="11" pageId="6" pageNumber="88">South Africa, Northern Cape Province</td>
</tr>
<tr box="[150,1818,1032,1055]" gridrow="12" pageId="6" pageNumber="88" rowspan-3="1">
<taxonomicName baseAuthorityName="Milne-edwards" baseAuthorityYear="1829" box="[150,435,1032,1055]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="87" phylum="Chordata" rank="species" species="lalandii">
<th box="[150,219,1032,1055]" gridcol="0" gridrow="12" pageId="6" pageNumber="88">
<emphasis box="[150,219,1032,1055]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1032,1055]" gridcol="1" gridrow="12" pageId="6" pageNumber="88">
<emphasis box="[360,435,1032,1055]" italics="true" pageId="6" pageNumber="88">lalandii</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1032,1055]" gridcol="2" gridrow="12" pageId="6" pageNumber="88">HB037</td>
<td box="[1000,1122,1032,1055]" gridcol="4" gridrow="12" pageId="6" pageNumber="88">HF951554</td>
<td box="[1147,1270,1032,1055]" gridcol="5" gridrow="12" pageId="6" pageNumber="88">HF951533</td>
<td box="[1295,1417,1032,1055]" gridcol="6" gridrow="12" pageId="6" pageNumber="88">HF951538</td>
<td box="[1442,1818,1032,1055]" gridcol="7" gridrow="12" pageId="6" pageNumber="88">South Africa, KwaZulu-Natal Province</td>
</tr>
<tr box="[150,1818,1070,1094]" gridrow="13" pageId="6" pageNumber="88" rowspan-3="1">
<taxonomicName baseAuthorityName="Milne-edwards" baseAuthorityYear="1829" box="[150,435,1070,1093]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="87" phylum="Chordata" rank="species" species="lalandii">
<th box="[150,219,1070,1094]" gridcol="0" gridrow="13" pageId="6" pageNumber="88">
<emphasis box="[150,219,1070,1093]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1070,1094]" gridcol="1" gridrow="13" pageId="6" pageNumber="88">
<emphasis box="[360,435,1070,1093]" italics="true" pageId="6" pageNumber="88">lalandii</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1070,1094]" gridcol="2" gridrow="13" pageId="6" pageNumber="88">HB124</td>
<td box="[1000,1122,1070,1094]" gridcol="4" gridrow="13" pageId="6" pageNumber="88">HF951553</td>
<td box="[1147,1270,1070,1094]" gridcol="5" gridrow="13" pageId="6" pageNumber="88">HF951532</td>
<td box="[1295,1417,1070,1094]" gridcol="6" gridrow="13" pageId="6" pageNumber="88">HF951537</td>
<td box="[1442,1818,1070,1094]" gridcol="7" gridrow="13" pageId="6" pageNumber="88">South Africa, Eastern Cape Province</td>
</tr>
<tr box="[150,1818,1109,1133]" gridrow="14" pageId="6" pageNumber="88" rowspan-3="1">
<taxonomicName baseAuthorityName="Milne-edwards" baseAuthorityYear="1829" box="[150,435,1109,1132]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="87" phylum="Chordata" rank="species" species="lalandii">
<th box="[150,219,1109,1133]" gridcol="0" gridrow="14" pageId="6" pageNumber="88">
<emphasis box="[150,219,1109,1132]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1109,1133]" gridcol="1" gridrow="14" pageId="6" pageNumber="88">
<emphasis box="[360,435,1109,1132]" italics="true" pageId="6" pageNumber="88">lalandii</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1109,1133]" gridcol="2" gridrow="14" pageId="6" pageNumber="88">HZ246</td>
<td box="[1000,1122,1109,1133]" gridcol="4" gridrow="14" pageId="6" pageNumber="88">HF951555</td>
<td box="[1147,1270,1109,1133]" gridcol="5" gridrow="14" pageId="6" pageNumber="88">HF951534</td>
<td box="[1295,1417,1109,1133]" gridcol="6" gridrow="14" pageId="6" pageNumber="88">HF951539</td>
<td box="[1442,1818,1109,1133]" gridcol="7" gridrow="14" pageId="6" pageNumber="88">South Africa, Eastern Cape Province</td>
</tr>
<tr box="[150,1818,1148,1171]" gridrow="15" pageId="6" pageNumber="88" rowspan-3="1">
<taxonomicName baseAuthorityName="Milne-edwards" baseAuthorityYear="1829" box="[150,435,1148,1171]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="87" phylum="Chordata" rank="species" species="lalandii">
<th box="[150,219,1148,1171]" gridcol="0" gridrow="15" pageId="6" pageNumber="88">
<emphasis box="[150,219,1148,1171]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1148,1171]" gridcol="1" gridrow="15" pageId="6" pageNumber="88">
<emphasis box="[360,435,1148,1171]" italics="true" pageId="6" pageNumber="88">lalandii</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1148,1171]" gridcol="2" gridrow="15" pageId="6" pageNumber="88">MB20982</td>
<td box="[1000,1122,1148,1171]" gridcol="4" gridrow="15" pageId="6" pageNumber="88">HG005197</td>
<td box="[1147,1270,1148,1171]" gridcol="5" gridrow="15" pageId="6" pageNumber="88">HG005225</td>
<td box="[1295,1417,1148,1171]" gridcol="6" gridrow="15" pageId="6" pageNumber="88">HG005245</td>
<td box="[1442,1818,1148,1171]" gridcol="7" gridrow="15" pageId="6" pageNumber="88">South Africa, Western Cape Province</td>
</tr>
<tr box="[150,1818,1186,1209]" gridrow="16" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Milne-edwards" baseAuthorityYear="1829" box="[150,435,1186,1209]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="87" phylum="Chordata" rank="species" species="lalandii">
<th box="[150,219,1186,1209]" gridcol="0" gridrow="16" pageId="6" pageNumber="88">
<emphasis box="[150,219,1186,1209]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1186,1209]" gridcol="1" gridrow="16" pageId="6" pageNumber="88">
<emphasis box="[360,435,1186,1209]" italics="true" pageId="6" pageNumber="88">lalandii</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1186,1209]" gridcol="2" gridrow="16" pageId="6" pageNumber="88">MBUR00414</td>
<td box="[788,970,1186,1209]" gridcol="3" gridrow="16" pageId="6" pageNumber="88">PEM R22815</td>
<td box="[1000,1122,1186,1209]" gridcol="4" gridrow="16" pageId="6" pageNumber="88">HG005195</td>
<td box="[1147,1270,1186,1209]" gridcol="5" gridrow="16" pageId="6" pageNumber="88">HG005223</td>
<td box="[1295,1417,1186,1209]" gridcol="6" gridrow="16" pageId="6" pageNumber="88">HG005243</td>
<td box="[1442,1818,1186,1209]" gridcol="7" gridrow="16" pageId="6" pageNumber="88">South Africa, Eastern Cape Province</td>
</tr>
<tr box="[150,1818,1225,1248]" gridrow="17" pageId="6" pageNumber="88" rowspan-3="1">
<taxonomicName baseAuthorityName="Milne-edwards" baseAuthorityYear="1829" box="[150,435,1225,1248]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="87" phylum="Chordata" rank="species" species="lalandii">
<th box="[150,219,1225,1248]" gridcol="0" gridrow="17" pageId="6" pageNumber="88">
<emphasis box="[150,219,1225,1248]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1225,1248]" gridcol="1" gridrow="17" pageId="6" pageNumber="88">
<emphasis box="[360,435,1225,1248]" italics="true" pageId="6" pageNumber="88">lalandii</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1225,1248]" gridcol="2" gridrow="17" pageId="6" pageNumber="88">MBUR00483</td>
<td box="[1000,1122,1225,1248]" gridcol="4" gridrow="17" pageId="6" pageNumber="88">HG005196</td>
<td box="[1147,1270,1225,1248]" gridcol="5" gridrow="17" pageId="6" pageNumber="88">HG005224</td>
<td box="[1295,1417,1225,1248]" gridcol="6" gridrow="17" pageId="6" pageNumber="88">HG005244</td>
<td box="[1442,1818,1225,1248]" gridcol="7" gridrow="17" pageId="6" pageNumber="88">South Africa, Eastern Cape Province</td>
</tr>
<tr box="[150,1818,1263,1286]" gridrow="18" pageId="6" pageNumber="88">
<th box="[150,219,1263,1286]" gridcol="0" gridrow="18" pageId="6" pageNumber="88">
<taxonomicName authorityName="Gray" authorityYear="1838" box="[150,219,1263,1286]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="genus">
<emphasis box="[150,219,1263,1286]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</taxonomicName>
</th>
<td box="[360,535,1263,1286]" gridcol="1" gridrow="18" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Branch and Bauer" baseAuthorityYear="1995" box="[360,415,1263,1286]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="species" species="livida">
<emphasis box="[360,415,1263,1286]" italics="true" pageId="6" pageNumber="88">livida</emphasis>
</taxonomicName>
</td>
<td box="[600,753,1263,1286]" gridcol="2" gridrow="18" pageId="6" pageNumber="88">KTH08-071</td>
<td box="[788,970,1263,1286]" gridcol="3" gridrow="18" pageId="6" pageNumber="88">PEM R18747</td>
<td box="[1000,1122,1263,1286]" gridcol="4" gridrow="18" pageId="6" pageNumber="88">HG005200</td>
<td box="[1147,1270,1263,1286]" gridcol="5" gridrow="18" pageId="6" pageNumber="88">HG005227</td>
<td box="[1295,1417,1263,1286]" gridcol="6" gridrow="18" pageId="6" pageNumber="88">HG005247</td>
<td box="[1442,1818,1263,1286]" gridcol="7" gridrow="18" pageId="6" pageNumber="88">South Africa, Western Cape Province</td>
</tr>
<tr box="[150,1818,1302,1325]" gridrow="19" pageId="6" pageNumber="88" rowspan-3="1">
<th box="[150,219,1302,1325]" gridcol="0" gridrow="19" pageId="6" pageNumber="88">
<taxonomicName authorityName="Gray" authorityYear="1838" box="[150,219,1302,1325]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="genus">
<emphasis box="[150,219,1302,1325]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</taxonomicName>
</th>
<td box="[360,535,1302,1325]" gridcol="1" gridrow="19" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Branch and Bauer" baseAuthorityYear="1995" box="[360,415,1302,1325]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="species" species="livida">
<emphasis box="[360,415,1302,1325]" italics="true" pageId="6" pageNumber="88">livida</emphasis>
</taxonomicName>
</td>
<td box="[600,753,1302,1325]" gridcol="2" gridrow="19" pageId="6" pageNumber="88">MB21176</td>
<td box="[1000,1122,1302,1325]" gridcol="4" gridrow="19" pageId="6" pageNumber="88">HG005201</td>
<td box="[1147,1270,1302,1325]" gridcol="5" gridrow="19" pageId="6" pageNumber="88">HG005228</td>
<td box="[1295,1417,1302,1325]" gridcol="6" gridrow="19" pageId="6" pageNumber="88">HG005248</td>
<td box="[1442,1818,1302,1325]" gridcol="7" gridrow="19" pageId="6" pageNumber="88">South Africa, Northern Cape Province</td>
</tr>
<tr box="[150,1818,1340,1364]" gridrow="20" pageId="6" pageNumber="88" rowspan-3="1">
<th box="[150,219,1340,1364]" gridcol="0" gridrow="20" pageId="6" pageNumber="88">
<taxonomicName authorityName="Gray" authorityYear="1838" box="[150,219,1340,1363]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="genus">
<emphasis box="[150,219,1340,1363]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</taxonomicName>
</th>
<td box="[360,535,1340,1364]" gridcol="1" gridrow="20" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Branch and Bauer" baseAuthorityYear="1995" box="[360,415,1340,1363]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="species" species="livida">
<emphasis box="[360,415,1340,1363]" italics="true" pageId="6" pageNumber="88">livida</emphasis>
</taxonomicName>
</td>
<td box="[600,753,1340,1364]" gridcol="2" gridrow="20" pageId="6" pageNumber="88">MB21225</td>
<td box="[1000,1122,1340,1364]" gridcol="4" gridrow="20" pageId="6" pageNumber="88">HG005202</td>
<td box="[1147,1270,1340,1364]" gridcol="5" gridrow="20" pageId="6" pageNumber="88">HG005229</td>
<td box="[1295,1417,1340,1364]" gridcol="6" gridrow="20" pageId="6" pageNumber="88">HG005249</td>
<td box="[1442,1818,1340,1364]" gridcol="7" gridrow="20" pageId="6" pageNumber="88">South Africa, Northern Cape Province</td>
</tr>
<tr box="[150,1818,1379,1402]" gridrow="21" pageId="6" pageNumber="88">
<th box="[150,219,1379,1402]" gridcol="0" gridrow="21" pageId="6" pageNumber="88">
<taxonomicName authorityName="Gray" authorityYear="1838" box="[150,219,1379,1402]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="genus">
<emphasis box="[150,219,1379,1402]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</taxonomicName>
</th>
<td box="[360,535,1379,1402]" gridcol="1" gridrow="21" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Branch and Bauer" baseAuthorityYear="1995" box="[360,415,1379,1402]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="species" species="livida">
<emphasis box="[360,415,1379,1402]" italics="true" pageId="6" pageNumber="88">livida</emphasis>
</taxonomicName>
</td>
<td box="[600,753,1379,1402]" gridcol="2" gridrow="21" pageId="6" pageNumber="88">MBUR00670</td>
<td box="[788,970,1379,1402]" gridcol="3" gridrow="21" pageId="6" pageNumber="88">PEM R22822</td>
<td box="[1000,1122,1379,1402]" gridcol="4" gridrow="21" pageId="6" pageNumber="88">HG005198</td>
<td box="[1147,1270,1379,1402]" gridcol="5" gridrow="21" pageId="6" pageNumber="88"></td>
<td box="[1295,1417,1379,1402]" gridcol="6" gridrow="21" pageId="6" pageNumber="88">HG005246</td>
<td box="[1442,1818,1379,1402]" gridcol="7" gridrow="21" pageId="6" pageNumber="88">South Africa, Eastern Cape Province</td>
</tr>
<tr box="[150,1818,1417,1441]" gridrow="22" pageId="6" pageNumber="88" rowspan-3="1">
<th box="[150,219,1417,1441]" gridcol="0" gridrow="22" pageId="6" pageNumber="88">
<taxonomicName authorityName="Gray" authorityYear="1838" box="[150,219,1417,1440]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="genus">
<emphasis box="[150,219,1417,1440]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</taxonomicName>
</th>
<td box="[360,535,1417,1441]" gridcol="1" gridrow="22" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Branch and Bauer" baseAuthorityYear="1995" box="[360,415,1417,1440]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="88" phylum="Chordata" rank="species" species="livida">
<emphasis box="[360,415,1417,1440]" italics="true" pageId="6" pageNumber="88">livida</emphasis>
</taxonomicName>
</td>
<td box="[600,753,1417,1441]" gridcol="2" gridrow="22" pageId="6" pageNumber="88">MBUR00687</td>
<td box="[1000,1122,1417,1441]" gridcol="4" gridrow="22" pageId="6" pageNumber="88">HG005199</td>
<td box="[1147,1270,1417,1441]" gridcol="5" gridrow="22" pageId="6" pageNumber="88">HG005226</td>
<td box="[1295,1417,1417,1441]" gridcol="6" gridrow="22" pageId="6" pageNumber="88"></td>
<td box="[1442,1818,1417,1441]" gridcol="7" gridrow="22" pageId="6" pageNumber="88">South Africa, Western Cape Province</td>
</tr>
<tr box="[150,1818,1456,1479]" gridrow="23" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Broadley" baseAuthorityYear="1965" box="[150,424,1456,1479]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="86" phylum="Chordata" rank="species" species="ornata">
<th box="[150,219,1456,1479]" gridcol="0" gridrow="23" pageId="6" pageNumber="88">
<emphasis box="[150,219,1456,1479]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1456,1479]" gridcol="1" gridrow="23" pageId="6" pageNumber="88">
<emphasis box="[360,424,1456,1479]" italics="true" pageId="6" pageNumber="88">ornata</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1456,1479]" gridcol="2" gridrow="23" pageId="6" pageNumber="88">AMB8635</td>
<td box="[788,970,1456,1479]" gridcol="3" gridrow="23" pageId="6" pageNumber="88">PEM R17591</td>
<td box="[1000,1122,1456,1479]" gridcol="4" gridrow="23" pageId="6" pageNumber="88">HG005206</td>
<td box="[1147,1270,1456,1479]" gridcol="5" gridrow="23" pageId="6" pageNumber="88"></td>
<td box="[1295,1417,1456,1479]" gridcol="6" gridrow="23" pageId="6" pageNumber="88">HG005252</td>
<td box="[1442,1818,1456,1479]" gridcol="7" gridrow="23" pageId="6" pageNumber="88">South Africa, Limpopo Province</td>
</tr>
<tr box="[150,1818,1494,1517]" gridrow="24" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Broadley" baseAuthorityYear="1965" box="[150,424,1494,1517]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="86" phylum="Chordata" rank="species" species="ornata">
<th box="[150,219,1494,1517]" gridcol="0" gridrow="24" pageId="6" pageNumber="88">
<emphasis box="[150,219,1494,1517]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1494,1517]" gridcol="1" gridrow="24" pageId="6" pageNumber="88">
<emphasis box="[360,424,1494,1517]" italics="true" pageId="6" pageNumber="88">ornata</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1494,1517]" gridcol="2" gridrow="24" pageId="6" pageNumber="88">MBUR01226</td>
<td box="[788,970,1494,1517]" gridcol="3" gridrow="24" pageId="6" pageNumber="88">NMB R10907</td>
<td box="[1000,1122,1494,1517]" gridcol="4" gridrow="24" pageId="6" pageNumber="88">HG005203</td>
<td box="[1147,1270,1494,1517]" gridcol="5" gridrow="24" pageId="6" pageNumber="88"></td>
<td box="[1295,1417,1494,1517]" gridcol="6" gridrow="24" pageId="6" pageNumber="88">HG005250</td>
<td box="[1442,1818,1494,1517]" gridcol="7" gridrow="24" pageId="6" pageNumber="88">South Africa, KwaZulu-Natal Province</td>
</tr>
<tr box="[150,1818,1532,1555]" gridrow="25" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Broadley" baseAuthorityYear="1965" box="[150,424,1532,1555]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="86" phylum="Chordata" rank="species" species="ornata">
<th box="[150,219,1532,1555]" gridcol="0" gridrow="25" pageId="6" pageNumber="88">
<emphasis box="[150,219,1532,1555]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1532,1555]" gridcol="1" gridrow="25" pageId="6" pageNumber="88">
<emphasis box="[360,424,1532,1555]" italics="true" pageId="6" pageNumber="88">ornata</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1532,1555]" gridcol="2" gridrow="25" pageId="6" pageNumber="88">MBUR01230</td>
<td box="[788,970,1532,1555]" gridcol="3" gridrow="25" pageId="6" pageNumber="88">NMB R10658</td>
<td box="[1000,1122,1532,1555]" gridcol="4" gridrow="25" pageId="6" pageNumber="88">HG005204</td>
<td box="[1147,1270,1532,1555]" gridcol="5" gridrow="25" pageId="6" pageNumber="88"></td>
<td box="[1295,1417,1532,1555]" gridcol="6" gridrow="25" pageId="6" pageNumber="88">HG005251</td>
<td box="[1442,1818,1532,1555]" gridcol="7" gridrow="25" pageId="6" pageNumber="88">South Africa, KwaZulu-Natal Province</td>
</tr>
<tr box="[150,1818,1570,1593]" gridrow="26" pageId="6" pageNumber="88" rowspan-3="1">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[150,457,1570,1593]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="taeniolata">
<th box="[150,219,1570,1593]" gridcol="0" gridrow="26" pageId="6" pageNumber="88">
<emphasis box="[150,219,1570,1593]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1570,1593]" gridcol="1" gridrow="26" pageId="6" pageNumber="88">
<emphasis box="[360,457,1570,1593]" italics="true" pageId="6" pageNumber="88">taeniolata</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1570,1593]" gridcol="2" gridrow="26" pageId="6" pageNumber="88">HZ250</td>
<td box="[1000,1122,1570,1593]" gridcol="4" gridrow="26" pageId="6" pageNumber="88">HG005207</td>
<td box="[1147,1270,1570,1593]" gridcol="5" gridrow="26" pageId="6" pageNumber="88"></td>
<td box="[1295,1417,1570,1593]" gridcol="6" gridrow="26" pageId="6" pageNumber="88">HG005253</td>
<td box="[1442,1818,1570,1593]" gridcol="7" gridrow="26" pageId="6" pageNumber="88">South Africa, Eastern Cape Province</td>
</tr>
<tr box="[150,1818,1609,1632]" gridrow="27" pageId="6" pageNumber="88" rowspan-3="1">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[150,457,1609,1632]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="taeniolata">
<th box="[150,219,1609,1632]" gridcol="0" gridrow="27" pageId="6" pageNumber="88">
<emphasis box="[150,219,1609,1632]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1609,1632]" gridcol="1" gridrow="27" pageId="6" pageNumber="88">
<emphasis box="[360,457,1609,1632]" italics="true" pageId="6" pageNumber="88">taeniolata</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1609,1632]" gridcol="2" gridrow="27" pageId="6" pageNumber="88">HZ251</td>
<td box="[1000,1122,1609,1632]" gridcol="4" gridrow="27" pageId="6" pageNumber="88">HG005208</td>
<td box="[1147,1270,1609,1632]" gridcol="5" gridrow="27" pageId="6" pageNumber="88">HG005230</td>
<td box="[1295,1417,1609,1632]" gridcol="6" gridrow="27" pageId="6" pageNumber="88">HG005254</td>
<td box="[1442,1818,1609,1632]" gridcol="7" gridrow="27" pageId="6" pageNumber="88">South Africa, Eastern Cape Province</td>
</tr>
<tr box="[150,1818,1648,1671]" gridrow="28" pageId="6" pageNumber="88" rowspan-3="1">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[150,457,1648,1671]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="taeniolata">
<th box="[150,219,1648,1671]" gridcol="0" gridrow="28" pageId="6" pageNumber="88">
<emphasis box="[150,219,1648,1671]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1648,1671]" gridcol="1" gridrow="28" pageId="6" pageNumber="88">
<emphasis box="[360,457,1648,1671]" italics="true" pageId="6" pageNumber="88">taeniolata</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1648,1671]" gridcol="2" gridrow="28" pageId="6" pageNumber="88">HZ252</td>
<td box="[1000,1122,1648,1671]" gridcol="4" gridrow="28" pageId="6" pageNumber="88">HG005209</td>
<td box="[1147,1270,1648,1671]" gridcol="5" gridrow="28" pageId="6" pageNumber="88"></td>
<td box="[1295,1417,1648,1671]" gridcol="6" gridrow="28" pageId="6" pageNumber="88">HG005255</td>
<td box="[1442,1818,1648,1671]" gridcol="7" gridrow="28" pageId="6" pageNumber="88">South Africa, Eastern Cape Province</td>
</tr>
<tr box="[150,1818,1686,1709]" gridrow="29" pageId="6" pageNumber="88">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[150,457,1686,1709]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="83" phylum="Chordata" rank="species" species="taeniolata">
<th box="[150,219,1686,1709]" gridcol="0" gridrow="29" pageId="6" pageNumber="88">
<emphasis box="[150,219,1686,1709]" italics="true" pageId="6" pageNumber="88">Nucras</emphasis>
</th>
<td box="[360,535,1686,1709]" gridcol="1" gridrow="29" pageId="6" pageNumber="88">
<emphasis box="[360,457,1686,1709]" italics="true" pageId="6" pageNumber="88">taeniolata</emphasis>
</td>
</taxonomicName>
<td box="[600,753,1686,1709]" gridcol="2" gridrow="29" pageId="6" pageNumber="88">PEM R18080</td>
<td box="[788,970,1686,1709]" gridcol="3" gridrow="29" pageId="6" pageNumber="88">PEM R18080</td>
<td box="[1000,1122,1686,1709]" gridcol="4" gridrow="29" pageId="6" pageNumber="88">HG005210</td>
<td box="[1147,1270,1686,1709]" gridcol="5" gridrow="29" pageId="6" pageNumber="88">HG005231</td>
<td box="[1295,1417,1686,1709]" gridcol="6" gridrow="29" pageId="6" pageNumber="88">HG005256</td>
<td box="[1442,1818,1686,1709]" gridcol="7" gridrow="29" pageId="6" pageNumber="88">South Africa, Eastern Cape Province</td>
</tr>
</table>
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,144,279]" pageId="7" pageNumber="89">Bloemfontein</paragraph>
<paragraph blockId="7.[142,703,141,1965]" pageId="7" pageNumber="89">
Museum Province Province Province Province Province Province Province Province Province National
<collectingRegion box="[261,284,386,435]" country="South Africa" name="Western Cape" pageId="7" pageNumber="89">Cape</collectingRegion>
<collectingRegion box="[299,322,395,444]" country="South Africa" name="Western Cape" pageId="7" pageNumber="89">Cape</collectingRegion>
<collectingRegion box="[337,360,386,435]" country="South Africa" name="Western Cape" pageId="7" pageNumber="89">Cape</collectingRegion>
<collectingRegion box="[375,398,386,435]" country="South Africa" name="Western Cape" pageId="7" pageNumber="89">Cape</collectingRegion>
<collectingRegion box="[413,436,386,435]" country="South Africa" name="Western Cape" pageId="7" pageNumber="89">Cape</collectingRegion>
<collectingRegion box="[528,551,395,444]" country="South Africa" name="Western Cape" pageId="7" pageNumber="89">Cape</collectingRegion>
<collectingRegion box="[566,589,395,444]" country="South Africa" name="Western Cape" pageId="7" pageNumber="89">Cape</collectingRegion>
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,476,531]" pageId="7" pageNumber="89">
<collectingRegion box="[142,165,476,531]" country="Angola" name="Namibe" pageId="7" pageNumber="89">NMB</collectingRegion>
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,460,472]" pageId="7" pageNumber="89"></paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,536,540]" pageId="7" pageNumber="89">,</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[261,665,441,568]" pageId="7" pageNumber="89">, Northern Western, Northern,, Northern Northern, Western,, Western Kamanjab,</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[642,703,377,528]" pageId="7" pageNumber="89">
<collectingRegion box="[642,703,377,528]" isEnumeration="true" pageId="7" pageNumber="89">
<collectingRegion box="[642,665,377,528]" country="South Africa" name="KwaZulu-Natal" pageId="7" pageNumber="89">KwaZulu-Natal</collectingRegion>
<collectingRegion box="[680,703,395,444]" country="South Africa" name="Western Cape" pageId="7" pageNumber="89">Cape</collectingRegion>
</collectingRegion>
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[680,703,450,537]" pageId="7" pageNumber="89">, Western</paragraph>
<paragraph blockId="7.[142,703,141,1965]" pageId="7" pageNumber="89">
Museum
<emphasis bold="true" box="[205,228,578,662]" pageId="7" pageNumber="89">Locality</emphasis>
<collectingCountry box="[261,284,537,662]" name="South Africa" pageId="7" pageNumber="89">South Africa</collectingCountry>
Africa South
<collectingCountry box="[337,436,537,662]" name="South Africa South Africa South Africa" pageId="7" pageNumber="89">South Africa South Africa South Africa</collectingCountry>
GenBank Africa South
<collectingCountry box="[566,627,537,662]" name="South Africa Namibia" pageId="7" pageNumber="89">South Africa Namibia</collectingCountry>
,
<collectingCountry box="[642,665,537,662]" name="South Africa" pageId="7" pageNumber="89">South Africa</collectingCountry>
Africa South Elizabeth Port
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,770,782]" pageId="7" pageNumber="89"></paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[205,703,703,781]" pageId="7" pageNumber="89">
<uuid box="[205,436,703,779]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[205,228,744,754]" pageId="7" pageNumber="89">1</emphasis>
005257 951543 951540 951542 951541
</uuid>
<uuid box="[489,665,703,781]" pageId="7" pageNumber="89">632229 871208 547691 871206 871207</uuid>
547718
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[205,228,754,810]" pageId="7" pageNumber="89">
<collectionCode box="[205,228,754,810]" pageId="7" pageNumber="89">RAG</collectionCode>
</emphasis>
<collectionCode box="[142,165,787,836]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="7" pageNumber="89" type="Museum">PEM</collectionCode>
<collectionCode box="[261,284,775,810]" pageId="7" pageNumber="89">HG</collectionCode>
HF HF HF HF
<collectionCode box="[489,512,781,809]" pageId="7" pageNumber="89">EF</collectionCode>
<collectionCode box="[528,551,774,809]" pageId="7" pageNumber="89">DQ</collectionCode>
HF
<collectionCode box="[604,627,775,810]" pageId="7" pageNumber="89">DQ</collectionCode>
<collectionCode box="[642,665,774,809]" pageId="7" pageNumber="89">DQ</collectionCode>
HF,
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" pageId="7" pageNumber="89">
Science
<uuid box="[205,589,850,927]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[205,228,910,921]" pageId="7" pageNumber="89">4</emphasis>
005232 951535 951536 751398 547725
</uuid>
547732 547733 547759 of
<emphasis bold="true" box="[205,228,922,957]" pageId="7" pageNumber="89">ND</emphasis>
<collectionCode box="[261,284,922,956]" pageId="7" pageNumber="89">HG</collectionCode>
— HF — HF — FR HF HF HF HF Academy
<uuid box="[261,436,998,1073]" pageId="7" pageNumber="89">005211 951559 951556 951558 951557</uuid>
<uuid box="[489,665,998,1073]" pageId="7" pageNumber="89">080358 871152 547772 871148 871149</uuid>
547800
<collectingRegion box="[142,165,1046,1143]" country="United States of America" name="California" pageId="7" pageNumber="89">California</collectingRegion>
<emphasis bold="true" box="[205,228,1067,1105]" pageId="7" pageNumber="89">S 16</emphasis>
<collectionCode box="[261,284,1069,1104]" pageId="7" pageNumber="89">HG</collectionCode>
HF HF HF HF
<collectionCode box="[489,512,1073,1104]" pageId="7" pageNumber="89">AF</collectionCode>
<collectionCode box="[528,551,1070,1104]" pageId="7" pageNumber="89">DQ</collectionCode>
HF
<collectionCode box="[604,627,1069,1104]" pageId="7" pageNumber="89">DQ</collectionCode>
<collectionCode box="[642,665,1070,1105]" pageId="7" pageNumber="89">DQ</collectionCode>
HF
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,1164,1211]" pageId="7" pageNumber="89">
<collectionCode box="[142,165,1164,1211]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="7" pageNumber="89" type="Museum">CAS</collectionCode>
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,1148,1160]" pageId="7" pageNumber="89"></paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,703,1134,1263]" pageId="7" pageNumber="89">
:
<emphasis bold="true" box="[223,246,1134,1215]" pageId="7" pageNumber="89">number</emphasis>
206725
<accessionNumber box="[299,322,1184,1260]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R18745" pageId="7" pageNumber="89">
<collectionCode box="[299,322,1244,1260]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="89">R</collectionCode>
18745
</accessionNumber>
<accessionNumber box="[337,360,1184,1260]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R16873" pageId="7" pageNumber="89">
<collectionCode box="[337,360,1244,1260]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="89">R</collectionCode>
16873
</accessionNumber>
<accessionNumber box="[375,398,1180,1255]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R11574" pageId="7" pageNumber="89">
<collectionCode box="[375,398,1239,1255]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="89">R</collectionCode>
11574
</accessionNumber>
<accessionNumber box="[413,436,1184,1260]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R16872" pageId="7" pageNumber="89">
<collectionCode box="[413,436,1244,1260]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="89">R</collectionCode>
16872
</accessionNumber>
209602
<accessionNumber box="[680,703,1184,1260]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R18376" pageId="7" pageNumber="89">
<collectionCode box="[680,703,1244,1260]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="89">R</collectionCode>
18376
</accessionNumber>
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" pageId="7" pageNumber="89">
abbreviations
<emphasis bold="true" box="[187,246,1221,1316]" pageId="7" pageNumber="89">Museum accession</emphasis>
<collectionCode box="[261,284,1269,1316]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="7" pageNumber="89" type="Museum">CAS</collectionCode>
<collectionCode box="[299,322,1266,1315]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="7" pageNumber="89" type="Museum">PEM</collectionCode>
<collectionCode box="[337,360,1266,1315]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="7" pageNumber="89" type="Museum">PEM</collectionCode>
<collectingRegion box="[375,398,1261,1316]" country="Angola" name="Namibe" pageId="7" pageNumber="89">NMB</collectingRegion>
<collectionCode box="[413,436,1266,1315]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="7" pageNumber="89" type="Museum">PEM</collectionCode>
NA NA
<collectionCode box="[604,627,1237,1316]" country="Namibia" httpUri="http://grbio.org/cool/y6s1-e7mm" name="National Museum of Namibia" pageId="7" pageNumber="89">NMNW</collectionCode>
<collectionCode box="[642,665,1269,1316]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="7" pageNumber="89" type="Museum">CAS</collectionCode>
<collectionCode box="[680,703,1266,1315]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="7" pageNumber="89" type="Museum">PEM</collectionCode>
Museum.
<emphasis bold="true" box="[187,210,1352,1447]" pageId="7" pageNumber="89">accession</emphasis>
5584 08 069 - 20650 21061 20687 0531
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[566,589,1440,1463]" pageId="7" pageNumber="89">08</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[604,703,1402,1456]" pageId="7" pageNumber="89">6001 6067 049</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,246,1448,1528]" pageId="7" pageNumber="89">
analysis
<emphasis bold="true" box="[187,246,1452,1505]" pageId="7" pageNumber="89">Field ID</emphasis>
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" pageId="7" pageNumber="89">
<collectionCode box="[261,284,1450,1504]" country="Bulgaria" httpUri="http://grbio.org/cool/rzgn-jz26" name="Asenovgrad Museum" pageId="7" pageNumber="89" type="Museum">AMB</collectionCode>
<collectionCode box="[299,322,1454,1505]" pageId="7" pageNumber="89">KTH</collectionCode>
<collectionCode box="[337,360,1468,1504]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="7" pageNumber="89">MB</collectionCode>
<collectionCode box="[375,398,1467,1504]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="7" pageNumber="89">MB</collectionCode>
<collectionCode box="[413,436,1468,1505]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="7" pageNumber="89">MB</collectionCode>
<collectionCode box="[528,551,1465,1504]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="7" pageNumber="89">MH</collectionCode>
GW
<collectionCode box="[604,627,1449,1504]" country="Bulgaria" httpUri="http://grbio.org/cool/rzgn-jz26" name="Asenovgrad Museum" pageId="7" pageNumber="89" type="Museum">AMB</collectionCode>
<collectionCode box="[642,665,1450,1505]" country="Bulgaria" httpUri="http://grbio.org/cool/rzgn-jz26" name="Asenovgrad Museum" pageId="7" pageNumber="89" type="Museum">AMB</collectionCode>
<collectionCode box="[680,703,1456,1504]" pageId="7" pageNumber="89">SVN</collectionCode>
genetic
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,1607,1626]" pageId="7" pageNumber="89">in</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,1678,1759]" pageId="7" pageNumber="89">Samples</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,1630,1674]" pageId="7" pageNumber="89">used</paragraph>
<paragraph blockId="7.[142,703,141,1965]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[205,228,1669,1744]" pageId="7" pageNumber="89">Species</emphasis>
<emphasis box="[261,703,1617,1744]" italics="true" pageId="7" pageNumber="89">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[261,284,1653,1744]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="tessellata">tessellata</taxonomicName>
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[299,322,1653,1744]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="tessellata">tessellata</taxonomicName>
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[337,360,1653,1744]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="tessellata">tessellata</taxonomicName>
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[375,398,1653,1744]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="tessellata">tessellata</taxonomicName>
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[413,436,1653,1744]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="tessellata">tessellata</taxonomicName>
longicaudata australis australis capensis capensis suborbitalis
</emphasis>
.
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,1770,1776]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[142,165,1770,1776]" pageId="7" pageNumber="89">)</emphasis>
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,1777,1881]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[142,165,1777,1881]" pageId="7" pageNumber="89">continued</emphasis>
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,1892,1904]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[142,165,1892,1904]" pageId="7" pageNumber="89">2</emphasis>
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,165,1909,1965]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[142,165,1909,1965]" pageId="7" pageNumber="89">Table</emphasis>
</paragraph>
<paragraph blockId="7.[142,703,141,1965]" box="[142,703,1803,1954]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[142,228,1881,1953]" pageId="7" pageNumber="89">(Genus</emphasis>
<emphasis box="[261,398,1884,1953]" italics="true" pageId="7" pageNumber="89">
<taxonomicName authorityName="Gray" authorityYear="1838" box="[261,284,1884,1953]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="genus">Nucras</taxonomicName>
<taxonomicName authorityName="Gray" authorityYear="1838" box="[299,322,1884,1953]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="genus">Nucras</taxonomicName>
<taxonomicName authorityName="Gray" authorityYear="1838" box="[337,360,1884,1953]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="genus">Nucras</taxonomicName>
<taxonomicName authorityName="Gray" authorityYear="1838" box="[375,398,1884,1953]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="genus">Nucras</taxonomicName>
</emphasis>
<taxonomicName authority="Outgroup" authorityName="Outgroup" box="[413,474,1862,1954]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="genus">
<emphasis box="[413,436,1884,1953]" italics="true" pageId="7" pageNumber="89">Nucras</emphasis>
Outgroup
</taxonomicName>
<emphasis box="[489,703,1803,1954]" italics="true" pageId="7" pageNumber="89">
<taxonomicName authorityName="Bedriaga" authorityYear="1884" box="[489,512,1875,1954]" class="Reptilia" family="Lacertidae" genus="Latastia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="genus">Latastia</taxonomicName>
<taxonomicName authorityName="Arnold" authorityYear="1989" box="[528,551,1803,1954]" class="Reptilia" family="Lacertidae" genus="Australolacerta" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="genus">Australolacerta</taxonomicName>
<taxonomicName authorityName="Arnold" authorityYear="1989" box="[566,589,1803,1954]" class="Reptilia" family="Lacertidae" genus="Australolacerta" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="genus">Australolacerta</taxonomicName>
<taxonomicName authorityName="W.Peters" authorityYear="1854" box="[604,627,1844,1954]" class="Reptilia" family="Lacertidae" genus="Ichnotropis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="genus">Ichnotropis</taxonomicName>
<taxonomicName authorityName="W.Peters" authorityYear="1854" box="[642,665,1844,1954]" class="Reptilia" family="Lacertidae" genus="Ichnotropis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="genus">Ichnotropis</taxonomicName>
<taxonomicName authorityName="Gray" authorityYear="1838" box="[680,703,1876,1954]" class="Reptilia" family="Lacertidae" genus="Meroles" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="genus">Meroles</taxonomicName>
</emphasis>
</paragraph>
<paragraph blockId="7.[791,1396,144,680]" pageId="7" pageNumber="89">
continuous light stripe with the outer edges of the parietals (similar to
<bibRefCitation author="Broadley DG" box="[910,1100,176,200]" journalOrPublisher="Arnoldia" pageId="7" pageNumber="89" pagination="1 - 35" part="20" refId="ref8654" refString="Broadley DG. 1972. A review of the Nucras tessellata group (Sauria: Lacertidae). Arnoldia 20: 1 - 35." title="A review of the Nucras tessellata group (Sauria: Lacertidae)" type="journal article" year="1972">Broadleys (1972)</bibRefCitation>
<taxonomicName authority="tessellata" box="[1110,1348,176,200]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="subSpecies" species="tessellata" subSpecies="tessellata">
<emphasis box="[1110,1348,176,200]" italics="true" pageId="7" pageNumber="89">N. tessellata tessellata</emphasis>
</taxonomicName>
var. “
<collectionCode box="[802,820,208,232]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="89">T</collectionCode>
;” differs from
<taxonomicName baseAuthorityName="Branch and Bauer" baseAuthorityYear="1995" box="[968,1056,208,232]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="livida">
<emphasis box="[968,1056,208,232]" italics="true" pageId="7" pageNumber="89">N. livida</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[1104,1231,208,232]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="tessellata">
<emphasis box="[1104,1231,208,232]" italics="true" pageId="7" pageNumber="89">N. tessellata</emphasis>
</taxonomicName>
where the outer stripes often do not form a continuous light stripe with the outer edges of the parietals; differs from
<taxonomicName box="[1225,1394,272,296]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="caeiscaudata">
<emphasis box="[1225,1394,272,296]" italics="true" pageId="7" pageNumber="89">N. caeiscaudata</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Broadley" baseAuthorityYear="1965" box="[841,947,305,328]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="ornata">
<emphasis box="[841,947,305,328]" italics="true" pageId="7" pageNumber="89">N. ornata</emphasis>
</taxonomicName>
where there are only three longitudinal stripes present on nape and sometimes the vertebral ones are absent), well defined occipital scale separating parietals (versus reduced or absent in northern
<collectingCountry box="[1192,1282,400,424]" name="Namibia" pageId="7" pageNumber="89">Namibia</collectingCountry>
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1882" box="[1290,1388,400,424]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="holubi">
<emphasis box="[1290,1388,400,424]" italics="true" pageId="7" pageNumber="89">N. holubi</emphasis>
</taxonomicName>
, which is referred to as
<taxonomicName authority="Parker" authorityName="Parker" box="[1036,1361,432,456]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="subSpecies" species="intertexta" subSpecies="damarana">
<emphasis box="[1036,1282,432,456]" italics="true" pageId="7" pageNumber="89">N. intertexta damarana</emphasis>
Parker
</taxonomicName>
; as well as absent in
<taxonomicName authorityName="Broadley" authorityYear="1972" box="[966,1132,464,488]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="caesicaudata">
<emphasis box="[966,1132,464,488]" italics="true" pageId="7" pageNumber="89">N. caesicaudata</emphasis>
</taxonomicName>
), parietal foramen absent (often present in all other species except
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[1241,1380,496,520]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="taeniolata">
<emphasis box="[1241,1380,496,520]" italics="true" pageId="7" pageNumber="89">N. taeniolata</emphasis>
</taxonomicName>
), and postnasals separated (usually fused in
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[1230,1365,528,552]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="taeniolata">
<emphasis box="[1230,1365,528,552]" italics="true" pageId="7" pageNumber="89">N. taeniolata</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph blockId="7.[791,1396,144,680]" pageId="7" pageNumber="89">
In the phylogenetic analysis, the uncorrected
<emphasis box="[1278,1291,561,584]" italics="true" pageId="7" pageNumber="89">p</emphasis>
-distances show that this clade differs by&gt;8% for 16S,&gt;14% for ND4, and&gt;1% for
<collectionCode box="[989,1049,624,648]" pageId="7" pageNumber="89">RAG</collectionCode>
1 sequence divergence from other members of the
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[958,1087,656,680]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="89" phylum="Chordata" rank="species" species="tessellata">
<emphasis box="[958,1087,656,680]" italics="true" pageId="7" pageNumber="89">N. tessellata</emphasis>
</taxonomicName>
clade.
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="91" pageId="7" pageNumber="89" type="description">
<paragraph blockId="7.[791,1396,720,1960]" pageId="7" pageNumber="89">
<emphasis bold="true" box="[791,1146,720,744]" pageId="7" pageNumber="89">
Description of
<typeStatus box="[953,1054,720,744]" pageId="7" pageNumber="89">Holotype</typeStatus>
(
<figureCitation box="[1068,1132,720,744]" captionStart="Fig" captionStartId="9.[141,176,1075,1097]" captionTargetBox="[141,1394,141,1063]" captionTargetId="figure@9.[141,1394,141,1066]" captionTargetPageId="9" captionText="Fig. 4. Nucras broadleyi sp. nov. A holotype, adult male, PEM R24005 (AG 18) in life; B general habitat photo of type locality, 10 km west of Lola, edge of Bentiaba River valley, Namibe Province, Angola; C lateral close-up of head of holotype; D dorsal close-up of head of holotype; E ventral close-up of head of holotype (Photos: Bill Branch)." figureDoi="http://doi.org/10.5281/zenodo.3731985" httpUri="https://zenodo.org/record/3731985/files/figure.png" pageId="7" pageNumber="89">Fig. 4</figureCitation>
).
</emphasis>
Body relatively slender (
<collectionCode box="[798,848,752,776]" pageId="7" pageNumber="89">SVL</collectionCode>
approx. 4.5 times the head length, tail truncated), with hindlimbs larger than forelimbs (femur of hind limb equal to length of tibia); head narrow and elongated (56% longer than wide) with narrow pointed but blunt snout, that is slightly longer than distance from back of eye to rear of ear opening. Rostrum protruding and visible from below. Nasals paired and in contact (
<quantity box="[1108,1192,944,968]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="89" unit="mm" value="0.2">0.2 mm</quantity>
suture length), not swollen, nostril directed backwards separating postnasals. Frontonasal single, wider than long (1.1 ×
<quantity box="[1291,1384,1008,1032]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="89" unit="mm" value="1.8">1.8 mm</quantity>
). Prefrontals paired and in broad median contact with one another (
<quantity box="[882,963,1072,1096]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="89" unit="mm" value="0.6">0.6 mm</quantity>
suture length), wider than long (1.1 ×
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="89" unit="mm" value="1.2">1.2 mm</quantity>
). Frontal entire, longer than wide (2.7 ×
<quantity box="[1249,1332,1104,1128]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="89" unit="mm" value="1.9">1.9 mm</quantity>
). Two large rounded supraoculars, both in contact with the frontal, with anterior supraocular preceded by a single large scale in contact with prefrontal, frontonasal, and posterior loreal, with posterior supraocular bordered by a single large scale in contact with parietal and frontoparietal. Paired frontoparietal in broad contact (
<quantity box="[1141,1228,1296,1320]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="89" unit="mm" value="1.3">1.3 mm</quantity>
suture length), nearly as wide as long (1.7 ×
<quantity box="[1107,1193,1328,1352]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="89" unit="mm" value="1.5">1.5 mm</quantity>
). Parietals twice as long as wide (3.1 ×
<quantity box="[1016,1103,1360,1384]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="89" unit="mm" value="1.8">1.8 mm</quantity>
), fully separate by a large, pentagonal interparietal (2.5 ×
<quantity box="[1104,1183,1392,1416]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="89" unit="mm" value="1.2">1.2 mm</quantity>
) that is twice as long as wide, slightly shorter than frontoparietals and nearly equal to length of frontonasal and prefrontal combined. Small subtriangular occipital (0.5 ×
<quantity box="[1166,1250,1488,1512]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="89" unit="mm" value="0.7">0.7 mm</quantity>
). Two loreals, second much larger than first. Six supraciliaries on each side, 1
<superScript attach="left" box="[865,875,1550,1564]" fontSize="6" pageId="7" pageNumber="89">st</superScript>
is the longest. A single minute granule scale between supraocular and supracilliares on right side, none on left side. Four supralabials anterior to subocular and three supralabials posterior to subocular, on both sides. Subocular slightly elevated medial and bordering the lip, its lower border being shorter than the upper. Three temporal scales, first longer than others, smooth. Tympanic shield as wide as long, border of ear opening. No ear lobes. Lower eyelid with transparent brille formed by five larger scales, surrounded by numerous smaller scales. Lower eyelid separated from subocular and enlarged temporal scales by a series of 10 smaller scales. Small scale above 3
<superScript attach="right" box="[804,817,1933,1948]" fontSize="6" pageId="7" pageNumber="89">rd</superScript>
supralabial separating the posterior loreal and subocular.
</paragraph>
<caption pageId="8" pageNumber="90">
<paragraph blockId="8.[142,1396,144,194]" pageId="8" pageNumber="90">
<emphasis bold="true" box="[142,225,144,165]" pageId="8" pageNumber="90">Table 3.</emphasis>
Pairwise uncorrected net
<emphasis box="[482,494,144,165]" italics="true" pageId="8" pageNumber="90">p</emphasis>
-distances for species of
<taxonomicName box="[739,808,144,165]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="genus">
<emphasis box="[739,808,144,165]" italics="true" pageId="8" pageNumber="90">Nucras</emphasis>
</taxonomicName>
: a) 16S, b) ND4, c) RAG-1. Comparisons not made due to missing data indicated by na.
</paragraph>
</caption>
<paragraph pageId="8" pageNumber="90">
<table box="[152,1367,212,1319]" gridcols="12" gridrows="29" pageId="8" pageNumber="90">
<tr box="[152,1367,212,234]" gridrow="0" pageId="8" pageNumber="90" rowspan-1="1" rowspan-2="1">
<th box="[152,170,212,234]" gridcol="0" gridrow="0" pageId="8" pageNumber="90">a)</th>
<th box="[437,492,212,234]" gridcol="3" gridrow="0" pageId="8" pageNumber="90">1</th>
<th box="[547,601,212,234]" gridcol="4" gridrow="0" pageId="8" pageNumber="90">2</th>
<th box="[656,711,212,234]" gridcol="5" gridrow="0" pageId="8" pageNumber="90">3</th>
<th box="[765,820,212,234]" gridcol="6" gridrow="0" pageId="8" pageNumber="90">4</th>
<th box="[875,930,212,234]" gridcol="7" gridrow="0" pageId="8" pageNumber="90">5</th>
<th box="[984,1039,212,234]" gridcol="8" gridrow="0" pageId="8" pageNumber="90">6</th>
<th box="[1094,1148,212,234]" gridcol="9" gridrow="0" pageId="8" pageNumber="90">7</th>
<th box="[1203,1258,212,234]" gridcol="10" gridrow="0" pageId="8" pageNumber="90">8</th>
<th box="[1312,1367,212,234]" gridcol="11" gridrow="0" pageId="8" pageNumber="90">9</th>
</tr>
<tr box="[152,1367,251,273]" gridrow="1" pageId="8" pageNumber="90" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td box="[188,201,251,273]" gridcol="1" gridrow="1" pageId="8" pageNumber="90">1</td>
<td box="[221,391,251,273]" gridcol="2" gridrow="1" pageId="8" pageNumber="90">
<taxonomicName authority="Branch &amp; Conradie &amp; Pinto &amp; Tolley, 2019" authorityName="Branch &amp; Conradie &amp; Pinto &amp; Tolley" authorityYear="2019" box="[221,312,251,272]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov">
<emphasis box="[221,312,251,272]" italics="true" pageId="8" pageNumber="90">broadleyi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[318,390,252,273]" pageId="8" pageNumber="90">
<taxonomicNameLabel box="[318,390,252,273]" pageId="8" pageNumber="90" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
</td>
<td box="[437,492,251,273]" gridcol="3" gridrow="1" pageId="8" pageNumber="90">0.048</td>
</tr>
<tr box="[152,1367,290,311]" gridrow="2" pageId="8" pageNumber="90" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td box="[188,201,290,311]" gridcol="1" gridrow="2" pageId="8" pageNumber="90">2</td>
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<td box="[188,201,406,429]" gridcol="1" gridrow="5" pageId="8" pageNumber="90">5</td>
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<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[221,312,407,428]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="tessellata">
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<taxonomicName authorityName="Neumann" authorityYear="1900" box="[222,326,445,466]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="boulengeri">
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<emphasis box="[999,1023,446,467]" italics="true" pageId="8" pageNumber="90">na</emphasis>
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<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1882" box="[222,283,484,505]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="holubi">
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<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[222,314,524,545]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="intertexta">
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<th box="[152,1367,600,622]" colspan="12" colspanRight="11" gridcol="0" gridrow="10" pageId="8" pageNumber="90">b)</th>
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<tr box="[152,1367,639,660]" gridrow="11" pageId="8" pageNumber="90" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
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<taxonomicName authorityName="Neumann" authorityYear="1900" box="[222,326,833,854]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="boulengeri">
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<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1882" box="[222,283,872,893]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="holubi">
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<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[222,314,911,932]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="intertexta">
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<td box="[1203,1258,910,932]" gridcol="10" gridrow="18" pageId="8" pageNumber="90">0.017</td>
</tr>
<tr box="[152,1367,950,971]" gridrow="19" pageId="8" pageNumber="90">
<th box="[152,1367,950,971]" colspan="12" colspanRight="11" gridcol="0" gridrow="19" pageId="8" pageNumber="90">c)</th>
</tr>
<tr box="[152,1367,988,1009]" gridrow="20" pageId="8" pageNumber="90" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td box="[188,201,988,1009]" gridcol="1" gridrow="20" pageId="8" pageNumber="90">1</td>
<td box="[221,391,988,1009]" gridcol="2" gridrow="20" pageId="8" pageNumber="90">
<taxonomicName authority="Branch &amp; Conradie &amp; Pinto &amp; Tolley, 2019" authorityName="Branch &amp; Conradie &amp; Pinto &amp; Tolley" authorityYear="2019" box="[222,313,988,1009]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov">
<emphasis box="[222,313,988,1009]" italics="true" pageId="8" pageNumber="90">broadleyi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[319,391,988,1009]" pageId="8" pageNumber="90">
<taxonomicNameLabel box="[319,391,988,1009]" pageId="8" pageNumber="90" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
</td>
<td box="[437,492,988,1009]" gridcol="3" gridrow="20" pageId="8" pageNumber="90">0.001</td>
</tr>
<tr box="[152,1367,1027,1048]" gridrow="21" pageId="8" pageNumber="90" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td box="[188,201,1027,1048]" gridcol="1" gridrow="21" pageId="8" pageNumber="90">2</td>
<td box="[221,391,1027,1048]" gridcol="2" gridrow="21" pageId="8" pageNumber="90">
<taxonomicName baseAuthorityName="Milne-edwards" baseAuthorityYear="1829" box="[222,297,1027,1048]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="lalandii">
<emphasis box="[222,297,1027,1048]" italics="true" pageId="8" pageNumber="90">lalandii</emphasis>
</taxonomicName>
</td>
<td box="[437,492,1027,1048]" gridcol="3" gridrow="21" pageId="8" pageNumber="90">0.012</td>
<td box="[547,601,1027,1048]" gridcol="4" gridrow="21" pageId="8" pageNumber="90">0.002</td>
</tr>
<tr box="[152,1367,1066,1087]" gridrow="22" pageId="8" pageNumber="90" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td box="[188,201,1066,1087]" gridcol="1" gridrow="22" pageId="8" pageNumber="90">3</td>
<td box="[221,391,1066,1087]" gridcol="2" gridrow="22" pageId="8" pageNumber="90">
<taxonomicName baseAuthorityName="Branch and Bauer" baseAuthorityYear="1995" box="[222,277,1066,1087]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="livida">
<emphasis box="[222,277,1066,1087]" italics="true" pageId="8" pageNumber="90">livida</emphasis>
</taxonomicName>
</td>
<td box="[437,492,1066,1087]" gridcol="3" gridrow="22" pageId="8" pageNumber="90">0.014</td>
<td box="[547,601,1066,1087]" gridcol="4" gridrow="22" pageId="8" pageNumber="90">0.009</td>
<td box="[656,711,1066,1087]" gridcol="5" gridrow="22" pageId="8" pageNumber="90">0.004</td>
</tr>
<tr box="[152,1367,1104,1125]" gridrow="23" pageId="8" pageNumber="90" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td box="[188,201,1104,1125]" gridcol="1" gridrow="23" pageId="8" pageNumber="90">4</td>
<td box="[221,391,1104,1125]" gridcol="2" gridrow="23" pageId="8" pageNumber="90">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[222,319,1104,1125]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="taeniolata">
<emphasis box="[222,319,1104,1125]" italics="true" pageId="8" pageNumber="90">taeniolata</emphasis>
</taxonomicName>
</td>
<td box="[437,492,1104,1125]" gridcol="3" gridrow="23" pageId="8" pageNumber="90">0.013</td>
<td box="[547,601,1104,1125]" gridcol="4" gridrow="23" pageId="8" pageNumber="90">0.007</td>
<td box="[656,711,1104,1125]" gridcol="5" gridrow="23" pageId="8" pageNumber="90">0.009</td>
<td box="[765,820,1104,1125]" gridcol="6" gridrow="23" pageId="8" pageNumber="90">0.004</td>
</tr>
<tr box="[152,1367,1143,1165]" gridrow="24" pageId="8" pageNumber="90" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-8="1" rowspan-9="1">
<td box="[188,201,1143,1165]" gridcol="1" gridrow="24" pageId="8" pageNumber="90">5</td>
<td box="[221,391,1143,1165]" gridcol="2" gridrow="24" pageId="8" pageNumber="90">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[222,313,1143,1164]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="tessellata">
<emphasis box="[222,313,1143,1164]" italics="true" pageId="8" pageNumber="90">tessellata</emphasis>
</taxonomicName>
</td>
<td box="[437,492,1143,1165]" gridcol="3" gridrow="24" pageId="8" pageNumber="90">0.012</td>
<td box="[547,601,1143,1165]" gridcol="4" gridrow="24" pageId="8" pageNumber="90">0.006</td>
<td box="[656,711,1143,1165]" gridcol="5" gridrow="24" pageId="8" pageNumber="90">0.008</td>
<td box="[765,820,1143,1165]" gridcol="6" gridrow="24" pageId="8" pageNumber="90">0.007</td>
<td box="[875,930,1143,1165]" gridcol="7" gridrow="24" pageId="8" pageNumber="90">0.008</td>
</tr>
<tr box="[152,1367,1182,1204]" gridrow="25" pageId="8" pageNumber="90" rowspan-0="1" rowspan-10="1" rowspan-11="1" rowspan-9="1">
<td box="[188,201,1182,1204]" gridcol="1" gridrow="25" pageId="8" pageNumber="90">6</td>
<td box="[221,391,1182,1204]" gridcol="2" gridrow="25" pageId="8" pageNumber="90">
<taxonomicName authorityName="Neumann" authorityYear="1900" box="[222,326,1182,1203]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="boulengeri">
<emphasis box="[222,326,1182,1203]" italics="true" pageId="8" pageNumber="90">boulengeri</emphasis>
</taxonomicName>
</td>
<td box="[437,492,1182,1204]" gridcol="3" gridrow="25" pageId="8" pageNumber="90">0.066</td>
<td box="[547,601,1182,1204]" gridcol="4" gridrow="25" pageId="8" pageNumber="90">0.059</td>
<td box="[656,711,1182,1204]" gridcol="5" gridrow="25" pageId="8" pageNumber="90">0.064</td>
<td box="[765,820,1182,1204]" gridcol="6" gridrow="25" pageId="8" pageNumber="90">0.064</td>
<td box="[875,930,1182,1204]" gridcol="7" gridrow="25" pageId="8" pageNumber="90">0.058</td>
<td box="[984,1039,1182,1204]" gridcol="8" gridrow="25" pageId="8" pageNumber="90">
<emphasis box="[1000,1024,1183,1204]" italics="true" pageId="8" pageNumber="90">na</emphasis>
</td>
</tr>
<tr box="[152,1367,1221,1243]" gridrow="26" pageId="8" pageNumber="90" rowspan-0="1" rowspan-10="1" rowspan-11="1">
<td box="[188,201,1221,1243]" gridcol="1" gridrow="26" pageId="8" pageNumber="90">7</td>
<td box="[221,391,1221,1243]" gridcol="2" gridrow="26" pageId="8" pageNumber="90">
<taxonomicName baseAuthorityName="Steindachner" baseAuthorityYear="1882" box="[222,283,1221,1242]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="holubi">
<emphasis box="[222,283,1221,1242]" italics="true" pageId="8" pageNumber="90">holubi</emphasis>
</taxonomicName>
</td>
<td box="[437,492,1221,1243]" gridcol="3" gridrow="26" pageId="8" pageNumber="90">0.020</td>
<td box="[547,601,1221,1243]" gridcol="4" gridrow="26" pageId="8" pageNumber="90">0.015</td>
<td box="[656,711,1221,1243]" gridcol="5" gridrow="26" pageId="8" pageNumber="90">0.017</td>
<td box="[765,820,1221,1243]" gridcol="6" gridrow="26" pageId="8" pageNumber="90">0.016</td>
<td box="[875,930,1221,1243]" gridcol="7" gridrow="26" pageId="8" pageNumber="90">0.016</td>
<td box="[984,1039,1221,1243]" gridcol="8" gridrow="26" pageId="8" pageNumber="90">0.068</td>
<td box="[1094,1148,1221,1243]" gridcol="9" gridrow="26" pageId="8" pageNumber="90">0.004</td>
</tr>
<tr box="[152,1367,1259,1281]" gridrow="27" pageId="8" pageNumber="90" rowspan-0="1" rowspan-11="1">
<td box="[188,201,1259,1281]" gridcol="1" gridrow="27" pageId="8" pageNumber="90">8</td>
<td box="[221,391,1259,1281]" gridcol="2" gridrow="27" pageId="8" pageNumber="90">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[222,314,1260,1281]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="intertexta">
<emphasis box="[222,314,1260,1281]" italics="true" pageId="8" pageNumber="90">intertexta</emphasis>
</taxonomicName>
</td>
<td box="[437,492,1259,1281]" gridcol="3" gridrow="27" pageId="8" pageNumber="90">0.017</td>
<td box="[547,601,1259,1281]" gridcol="4" gridrow="27" pageId="8" pageNumber="90">0.012</td>
<td box="[656,711,1259,1281]" gridcol="5" gridrow="27" pageId="8" pageNumber="90">0.013</td>
<td box="[765,820,1259,1281]" gridcol="6" gridrow="27" pageId="8" pageNumber="90">0.012</td>
<td box="[875,930,1259,1281]" gridcol="7" gridrow="27" pageId="8" pageNumber="90">0.013</td>
<td box="[984,1039,1259,1281]" gridcol="8" gridrow="27" pageId="8" pageNumber="90">0.064</td>
<td box="[1094,1148,1259,1281]" gridcol="9" gridrow="27" pageId="8" pageNumber="90">0.015</td>
<td box="[1203,1258,1259,1281]" gridcol="10" gridrow="27" pageId="8" pageNumber="90">0.001</td>
</tr>
<tr box="[152,1367,1297,1319]" gridrow="28" pageId="8" pageNumber="90" rowspan-0="1">
<td box="[188,201,1297,1319]" gridcol="1" gridrow="28" pageId="8" pageNumber="90">9</td>
<td box="[221,391,1297,1319]" gridcol="2" gridrow="28" pageId="8" pageNumber="90">
<taxonomicName baseAuthorityName="Broadley" baseAuthorityYear="1965" box="[222,286,1298,1319]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="90" phylum="Chordata" rank="species" species="ornata">
<emphasis box="[222,286,1298,1319]" italics="true" pageId="8" pageNumber="90">ornata</emphasis>
</taxonomicName>
</td>
<td box="[437,492,1297,1319]" gridcol="3" gridrow="28" pageId="8" pageNumber="90">0.021</td>
<td box="[547,601,1297,1319]" gridcol="4" gridrow="28" pageId="8" pageNumber="90">0.016</td>
<td box="[656,711,1297,1319]" gridcol="5" gridrow="28" pageId="8" pageNumber="90">0.016</td>
<td box="[765,820,1297,1319]" gridcol="6" gridrow="28" pageId="8" pageNumber="90">0.016</td>
<td box="[875,930,1297,1319]" gridcol="7" gridrow="28" pageId="8" pageNumber="90">0.017</td>
<td box="[984,1039,1297,1319]" gridcol="8" gridrow="28" pageId="8" pageNumber="90">0.064</td>
<td box="[1094,1148,1297,1319]" gridcol="9" gridrow="28" pageId="8" pageNumber="90">0.019</td>
<td box="[1203,1258,1297,1319]" gridcol="10" gridrow="28" pageId="8" pageNumber="90">0.004</td>
<td box="[1312,1367,1297,1319]" gridcol="11" gridrow="28" pageId="8" pageNumber="90">0.001</td>
</tr>
</table>
</paragraph>
<paragraph blockId="8.[141,746,1342,1945]" lastBlockId="8.[791,1396,1345,1945]" pageId="8" pageNumber="90">
Enlarged scale bordering 1
<superScript attach="left" box="[433,443,1342,1356]" fontSize="6" pageId="8" pageNumber="90">st</superScript>
post subocular, supralabial, and the subocular. Six infralabials on both sides, with 3
<superScript attach="left" box="[154,167,1405,1420]" fontSize="6" pageId="8" pageNumber="90">rd</superScript>
being longest; four enlarged pairs of chin shields, last largest and first three in broad contact. Twenty-four gular scales in a straight line between symphysis of chin shields and median collar plate, equal in size except last 45 larger. Collar free, comprising seven enlarged plates (median subtriangular) and extending slightly onto side of neck as a crease, bordered by 23 smaller scales. Dorsal scales small, juxtaposed, granular, smooth, larger on sides toward ventrals. Midbody scales 42. Ventral plates eight longitudinal and 28 transverse rows (from collar to groin), plates of the innermost rows longer than broad, with outer row notably smaller than other rows, transverse row of ventrals across chest just behind collar longer than broad; preanal scales irregular, median ones larger. Scales on upper surface of forearm large, smooth or slightly keeled. Scales on lower surface of forearm with eight enlarged plates, at least twice the width of scales on upper forearm. Scales on upper surface of tibia rhombic, subimbricate, smooth, and much larger than dorsal scales. Tibia below with a series of large plates. Subdigital lamellae under fourth toe 23
<collectionCode box="[922,940,1440,1464]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="90">R</collectionCode>
/
<specimenCount box="[949,996,1440,1464]" pageId="8" pageNumber="90" type="generic">25L.</specimenCount>
Femoral pores 13
<collectionCode box="[1179,1198,1440,1464]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="90">R</collectionCode>
/
<specimenCount box="[1207,1253,1440,1464]" pageId="8" pageNumber="90" type="generic">15L.</specimenCount>
Dorsal scales on tail oblique, strongly keeled diagonally, and truncate behind, ventral scales on tail obtusely keeled.
</paragraph>
<paragraph blockId="8.[791,1396,1345,1945]" pageId="8" pageNumber="90">
<emphasis box="[823,944,1537,1561]" italics="true" pageId="8" pageNumber="90">Coloration.</emphasis>
Dorsum with eight pale cream to white dorsolateral longitudinal stripes, separated by dark brown to black stripes. These stripes are more boldly patterned anteriorly, fading posteriorly. No light vertebral stripe. The two pale paravertebral stripes are separated by a very narrow strip of darker scales that starts on the interparietal through the occipital scale and fades posteriorly onto body and tail. The dorsolateral stripe extending along outer borders of parietals continues onto the tail. It is followed by the upper lateral stripe extending from posterior of the eye onto the head through the mid-temporal with a brief break above the ear opening, and continues onto the tail. The lower lateral stripe starts at the subocular, through the
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3731985" ID-Zenodo-Dep="3731985" httpUri="https://zenodo.org/record/3731985/files/figure.png" pageId="9" pageNumber="91" startId="9.[141,176,1075,1097]" targetBox="[141,1394,141,1063]" targetPageId="9">
<paragraph blockId="9.[141,1395,1075,1155]" pageId="9" pageNumber="91">
<emphasis bold="true" box="[141,204,1075,1097]" pageId="9" pageNumber="91">Fig. 4.</emphasis>
<taxonomicName authority="Branch &amp; Conradie &amp; Pinto &amp; Tolley, 2019" authorityName="Branch &amp; Conradie &amp; Pinto &amp; Tolley" authorityYear="2019" box="[210,376,1076,1097]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="91" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
<emphasis box="[210,376,1076,1097]" italics="true" pageId="9" pageNumber="91">Nucras broadleyi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[381,457,1076,1097]" pageId="9" pageNumber="91">
<taxonomicNameLabel box="[381,457,1076,1097]" pageId="9" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A holotype, adult male, PEM R24005 (AG 18) in life; B general habitat photo of type locality, 10 km west of Lola, edge of Bentiaba River valley, Namibe Province, Angola; C lateral close-up of head of holotype; D dorsal close-up of head of holotype; E ventral close-up of head of holotype (
<emphasis box="[828,1021,1133,1154]" italics="true" pageId="9" pageNumber="91">Photos: Bill Branch</emphasis>
).
</paragraph>
</caption>
<paragraph blockId="9.[141,746,1181,1365]" pageId="9" pageNumber="91">ear opening, broken briefly above the arm, after which it continues all the way onto the tail. Ventrum white and lower limbs oblique white. Fore limbs upper surface black with scattered pale blotches. Hind limbs light brown with pale blotches. Upper surface of tail red-brown, similar to hind limbs. Scales bordering the orbit are black edged.</paragraph>
<paragraph blockId="9.[141,746,1405,1749]" pageId="9" pageNumber="91">
<emphasis bold="true" box="[141,384,1405,1429]" pageId="9" pageNumber="91">
Variation (
<figureCitation box="[265,371,1405,1429]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="10.[141,176,1072,1094]" captionStartId-1="11.[139,174,843,865]" captionTargetBox-0="[141,1394,141,1060]" captionTargetBox-1="[146,1391,140,834]" captionTargetId-0="figure@10.[141,1394,139,1064]" captionTargetId-1="figure@11.[139,1392,133,837]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Fig. 5. Nucras broadleyi sp. nov. A paratype, adult female, PEM R24157 (AG 166) dorsal view; B ventral view; C dorsal close-up of head of paratype; D ventral close-up of head of paratype; E lateral close-up of head of paratype; F general habitat photo of type locality, 8.8 km southwest of Farm Mucongo, Namibe Province, Angola (Photos: Bill Branch)." captionText-1="Fig. 6. Variation of Nucras broadleyi nov. sp. dorsal color pattern.A TM 40392 from “34 km S of Moçâmedes to Porto Alexandre;” B BM 1970.6.29.10 from Ponang Kuma (=Donquena); C MHNC 91.0524 from Capelongo; D MD 1967 from “km 34 de la route de Moçâmedes à Sa da Bandeira” (Photos: A,B Bill Branch, C, D Luis Ceríaco)." figureDoi-0="http://doi.org/10.5281/zenodo.3731987" figureDoi-1="http://doi.org/10.5281/zenodo.3731989" httpUri-0="https://zenodo.org/record/3731987/files/figure.png" httpUri-1="https://zenodo.org/record/3731989/files/figure.png" pageId="9" pageNumber="91">Figs. 56</figureCitation>
).
</emphasis>
Meristic and escalation data are summarized in
<tableCitation box="[309,391,1437,1462]" captionStart="Table 1" captionStartId="5.[139,195,316,339]" captionText="Table 1. Meristic and scalation data for Angolan Nucras broadleyi sp. nov. PEM Port Elizabeth Museum, BM Natural History Museum, London (formerly British Museum), MD Museu Dundo Regional do Dundo, Angola, TM Ditsong Museum of Natural History, South Africa (formerly Transvaal Museum), MBL Museu Bocage Lisboa, Portugal (material destroyed), MHNC Musée d' Histoire Naturelle de La Chaux-de-Fonds, Chaux-de-Fonds, Switzerland." pageId="9" pageNumber="91">Table 1</tableCitation>
. The largest specimen examined is (
<collectionCode box="[179,221,1470,1494]" country="United Kingdom" name="Bristol Museum" pageId="9" pageNumber="91">BM</collectionCode>
1907.6.29.10) 74 +
<quantity box="[447,541,1470,1494]" metricMagnitude="-1" metricUnit="m" metricValue="1.44" pageId="9" pageNumber="91" unit="mm" value="144.0">144 mm</quantity>
(tail regenerated). Regarding coloration, there seem to be three main variations among material examined: 1) 89 longitudinal stripes as in
<typeStatus box="[170,261,1565,1589]" pageId="9" pageNumber="91">holotype</typeStatus>
(in
<collectionCode box="[308,362,1566,1589]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="9" pageNumber="91" type="Museum">PEM</collectionCode>
<accessionNumber box="[371,455,1565,1589]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R24005" pageId="9" pageNumber="91">
<collectionCode box="[371,389,1565,1589]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="91">R</collectionCode>
24005
</accessionNumber>
,
<collectionCode box="[468,524,1566,1589]" country="Portugal" httpUri="http://grbio.org/cool/pyk6-nt7k" name="Museu Nacional de Historia Natural" pageId="9" pageNumber="91">MBL</collectionCode>
647a, 647b,
<collectionCode box="[666,745,1566,1590]" country="0" httpUri="http://grbio.org/cool/25c9-tsqt" name="Museo de Historia Natural de Concepcion (Chile)" pageId="9" pageNumber="91">MHNC</collectionCode>
91.05245), 2) 45 pale longitudinal stripes broken up posteriorly with flanks spotted (in
<collectionCode box="[514,555,1630,1653]" country="United Kingdom" name="Bristol Museum" pageId="9" pageNumber="91">BM</collectionCode>
1970.6.29.1011,
<collectionCode box="[141,180,1662,1685]" country="Germany" name="Teylers Museum, Paleontologische" pageId="9" pageNumber="91">TM</collectionCode>
40392,
<collectionCode box="[263,305,1662,1685]" country="Germany" name="Museum Donaueschingen" pageId="9" pageNumber="91">MD</collectionCode>
1967), and 3) broken paravertebral stripes, continuous dorsolateral line and barred flanks (in
<collectionCode box="[691,745,1694,1717]" country="South Africa" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="9" pageNumber="91" type="Museum">PEM</collectionCode>
<accessionNumber box="[141,227,1725,1749]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R24157" pageId="9" pageNumber="91">
<collectionCode box="[141,159,1725,1749]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="91">R</collectionCode>
24157
</accessionNumber>
), similar to
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1838" box="[350,479,1726,1749]" class="Reptilia" family="Lacertidae" genus="Nucras" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="91" phylum="Chordata" rank="species" species="intertexta">
<emphasis box="[350,479,1726,1749]" italics="true" pageId="9" pageNumber="91">N. intertexta</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="91" type="distribution">
<paragraph blockId="9.[141,746,1789,1942]" lastBlockId="9.[791,1395,1181,1430]" pageId="9" pageNumber="91">
<emphasis bold="true" box="[141,283,1789,1813]" pageId="9" pageNumber="91">Distribution.</emphasis>
Found only in semi-arid south-western
<collectingCountry box="[141,220,1821,1845]" name="Angola" pageId="9" pageNumber="91">Angola</collectingCountry>
, throughout much of
<collectingRegion box="[494,688,1821,1845]" country="Angola" name="Namibe" pageId="9" pageNumber="91">Namibe Province</collectingRegion>
and extending onto the escarpment of southern Hu
<emphasis bold="true" box="[668,675,1853,1877]" pageId="9" pageNumber="91">í</emphasis>
la and
<collectingRegion box="[141,221,1886,1910]" country="Angola" name="Cunene" pageId="9" pageNumber="91">Cunene</collectingRegion>
Provinces (
<figureCitation box="[361,430,1885,1910]" captionStart="Fig" captionStartId="2.[141,179,1119,1141]" captionTargetBox="[141,1394,141,1105]" captionTargetId="figure@2.[141,1394,141,1105]" captionTargetPageId="2" captionText="Fig.1. Map showing distribution of Nucras broadleyi sp. nov. in Angola." figureDoi="http://doi.org/10.5281/zenodo.3731979" httpUri="https://zenodo.org/record/3731979/files/figure.png" pageId="9" pageNumber="91">Fig. 1</figureCitation>
). Known localities include: Maconjo (
<bibRefCitation author="Bocage JVB" bookContentInfo="203 p." box="[248,430,1917,1942]" journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="9" pageNumber="91" refId="ref8226" refString="Bocage JVB. 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal. 203 p., 20 pls." title="Herpetologie d'Angola et du Congo" type="book" year="1895">Bocage 1895: 30</bibRefCitation>
), Ponang Kuma (=Donguena) (
<bibRefCitation author="Boulenger GA" box="[798,1024,1181,1206]" journalOrPublisher="Annals of the South African Museum" pageId="9" pageNumber="91" pagination="455 - 538" part="5" refId="ref8255" refString="Boulenger GA. 1910. A revised list of South African reptiles and batrachians, with synoptic tables, special reference of specimens in the South African Museum, and descriptions of new species. Annals of the South African Museum 5: 455 - 538." title="A revised list of South African reptiles and batrachians, with synoptic tables, special reference of specimens in the South African Museum, and descriptions of new species" type="journal article" year="1910">Boulenger 1910: 472</bibRefCitation>
),
<quantity box="[1044,1111,1181,1206]" metricMagnitude="4" metricUnit="m" metricValue="3.4" pageId="9" pageNumber="91" unit="km" value="34.0">34 km</quantity>
from
<collectingRegion box="[1178,1261,1181,1205]" country="Angola" name="Namibe" pageId="9" pageNumber="91">Namibe</collectingRegion>
on Lubango road (
<bibRefCitation author="Laurent RF" box="[854,1045,1213,1238]" journalOrPublisher="Publicacoes Culturais Com- panhia de Diamantes de Angola" pageId="9" pageNumber="91" pagination="1 - 165" part="67" refId="ref9224" refString="Laurent RF. 1964. Reptiles et amphibiens de l'Angola (troiseme contribution). Publicacoes Culturais Com- panhia de Diamantes de Angola 67: 1 - 165." title="Reptiles et amphibiens de l'Angola (troiseme contribution)" type="journal article" year="1964">Laurent 1964: 56</bibRefCitation>
),
<quantity box="[1067,1137,1213,1238]" metricMagnitude="4" metricUnit="m" metricValue="3.4" pageId="9" pageNumber="91" unit="km" value="34.0">34 km</quantity>
south of Tombwa (
<collectionCode box="[1354,1395,1214,1238]" country="Germany" name="Teylers Museum, Paleontologische" pageId="9" pageNumber="91">TM</collectionCode>
40397),
<quantity box="[878,951,1245,1270]" metricMagnitude="3" metricUnit="m" metricValue="8.8" pageId="9" pageNumber="91" unit="km" value="8.8">8.8 km</quantity>
southwest of Farm Mucungo (this study),
<quantity box="[791,857,1277,1302]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="91" unit="km" value="10.0">10 km</quantity>
west of Lola (this study), and Capelongo (
<bibRefCitation author="Monard A." journalOrPublisher="Arquivos do Museu Bocage" pageId="9" pageNumber="91" pagination="19 - 154" part="8" refId="ref9569" refString="Monard A. 1937. Contribution a l'herpetologie d'Angola. Arquivos do Museu Bocage 8: 19 - 154." title="Contribution a l'herpetologie d'Angola" type="journal article" year="1937">Monard 1937: 73</bibRefCitation>
). The locality of Caconda (
<bibRefCitation author="Bocage JVB" bookContentInfo="203 p." box="[1163,1302,1309,1334]" journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="9" pageNumber="91" refId="ref8226" refString="Bocage JVB. 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal. 203 p., 20 pls." title="Herpetologie d'Angola et du Congo" type="book" year="1895">Bocage 1895</bibRefCitation>
) extends the species distribution further north into
<collectingRegion box="[1230,1391,1341,1365]" country="Angola" name="Huila" pageId="9" pageNumber="91">Huíla Province</collectingRegion>
, but the specimens could not be critically evaluated by
<bibRefCitation author="Broadley DG" box="[791,961,1405,1430]" journalOrPublisher="Arnoldia" pageId="9" pageNumber="91" pagination="1 - 35" part="20" refId="ref8654" refString="Broadley DG. 1972. A review of the Nucras tessellata group (Sauria: Lacertidae). Arnoldia 20: 1 - 35." title="A review of the Nucras tessellata group (Sauria: Lacertidae)" type="journal article" year="1972">Broadley (1972)</bibRefCitation>
and are now presumably lost.
</paragraph>
<paragraph blockId="9.[791,1395,1469,1878]" pageId="9" pageNumber="91">
<emphasis bold="true" box="[791,882,1469,1493]" pageId="9" pageNumber="91">Habitat.</emphasis>
The species appears to be associated with mopane woodlands, dry savannas, and semi-desert shrublands (
<bibRefCitation author="Barbosa LAG" bookContentInfo="323 p." box="[919,1070,1533,1558]" journalOrPublisher="Instituto de Investigacao Cientifica de Angola, Luanda, Angola" pageId="9" pageNumber="91" refId="ref8156" refString="Barbosa LAG. 1970. Carta Fitogeografica de Angola. Instituto de Investigacao Cientifica de Angola, Luanda, Angola. 323 p." title="Carta Fitogeografica de Angola" type="book" year="1970">Barbosa 1970</bibRefCitation>
). The new material was found in sandy plains with scattered low granite outcrops, with varying degrees of short grass cover and scattered bushes. Vegetation included
<taxonomicName box="[1005,1281,1629,1653]" class="Magnoliopsida" family="Fabaceae" genus="Colophospermum" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="91" phylum="Tracheophyta" rank="species" species="mopane">
<emphasis box="[1005,1281,1629,1653]" italics="true" pageId="9" pageNumber="91">Colophospermum mopane</emphasis>
</taxonomicName>
,
<taxonomicName box="[1294,1352,1630,1653]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="91" phylum="Mollusca" rank="genus">
<emphasis box="[1294,1352,1630,1653]" italics="true" pageId="9" pageNumber="91">Ficus</emphasis>
</taxonomicName>
sp.,
<taxonomicName authorityName="Rafinesque" authorityYear="1838" box="[791,893,1661,1685]" class="Magnoliopsida" family="Fabaceae" genus="Senegalia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="91" phylum="Tracheophyta" rank="genus">
<emphasis box="[791,893,1661,1685]" italics="true" pageId="9" pageNumber="91">Senegalia</emphasis>
</taxonomicName>
(
<emphasis box="[912,1002,1662,1685]" italics="true" pageId="9" pageNumber="91">=Acacia</emphasis>
)
<emphasis box="[1020,1113,1661,1685]" italics="true" pageId="9" pageNumber="91">mellifera</emphasis>
,
<taxonomicName authorityName="N.J.Jacquin" authorityYear="1797" box="[1130,1268,1661,1685]" class="Magnoliopsida" family="Burseraceae" genus="Commiphora" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="91" phylum="Tracheophyta" rank="genus">
<emphasis box="[1130,1268,1661,1685]" italics="true" pageId="9" pageNumber="91">Commiphora</emphasis>
</taxonomicName>
sp.,
<taxonomicName authorityName="Schinz" authorityYear="1888" class="Magnoliopsida" family="Capparaceae" genus="Boscia" kingdom="Animalia" order="Capparales" pageId="9" pageNumber="91" phylum="Tracheophyta" rank="species" species="foetida">
<emphasis italics="true" pageId="9" pageNumber="91">Boscia foetida</emphasis>
</taxonomicName>
, and
<taxonomicName box="[925,1118,1693,1717]" class="Reptilia" family="Colubridae" genus="Salvadora" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="91" phylum="Chordata" rank="species" species="persica">
<emphasis box="[925,1118,1693,1717]" italics="true" pageId="9" pageNumber="91">Salvadora persica</emphasis>
</taxonomicName>
. The confirmed historical records were also obtained within the dry woodland zone, even though the possible occurrence of the species in Caconda would place the species above
<quantity box="[1223,1310,1789,1813]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="91" unit="m" value="1500.0">1,500 m</quantity>
asl and well into the mesic conditions of
<taxonomicName box="[1166,1303,1821,1845]" class="Magnoliopsida" family="Fabaceae" genus="Brachystegia" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="91" phylum="Tracheophyta" rank="genus">
<emphasis box="[1166,1303,1821,1845]" italics="true" pageId="9" pageNumber="91">Brachystegia</emphasis>
</taxonomicName>
habitats (
<bibRefCitation author="Barbosa LAG" bookContentInfo="323 p." box="[798,947,1853,1878]" journalOrPublisher="Instituto de Investigacao Cientifica de Angola, Luanda, Angola" pageId="9" pageNumber="91" refId="ref8156" refString="Barbosa LAG. 1970. Carta Fitogeografica de Angola. Instituto de Investigacao Cientifica de Angola, Luanda, Angola. 323 p." title="Carta Fitogeografica de Angola" type="book" year="1970">Barbosa 1970</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>