283 lines
23 KiB
XML
283 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.46.32823" ID-GBIF-Dataset="e546c2a6-4668-4253-b2e2-b462f23dab8c" ID-PMC="PMC6379322" ID-Pensoft-Pub="1314-4049-46-1" ID-PubMed="30787668" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-4049-46-1" ModsDocOrigin="MycoKeys 46" ModsDocTitle="The genus Hebeloma in the Rocky Mountain Alpine Zone" checkinTime="1555333436509" checkinUser="pensoft" docAuthor="Cripps, Cathy L., Eberhardt, Ursula, Schuetz, Nicole, Beker, Henry J., Vera S. Evenson, & Horak, Egon" docDate="2019" docId="3FA64197F2E65BA4BCD9CE8ED39F2192" docLanguage="en" docName="MycoKeys 46: 1-54" docOrigin="MycoKeys 46" docSource="http://dx.doi.org/10.3897/mycokeys.46.32823" docTitle="Hebeloma marginatulum Bruchet, Bull. Mens. Soc. Linn. Lyon 39 (6, suppl.): 43 1970" docType="treatment" docVersion="4" lastPageNumber="28" masterDocId="AC18AA047D78FFBFD06BFFFDFF948577" masterDocTitle="The genus Hebeloma in the Rocky Mountain Alpine Zone" masterLastPageNumber="54" masterPageNumber="1" pageNumber="25" updateTime="1668136151891" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Hebeloma in the Rocky Mountain Alpine Zone</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cripps, Cathy L.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Eberhardt, Ursula</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schuetz, Nicole</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Beker, Henry J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vera S. Evenson,</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Horak, Egon</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>MycoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>46</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>54</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.46.32823</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.46.32823</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-4049-46-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="156202109" LSID="urn:lsid:plazi:treatment:3FA64197F2E65BA4BCD9CE8ED39F2192" httpUri="http://treatment.plazi.org/id/3FA64197F2E65BA4BCD9CE8ED39F2192" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25">
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<subSubSection pageId="24" pageNumber="25" type="nomenclature">
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<paragraph pageId="24" pageNumber="25">
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8.
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<taxonomicName authority="J. Favre) Bruchet, Bull. Mens. Soc. Linn. Lyon 39 (6, suppl.): 43 (1970" authorityYear="1970" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma marginatulum" order="Agaricales" pageId="24" pageNumber="25" phylum="Basidiomycota" rank="species" species="marginatulum">Hebeloma marginatulum (J. Favre) Bruchet, Bull. Mens. Soc. Linn. Lyon 39 (6, suppl.): 43 (1970)</taxonomicName>
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Figures 6B, 14, 23 (8)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="26" type="etymology">
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<paragraph pageId="25" pageNumber="26">
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<pageBreakToken pageId="25" pageNumber="26" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="25" pageNumber="26">From marginatus, with a margin or border, emphasizing a thin line of tissue near the margin.</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="26" type="description">
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<paragraph pageId="25" pageNumber="26">Description.</paragraph>
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<paragraph pageId="25" pageNumber="26">
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Cortina present, remnants distinctly present in some. Pileus 15
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<normalizedToken originalValue="–40(–">-40(-</normalizedToken>
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50) mm in diameter, slightly conic-convex, domed convex, irregular, sometimes with a flat center that can even be dished, smooth or rough due to velipellus, shiny, strongly canescent, underneath dark brown, dark chestnut, to dark caramel color, mostly uniform but two-toned in some and then lighter at margin (more hoary, dingy whitish, or ochraceous in one), with a fine white border around the pileus perimeter a few mm in from margin, not hygrophanous; margin turned down or in, rather persistently so, and then covered with copious veil, often irregular, wavy, fragile. In one collection, the cuticle is rather thick and rubbery. Lamellae deeply emarginate and squared off, some pulling away, somewhat broad, L = 30-40 plus lamellulae, cream, then pinkish buff, darkening to medium coffee brown; edges fimbriate. Stipe 20
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<normalizedToken originalValue="–40(–">-40(-</normalizedToken>
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45) mm
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<normalizedToken originalValue="×">x</normalizedToken>
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2
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<normalizedToken originalValue="–6(–">-6(-</normalizedToken>
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10) mm, equal, undulating or not, pale buff (some with possible yellow tint), and dark (up to black) at base, pruinose at apex, longitudinally fibrous lower, with a few longitudinal fibrils. Context dingy whitish, some with yellowish tones and dark at base. Odor raphanoid or sourish, sometimes faint. Exsiccate: pileus pale brown to dark brown, some obviously canescent; lamellae medium brown; stipe buff or ocher, darker at base.
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</paragraph>
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<paragraph pageId="25" pageNumber="26">
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Basidiospores yellowish gray, pale in Melzers, elliptical with rounded end, inequilateral in side view, no big apiculus, not guttulate, smooth to slightly punctate or rough (O1, O2), indextrinoid (D0, D1), perispore not loosening (P0), 9
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<normalizedToken originalValue="–12(–">-12(-</normalizedToken>
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13)
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<normalizedToken originalValue="×">x</normalizedToken>
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5.5
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<normalizedToken originalValue="–7(–">-7(-</normalizedToken>
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8)
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, on average 10.1
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<normalizedToken originalValue="×">x</normalizedToken>
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6.4
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, Q = 1.59. Basidia 25-35
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<normalizedToken originalValue="×">x</normalizedToken>
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8-9
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, clavate, two and four-spored. Cheilocystidia lageniform, ventricose, often with very long equal neck, and somewhat gradually swollen base, occasionally clavate at apex, sometimes cylindrical, 35-80
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long
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<normalizedToken originalValue="×">x</normalizedToken>
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4-7
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<normalizedToken originalValue="µm">µm</normalizedToken>
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at apex, 4-6 in middle, and 7-12 (13) at base, no thickening noticed. Pleurocystidia absent. Epicutis thickness 40-100
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, with some encrusted hyphae.
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</paragraph>
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<caption pageId="25" pageNumber="26">
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<paragraph pageId="25" pageNumber="26">
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Figure 14.
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma marginatulum" order="Agaricales" pageId="25" pageNumber="26" phylum="Basidiomycota" rank="species" species="marginatulum">Hebeloma marginatulum</taxonomicName>
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DBG-F-020841 and CLC3545.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="rocky mountain ecology">
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<paragraph pageId="26" pageNumber="27">
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<pageBreakToken pageId="26" pageNumber="27" start="start">Rocky</pageBreakToken>
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Mountain ecology.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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In the Rocky Mountain alpine zone, with various willows, including dwarf willows
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix arctica" order="Malpighiales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="arctica">Salix arctica</taxonomicName>
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and
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<taxonomicName lsidName="S. reticulata" pageId="26" pageNumber="27" rank="species" species="reticulata">S. reticulata</taxonomicName>
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, and shrub willow
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<taxonomicName lsidName="S. planifolia" pageId="26" pageNumber="27" rank="species" species="planifolia">S. planifolia</taxonomicName>
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. Known from both Colorado and Montana.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="materials_examined">
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<paragraph pageId="26" pageNumber="27">Rocky Mountain specimens examined.</paragraph>
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<paragraph pageId="26" pageNumber="27">
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U.S.A. COLORADO: Front Range, Loveland Pass, 12 Aug 2013, in
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<taxonomicName class="Insecta" family="Nymphalidae" genus="Dryas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dryas" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">Dryas</taxonomicName>
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, DBG-F-027694, C. Cripps; 12 Aug 2013, with
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., DBG-F-027695, C. Cripps; 25 Aug 2000, with
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., DBG-F-020708, V.S. Evenson; 21 Aug 2003, with
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., DBG-F-021388, V.S. Evenson; 20 Aug 2013, DBG-F-027682, L. Gillman; 21 Aug 2003, with
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., DBG-F-021405, O.K. Miller, Jr; San Juan County, Cinnamon Pass, 3700 m, with
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix arctica" order="Malpighiales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="arctica">Salix arctica</taxonomicName>
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, 29 July 2000, CLC 1413 (MONT), C. Cripps, 3700 m, with
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix arctica" order="Malpighiales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="arctica">Salix arctica</taxonomicName>
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, 27 July 2002, CLC1811 (MONT), C. Cripps; 29 July 2000, with
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<taxonomicName lsidName="S. reticulata" pageId="26" pageNumber="27" rank="species" species="reticulata">S. reticulata</taxonomicName>
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and
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., ZT9002 (ETH), E. Horak; Black Bear Basin, 2 Aug 2000, 3830 m, with
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<taxonomicName lsidName="S. planifolia" pageId="26" pageNumber="27" rank="species" species="planifolia">S. planifolia</taxonomicName>
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, CLC1448 (MONT), C. Cripps; 8 Aug 2000, with
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<taxonomicName lsidName="S. arctica" pageId="26" pageNumber="27" rank="species" species="arctica">S. arctica</taxonomicName>
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, CLC1449 (MONT), C. Cripps; 11 Aug 2001, with
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<taxonomicName lsidName="S. reticulata" pageId="26" pageNumber="27" rank="species" species="reticulata">S. reticulata</taxonomicName>
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, ZT9813 (ETH), E. Horak; 3760 m, with
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix arctica" order="Malpighiales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="arctica">Salix arctica</taxonomicName>
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, 11 Aug 2001, CLC1718 (MONT), C. Cripps; Emma Lake/Horseshoe Basin, 3688 m, with
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<taxonomicName lsidName="S. arctica" pageId="26" pageNumber="27" rank="species" species="arctica">S. arctica</taxonomicName>
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, 31 July 2002, CLC1874 (MONT), C. Cripps; 31 July 2002, with
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<taxonomicName lsidName="S. arctica" pageId="26" pageNumber="27" rank="species" species="arctica">S. arctica</taxonomicName>
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, CLC1880 (MONT), C. Cripps; Imogene Pass, 29 July 2002, 3850 m, with
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<taxonomicName lsidName="S. arctica" pageId="26" pageNumber="27" rank="species" species="arctica">S. arctica</taxonomicName>
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, CLC1836 (MONT), C. Cripps; Mineral Basin, 3850 m, with
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<taxonomicName lsidName="S. arctica" pageId="26" pageNumber="27" rank="species" species="arctica">S. arctica</taxonomicName>
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, 29 July 2002, with
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<taxonomicName lsidName="S. arctica" pageId="26" pageNumber="27" rank="species" species="arctica">S. arctica</taxonomicName>
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, CLC1840 (MONT), C. Cripps; without obvious host, although
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Salix</taxonomicName>
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in the vicinity, 30 July 2002, CLC1860 (MONT), C. Cripps; with
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<taxonomicName lsidName="S. arctica" pageId="26" pageNumber="27" rank="species" species="arctica">S. arctica</taxonomicName>
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and
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<taxonomicName lsidName="S. planifolia" pageId="26" pageNumber="27" rank="species" species="planifolia">S. planifolia</taxonomicName>
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, 30 July 2002, CLC1861 (MONT), C. Cripps; 3835 m, with
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<taxonomicName lsidName="S. arctica" pageId="26" pageNumber="27" rank="species" species="arctica">S. arctica</taxonomicName>
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, 7 Aug 2001, CLC1667 (MONT), C. Cripps; Stony Pass, 3840 m, with
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<taxonomicName lsidName="S. arctica" pageId="26" pageNumber="27" rank="species" species="arctica">S. arctica</taxonomicName>
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, 28 July 2002, CLC1824 (MONT), C. Cripps; 3840 m, with
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<taxonomicName lsidName="S. arctica" pageId="26" pageNumber="27" rank="species" species="arctica">S. arctica</taxonomicName>
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, 28 July 2002, CLC1826 (MONT), C. Cripps. Sawatch Range, Independence, 3 Aug 2000, with
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., DBG-F-020841, DBG-F-020856, V.S. Evenson; 3 Aug 2000 with
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">Salix</taxonomicName>
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sp., DBG-F-020843, V.S. Evenson; 3760 m, with
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<taxonomicName lsidName="S. planifolia" pageId="26" pageNumber="27" rank="species" species="planifolia">S. planifolia</taxonomicName>
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, 7 Aug 2000, CLC1478 (MONT), C. Cripps. MONTANA: Carbon County, Beartooth Plateau, site 1, 9 Sept 2000, with
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<taxonomicName lsidName="S. planifolia" pageId="26" pageNumber="27" rank="species" species="planifolia">S. planifolia</taxonomicName>
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, CLC1545 (MONT), C. Cripps; Quad Creek, 8 Aug 2008, with
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<taxonomicName lsidName="S. planifolia" pageId="26" pageNumber="27" rank="species" species="planifolia">S. planifolia</taxonomicName>
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, HJB12458, A. and M. Ronikier; 11 Aug 2017; with
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix reticulata" order="Malpighiales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="reticulata">Salix reticulata</taxonomicName>
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and
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<taxonomicName lsidName="S. planifolia" pageId="26" pageNumber="27" rank="species" species="planifolia">S. planifolia</taxonomicName>
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, 11 Aug 2017, CLC3545 (MONT), C. Cripps.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="discussion">
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<paragraph pageId="27" pageNumber="28">
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<pageBreakToken pageId="27" pageNumber="28" start="start">Discussion</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="27" pageNumber="28">
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma marginatulum" order="Agaricales" pageId="27" pageNumber="28" phylum="Basidiomycota" rank="species" species="marginatulum">Hebeloma marginatulum</taxonomicName>
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is distinct from other species of the
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<taxonomicName lsidName="H. mesophaeum" pageId="27" pageNumber="28" rank="species" species="mesophaeum">H. mesophaeum</taxonomicName>
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complex, but not by much as to molecular distance (Fig. 6B). The species is paraphyletic in relation to the monophylum including the other taxa of the complex. With 0-19 [0-2] bp, the intraspecific variation is quite extensive in
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<taxonomicName lsidName="H. marginatulum" pageId="27" pageNumber="28" rank="species" species="marginatulum">H. marginatulum</taxonomicName>
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in terms of total differences. Within each dataset, the ITS variation is also quite large, 0-14 [0-2] bp for the RM (29 sequences) and 0-17 [0-2] bp for the FE dataset (21 sequences). However, the total number of considered sequences is also larger than for other species.
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</paragraph>
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<paragraph pageId="27" pageNumber="28">
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This taxon was first described as
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<taxonomicName infraspecific-rank="var." lsidName="H. versipelle var. marginatulum" pageId="27" pageNumber="28" rank="variety" species="versipelle" variety="marginatulum">H. versipelle var. marginatulum</taxonomicName>
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by
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<bibRefCitation author="Favre, J" journalOrPublisher="Ergebnisse der Wissenschaftlichen Untersuchungen des Schweizerischen Nationalparks" pageId="43" pageNumber="44" pagination="1 - 212" title="Les champinons superieurs de la zone alpine du Parc National Suisse." volume="5" year="1955">Favre (1955)</bibRefCitation>
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from the alpine region of the Swiss Alps and was later raised to species level by
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<bibRefCitation pageId="27" pageNumber="28">Bruchet (1970)</bibRefCitation>
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. It is now considered to be restricted to arctic and alpine habitats primarily with dwarf willows (
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<bibRefCitation pageId="27" pageNumber="28">Beker et al. 2016</bibRefCitation>
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,
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<bibRefCitation pageId="27" pageNumber="28">2018</bibRefCitation>
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). Confirmed records show it to be present in these habitats in Canada, Greenland, Iceland, Scandinavia, Svalbard as well as the European Alps and the Carpathians and Rocky Mountains (
|
||
<bibRefCitation author="Eberhardt, U" journalOrPublisher="Mycologia" pageId="43" pageNumber="44" pagination="1285 - 1303" title="The genus Hebeloma in the alpine belt of the Carpathians including two new species." url="https://doi.org/10.3852/15-097" volume="107" year="2015 b">Eberhardt et al. 2015b</bibRefCitation>
|
||
;
|
||
<bibRefCitation pageId="27" pageNumber="28">Beker et al. 2016</bibRefCitation>
|
||
).
|
||
<bibRefCitation author="Vesterholt, J" journalOrPublisher="Svampetryk, Tilst, Denmark" pageId="45" pageNumber="46" title="The genus Hebeloma. Fungi of Northern Europe - Vol. 3." year="2005">Vesterholt (2005)</bibRefCitation>
|
||
described
|
||
<taxonomicName lsidName="H. polare" pageId="27" pageNumber="28" rank="species" species="polare">H. polare</taxonomicName>
|
||
as a darker brown closely related species, but this has been synonymized with
|
||
<taxonomicName lsidName="H. marginatulum" pageId="27" pageNumber="28" rank="species" species="marginatulum">H. marginatulum</taxonomicName>
|
||
(
|
||
<bibRefCitation pageId="27" pageNumber="28">Beker et al. 2016</bibRefCitation>
|
||
). The Rocky Mountain specimens are also mostly uniformly dark brown with a canescent sheen.
|
||
</paragraph>
|
||
<paragraph pageId="27" pageNumber="28">
|
||
Collections from the alpine that are very hoary and dark brown have been misinterpreted as
|
||
<taxonomicName lsidName="H. bruchetii" pageId="27" pageNumber="28" rank="species" species="bruchetii">H. bruchetii</taxonomicName>
|
||
Bon (
|
||
<bibRefCitation author="Miller, Jr OK" journalOrPublisher="Harvard Papers in Botany" pageId="44" pageNumber="45" pagination="155 - 162" title="Observations on the alpine tundra species of Hebeloma in Colorado." volume="6" year="2001">Miller and Evenson 2001</bibRefCitation>
|
||
) before molecular techniques;
|
||
<taxonomicName lsidName="H. bruchetii" pageId="27" pageNumber="28" rank="species" species="bruchetii">H. bruchetii</taxonomicName>
|
||
, first described as an alpine species, has now been synonymized with
|
||
<taxonomicName lsidName="H. mesophaeum" pageId="27" pageNumber="28" rank="species" species="mesophaeum">H. mesophaeum</taxonomicName>
|
||
and should have smaller spores.
|
||
<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma marginatulum" order="Agaricales" pageId="27" pageNumber="28" phylum="Basidiomycota" rank="species" species="marginatulum">Hebeloma marginatulum</taxonomicName>
|
||
is mentioned as a subalpine species (in Idaho) by
|
||
<bibRefCitation author="Smith, AH" journalOrPublisher="University of Michigan Press, Ann Arbor" pageId="45" pageNumber="46" title="Veiled Species of Hebeloma in the Western United States." year="1983">Smith et al. (1983)</bibRefCitation>
|
||
who described two varieties (ver.
|
||
<taxonomicName genus="Velutipes" lsidName="Velutipes var. fallax" pageId="27" pageNumber="28" rank="variety" variety="fallax">fallax</taxonomicName>
|
||
, var.
|
||
<taxonomicName genus="Velutipes" lsidName="Velutipes var. proximum" pageId="27" pageNumber="28" rank="variety" variety="proximum">proximum</taxonomicName>
|
||
) from the subalpine in Colorado.
|
||
<normalizedToken originalValue="Smith’s">Smith's</normalizedToken>
|
||
spore descriptions (dextrinoid with sharp ends) for his varieties may not fit this species, but the authors recognize that these varieties of
|
||
<taxonomicName lsidName="H. marginatulum" pageId="27" pageNumber="28" rank="species" species="marginatulum">H. marginatulum</taxonomicName>
|
||
, and indeed other closely related species, need more study in North America.
|
||
</paragraph>
|
||
<paragraph pageId="27" pageNumber="28">
|
||
This species is in
|
||
<taxonomicName infraspecific-rank="sect." lsidName="H." pageId="27" pageNumber="28" rank="section" section="Hebeloma">H. sect. Hebeloma</taxonomicName>
|
||
because of basidiomes with a cortina and the ventricose cheilocystidia together with the non-dextrinoid, or barely dextrinoid, spores that are primarily elliptical; within this group, it has an arctic-alpine habitat and relatively large spores (greater than 10
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
6
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |