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<document id="4D5C05FA1F269369A964C0E29827CF8D" ID-CLB-Dataset="24222" ID-DOI="10.11646/zootaxa.4729.2.2" ID-GBIF-Dataset="b44745ed-b90a-4b62-b636-23c48c56471a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3632249" ID-ZooBank="EC42F789-C869-4551-998E-CC819044C775" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580456035197" checkinUser="plazi" docAuthor="Coimbra, João Carlos, Bergue, Cristianini Trescastro &amp; Ramos, Maria Inês Feijó" docDate="2020" docId="8C0B878F3926577E49CCF959FCC5FE09" docLanguage="en" docName="zootaxa.4729.2.2.pdf" docOrigin="Zootaxa 4729 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copytus novaezealandiae" docType="treatment" docVersion="8" lastPageNumber="183" masterDocId="7032FFF739235778495BFF89FFE0FFA6" masterDocTitle="Is Copytus Skogsberg, 1939 (Crustacea: Ostracoda) a neocytherideid? With description of a new family and two new species" masterLastPageNumber="194" masterPageNumber="177" pageNumber="182" updateTime="1698779709062" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="19F81CD62A8576F0FC1E8121A7B37A82">Is Copytus Skogsberg, 1939 (Crustacea: Ostracoda) a neocytherideid? With description of a new family and two new species</mods:title>
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<mods:namePart id="1140517EDC45E0BD5D5504C6AC0B3843">Coimbra, João Carlos</mods:namePart>
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<mods:namePart id="98FFCE105594A77B1E70994524D47F81">Bergue, Cristianini Trescastro</mods:namePart>
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<mods:namePart id="5B704759E794556172DEFA5F14C6FC53">Ramos, Maria Inês Feijó</mods:namePart>
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<treatment id="8C0B878F3926577E49CCF959FCC5FE09" ID-GBIF-Taxon="161926163" LSID="urn:lsid:plazi:treatment:8C0B878F3926577E49CCF959FCC5FE09" httpUri="http://treatment.plazi.org/id/8C0B878F3926577E49CCF959FCC5FE09" lastPageId="6" lastPageNumber="183" pageId="5" pageNumber="182">
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<paragraph id="041D36993926577D49CCF959FB54F94C" blockId="5.[151,1437,1743,2022]" box="[151,1204,1743,1770]" pageId="5" pageNumber="182">
<heading id="5F5581F53926577D49CCF959FB54F94C" bold="true" box="[151,1204,1743,1770]" fontSize="11" level="1" pageId="5" pageNumber="182" reason="1">
<emphasis id="36D6EA8B3926577D49CCF959FB54F94C" bold="true" box="[151,1204,1743,1770]" pageId="5" pageNumber="182">
<taxonomicName id="C3A24D1A3926577D49CCF959FD84F94C" authority="(Brady, 1898)" baseAuthorityName="Brady" baseAuthorityYear="1898" box="[151,612,1743,1770]" class="Ostracoda" family="Neocytherideidae" genus="Copytus" kingdom="Animalia" order="Podocopida" pageId="5" pageNumber="182" phylum="Arthropoda" rank="species" species="novaezealandiae">
<emphasis id="36D6EA8B3926577D49CCF959FE5BF94C" bold="true" box="[151,443,1743,1770]" italics="true" pageId="5" pageNumber="182">Copytus novaezealandiae</emphasis>
(
<bibRefCitation id="60334B683926577D4891F946FDBBF94C" author="Brady, G. S." box="[458,603,1743,1770]" pageId="5" pageNumber="182" pagination="429 - 452" refId="ref12326" refString="Brady, G. S. (1898) On new or imperfectly known species of Ostracoda, chiefly from New Zealand. Transactions of the Zoological Society of London, 14, 429 - 452." type="journal article" year="1898">Brady, 1898</bibRefCitation>
)
</taxonomicName>
or
<taxonomicName id="C3A24D1A3926577D4BD7F946FB48F94C" authority="(Brady, 1898)" baseAuthorityName="Brady" baseAuthorityYear="1898" box="[652,1192,1743,1770]" class="Ostracoda" family="Neocytherideidae" genus="Neocytherideis" kingdom="Animalia" order="Podocopida" pageId="5" pageNumber="182" phylum="Arthropoda" rank="species" species="novaezealandiae">
<emphasis id="36D6EA8B3926577D4BD7F946FC1DF94C" bold="true" box="[652,1021,1743,1770]" italics="true" pageId="5" pageNumber="182">Neocytherideis novaezealandiae</emphasis>
(
<bibRefCitation id="60334B683926577D4D57F946FB7FF94C" author="Brady, G. S." box="[1036,1183,1743,1770]" pageId="5" pageNumber="182" pagination="429 - 452" refId="ref12326" refString="Brady, G. S. (1898) On new or imperfectly known species of Ostracoda, chiefly from New Zealand. Transactions of the Zoological Society of London, 14, 429 - 452." type="journal article" year="1898">Brady, 1898</bibRefCitation>
)
</taxonomicName>
?
</emphasis>
</heading>
</paragraph>
<paragraph id="041D36993926577D49CCF97AFBCEF840" blockId="5.[151,1437,1743,2022]" pageId="5" pageNumber="182">
This species has been synonymised by
<bibRefCitation id="60334B683926577D4B00F97DFCEDF8AB" author="Swanson, K. M." box="[603,781,1779,1806]" pageId="5" pageNumber="182" pagination="33 - 48" refId="ref15120" refString="Swanson, K. M. (1969) Some Lower Miocene Ostracoda from the Middle Waipara District, New Zealand. Transactions of the Royal Society of New Zealand, Earth Sciences, 7, 33 - 48." type="journal article" year="1969">Swanson (1969</bibRefCitation>
,
<bibRefCitation id="60334B683926577D4A41F97AFC82F8A8" author="Swanson, K. M." box="[794,866,1779,1806]" pageId="5" pageNumber="182" pagination="1 - 56" refId="ref15160" refString="Swanson, K. M. (1979 a) The marine fauna of New Zealand: ostracods of the Otago Shelf. New Zealand Oceanographic Institute Memoir, 78, 1 - 56." type="journal article" year="1979">1979a</bibRefCitation>
,b) and
<bibRefCitation id="60334B683926577D4AE3F97DFBADF8A8" author="Eagar, S. H." box="[952,1101,1780,1806]" pageId="5" pageNumber="182" pagination="53 - 64" refId="ref12934" refString="Eagar, S. H. (1971) A checklist of the Ostracoda of New Zealand. Journal of the Royal Society of New Zealand, 1, 53 - 64. https: // doi. org / 10.1080 / 03036758.1971.10419355" type="journal article" year="1971">Eagar (1971)</bibRefCitation>
without reason or discussion with
<taxonomicName id="C3A24D1A3926577D498AF891FDCDF897" authority="McKenzie, 1967" authorityName="McKenzie" authorityYear="1967" box="[209,557,1815,1842]" class="Ostracoda" family="Neocytherideidae" genus="Copytus" kingdom="Animalia" order="Podocopida" pageId="5" pageNumber="182" phylum="Arthropoda" rank="species" species="rara">
<emphasis id="36D6EA8B3926577D498AF891FE87F897" box="[209,359,1816,1842]" italics="true" pageId="5" pageNumber="182">Copytus rara</emphasis>
<bibRefCitation id="60334B683926577D4835F89EFDCDF897" author="McKenzie, K. G." box="[366,557,1815,1841]" pageId="5" pageNumber="182" pagination="61 - 106" refId="ref14559" refString="McKenzie, K. G. (1967) Recent Ostracoda from Port Phillip Bay, Victoria. Proceedings of the Royal Society of Victoria, 80, 61 - 106." type="journal article" year="1967">McKenzie, 1967</bibRefCitation>
</taxonomicName>
. The first species (l =
<quantity id="C35A9B7C3926577D4A70F891FC74F894" box="[811,916,1816,1842]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="5" pageNumber="182" unit="mm" value="0.88">0.88 mm</quantity>
), originally
<taxonomicName id="C3A24D1A3926577D4D47F891FA82F894" authorityName="Brady" authorityYear="1898" box="[1052,1378,1816,1842]" class="Ostracoda" family="Cushmanideidae" genus="Cytherideis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="5" pageNumber="182" phylum="Arthropoda" rank="species" species="novaezealandiae">
<emphasis id="36D6EA8B3926577D4D47F891FA82F894" box="[1052,1378,1816,1842]" italics="true" pageId="5" pageNumber="182">Cytherideis novaezealandiae</emphasis>
</taxonomicName>
, was recovered from Lyttleton Harbour (
<collectingCountry id="7CB576093926577D4B75F8B5FD2BF8F0" box="[558,715,1851,1878]" name="New Zealand" pageId="5" pageNumber="182">New Zealand</collectingCountry>
) and Bradys illustrations (figs. 1, 2) show that it is relatively well-rounded anteriorly and very well-rounded posteriorly. Thus, it does not seem to be conspecific with
<taxonomicName id="C3A24D1A3926577D4C19F8E9FA75F8DF" authorityName="McKenzie" authorityYear="1967" box="[1346,1429,1888,1914]" class="Ostracoda" family="Neocytherideidae" genus="Copytus" kingdom="Animalia" order="Podocopida" pageId="5" pageNumber="182" phylum="Arthropoda" rank="species" species="rara">
<emphasis id="36D6EA8B3926577D4C19F8E9FA75F8DF" box="[1346,1429,1888,1914]" italics="true" pageId="5" pageNumber="182">C. rara</emphasis>
</taxonomicName>
, a species that is pointed anteroventrally and larger (l =
<quantity id="C35A9B7C3926577D4A5DF80AFC8FF838" box="[774,879,1923,1950]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="182" unit="mm" value="0.95">0.95 mm</quantity>
). John Neil (lit. comm. to J.C.C. 2009), however, comments: “My own specimens of
<taxonomicName id="C3A24D1A3926577D4B76F821FCEBF864" authorityName="Brady" authorityYear="1898" box="[557,779,1960,1986]" class="Ostracoda" family="Cushmanideidae" genus="Cytherideis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="5" pageNumber="182" phylum="Arthropoda" rank="species" species="novaezealandiae">
<emphasis id="36D6EA8B3926577D4B76F821FCEBF864" box="[557,779,1960,1986]" italics="true" pageId="5" pageNumber="182">C. novaezealandiae</emphasis>
</taxonomicName>
(
<taxonomicName id="C3A24D1A3926577D4A45F821FC92F867" authorityName="McKenzie" authorityYear="1967" box="[798,882,1960,1986]" class="Ostracoda" family="Neocytherideidae" genus="Copytus" kingdom="Animalia" order="Podocopida" pageId="5" pageNumber="182" phylum="Arthropoda" rank="species" species="rara">
<emphasis id="36D6EA8B3926577D4A45F821FC92F867" box="[798,882,1960,1986]" italics="true" pageId="5" pageNumber="182">C. rara</emphasis>
</taxonomicName>
) are definitely adont, and have a
<taxonomicName id="C3A24D1A3926577D4DAFF821FA7CF864" authorityName="Puri" authorityYear="1952" box="[1268,1436,1960,1986]" class="Ostracoda" family="Neocytherideidae" genus="Neocytherideis" kingdom="Animalia" order="Podocopida" pageId="5" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="36D6EA8B3926577D4DAFF821FA7CF864" box="[1268,1436,1960,1986]" italics="true" pageId="5" pageNumber="182">Neocytherideis</emphasis>
</taxonomicName>
muscle scar pattern of four adductors in a vertical row with a curved frontal scar”.
</paragraph>
</subSubSection>
<subSubSection id="4CB865123925577E499CFF10FCC5FE09" pageId="6" pageNumber="183" type="description">
<paragraph id="041D36993925577E499CFF10FDF3FEE5" blockId="6.[151,1436,153,431]" pageId="6" pageNumber="183">
<bibRefCitation id="60334B683925577E499CFF10FDE0FF15" author="Morley, M. S. &amp; Hayward, B. W." box="[199,512,153,179]" pageId="6" pageNumber="183" pagination="51 - 75" refId="ref14762" refString="Morley, M. S. &amp; Hayward, B. W. (2012) Ostracoda of the Hauraki Gulf, New Zealand. Records of the Auckland Museum, 48, 51 - 75." type="journal article" year="2012">Morley &amp; Hayward (2012)</bibRefCitation>
, in a study of ostracods from the Hauraki Gulf (
<collectingCountry id="7CB576093925577E4D68FF10FB33FF12" box="[1075,1235,153,180]" name="New Zealand" pageId="6" pageNumber="183">New Zealand</collectingCountry>
), recorded
<taxonomicName id="C3A24D1A3925577E4C0FFF10FED7FF71" authorityName="Brady" authorityYear="1898" class="Ostracoda" family="Cushmanideidae" genus="Cytherideis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="6" pageNumber="183" phylum="Arthropoda" rank="species" species="novaezealandiae">
<emphasis id="36D6EA8B3925577E4C0FFF10FED7FF71" italics="true" pageId="6" pageNumber="183">C. novaezealandiae</emphasis>
</taxonomicName>
and
<taxonomicName id="C3A24D1A3925577E482BFF34FD9BFF71" authorityName="Brady" authorityYear="1898" box="[368,635,189,215]" class="Ostracoda" family="Cushmanideidae" genus="Cytherideis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="6" pageNumber="183" phylum="Arthropoda" rank="species" species="novaezealandiae">
<emphasis id="36D6EA8B3925577E482BFF34FE6AFF71" box="[368,394,189,215]" italics="true" pageId="6" pageNumber="183">C.</emphasis>
aff.
<emphasis id="36D6EA8B3925577E48E4FF34FD9BFF71" box="[447,635,189,215]" italics="true" pageId="6" pageNumber="183">novaezealandiae</emphasis>
</taxonomicName>
. However, in their drawings, it can be seen that fig. 8c does not represent a specimen of
<taxonomicName id="C3A24D1A3925577E482FFF68FDB3FF5D" authorityName="Brady" authorityYear="1898" box="[372,595,225,251]" class="Ostracoda" family="Cushmanideidae" genus="Cytherideis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="6" pageNumber="183" phylum="Arthropoda" rank="species" species="novaezealandiae">
<emphasis id="36D6EA8B3925577E482FFF68FDB3FF5D" box="[372,595,225,251]" italics="true" pageId="6" pageNumber="183">C. novaezealandiae</emphasis>
</taxonomicName>
, while fig. 8d shows another different and very ornate species.
<bibRefCitation id="60334B683925577E4C70FF68FEA8FEB9" author="Morley, M. S. &amp; Hayward, B. W." pageId="6" pageNumber="183" pagination="24 - 33" refId="ref14684" refString="Morley, M. S. &amp; Hayward, B. W. (2006) Molluscs and ostracods in a tidal transect from Whangapoua Estuary, Coromandel Peninsula. Poirieria, 32, 24 - 33." type="journal article" year="2006">Morley &amp; Hayward (2006</bibRefCitation>
,
<bibRefCitation id="60334B683925577E4808FE8CFE6DFEB9" author="Morley, M. S. &amp; Hayward, B. W." box="[339,397,261,287]" pageId="6" pageNumber="183" pagination="17 - 32" refId="ref14722" refString="Morley, M. S. &amp; Hayward, B. W. (2007) Intertidal and shallow-water Ostracoda of the Waitemata Harbour, New Zealand. Records of the Auckland Museum, 44, 17 - 32." type="journal article" year="2007">2007</bibRefCitation>
) cited this species in a tidal transect from Whangapoua Estuary and Waitemata Harbour,
<collectingCountry id="7CB576093925577E4C3CFE8CFF14FEE5" name="New Zealand" pageId="6" pageNumber="183">New Zealand</collectingCountry>
, but without illustrations.
</paragraph>
<paragraph id="041D36993925577E499CFEC4FCC5FE09" blockId="6.[151,1436,153,431]" pageId="6" pageNumber="183">
Regarding its stratigraphic distribution,
<bibRefCitation id="60334B683925577E4BDDFEC4FCDCFECE" author="Swanson, K. M." box="[646,828,333,360]" pageId="6" pageNumber="183" pagination="33 - 48" refId="ref15120" refString="Swanson, K. M. (1969) Some Lower Miocene Ostracoda from the Middle Waipara District, New Zealand. Transactions of the Royal Society of New Zealand, Earth Sciences, 7, 33 - 48." type="journal article" year="1969">Swanson (1969)</bibRefCitation>
found many specimens in the Middle Waipara section (Lower Miocene) in
<collectingCountry id="7CB576093925577E48DFFEF8FDFFFE2D" box="[388,543,369,395]" name="New Zealand" pageId="6" pageNumber="183">New Zealand</collectingCountry>
. Previously,
<bibRefCitation id="60334B683925577E4BEBFEF8FC66FE2D" author="Hornibrook, N. B." box="[688,902,368,395]" pageId="6" pageNumber="183" pagination="1 - 82" refId="ref13924" refString="Hornibrook, N. B. (1952) Tertiary and Recent marine Ostracoda of New Zealand: their origin, affinities, and distribution. New Zealand Geological Survey, Palaeontological Bulletin, 18, 1 - 82." type="journal article" year="1952">Hornibrook (1952)</bibRefCitation>
stated that this species occurs in
<collectingCountry id="7CB576093925577E4C5EFEF8FA7CFE2D" box="[1285,1436,369,395]" name="New Zealand" pageId="6" pageNumber="183">New Zealand</collectingCountry>
since the Eocene; however, the species was not illustrated.
</paragraph>
</subSubSection>
</treatment>
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