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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.898.39265" ID-GBIF-Dataset="9dbc167e-c273-42f5-82a0-dedb1dfce5bc" ID-PMC="PMC6917625" ID-Pensoft-Pub="1313-2970-898-121" ID-Pensoft-UUID="2A2FB0D263AA5E498C5C85EA4031053A" ID-PubMed="31866734" ID-ZooBank="9B537DC38DB9459E97717687AFA19244" ModsDocID="1313-2970-898-121" checkinTime="1575997087753" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I., Semenyuk, Irina, Efeykin, Boris D. &amp; Panha, Somsak" docDate="2019" docId="4BDFE925C1D45CD29537931853201D5D" docLanguage="en" docName="ZooKeys 898: 121-158" docOrigin="ZooKeys 898" docSource="http://dx.doi.org/10.3897/zookeys.898.39265" docTitle="Orthomorpha scabra Jeekel 1964" docType="treatment" docVersion="4" id="2A2FB0D263AA5E498C5C85EA4031053A" lastPageNumber="121" masterDocId="2A2FB0D263AA5E498C5C85EA4031053A" masterDocTitle="Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Vietnam, with several new records and descriptions of two new species" masterLastPageNumber="158" masterPageNumber="121" pageNumber="121" updateTime="1668168219523" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Review of the millipede genus Orthomorpha Bollman, 1893 (Diplopoda, Polydesmida, Paradoxosomatidae) in Vietnam, with several new records and descriptions of two new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Golovatch, Sergei I.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Semenyuk, Irina</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Efeykin, Boris D.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>898</mods:number>
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<mods:start>121</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.898.39265</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.898.39265</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-898-121</mods:identifier>
<mods:identifier type="ZooBank">9B537DC38DB9459E97717687AFA19244</mods:identifier>
<mods:identifier type="Pensoft-UUID">2A2FB0D263AA5E498C5C85EA4031053A</mods:identifier>
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<treatment ID-GBIF-Taxon="160689313" LSID="urn:lsid:plazi:treatment:4BDFE925C1D45CD29537931853201D5D" httpUri="http://treatment.plazi.org/id/4BDFE925C1D45CD29537931853201D5D" lastPageNumber="121" pageId="0" pageNumber="121">
<subSubSection pageId="0" pageNumber="121" type="nomenclature">
<paragraph pageId="0" pageNumber="121">
<taxonomicName LSID="4BDFE925-C1D4-5CD2-9537-931853201D5D" authority="Jeekel, 1964" authorityName="Jeekel" authorityYear="1964" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha scabra" order="Polydesmida" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="scabra">Orthomorpha scabra Jeekel, 1964</taxonomicName>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Orthomorpha scabra Jeekel, 1964, ♂ from Kon Ka Kinh National Park A habitus and live colouration B, C anterior part of body, lateral and dorsal views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure6" httpUri="https://binary.pensoft.net/fig/363861" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure6">Figs 6</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Orthomorpha scabra Jeekel, 1964, left gonopod A, B mesal and lateral views, respectively. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.898.39265.figure7" httpUri="https://binary.pensoft.net/fig/363862" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure7">7</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Orthomorpha scabra Jeekel, 1964, left gonopod A, B lateral and mesal views, respectively C-F distal part, sublateral, subcaudal, caudal and oral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure8" httpUri="https://binary.pensoft.net/fig/363863" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure8">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="reference_group">
<paragraph pageId="0" pageNumber="121">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus granosus" order="Polydesmida" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="granosus">Pratinus granosus</taxonomicName>
Attems, 1953: 166 (D).
</paragraph>
<paragraph pageId="0" pageNumber="121">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha granosa" order="Polydesmida" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="granosa">Orthomorpha granosa</taxonomicName>
-
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="121" pagination="205 - 283" refId="B19" refString="Jeekel, CAW, 1963. Paradoxosomatidae from Borneo (Diplopoda, Polydesmida). Tijdschrift voor Entomologie 106: 205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
: 265 (M).
</paragraph>
<paragraph pageId="0" pageNumber="121">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha scabra" order="Polydesmida" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="scabra">Orthomorpha scabra</taxonomicName>
Jeekel, 1964: 361 (N, D). Renamed by
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="121" pagination="355 - 364" refId="B20" refString="Jeekel, CAW, 1964. A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda). Tijdschrift voor Entomologie 107: 355 - 364" title="A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda)." volume="107" year="1964">Jeekel (1964)</bibRefCitation>
to avoid homonymy.
</paragraph>
<paragraph pageId="0" pageNumber="121">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha scabra" order="Polydesmida" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="scabra">Orthomorpha scabra</taxonomicName>
-
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift (PhD Thesis, privately published), Rotterdam" pageId="0" pageNumber="121" refId="B21" refString="Jeekel, CAW, 1968. On the Classification and Geographical Distribution of the Family Paradoxosomatidae (Diplopoda, Polydesmida). Academisch Proefschrift (PhD Thesis, privately published), Rotterdam" title="On the Classification and Geographical Distribution of the Family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">Jeekel 1968</bibRefCitation>
: 56 (M);
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.91419" author="Hoffman, RL" journalOrPublisher="Revue suisse de Zoologie" pageId="0" pageNumber="121" pagination="699 - 719" refId="B17" refString="Hoffman, RL, 1977. Diplopoda from Malayan caves, collected by M. Pierre Strinati. Revue suisse de Zoologie 84: 699 - 719" title="Diplopoda from Malayan caves, collected by M. Pierre Strinati." url="https://doi.org/10.5962/bhl.part.91419" volume="84" year="1977">Hoffman 1977</bibRefCitation>
: 700 (M);
<bibRefCitation author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="121" refId="B13" refString="Golovatch, SI, 1998. On several new or poorly-known Oriental Paradoxosomatidae (DiplopodaPolydesmida), VI. Arthropoda Selecta 6(3-4): 35-46 [for 1997]." title="On several new or poorly-known Oriental Paradoxosomatidae (DiplopodaPolydesmida), VI. Arthropoda Selecta 6 (3 - 4): 35 - 46 [for 1997]." year="1998">Golovatch 1998</bibRefCitation>
: 42 (M, D);
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.131.1921" author="Likhitrakarn, N" journalOrPublisher="ZooKeys" pageId="0" pageNumber="121" pagination="1 - 161" refId="B23" refString="Likhitrakarn, N, Golovatch, SI, Panha, S, 2011. Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 131: 1 - 161" title="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)." url="https://doi.org/10.3897/zookeys.131.1921" volume="131" year="2011">Likhitrakarn et al. 2011</bibRefCitation>
: 58 (D).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="old records">
<paragraph pageId="0" pageNumber="121">Old records.</paragraph>
<paragraph pageId="0" pageNumber="121">
Vietnam, Lam Dong Province, Peak Lang Biang; Laos, Luang Prabang Province, Luang Prabang; Xiang Khouang Province, Xieng Khouang (
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, serie A, Zoologie" pageId="0" pageNumber="121" pagination="133 - 230" refId="B5" refString="Attems, C, 1953. Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938-1939). Memoires du Museum national d'Histoire naturelle, serie A, Zoologie 5: 133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">Attems 1953</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="materials_examined">
<paragraph pageId="0" pageNumber="121">New material examined.</paragraph>
<paragraph pageId="0" pageNumber="121">
1 ♂ (ZMUM Rd 4202), Vietnam, Gia Lai Province, Kon Ka Kinh National Park, forest around headquarters,
<geoCoordinate degrees="14" direction="north" minutes="13" orientation="latitude" precision="15" seconds="12" value="14.219999">14°13'12&quot;N</geoCoordinate>
,
<geoCoordinate degrees="108" direction="east" minutes="19" orientation="longitude" precision="15" seconds="54" value="108.331665">108°19'54&quot;E</geoCoordinate>
, 1,400 m a.s.l., mixed tropical forest on hill slope, under log, daytime, 26.V.2017; 2 ♂ (ZMUM Rd 4213), same locality, but near the village of Krong,
<geoCoordinate degrees="14" direction="north" minutes="17" orientation="latitude" precision="15" seconds="46" value="14.296111">14°17'46&quot;N</geoCoordinate>
,
<geoCoordinate degrees="108" direction="east" minutes="26" orientation="longitude" precision="15" seconds="41" value="108.444725">108°26'41&quot;E</geoCoordinate>
, 800 m a.s.l., disturbed tropical forest on hills with bamboo, on bushes, daytime, 18.V.2017; 3 ♀ (ZMUM), same locality,
<geoCoordinate degrees="14" direction="north" minutes="18" orientation="latitude" precision="15" seconds="59" value="14.316389">14°18'59&quot;N</geoCoordinate>
,
<geoCoordinate degrees="108" direction="east" minutes="26" orientation="longitude" precision="15" seconds="26" value="108.44056">108°26'26&quot;E</geoCoordinate>
, 750 m a.s.l., mixed tropical forest on slopes, on tree trunk, daytime, 9.05.2017; 2 ♂ (ZMUM), Vietnam, Lam Dong Province, Bidoup Nui Ba National Park, forest near Giang Ly ranger station,
<geoCoordinate degrees="12" direction="north" minutes="10" orientation="latitude" precision="15" seconds="23" value="12.173056">12°10'23&quot;N</geoCoordinate>
,
<geoCoordinate degrees="108" direction="east" minutes="40" orientation="longitude" precision="15" seconds="59" value="108.68305">108°40'59&quot;E</geoCoordinate>
, 1,550 m a.sl., mixed forest on slopes, in
<taxonomicName class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium nidus" order="Polypodiales" pageId="0" pageNumber="121" phylum="Tracheophyta" rank="species" species="nidus">
<emphasis italics="true" pageId="0" pageNumber="121">Asplenium nidus</emphasis>
</taxonomicName>
, daytime, 11.I.2018; 2 ♀ (ZMUM), same locality, mixed forest,
<geoCoordinate degrees="12" direction="north" minutes="11" orientation="latitude" precision="15" seconds="59" value="12.199722">12°11'59&quot;N</geoCoordinate>
,
<geoCoordinate degrees="108" direction="east" minutes="40" orientation="longitude" precision="15" seconds="34" value="108.67611">108°40'34&quot;E</geoCoordinate>
, 1,500 m a.s.l., on log, daytime, 13.I.2018; 3 ♂, 2 ♀ (ZMUM), same locality,
<geoCoordinate degrees="12" direction="north" minutes="11" orientation="latitude" precision="15" seconds="09" value="12.185833">12°11'09&quot;N</geoCoordinate>
,
<geoCoordinate degrees="108" direction="east" minutes="40" orientation="longitude" precision="15" seconds="43" value="108.67861">108°40'43&quot;E</geoCoordinate>
, 1,430 m a.s.l., mixed forest in the river valley, on forest floor, night time, 12 &amp; 14.VI.2018, all leg. I. Semenyuk.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="descriptive notes">
<paragraph pageId="0" pageNumber="121">Descriptive notes.</paragraph>
<paragraph pageId="0" pageNumber="121">Length 26-46 mm (♂) or 21.5-42 mm (♀), width of midbody pro- and metazonae 2.4-4.0 and 3.6-5.8 mm (♂) or 3.5-4.2 and 3.7-6.2 mm (♀), respectively.</paragraph>
<paragraph pageId="0" pageNumber="121">
Colouration of live animals (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Orthomorpha scabra Jeekel, 1964, ♂ from Kon Ka Kinh National Park A habitus and live colouration B, C anterior part of body, lateral and dorsal views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure6" httpUri="https://binary.pensoft.net/fig/363861" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure6">Fig. 6A</figureCitation>
) black-brownish with contrasting dark yellow to orange paraterga and epiproct, head and antennae brownish, legs pale brownish; colouration in alcohol, after one year of preservation, faded to black-brown (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Orthomorpha scabra Jeekel, 1964, ♂ from Kon Ka Kinh National Park A habitus and live colouration B, C anterior part of body, lateral and dorsal views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F-H posterior part of body, dorsal, ventral and lateral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.898.39265.figure6" httpUri="https://binary.pensoft.net/fig/363861" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure6">
Fig. 6
<normalizedToken originalValue="BH">B-H</normalizedToken>
</figureCitation>
), paraterga and epiproct pale whitish yellow or pale brown, legs whitish to pale brown distally.
</paragraph>
<caption doi="10.3897/zookeys.898.39265.figure6" httpUri="https://binary.pensoft.net/fig/363861" pageId="0" pageNumber="121" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="121">
<emphasis bold="true" pageId="0" pageNumber="121">Figure 6.</emphasis>
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha scabra" order="Polydesmida" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="scabra">
<emphasis italics="true" pageId="0" pageNumber="121">Orthomorpha scabra</emphasis>
</taxonomicName>
Jeekel, 1964, ♂ from Kon Ka Kinh National Park
<emphasis bold="true" pageId="0" pageNumber="121">A</emphasis>
habitus and live colouration
<emphasis bold="true" pageId="0" pageNumber="121">B, C</emphasis>
anterior part of body, lateral and dorsal views, respectively
<emphasis bold="true" pageId="0" pageNumber="121">D, E</emphasis>
segments 10 and 11, dorsal and lateral views, respectively
<emphasis bold="true" pageId="0" pageNumber="121">
<normalizedToken originalValue="FH">F-H</normalizedToken>
</emphasis>
posterior part of body, dorsal, ventral and lateral views, respectively
<emphasis bold="true" pageId="0" pageNumber="121">I, J</emphasis>
sternal cones between coxae 4, subcaudal and sublateral views, respectively.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="121" type="remarks">
<paragraph pageId="0" pageNumber="121">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="121">
This species has recently been redescribed (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.131.1921" author="Likhitrakarn, N" journalOrPublisher="ZooKeys" pageId="0" pageNumber="121" pagination="1 - 161" refId="B23" refString="Likhitrakarn, N, Golovatch, SI, Panha, S, 2011. Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 131: 1 - 161" title="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)." url="https://doi.org/10.3897/zookeys.131.1921" volume="131" year="2011">Likhitrakarn et al. 2011</bibRefCitation>
), based on type material. The new specimens agree nearly fully with the available descriptions except for the ♂ tarsal brushes being present until legs 7 or 8 vs. legs 5. One variety,
<taxonomicName lsidName="grandis" pageId="0" pageNumber="121" rank="species" species="grandis">
<emphasis italics="true" pageId="0" pageNumber="121">grandis</emphasis>
</taxonomicName>
var. nov., is more disjunct morphologically, but not genetically, being treated separately below.
</paragraph>
<caption doi="10.3897/zookeys.898.39265.figure7" httpUri="https://binary.pensoft.net/fig/363862" pageId="0" pageNumber="121" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="121">
<emphasis bold="true" pageId="0" pageNumber="121">Figure 7.</emphasis>
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha scabra" order="Polydesmida" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="scabra">
<emphasis italics="true" pageId="0" pageNumber="121">Orthomorpha scabra</emphasis>
</taxonomicName>
Jeekel, 1964, left gonopod
<emphasis bold="true" pageId="0" pageNumber="121">A, B</emphasis>
mesal and lateral views, respectively. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="121">
This species seems to be widespread, occurring in Vietnam and northern Laos (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distributions of Orthomorpha species (nine species or varieties, excluding the pantropical anthropochore O. coarctata (de Saussure, 1860 )) recorded in Vietnam. Open triangle O. hydrobiologica Attems, 1930 Filled circle O. scabra Jeekel, 1964 Open circle O. scabra, grandis var. nov., O. scabra Jeekel, 1964 and O. rotundicollis (Attems, 1937) Crossed square O. rotundicollis, subrotundicollis var. nov., O. caramel sp. nov. and O. vietnamica sp. nov. Filled Inverted triangle O. glandulosa (Attems, 1937) Opened Inverted triangle O. scabra Jeekel, 1964 and O. scabra, grandis var. nov. Open diamond O. glandulosa (Attems, 1937) and O. hydrobiologica Attems, 1930 Filled diamond O. glandulosa (Attems, 1937) Open square O. arboricola (Attems, 1937) and O. rotundicollis (Attems, 1937) Filled square O. rotundicollis (Attems, 1937) and O. coarctata (de Saussure, 1860)." figureDoi="10.3897/zookeys.898.39265.figure19" httpUri="https://binary.pensoft.net/fig/363874" pageId="0" pageNumber="121" tableDoi="10.3897/zookeys.898.39265.figure19">Fig. 19</figureCitation>
), and in two places it was found coexisting with
<taxonomicName lsidName="O. rotundicollis" pageId="0" pageNumber="121" rank="species" species="rotundicollis">
<emphasis italics="true" pageId="0" pageNumber="121">O. rotundicollis</emphasis>
</taxonomicName>
(Attems, 1937) (see also above).
</paragraph>
<paragraph pageId="0" pageNumber="121">
According to
<normalizedToken originalValue="IS">IS'</normalizedToken>
field observations, the millipedes occupy a wide range of habitats, including riparian forests with flooding occurring every year and non-flooding mixed forests on slopes. They also appear in a forestless farm area, usually being found on the forest floor and at bases of tree trunks. They occupy many microhabitats in lower forest strata. The activity is mainly restricted to the night time, but some individuals keep it up in daylight as well.
</paragraph>
<caption doi="10.3897/zookeys.898.39265.figure8" httpUri="https://binary.pensoft.net/fig/363863" pageId="0" pageNumber="121" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="121">
<emphasis bold="true" pageId="0" pageNumber="121">Figure 8.</emphasis>
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha scabra" order="Polydesmida" pageId="0" pageNumber="121" phylum="Arthropoda" rank="species" species="scabra">
<emphasis italics="true" pageId="0" pageNumber="121">Orthomorpha scabra</emphasis>
</taxonomicName>
Jeekel, 1964, left gonopod
<emphasis bold="true" pageId="0" pageNumber="121">A, B</emphasis>
lateral and mesal views, respectively
<emphasis bold="true" pageId="0" pageNumber="121">
<normalizedToken originalValue="CF">C-F</normalizedToken>
</emphasis>
distal part, sublateral, subcaudal, caudal and oral views, respectively.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>