402 lines
25 KiB
XML
402 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.429.7755" ID-GBIF-Dataset="508c3f12-f203-44a4-8784-04d55275dcd2" ID-PMC="PMC4137295" ID-Pensoft-Pub="1313-2970-429-1" ID-PubMed="25147467" ID-ZBK="45B807380F9540E6846C18CA5E9DA09B" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-429-1" ModsDocOrigin="ZooKeys 429" ModsDocTitle="Milnesium berladnicorum sp. n. (Eutardigrada, Apochela, Milnesiidae), a new species of water bear from Romania" checkinTime="1451245567636" checkinUser="pensoft" docAuthor="Ciobanu, Daniel Adrian, Zawierucha, Krzysztof, Moglan, Ioan & Kaczmarek, Lukasz" docDate="2014" docId="650E4AD95664BAA02E1C4F279AF69C03" docLanguage="en" docName="ZooKeys 429: 1-11" docOrigin="ZooKeys 429" docSource="http://dx.doi.org/10.3897/zookeys.429.7755" docTitle="Milnesium berladnicorum Ciobanu, Zawierucha, Moglan & Kaczmarek, 2014, sp. n." docType="treatment" docUuid="FBF8C785-2B53-48B2-B696-D442BAD89A0F" docUuidSource="ZooBank" docVersion="4" lastPageNumber="6" masterDocId="9A7CF96B0461FFF0FFDBFF93FF83C748" masterDocTitle="Milnesium berladnicorum sp. n. (Eutardigrada, Apochela, Milnesiidae), a new species of water bear from Romania" masterLastPageNumber="11" masterPageNumber="1" pageNumber="3" updateTime="1668159043696" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Milnesium berladnicorum sp. n. (Eutardigrada, Apochela, Milnesiidae), a new species of water bear from Romania</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ciobanu, Daniel Adrian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zawierucha, Krzysztof</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moglan, Ioan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kaczmarek, Lukasz</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>429</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>11</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.429.7755</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.429.7755</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-429-1</mods:identifier>
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<mods:identifier type="ZBK">45B807380F9540E6846C18CA5E9DA09B</mods:identifier>
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<mods:identifier type="ZooBank">45B807380F9540E6846C18CA5E9DA09B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152054629" LSID="urn:lsid:zoobank.org:act:FBF8C785-2B53-48B2-B696-D442BAD89A0F" httpUri="http://treatment.plazi.org/id/650E4AD95664BAA02E1C4F279AF69C03" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
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<subSubSection pageId="2" pageNumber="3" type="multiple">
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<paragraph pageId="2" pageNumber="3">Taxon classification Animalia Apochela Milnesiidae</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="nomenclature">
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<paragraph pageId="2" pageNumber="3">
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<taxonomicName LSID="http://zoobank.org/FBF8C785-2B53-48B2-B696-D442BAD89A0F" class="Eutardigrada" family="Milnesiidae" genus="Milnesium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Milnesium berladnicorum" order="Apochela" pageId="2" pageNumber="3" phylum="Tardigrada" rank="species" species="berladnicorum">Milnesium berladnicorum</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="3">sp. n.</taxonomicNameLabel>
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Figs 1-6, Table 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="material examined">
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<paragraph pageId="2" pageNumber="3">Material examined.</paragraph>
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<paragraph pageId="2" pageNumber="3">Holotype (female), 52 paratypes and 2 exuvia with 7 and 9 smooth eggs.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
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<paragraph pageId="2" pageNumber="3">Description</paragraph>
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<paragraph lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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(measurements and statistics in Table 1). Body brownish (in live specimens) or transparent (in fixed specimens) with eyes (visible before and after mounting in
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<normalizedToken originalValue="Hoyer’s">Hoyer's</normalizedToken>
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medium - 90% of fixed specimens had eyes). Six peribuccal papillae (ventral papilla smallest) and six peribuccal lamellae (of equal size) around the mouth opening present. Two cephalic papillae positioned laterally. The cuticle is covered with numerous tiny, shallow and rounded depressions (pseudopores) (Figs 4-5). Under PCM these pseudopores are visible as light spots, placing the species within the
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<pageBreakToken pageId="3" pageNumber="4" start="start">granulatum</pageBreakToken>
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group. Bucco-pharyngeal apparatus of the
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<taxonomicName class="Eutardigrada" family="Milnesiidae" genus="Milnesium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Milnesium" order="Apochela" pageId="3" pageNumber="4" phylum="Tardigrada" rank="genus">Milnesium</taxonomicName>
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type (Fig. 6). Buccal tube funnel-shaped, wider anteriorly (on average the posterior diameter is 73% of the anterior diameter). Pharyngeal bulb elongated, pear-shaped and without placoids or septulum. Claws of the
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<taxonomicName class="Eutardigrada" family="Milnesiidae" genus="Milnesium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Milnesium" order="Apochela" pageId="3" pageNumber="4" phylum="Tardigrada" rank="genus">Milnesium</taxonomicName>
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type, slender (Figs 2-3). Primary branches on all legs with small accessory points on the top of the branch. Secondary claws of all legs with rounded basal thickenings (lunules) (sometimes barely visible) (Fig. 3). Secondary branches of external claws
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<normalizedToken originalValue="I–III">I-III</normalizedToken>
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and posterior and anterior claws IV with two points. Secondary branches of internal claws
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<normalizedToken originalValue="I–III">I-III</normalizedToken>
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with three points (i.e. claw configuration: [2-3]-[2-2]) (Figs 2-3). Single, long transverse, cuticular bars under claws
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<normalizedToken originalValue="I–III">I-III</normalizedToken>
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present (Fig. 2).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="eggs">
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<paragraph pageId="3" pageNumber="4">Eggs.</paragraph>
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<paragraph pageId="3" pageNumber="4">Smooth, deposited in exuvia.</paragraph>
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<paragraph pageId="3" pageNumber="4">No males were found.</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="4" type="locus typicus">
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<paragraph pageId="3" pageNumber="4">Locus typicus.</paragraph>
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<paragraph pageId="3" pageNumber="4">
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46°
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="14.74167">14.74167N</geoCoordinate>
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, 27°
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="40.27333">40.27333E</geoCoordinate>
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; 99 m asl: Romania, Vaslui County,
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<normalizedToken originalValue="Bârlad">Barlad</normalizedToken>
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town, coppice, lichens (
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<taxonomicName class="Lecanoromycetes" family="Teloschistaceae" genus="Xanthoria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Xanthoria parietina" order="Teloschistales" pageId="3" pageNumber="4" phylum="Ascomycota" rank="species" species="parietina">Xanthoria parietina</taxonomicName>
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(L.) Th. Fr. (1860)) from tree.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="etymology">
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<paragraph pageId="4" pageNumber="5">
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<pageBreakToken pageId="4" pageNumber="5" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="4" pageNumber="5">
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This new species is named after the Berladnici, an ancient population with a controversial origin (most probably Slavs) who previously lived in the area of the present
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<normalizedToken originalValue="Bârlad">Barlad</normalizedToken>
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town.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="type depositories">
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<paragraph pageId="5" pageNumber="6">
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<pageBreakToken pageId="5" pageNumber="6" start="start">Type</pageBreakToken>
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depositories.
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</paragraph>
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<paragraph pageId="5" pageNumber="6">
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Holotype (female; slide: P8-8) and 29 paratypes (females) and 1 exuvium with eggs (slides: P8-4, P8-5, P8-6, P8-9, P8-13, P8-14, P8-15, P8-17, P8-19) are preserved at the Department of Animal Taxonomy and Ecology, A. Mickiewicz University in
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<normalizedToken originalValue="Poznań">Poznan</normalizedToken>
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, Umultowska 89, 61-614
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<normalizedToken originalValue="Poznań">Poznan</normalizedToken>
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, Poland. Additionally, 14 paratypes (females) and 1 exuvium with eggs (slides: P8-1, P8-3, P8-16, P8-18) are deposited at Natural History Museum of "Alexandru Ioan Cuza" University from
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<normalizedToken originalValue="Iași">Iași</normalizedToken>
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(Bd. Independentei No.16, 700101), 4 paratypes (females; slides: P8-7, P8-12) are deposited at collection of Binda and Pilato (Museum of the Department of Animal Biology "Marcello La Greca", University of Catania, Italy) and 5 paratypes (females; slides: P8-2, P8-10, P8-11) are deposited at the Natural History Museum, University of Copenhagen Universitetsparken 15, DK-2100 Copenhagen, Denmark.
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</paragraph>
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<caption pageId="5" pageNumber="6">
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<paragraph pageId="5" pageNumber="6">
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Figures 1.
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<taxonomicName class="Eutardigrada" family="Milnesiidae" genus="Milnesium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Milnesium berladnicorum" order="Apochela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="berladnicorum">Milnesium berladnicorum</taxonomicName>
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sp. n. Habitus (ventral view).
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="6">
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<paragraph pageId="5" pageNumber="6">
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Figures 2-6.
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<taxonomicName class="Eutardigrada" family="Milnesiidae" genus="Milnesium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Milnesium berladnicorum" order="Apochela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="berladnicorum">Milnesium berladnicorum</taxonomicName>
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sp. n.: 2 claws III 3 claws IV 4 sculpture on dorsal cuticle above
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<normalizedToken originalValue="II–III">II-III</normalizedToken>
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pair of legs 5 sculpture on dorsal cuticle above IV pair of legs 6 buccal apparatus (ventral view).
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="6">
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<paragraph pageId="5" pageNumber="6">
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Figure 7. The administrative map of Romania with 13 highlighted counties in which species of the genus
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<taxonomicName class="Eutardigrada" family="Milnesiidae" genus="Milnesium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Milnesium" order="Apochela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="genus">Milnesium</taxonomicName>
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were reported:
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<taxonomicName class="Eutardigrada" family="Milnesiidae" genus="Milnesium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Milnesium tardigradum" order="Apochela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="tardigradum">Milnesium tardigradum</taxonomicName>
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sensu lato (according with
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<bibRefCitation author="Rudescu, L" journalOrPublisher="7, Editura Academiei Republicii Populare Romane, București 4" pageId="10" pageNumber="11" title="Tardigrada. Fauna Republicii Populare Romane, Arthropoda, Fasc." year="1964">Rudescu 1964</bibRefCitation>
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; see Discussion): 1
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<normalizedToken originalValue="Argeş">Arges</normalizedToken>
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2
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<normalizedToken originalValue="Bistriţa-Năsăud">Bistrita-Năsăud</normalizedToken>
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3
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<normalizedToken originalValue="Caraş-Severin">Caras-Severin</normalizedToken>
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4 Cluj 5
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<normalizedToken originalValue="Dâmboviţa">Dambovita</normalizedToken>
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6 Harghita 7 Ilfov County and Bucharest City 8
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<normalizedToken originalValue="Maramureş">Maramures</normalizedToken>
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9
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<normalizedToken originalValue="Mehedinţi">Mehedinti</normalizedToken>
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11 Suceava 12 Tulcea.
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<taxonomicName class="Eutardigrada" family="Milnesiidae" genus="Milnesium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Milnesium granulatum" order="Apochela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="granulatum">Milnesium granulatum</taxonomicName>
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and
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<taxonomicName class="Eutardigrada" family="Milnesiidae" genus="Milnesium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Milnesium asiaticum" order="Apochela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="asiaticum">Milnesium asiaticum</taxonomicName>
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(according to
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<bibRefCitation author="Ciobanu, DA" journalOrPublisher="Zoologischer Anzeiger" pageId="9" pageNumber="10" url="http://biozoojournals.ro/nwjz/content/v10s1/nwjz_140301_Ciobanu.pdf" year="2014">Ciobanu et al. 2014</bibRefCitation>
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): 10
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<normalizedToken originalValue="Neamţ">Neamt</normalizedToken>
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(in green).
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<taxonomicName class="Eutardigrada" family="Milnesiidae" genus="Milnesium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Milnesium berladnicorum" order="Apochela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="berladnicorum">Milnesium berladnicorum</taxonomicName>
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sp. n. (present study): 13 Vaslui (in blue). Map outline according to Wikipedia: http://ro.wikipedia.org/wiki/Fi%C8%99ier:Romania_counties_blank_big.png
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</paragraph>
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</caption>
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<caption pageId="5" pageNumber="6">
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<paragraph pageId="5" pageNumber="6">
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Table 1. Measurements and pt values of selected morphological structures of fifteen females from the type population of
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<taxonomicName class="Eutardigrada" family="Milnesiidae" genus="Milnesium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Milnesium berladnicorum" order="Apochela" pageId="5" pageNumber="6" phylum="Tardigrada" rank="species" species="berladnicorum">Milnesium berladnicorum</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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<paragraph pageId="5" pageNumber="6">
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<table pageId="5" pageNumber="6">
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<tr pageId="5" pageNumber="6">
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<th colspan="1" pageId="5" pageNumber="6" rowspan="2" style="text-align:left">CHARACTER</th>
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<th colspan="1" pageId="5" pageNumber="6" rowspan="2" style="text-align:center">N</th>
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<th colspan="6" pageId="5" pageNumber="6" rowspan="1" style="text-align:center">RANGE</th>
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<th colspan="2" pageId="5" pageNumber="6" rowspan="1" style="text-align:center">MEAN</th>
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<th colspan="2" pageId="5" pageNumber="6" rowspan="1" style="text-align:center">SD</th>
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<th colspan="2" pageId="5" pageNumber="6" rowspan="1" style="text-align:center">Holotype</th>
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</tr>
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<tr pageId="5" pageNumber="6">
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<th colspan="3" pageId="5" pageNumber="6" rowspan="1" style="text-align:left">
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<normalizedToken originalValue="µm">µm</normalizedToken>
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</th>
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<th colspan="3" pageId="5" pageNumber="6" rowspan="1" style="text-align:center">pt</th>
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<th colspan="1" pageId="5" pageNumber="6" rowspan="1" style="text-align:center">
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<normalizedToken originalValue="µm">µm</normalizedToken>
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</th>
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<th colspan="1" pageId="5" pageNumber="6" rowspan="1" style="text-align:center">pt</th>
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<th colspan="1" pageId="5" pageNumber="6" rowspan="1" style="text-align:center">
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<normalizedToken originalValue="µm">µm</normalizedToken>
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</th>
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<th colspan="1" pageId="5" pageNumber="6" rowspan="1" style="text-align:center">pt</th>
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<th colspan="1" pageId="5" pageNumber="6" rowspan="1" style="text-align:center">
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<normalizedToken originalValue="µm">µm</normalizedToken>
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</th>
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<th colspan="1" pageId="5" pageNumber="6" rowspan="1" style="text-align:center">pt</th>
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</tr>
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<tr pageId="5" pageNumber="6">
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">1557</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">1942</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">1742</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">127</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">1619</td>
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||
</tr>
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<tr pageId="5" pageNumber="6">
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">14.8</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">20.5</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">17.4</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">1.6</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">18.0</td>
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</tr>
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<tr pageId="5" pageNumber="6">
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">12.7</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">17.9</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">14.5</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">1.6</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">14.4</td>
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</tr>
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||
<tr pageId="5" pageNumber="6">
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">66.6</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">71.2</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">69.2</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">1.3</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">70.4</td>
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</tr>
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<tr pageId="5" pageNumber="6">
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">35.0</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">47.1</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">41.3</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">3.4</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">42.2</td>
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</tr>
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<tr pageId="5" pageNumber="6">
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">30.6</td>
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<td colspan="1" pageId="5" pageNumber="6" rowspan="1">38.9</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">33.5</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">2.3</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">32.6</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">27.7</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">36.0</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">30.3</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">2.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">29.6</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">40.5</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">47.6</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">43.9</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">2.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">45.2</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">32.9</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">45.3</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">39.3</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">3.0</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">37.5</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">38.1</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">45.6</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">41.8</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">2.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">40.1</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">33.9</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">41.7</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">37.3</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">2.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">33.9</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">9.0</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">12.5</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">11.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">1.1</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">9.0</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">43.3</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">53.5</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">48.4</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">3.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">51.4</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">36.9</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">44.4</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">40.6</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">2.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">40.6</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">39.9</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">51.8</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">45.4</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">3.1</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">44.2</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">34.4</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">42.5</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">38.0</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">2.7</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">37.0</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">10.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">15.6</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">12.4</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">2.2</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">44.9</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">54.8</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">49.5</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">3.1</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">46.0</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">37.8</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">44.5</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">40.9</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">2.1</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">42.2</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">39.4</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">53.7</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">46.4</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">4.5</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">39.8</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">33.6</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">45.6</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">38.9</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">3.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">39.1</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">10.6</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">14.0</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">12.0</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">1.4</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">57.9</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">74.8</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">63.8</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">4.7</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">63.8</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">40.0</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">50.0</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">46.9</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">3.0</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">47.6</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">54.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">70.4</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">62.0</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">4.6</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">61.7</td>
|
||
</tr>
|
||
<tr pageId="5" pageNumber="6">
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">38.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">52.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">46.2</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">4.0</td>
|
||
<td colspan="1" pageId="5" pageNumber="6" rowspan="1">44.7</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |