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<heading id="7E66C147B956FF89FF70ED76FDC4E864" bold="true" box="[151,620,1325,1351]" fontSize="11" level="1" pageId="65" pageNumber="66" reason="1">
<taxonomicName id="E2910DA8B956FF89FF70ED76FDC4E864" ID-CoL="3CMK7" authority="Ciplak, 1999" authorityName="Ciplak" authorityYear="1999" box="[151,620,1325,1351]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="17E5AA39B956FF89FF70ED76FDC4E864" bold="true" box="[151,620,1325,1351]" pageId="65" pageNumber="66">
<emphasis id="17E5AA39B956FF89FF70ED76FE60E864" bold="true" box="[151,456,1325,1351]" italics="true" pageId="65" pageNumber="66">Eupholidoptera femorata</emphasis>
Çıplak, 1999
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="6D8B25A0B956FF89FF70ED0BFC98E849" box="[151,816,1360,1386]" pageId="65" pageNumber="66" type="description">
<paragraph id="252E762BB956FF89FF70ED0BFC98E849" blockId="65.[151,816,1325,1386]" box="[151,816,1360,1386]" pageId="65" pageNumber="66">
(
<figureCitation id="BDAA6AAEB956FF89FF78ED0BFEB7E849" box="[159,287,1360,1386]" captionStart="FIGURES 26–51" captionStartId="14.[151,275,1830,1854]" captionTargetBox="[367,1217,189,1786]" captionTargetId="figure-127@14.[316,1272,179,1800]" captionTargetPageId="14" captionText="FIGURES 26–51. Male cerci of Eupholidoptera species (Scale= 2 mm). 26—E. giuliae, 27—E. latens, 28—E. mariannae, 29—E. rammei, 30—E. astyla, 31—E. annamariae, 32—E. tasheliensis, 33—E. cretica, 34—E. anatolica, 35—E. tahtalica, 36—E. forcipata, 37—E. jacquelinae, 38—E. sevketi, 39—E. excisa, 40—E. demirsoyi, 41—E. krueperi, 42—E. tauricola, 43—E. mersinensis, 44—E. kykladica, 45—E. spinigera, 46—E. prasina, 47—E. tucherti, 48—E. karabagi, 49—E. unimacula, 50—E. femorata, 51—E. icariensis (Of the figures given above: 26–32, 36 from Willemse &amp; Heller, 2001; 46B–C, 51 from Willemse, 1980, 37 from Tilmans, 2002)." pageId="65" pageNumber="66">Figures 50</figureCitation>
, 
<figureCitation id="BDAA6AAEB956FF89FECBED0BFEE4E849" box="[300,332,1360,1386]" captionStart="FIGURES 83–96" captionStartId="20.[151,272,1461,1485]" captionTargetBox="[220,1366,186,1418]" captionTargetId="figure-163@20.[188,1398,132,1493]" captionTargetPageId="20" captionText="FIGURES 83–96. Male anal tergite of Eupholidoptera species (d— dorsal, c—caudal, l— lateral) (Scale = 2 mm). 83—E. sevketi, 84—E. excisa, 85—E. demirsoyi, 86—E. krueperi, 87—E. tauricola (A, from Salman, 1983; B, from Ramme, 1951), 88—E. mersinensis (A, Mersin; B, Konya), 89—E. kykladica, 90—E. spinigera, 91—E. prasina, 92—E. tucherti, 93—E. karabagi, 94—E. unimacula, 95—E. femorata, 96—E. icariensis (from Willemse, 1980)." pageId="65" pageNumber="66">95</figureCitation>
, 
<figureCitation id="BDAA6AAEB956FF89FEBFED0BFE2EE849" box="[344,390,1360,1386]" captionStart="FIGURES 129–142" captionStartId="31.[151,269,1863,1887]" captionTargetBox="[265,1302,194,1834]" captionTargetId="figure-90@31.[224,1376,145,1894]" captionTargetPageId="31" captionText="FIGURES 129–142. Male subgenital plate (and spines under the styli) of Eupholidoptera species (L— lateral view, V— ventral view) (Scale = 2 mm). 129—E. sevketi, 130—E. excisa, 131—E. demirsoyi, 132—E. krueperi, 133—E. tauricola, 134—E. mersinensis, 135—E. kykladica, 136—E. spinigera, 137—E. prasina, 138—E. tucherti, 139—E. karabagi, 140—E. unimacula, 141—E. femorata, 142—E. icariensis (Of the figures given above:133 from Salman, 1983)." pageId="65" pageNumber="66">141</figureCitation>
, 
<figureCitation id="BDAA6AAEB956FF89FE75ED0BFE68E849" box="[402,448,1360,1386]" captionStart="FIGURES 180–188" captionStartId="44.[151,269,1899,1923]" captionTargetBox="[244,1308,186,1865]" captionTargetId="figure-74@44.[213,1384,114,1892]" captionTargetPageId="44" captionText="FIGURES 180–188. Titillators of Eupholidoptera species (d— Dorsal view, L— lateral view, V— ventral view) (Scale= 2 mm). 180—E. mersinensis, 181—E. kykladica, 182—E. spinigera, 183—E. prasina, 184—E. tucherti, 185—E. karabagi, 186—E. unimacula, 187—E. femorata, 188—E. icariensis (Of the figures given above; 182 from Tilmans, 2002; 188 from Willemse, 1980)." pageId="65" pageNumber="66">187</figureCitation>
, 
<figureCitation id="BDAA6AAEB956FF89FE2AED0BFE54E849" box="[461,508,1360,1386]" captionStart="FIGURES 218–228" captionStartId="49.[151,269,1780,1804]" captionTargetBox="[220,1294,196,1731]" captionTargetId="figure-83@49.[188,1398,124,1789]" captionTargetPageId="49" captionText="FIGURES 218–228. Female subgenital plate of Eupholidoptera species (L— lateral view, V— ventral view) (Scale= 2 mm). 218—E. krueperi, 219—E. tauricola, 220—E. mersinensis (A, Mersin; B, Konya), 221—E. kykladica, 222—E. spinigera, 223—E. prasina, 224—E. tucherti, 225—E. karabagi, 226—E. unimacula, 227—E. femorata, 228—E. icariensis (Of the figures given above; 219 from Salman, 1983; 228 from Willemse, 1980)." pageId="65" pageNumber="66">227</figureCitation>
, 
<figureCitation id="BDAA6AAEB956FF89FDE0ED0BFD9DE849" box="[519,565,1360,1386]" captionStart="FIGURE 231" captionStartId="51.[151,257,1895,1919]" captionTargetBox="[220,1358,464,1855]" captionTargetId="figure-149@51.[200,1391,423,1863]" captionTargetPageId="51" captionText="FIGURE 231. Oscillograms of stridulatory movements [E. chabrieri only; synchronous registration of left tegmen movement –(upward deflection represents opening, downward closing)] and song of Eupholidoptera species: sequence of syllables. In E. helina beginning of a sequence." pageId="65" pageNumber="66">231</figureCitation>
, 
<figureCitation id="BDAA6AAEB956FF89FDA5ED0BFDD9E849" box="[578,625,1360,1386]" captionStart="FIGURE 234" captionStartId="53.[151,260,1854,1878]" captionTargetBox="[298,1301,196,1813]" captionTargetId="figure-93@53.[226,1360,165,1831]" captionTargetPageId="53" captionText="FIGURE 234. Comparative data of the calling songs in the genus Eupholidoptera. Different symbols indicate phylogenetic relationships (filled square: E. chabrieri group, open square: E. prasina group; triangles mark E. prasina group species living in Crete). Error bars indicate 95% confidence intervals Numbers indicate species occurring syntopically (1— Turkey, Mersin, Güzeloluk; 2— Turkey, Antalya, Termessos; 3— Bulgaria, near border to Greece (see Figure 2 in Ciplak et al. 2007); 4— Greece, Crete, Ida mountain)." pageId="65" pageNumber="66">234</figureCitation>
, 
<figureCitation id="BDAA6AAEB956FF89FD9BED0BFD01E849" box="[636,681,1360,1386]" captionStart="FIGURES 239–240" captionStartId="56.[151,273,1922,1946]" captionTargetBox="[152,1429,209,1885]" captionTargetPageId="56" captionText="FIGURES 239–240. Distribution maps of the species belonging to the E. prasina group. 239— Distribution of 13 species in southern Aegean islands and southern Anatolia, 240— Distribution of 13 species in central Aegean islands and southern Anatolia." pageId="65" pageNumber="66">240</figureCitation>
, Appendix)
</paragraph>
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<paragraph id="252E762BB956FF89FF70EDC6FDCDE897" blockId="65.[151,613,1437,1460]" box="[151,613,1437,1460]" pageId="65" pageNumber="66">
<taxonomicName id="E2910DA8B956FF89FF70EDC6FDC9E897" ID-CoL="3CMK7" authority="Ciplak, 1999: 77" authorityName="Ciplak" authorityPageNumber="77" authorityYear="1999" box="[151,609,1437,1460]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="17E5AA39B956FF89FF70EDC6FE0DE897" box="[151,421,1437,1460]" italics="true" pageId="65" pageNumber="66">Eupholidoptera femorata</emphasis>
Çıplak, 1999: 77
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.
</paragraph>
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<paragraph id="252E762BB956FF89FF70EDB8FCBAEB07" blockId="65.[151,1436,1507,2036]" pageId="65" pageNumber="66">
<emphasis id="17E5AA39B956FF89FF70EDB8FED6E8DE" bold="true" box="[151,382,1507,1533]" pageId="65" pageNumber="66">Type information:</emphasis>
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M, 
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: 
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, 
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, 
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, 
<quantity id="E269DBCEB956FF89FBC4EDBFFB2AE8DD" box="[1059,1154,1508,1534]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="65" pageNumber="66" unit="m" value="1600.0">
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, 
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(
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) (
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; 
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<typeStatus id="FA2AC889B956FF89FF19EE51FECCEB07" box="[254,356,1546,1572]" pageId="65" pageNumber="66" type="paratype">Paratype</typeStatus>
1F, same data as holotype (
<collectionCode id="4380EEEEB956FF89FD4FEE51FCADEB07" box="[680,773,1546,1572]" pageId="65" pageNumber="66">AUZM</collectionCode>
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.
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<emphasis id="17E5AA39B956FF89FF22EE6BFE1CEB69" bold="true" box="[197,436,1584,1610]" pageId="65" pageNumber="66">Material examined</emphasis>
: 
<materialsCitation id="95F97C76B956FF89FE22EE6AFD00EB68" box="[453,680,1585,1611]" country="Turkey" location="Mersin" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Mersin">
<collectingCountry id="5D8636BBB956FF89FE22EE6AFDE9EB68" box="[453,577,1585,1611]" name="Turkey" pageId="65" pageNumber="66">TURKEY</collectingCountry>
: 
<collectingRegion id="E755B8C9B956FF89FDA9EE6AFD00EB68" box="[590,680,1585,1611]" country="Turkey" name="Mersin" pageId="65" pageNumber="66">Mersin</collectingRegion>
</materialsCitation>
: 
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holo- and 
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; 3M, 1F, unterhalb 
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<collectorName id="886413FDB956FF89FA88EE6AFF50EB51" pageId="65" pageNumber="66">von Erdemli</collectorName>
) (
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<collectingDate id="416BA903B956FF89FDBFEE03FD68EB51" box="[600,704,1624,1650]" pageId="65" pageNumber="66" value="2002-07-06">6.7.2002</collectingDate>
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(
<collectorName id="886413FDB956FF89FD36EE03FCCEEB51" box="[721,870,1624,1650]" pageId="65" pageNumber="66">K.-G. Heller</collectorName>
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) (sound record); 1M, 2F, bei 
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(= 
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nach 
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), 
<date id="512F50EBB956FF89FCCAEE25FC3EEBBB" box="[813,918,1662,1688]" pageId="65" pageNumber="66" value="2002-07-05">
<collectingDate id="416BA903B956FF89FCCAEE25FC3EEBBB" box="[813,918,1662,1688]" pageId="65" pageNumber="66" value="2002-07-05">5.7.2002</collectingDate>
</date>
(
<collectorName id="886413FDB956FF89FC40EE25FB95EBBB" box="[935,1085,1662,1688]" pageId="65" pageNumber="66">K.-G. Heller</collectorName>
) (
<collectingCountry id="5D8636BBB956FF89FBB0EE25FB2EEBBB" box="[1111,1158,1662,1688]" name="Switzerland" pageId="65" pageNumber="66">CH</collectingCountry>
); 1F, 
<collectorName id="886413FDB956FF89FB2EEE25FAB7EBBB" box="[1225,1311,1662,1688]" pageId="65" pageNumber="66">Alahan</collectorName>
(
<geoCoordinate id="40A510ECB956FF89FAD6EE25FA3FEBBB" box="[1329,1431,1662,1688]" degrees="36" direction="north" minutes="52" orientation="latitude" pageId="65" pageNumber="66" precision="925" value="36.866665">36°52'N</geoCoordinate>
, 
<geoCoordinate id="40A510ECB956FF89FF70EEFEFF5FEB9C" box="[151,247,1701,1727]" degrees="33" direction="east" minutes="18" orientation="longitude" pageId="65" pageNumber="66" precision="925" value="33.3">33°18'E</geoCoordinate>
), 
<date id="512F50EBB956FF89FEEAEEFEFEDAEB9C" box="[269,370,1701,1727]" pageId="65" pageNumber="66" value="2002-07-07">
<collectingDate id="416BA903B956FF89FEEAEEFEFEDAEB9C" box="[269,370,1701,1727]" pageId="65" pageNumber="66" value="2002-07-07">7.7.2002</collectingDate>
</date>
(
<collectorName id="886413FDB956FF89FE64EEFEFDBEEB9C" box="[387,534,1701,1727]" pageId="65" pageNumber="66">K.-G. Heller</collectorName>
) (
<collectingCountry id="5D8636BBB956FF89FDC9EEFEFDF4EB9C" box="[558,604,1701,1727]" name="Switzerland" pageId="65" pageNumber="66">CH</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6D8B25A0B956FF89FF22EE90FF58EA2F" pageId="65" pageNumber="66" type="distribution">
<paragraph id="252E762BB956FF89FF22EE90FF58EA2F" blockId="65.[151,1436,1507,2036]" pageId="65" pageNumber="66">
<emphasis id="17E5AA39B956FF89FF22EE90FEC1EBC6" bold="true" box="[197,361,1739,1765]" pageId="65" pageNumber="66">Distribution:</emphasis>
This species is restricted to the western part of 
<collectingRegion id="E755B8C9B956FF89FC43EE97FBCFEBC5" box="[932,1127,1740,1766]" country="Turkey" name="Mersin" pageId="65" pageNumber="66">Mersin province</collectingRegion>
in Eastern Mediterranean 
<collectingCountry id="5D8636BBB956FF89FF70EEA9FF45EA2F" box="[151,237,1778,1804]" name="Turkey" pageId="65" pageNumber="66">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6D8B25A0B956FF8AFF22EF43FB3BEC6F" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="discussion">
<paragraph id="252E762BB956FF8AFF22EF43FB3BEC6F" blockId="65.[151,1436,1507,2036]" lastBlockId="66.[151,1436,152,332]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
<emphasis id="17E5AA39B956FF89FF22EF43FE9FEA11" bold="true" box="[197,311,1816,1842]" pageId="65" pageNumber="66">Remarks</emphasis>
: 
<bibRefCitation id="41000BDAB956FF89FEA2EF42FE7BEA10" author="Unal, M." box="[325,467,1817,1843]" pageId="65" pageNumber="66" pagination="157 - 203" refId="ref44688" refString="Unal, M. (2006) Tettigoniidae (Orthoptera) from Turkey and the Middle East. Transactions of the American Entomological Society, 132, 157 - 203." type="journal article" year="2006">Ünal (2006)</bibRefCitation>
reported that 
<taxonomicName id="E2910DA8B956FF89FD93EF42FD56EA11" authorityName="Ciplak" authorityYear="1999" box="[628,766,1817,1842]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="17E5AA39B956FF89FD93EF42FD56EA11" box="[628,766,1817,1842]" italics="true" pageId="65" pageNumber="66">E. femorata</emphasis>
</taxonomicName>
may be a synonym of 
<taxonomicName id="E2910DA8B956FF89FBE3EF42FB08EA11" authorityName="Karabag" authorityYear="1956" box="[1028,1184,1817,1842]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="unimacula">
<emphasis id="17E5AA39B956FF89FBE3EF42FB08EA11" box="[1028,1184,1817,1842]" italics="true" pageId="65" pageNumber="66">E. unimacula</emphasis>
</taxonomicName>
. This conclusion was mainly based on the probability of the presence of styli at the hind margin of the subgenital plate in the 
<typeStatus id="FA2AC889B956FF89FF70EF3DFEACEAA3" box="[151,260,1894,1920]" pageId="65" pageNumber="66" type="holotype">holotype</typeStatus>
of 
<emphasis id="17E5AA39B956FF89FED5EF3CFE4AEAA3" box="[306,482,1895,1920]" italics="true" pageId="65" pageNumber="66">
<taxonomicName id="E2910DA8B956FF89FED5EF3CFE75EAA3" authorityName="Karabag" authorityYear="1956" box="[306,477,1895,1920]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="unimacula">E. unimacula</taxonomicName>
,
</emphasis>
since presence/absence of styli was considered to be one of the diagnostic characters of these species by Çıplak (1999). During this study, type material and new specimens of these two species were examined. Although both species share numerous similarities and there is a possibility of loosing of styli in 
<emphasis id="17E5AA39B956FF89FEF3EF80FE14EAD7" box="[276,444,2011,2036]" italics="true" pageId="65" pageNumber="66">
<taxonomicName id="E2910DA8B956FF89FEF3EF80FE10EAD7" authorityName="Karabag" authorityYear="1956" box="[276,440,2011,2036]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="unimacula">E. unimacula</taxonomicName>
,
</emphasis>
there are further diagnostic characters that support the assumption that each is an independent evolutionary unit. Especially, the long unfused part of apical arms of the titillators (longer than fused part) and the shallow incision of the female subgenital plate (less than half the medial length) in 
<taxonomicName id="E2910DA8B955FF8AFA65E8E4FEAAEDDD" authorityName="Ciplak" authorityYear="1999" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="femorata">
<emphasis id="17E5AA39B955FF8AFA65E8E4FEAAEDDD" italics="true" pageId="66" pageNumber="67">E. femorata</emphasis>
</taxonomicName>
separates it distinctly from 
<emphasis id="17E5AA39B955FF8AFDA8E8BEFD5CEDDD" box="[591,756,229,254]" italics="true" pageId="66" pageNumber="67">
<taxonomicName id="E2910DA8B955FF8AFDA8E8BEFD58EDDD" authorityName="Karabag" authorityYear="1956" box="[591,752,229,254]" class="Insecta" family="Tettigoniidae" genus="Eupholidoptera" kingdom="Animalia" order="Orthoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="unimacula">E. unimacula</taxonomicName>
,
</emphasis>
in which the unfused parts of the apical arms are shorter than the fused part and the incision of the female subgenital plate is deeper than half the medial length (compare 
<figureCitation id="BDAA6AAEB955FF8AFEEAE969FE7EEC6F" box="[269,470,306,332]" captionStart="FIGURES 218–228" captionStartId="49.[151,269,1780,1804]" captionTargetBox="[220,1294,196,1731]" captionTargetId="figure-83@49.[188,1398,124,1789]" captionTargetPageId="49" captionText="FIGURES 218–228. Female subgenital plate of Eupholidoptera species (L— lateral view, V— ventral view) (Scale= 2 mm). 218—E. krueperi, 219—E. tauricola, 220—E. mersinensis (A, Mersin; B, Konya), 221—E. kykladica, 222—E. spinigera, 223—E. prasina, 224—E. tucherti, 225—E. karabagi, 226—E. unimacula, 227—E. femorata, 228—E. icariensis (Of the figures given above; 219 from Salman, 1983; 228 from Willemse, 1980)." pageId="66" pageNumber="67">Figures 226, 227</figureCitation>
). Thus, each of them was considered to be a separate taxon.
</paragraph>
</subSubSection>
</treatment>
</document>