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<document id="DEC9DB196961143979D9A7D0ED7286D0" ID-CLB-Dataset="32076" ID-DOI="10.11646/zootaxa.4320.3.6" ID-GBIF-Dataset="e80fc464-0c2a-4003-a867-ab8b5b3b399e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1051951" ID-ZooBank="Ca50E5A5-B417-4000-B67D-5F6028Da9154" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505712517045" checkinUser="plazi" docAuthor="Bocak, Ladislav" docDate="2017" docId="A312190FEC10FF85509F3895BF03FE5C" docLanguage="en" docName="zootaxa.4320.3.6.pdf" docOrigin="Zootaxa 4320 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladophorus boceki Bocak 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="511" masterDocId="5F2B6177EC15FF8350083D11BB5CFFD4" masterDocTitle="Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea" masterLastPageNumber="522" masterPageNumber="505" pageNumber="510" updateTime="1698468596363" updateUser="ExternalLinkService">
<mods:mods id="35FCECB72C37378AB21B418122BD0752" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7DCFCC08F0F0B888D56A9BEF38C93629">
<mods:title id="C874962F6F4BC0CF87D4DE9B7AF4FC50">Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="D08FBA9D4956E4F4DD15710C17C93E4C">Bocak, Ladislav</mods:namePart>
</mods:name>
<mods:typeOfResource id="78A67A527DE87D871B3F572224159888">text</mods:typeOfResource>
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<mods:title id="ECC2B7EA2ED11C95E178B8B6D23F4558">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="85903C8DB477A117DA09910BBA545692">
<mods:date id="00F8B9717ABF6007B3659262849758AD">2017</mods:date>
<mods:detail id="141B489C56AB07FF070BC1EB83555CBE" type="pubDate">
<mods:number id="EC8A8D488931C792BDC2234B2EF0AE13">2017-09-18</mods:number>
</mods:detail>
<mods:detail id="70D1E17B46CEE4F6A6E61D3BE98D7762" type="volume">
<mods:number id="A1F8C5539F1E442C8A695643D66244A8">4320</mods:number>
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<mods:detail id="1B7CE5912C39552ED802EDA1A6819ACE" type="issue">
<mods:number id="2B60762367D0BD1816838FE1D7420BA6">3</mods:number>
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<mods:classification id="FAA6722A68307BBDE3260DA083CEBF81">journal article</mods:classification>
<mods:identifier id="A088A765233E5FE375B20BD6FD5F2849" type="CLB-Dataset">32076</mods:identifier>
<mods:identifier id="40A6B25AA770341261A2ABD46E13FE32" type="DOI">10.11646/zootaxa.4320.3.6</mods:identifier>
<mods:identifier id="86DC8651A5C24E52E4DCDB64188240F3" type="GBIF-Dataset">e80fc464-0c2a-4003-a867-ab8b5b3b399e</mods:identifier>
<mods:identifier id="EC9E9A5DE73B6E9536063764BFBF360F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DBB6CBECAB9F28B1B490A7019990214A" type="Zenodo-Dep">1051951</mods:identifier>
<mods:identifier id="BBD183FDA1C7528522CF876D8F689307" type="ZooBank">Ca50E5A5-B417-4000-B67D-5F6028Da9154</mods:identifier>
</mods:mods>
<treatment id="A312190FEC10FF85509F3895BF03FE5C" ID-DOI="http://doi.org/10.5281/zenodo.6018016" ID-GBIF-Taxon="134090582" ID-Zenodo-Dep="6018016" LSID="urn:lsid:plazi:treatment:A312190FEC10FF85509F3895BF03FE5C" httpUri="http://treatment.plazi.org/id/A312190FEC10FF85509F3895BF03FE5C" lastPageId="6" lastPageNumber="511" pageId="5" pageNumber="510">
<subSubSection id="63A1FB92EC10FF86509F3895BAC2FA6B" pageId="5" pageNumber="510" type="nomenclature">
<paragraph id="2B04A819EC10FF86509F3895BAB1FA4A" blockId="5.[151,493,1412,1471]" box="[151,493,1412,1438]" pageId="5" pageNumber="510">
<heading id="704C1F75EC10FF86509F3895BAB1FA4A" bold="true" box="[151,493,1412,1438]" fontSize="11" level="1" pageId="5" pageNumber="510" reason="1">
<emphasis id="19CF740BEC10FF86509F3895BAB1FA4A" bold="true" box="[151,493,1412,1438]" pageId="5" pageNumber="510">
<taxonomicName id="ECBBD39AEC10FF86509F3895BAD5FA4A" authority="Bocak, 2017" authorityName="Bocak" authorityYear="2017" box="[151,393,1412,1438]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="boceki" status="sp. nov.">
<emphasis id="19CF740BEC10FF86509F3895BAD5FA4A" bold="true" box="[151,393,1412,1438]" italics="true" pageId="5" pageNumber="510">Cladophorus boceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02FCC970EC10FF8651983895BAB1FA4A" box="[400,493,1412,1438]" pageId="5" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2B04A819EC10FF86509F38B7BAC2FA6B" blockId="5.[151,493,1412,1471]" box="[151,414,1446,1471]" pageId="5" pageNumber="510">
(
<figureCitation id="B380B49CEC10FF86509738B7BA44FA6B" box="[159,280,1446,1471]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="5" pageNumber="510">Figs. 7, 21</figureCitation>
,
<figureCitation id="B380B49CEC10FF86512D38B6BA1FFA6B" box="[293,323,1447,1471]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="5" pageNumber="510">37</figureCitation>
,
<figureCitation id="B380B49CEC10FF86514638B7BAC9FA6B" box="[334,405,1446,1471]" captionStart="FIGURES 53 – 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 – 78. Male genitalia ventrally and laterally. 53 – 54, Cladophorus bicolor Kleine; 55 – 56, C. boceki sp. nov.; 57 – 58, C. craterensis sp. nov.; 59 – 60, C. haiaensis sp. nov.; 61 – 62, C. humeralis Kleine; 63 – 64, C. kailakiensis sp. nov.; 65 – 66, C. manokwarensis sp. nov.; 67 – 68, C. mindikensis sp. nov.; 69 – 70, C. motykai sp. nov.; 71 – 72, C. pallescens sp. nov.; 73 – 74, C. praecipuus Kleine; 75 – 76, C. riedeli sp. nov.; 77 – 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="5" pageNumber="510">55–56</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC10FF86509F38FFBE50F9FF" pageId="5" pageNumber="510" type="materials_examined">
<paragraph id="2B04A819EC10FF86509F38FFBE50F9FF" blockId="5.[151,1437,1518,2011]" pageId="5" pageNumber="510">
<materialsCitation id="9BD3A244EC10FF86509F38FFB91AF9FF" ID-GBIF-Occurrence="1638540605" collectionCode="LMBC" country="Indonesia" elevation="650" latitude="-1.75" location="Road Konti Unai" longLatPrecision="1308" longitude="136.31667" municipality="Yapen Island" pageId="5" pageNumber="510" specimenCount="2" stateProvince="Papua" typeStatus="holotype">
<emphasis id="19CF740BEC10FF86509F38FFBA16F9D3" bold="true" box="[151,330,1518,1543]" pageId="5" pageNumber="510">
<typeStatus id="F40016BBEC10FF86509F38FFBB8EF9D3" box="[151,210,1518,1543]" pageId="5" pageNumber="510">Type</typeStatus>
material.
</emphasis>
<typeStatus id="F40016BBEC10FF86515F38FFBA9EF9D3" box="[343,450,1518,1543]" pageId="5" pageNumber="510" type="holotype">Holotype</typeStatus>
, male.
<collectingCountry id="53ACE889EC10FF86521638FFB9D2F9D3" box="[542,654,1518,1543]" name="Indonesia" pageId="5" pageNumber="510">Indonesia</collectingCountry>
,
<collectingRegion id="E97F66FBEC10FF86529738FEB9BFF9D2" box="[671,739,1519,1542]" country="Indonesia" name="Papua Barat" pageId="5" pageNumber="510">Papua</collectingRegion>
Prov.,
<collectingMunicipality id="CB603263EC10FF86533638FEB884F9D3" box="[830,984,1518,1543]" pageId="5" pageNumber="510">Yapen Island</collectingMunicipality>
,
<location id="2E64FEC2EC10FF8653E138FFBFE9F9D3" LSID="urn:lsid:plazi:treatment:A312190FEC10FF85509F3895BF03FE5C:2E64FEC2EC10FF8653E138FFBFE9F9D3" box="[1001,1205,1518,1543]" country="Indonesia" latitude="-1.75" longLatPrecision="1308" longitude="136.31667" municipality="Yapen Island" name="Road Konti Unai" pageId="5" pageNumber="510" stateProvince="Papua">Road Konti Unai</location>
–Ambaid,
<geoCoordinate id="4E8FCEDEEC10FF86553938FFBEC0F9D3" box="[1329,1436,1518,1543]" direction="east" orientation="longitude" pageId="5" pageNumber="510" precision="925" value="136.31667">136°19´E</geoCoordinate>
<geoCoordinate id="4E8FCEDEEC10FF86509F3B03BBBBF9FF" box="[151,231,1554,1579]" direction="south" orientation="latitude" pageId="5" pageNumber="510" precision="925" value="-1.75">1°45´S</geoCoordinate>
,
<elevation id="A0964F2AEC10FF8650FB3B03BA2EF9FE" box="[243,370,1554,1579]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="5" pageNumber="510" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">
<quantity id="EC4305FCEC10FF8650FB3B03BA2EF9FE" box="[243,370,1554,1579]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="5" pageNumber="510" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">600–700 m</quantity>
</elevation>
(
<accessionNumber id="34E835FAEC10FF8651883B02BA83F9FF" box="[384,479,1555,1579]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A00407" pageId="5" pageNumber="510" type="EnaNcbi">A00407</accessionNumber>
,
<collectionCode id="4DAA30DCEC10FF8651E23B02B962F9FF" box="[490,574,1555,1579]" pageId="5" pageNumber="510">LMBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9BD3A244EC10FF86525A3B02BE54F9FF" ID-GBIF-Occurrence="1638540593" box="[594,1288,1554,1579]" collectionCode="LMBC" country="Indonesia" elevation="650" latitude="-1.75" location="Road Konti Unai" longLatPrecision="1308" longitude="136.31667" municipality="Yapen Island" pageId="5" pageNumber="510" specimenCount="2" specimenCount-male="2" stateProvince="Papua" typeStatus="paratype">
<typeStatus id="F40016BBEC10FF86525A3B02B99EF9FF" box="[594,706,1555,1579]" pageId="5" pageNumber="510" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="3DBD6390EC10FF8652C53B02B87AF9FF" box="[717,806,1554,1579]" pageId="5" pageNumber="510" type="male">2 males</specimenCount>
, same locality data (A00406, 408,
<collectionCode id="4DAA30DCEC10FF8654A43B02BE5CF9FF" box="[1196,1280,1555,1579]" pageId="5" pageNumber="510">LMBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC10FF8650CF3B27BEC9F90B" pageId="5" pageNumber="510" type="diagnosis">
<paragraph id="2B04A819EC10FF8650CF3B27BEC9F90B" blockId="5.[151,1437,1518,2011]" pageId="5" pageNumber="510">
<emphasis id="19CF740BEC10FF8650CF3B27BA1DF99B" bold="true" box="[199,321,1590,1615]" pageId="5" pageNumber="510">Diagnosis.</emphasis>
<taxonomicName id="ECBBD39AEC10FF8651413B27B970F99A" authorityName="Bocak" authorityYear="2017" box="[329,556,1590,1614]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="boceki">
<emphasis id="19CF740BEC10FF8651413B27B970F99A" box="[329,556,1590,1614]" italics="true" pageId="5" pageNumber="510">Cladophorus boceki</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="ECBBD39AEC10FF86531F3B26B8EAF99A" box="[791,950,1591,1614]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="praecipuus">
<emphasis id="19CF740BEC10FF86531F3B26B8EAF99A" box="[791,950,1591,1614]" italics="true" pageId="5" pageNumber="510">C. praecipuus</emphasis>
</taxonomicName>
(
<figureCitation id="B380B49CEC10FF8653CD3B27BF56F99B" box="[965,1034,1590,1615]" captionStart="FIGURES 4 – 5. 4" captionStartId="3.[151,264,1220,1243]" captionTargetBox="[159,1417,255,1198]" captionTargetId="figure@3.[151,1436,255,1210]" captionTargetPageId="3" captionText="FIGURES 4 – 5. 4, Phylogenetic hypothesis of Papuan Cladophorus inferred from the cox 1 mtDNA dataset using maximum likelihood criterion. 5, Cladophorus sp., general appearance." httpUri="https://zenodo.org/record/1051955/files/figure.png" pageId="5" pageNumber="510">Fig. 4</figureCitation>
) and both species are similar in the general appearance (
<figureCitation id="B380B49CEC10FF86518D3B4BB949F9A7" box="[389,533,1626,1651]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="5" pageNumber="510">Figs. 21, 30</figureCitation>
,
<figureCitation id="B380B49CEC10FF86522D3B4AB92FF9A7" box="[549,627,1627,1651]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="5" pageNumber="510">37, 47</figureCitation>
). Despite their very low genetic divergence, these species are clearly defined by the length of antennal lamella 3 (
<figureCitation id="B380B49CEC10FF86529B3B6FB853F943" box="[659,783,1662,1687]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="5" pageNumber="510">Figs. 7, 14</figureCitation>
), the relative size of male eyes (ratio 1.55 and
<quantity id="EC4305FCEC10FF8655223B6EBE26F943" box="[1322,1402,1662,1687]" metricMagnitude="-2" metricUnit="m" metricValue="5.1308" pageId="5" pageNumber="510" unit="in" value="2.02">2.02 in</quantity>
<taxonomicName id="ECBBD39AEC10FF86558A3B6EBBBCF96F" authorityName="Bocak" authorityYear="2017" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="boceki">
<emphasis id="19CF740BEC10FF86558A3B6EBBBCF96F" italics="true" pageId="5" pageNumber="510">C. boceki</emphasis>
</taxonomicName>
and
<taxonomicName id="ECBBD39AEC10FF8651113BB2BAE4F96E" box="[281,440,1699,1722]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="praecipuus">
<emphasis id="19CF740BEC10FF8651113BB2BAE4F96E" box="[281,440,1699,1722]" italics="true" pageId="5" pageNumber="510">C. praecipuus</emphasis>
</taxonomicName>
, respectively) and
<taxonomicName id="ECBBD39AEC10FF86529A3BB2B9A1F96F" authorityName="Bocak" authorityYear="2017" box="[658,765,1699,1723]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="boceki">
<emphasis id="19CF740BEC10FF86529A3BB2B9A1F96F" box="[658,765,1699,1723]" italics="true" pageId="5" pageNumber="510">C. boceki</emphasis>
</taxonomicName>
has a robust phallus with the gradually pointed apical part unlike
<taxonomicName id="ECBBD39AEC10FF8650EC3BD6BADCF90A" box="[228,384,1735,1758]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="praecipuus">
<emphasis id="19CF740BEC10FF8650EC3BD6BADCF90A" box="[228,384,1735,1758]" italics="true" pageId="5" pageNumber="510">C. praecipuus</emphasis>
</taxonomicName>
, which has the slender phallus, parallel-sided in four-fifths of its length (
<figureCitation id="B380B49CEC10FF8654A43BD7BED5F90B" box="[1196,1417,1734,1759]" captionStart="FIGURES 53 – 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 – 78. Male genitalia ventrally and laterally. 53 – 54, Cladophorus bicolor Kleine; 55 – 56, C. boceki sp. nov.; 57 – 58, C. craterensis sp. nov.; 59 – 60, C. haiaensis sp. nov.; 61 – 62, C. humeralis Kleine; 63 – 64, C. kailakiensis sp. nov.; 65 – 66, C. manokwarensis sp. nov.; 67 – 68, C. mindikensis sp. nov.; 69 – 70, C. motykai sp. nov.; 71 – 72, C. pallescens sp. nov.; 73 – 74, C. praecipuus Kleine; 75 – 76, C. riedeli sp. nov.; 77 – 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="5" pageNumber="510">Figs. 55–56, 73–74</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC10FF8550CF3BFBB8A1FEEB" lastPageId="6" lastPageNumber="511" pageId="5" pageNumber="510" type="description">
<paragraph id="2B04A819EC10FF8550CF3BFBBA86FF2C" blockId="5.[151,1437,1518,2011]" lastBlockId="6.[151,1436,151,392]" lastPageId="6" lastPageNumber="511" pageId="5" pageNumber="510">
<emphasis id="19CF740BEC10FF8650CF3BFBBA05F8D7" bold="true" box="[199,345,1770,1795]" pageId="5" pageNumber="510">Description.</emphasis>
Male. Body large,
<quantity id="EC4305FCEC10FF8652273BFBB98EF8D6" box="[559,722,1770,1795]" metricMagnitude="-2" metricUnit="m" metricValue="1.385" metricValueMax="1.41" metricValueMin="1.3599999999999999" pageId="5" pageNumber="510" unit="mm" value="13.85" valueMax="14.1" valueMin="13.6">13.6–14.1 mm</quantity>
long, dark brown to black, elytra yellow, only apical 1/12 black (
<figureCitation id="B380B49CEC10FF8650973A1FBBAFF8F3" box="[159,243,1806,1831]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="5" pageNumber="510">Fig. 37</figureCitation>
), transition between dark and light coloured parts of elytra abrupt, only elytral costae with slightly larger dark part (
<figureCitation id="B380B49CEC10FF86511D3A23BA31F89F" box="[277,365,1842,1867]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="5" pageNumber="510">Fig. 37</figureCitation>
). Head small, with hemispherically prominent eyes, frontal interocular distance 1.52 times maximum diameter; antennae long, reaching two thirds of elytral length, flabellate, antennomere 3 with lamella 2.4 times longer than stem of antennomere, lamella attached to apical part of antennomere (
<figureCitation id="B380B49CEC10FF8654CE3A6BBE4EF847" box="[1222,1298,1914,1939]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="5" pageNumber="510">Fig. 7</figureCitation>
). Pronotum transverse, 1.5 times wider than long at midline, lateral margins slightly elevated, frontal margin rounded, lateral margins bi-sinuate, frontal angles well marked, posterior angles sharply prominent, postero-lateral carinae weak (
<figureCitation id="B380B49CEC13FF8550973D86BBA9FF64" box="[159,245,151,176]" captionStart="FIGURES 6 – 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 – 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="6" pageNumber="511">Fig. 21</figureCitation>
). Elytra flat, slightly widened posteriorly, transverse costae weak, partly covered by pubescence, elytral areoles slightly transverse (
<figureCitation id="B380B49CEC13FF8551CF3DADB946FF00" box="[455,538,188,213]" captionStart="FIGURES 33 – 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 – 52. Elytron. 33 – 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 – 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="6" pageNumber="511">Fig. 37</figureCitation>
). Phallus robust, widest in apical third, gradually acuminate apically, with robust apical process (
<figureCitation id="B380B49CEC13FF85514C3DCEBA92FF2C" box="[324,462,223,248]" captionStart="FIGURES 53 – 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 – 78. Male genitalia ventrally and laterally. 53 – 54, Cladophorus bicolor Kleine; 55 – 56, C. boceki sp. nov.; 57 – 58, C. craterensis sp. nov.; 59 – 60, C. haiaensis sp. nov.; 61 – 62, C. humeralis Kleine; 63 – 64, C. kailakiensis sp. nov.; 65 – 66, C. manokwarensis sp. nov.; 67 – 68, C. mindikensis sp. nov.; 69 – 70, C. motykai sp. nov.; 71 – 72, C. pallescens sp. nov.; 73 – 74, C. praecipuus Kleine; 75 – 76, C. riedeli sp. nov.; 77 – 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="6" pageNumber="511">Figs. 55–56</figureCitation>
).
</paragraph>
<paragraph id="2B04A819EC13FF8550CF3C15B8A1FEEB" blockId="6.[151,1436,151,392]" pageId="6" pageNumber="511">
<emphasis id="19CF740BEC13FF8550CF3C15BA21FEC9" bold="true" box="[199,381,260,285]" pageId="6" pageNumber="511">Measurements.</emphasis>
BL
<quantity id="EC4305FCEC13FF8551A63C15B90FFEC8" box="[430,595,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.385" metricValueMax="1.41" metricValueMin="1.3599999999999999" pageId="6" pageNumber="511" unit="mm" value="13.85" valueMax="14.1" valueMin="13.6">13.6–14.1 mm</quantity>
, PL 1.95–2.0 mm, PW
<quantity id="EC4305FCEC13FF8553523C15B8A3FEC8" box="[858,1023,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="2.95" metricValueMin="2.85" pageId="6" pageNumber="511" unit="mm" value="2.9" valueMax="2.95" valueMin="2.85">2.85–2.95 mm</quantity>
, WH
<quantity id="EC4305FCEC13FF8554483C15BFB9FEC8" box="[1088,1253,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.625" metricValueMax="1.65" metricValueMin="1.6" pageId="6" pageNumber="511" unit="mm" value="1.625" valueMax="1.65" valueMin="1.6">1.60–1.65 mm</quantity>
, Edist
<quantity id="EC4305FCEC13FF8555393C15BEC4FEC8" box="[1329,1432,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="1.02" pageId="6" pageNumber="511" unit="mm" value="1.02">1.02 mm</quantity>
, Ediam
<quantity id="EC4305FCEC13FF8550EF3C36BA11FEEB" box="[231,333,295,320]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="6" pageNumber="511" unit="mm" value="0.66">0.66 mm</quantity>
, LE
<quantity id="EC4305FCEC13FF85518A3C36BABBFEEB" box="[386,487,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="6" pageNumber="511" unit="mm" value="11.6">11.6 mm</quantity>
, antennomere 3: stem
<quantity id="EC4305FCEC13FF8552EB3C39B867FEEB" box="[739,827,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="511" unit="mm" value="1.1">1.1 mm</quantity>
, lamella
<quantity id="EC4305FCEC13FF8553A83C36B8A5FEEB" box="[928,1017,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="511" unit="mm" value="2.6">2.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC13FF8550CF3C5DB906FEB0" box="[199,602,332,357]" pageId="6" pageNumber="511" type="distribution">
<paragraph id="2B04A819EC13FF8550CF3C5DB906FEB0" blockId="6.[151,1436,151,392]" box="[199,602,332,357]" pageId="6" pageNumber="511">
<emphasis id="19CF740BEC13FF8550CF3C5DBA3CFEB1" bold="true" box="[199,352,332,357]" pageId="6" pageNumber="511">Distribution.</emphasis>
Yapen Island (
<figureCitation id="B380B49CEC13FF8552003C5DB911FEB0" box="[520,589,332,357]" captionStart="FIGURES 1 – 3. 1 – 2" captionStartId="2.[151,264,1396,1419]" captionTargetBox="[151,1436,339,1375]" captionTargetId="figure@2.[151,1436,339,1375]" captionTargetPageId="2" captionText="FIGURES 1 – 3. 1 – 2, General appearance of Cladophorus bicolor in the Central mountains of New Guinea. 3, Sampled localities and the distribution of Cladophorus species" httpUri="https://zenodo.org/record/1051953/files/figure.png" pageId="6" pageNumber="511">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC13FF8550CF3C7EBF03FE5C" box="[199,1119,367,392]" pageId="6" pageNumber="511" type="etymology">
<paragraph id="2B04A819EC13FF8550CF3C7EBF03FE5C" blockId="6.[151,1436,151,392]" box="[199,1119,367,392]" pageId="6" pageNumber="511">
<emphasis id="19CF740BEC13FF8550CF3C7EBA12FE5C" bold="true" box="[199,334,367,392]" pageId="6" pageNumber="511">Etymology.</emphasis>
The species name is a patronym in honor of Matej Bocek (Olomouc).
</paragraph>
</subSubSection>
</treatment>
</document>
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