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<document id="AB7C78D323E991BCCE14D1E0C76D28E9" ID-DOI="10.21248/contrib.entomol.67.2.275-318" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742497" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638268338673" checkinUser="felipe" docAuthor="Irmler, Ulrich" docDate="2017" docId="03F6C753FFE1FFA6FC83DC66FE1FF824" docLanguage="en" docName="ContribEntomol.67.2.275-318.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Eleusis boliviana Irmler 2017, spec. nov." docType="treatment" docUuid="63DF628A-E08F-4D9E-9518-B5C00599B8D8" docUuidSource="ZooBank" docVersion="4" lastPageNumber="280" masterDocId="FFCFBF2BFFE5FFA3FFFFD92CFFDAFF9C" masterDocTitle="The Neotropical species of the genus Eleusis LAPORTE, 1835 with description of the new genus Eleusima (Coleoptera: Staphylinidae: Osoriinae)" masterLastPageNumber="318" masterPageNumber="275" pageNumber="279" updateTime="1699437037312" updateUser="plazi">
<mods:mods id="2AB0F04631E2B4B885D01D319B448EE8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="ED6C3B1CC5691CCE384ACA7BF2AB4EEE">
<mods:title id="E105A01CA6CEF45C73572BBF88EDB0DA">The Neotropical species of the genus Eleusis LAPORTE, 1835 with description of the new genus Eleusima (Coleoptera: Staphylinidae: Osoriinae)</mods:title>
</mods:titleInfo>
<mods:name id="F556DAE360B979282DB36907E8FDDA62" type="personal">
<mods:role id="7FF11EE3C3664F3EF5D75E368CDA25A0">
<mods:roleTerm id="38DDC92536A7A41820B1977373695825">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3E1934E4F218F6B5727E7144CFDBDF4">Irmler, Ulrich</mods:namePart>
</mods:name>
<mods:typeOfResource id="555E97B282AEF5FBDF79C884C56A097D">text</mods:typeOfResource>
<mods:relatedItem id="9FFED793BFE2962BC1F2F8D59EF730DD" type="host">
<mods:titleInfo id="DEAA3DEB5D107129B2844B04C57C7FDF">
<mods:title id="3CA14A554B6076E47B32C3203D7BBDA3">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="93A2A8F5BAC236387D6770D6D71A93D6">
<mods:date id="3669FD6F63D655F4C29F570DD92F5CE5">2017</mods:date>
<mods:detail id="440B10527D1BC6AEFF08EBA46CCD1D99" type="volume">
<mods:number id="2EF15F8AF8F1F080DAF35F756E18B5F4">67</mods:number>
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<mods:detail id="F86FCBBA0A479B29D23FEC206B30445A" type="issue">
<mods:number id="0036BD8A1604F231FB1FDA5F068A4D4F">2</mods:number>
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<mods:start id="A39038CEB6807DF1295A407F5494A166">275</mods:start>
<mods:end id="31CEF3510DF1FB37B2EC718700457336">318</mods:end>
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<mods:classification id="1D5F0F48E9B68E8B411BC7C4CAFC2A85">journal article</mods:classification>
<mods:identifier id="FD16CBA9F3A8473820C15448F83AC01B" type="DOI">10.21248/contrib.entomol.67.2.275-318</mods:identifier>
<mods:identifier id="DE5252619ED01450A4F1AE011EB372F5" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="07C9556C80E69C9374E258ECA89AC11A" type="Zenodo-Dep">5742497</mods:identifier>
<mods:identifier id="4B5569BBB21BF4A45D46B8A659D7BC54" type="ZooBank">98BC4B71-EE31-4BA0-855A-AAC48C955B38</mods:identifier>
</mods:mods>
<treatment id="03F6C753FFE1FFA6FC83DC66FE1FF824" LSID="urn:lsid:zoobank.org:act:63DF628A-E08F-4D9E-9518-B5C00599B8D8" httpUri="http://treatment.plazi.org/id/03F6C753FFE1FFA6FC83DC66FE1FF824" lastPageId="5" lastPageNumber="280" pageId="4" pageNumber="279">
<subSubSection id="C34525CEFFE1FFA7FC83DC66FA59FA1A" pageId="4" pageNumber="279" type="nomenclature">
<paragraph id="8BE07645FFE1FFA7FC83DC66FB4CFAF9" blockId="4.[892,1411,1354,1446]" box="[892,1174,1354,1381]" pageId="4" pageNumber="279">
<heading id="D0A8C129FFE1FFA7FC83DC66FB4CFAF9" box="[892,1174,1354,1381]" fontSize="12" level="2" pageId="4" pageNumber="279" reason="2">
<taxonomicName id="4C5F0DC6FFE1FFA7FC83DC66FBFCFAF9" ID-CoL="398M6" authority="Irmler, 2017" authorityName="Irmler" authorityYear="2017" box="[892,1062,1354,1381]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="boliviana" status="sp. nov.">
<emphasis id="B92BAA57FFE1FFA7FC83DC66FBFCFAF9" box="[892,1062,1354,1381]" italics="true" pageId="4" pageNumber="279">Eleusis boliviana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218172CFFE1FFA7FBD3DC66FB4CFAF9" box="[1068,1174,1354,1381]" pageId="4" pageNumber="279" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BE07645FFE1FFA7FC83DC5EFA59FA1A" blockId="4.[892,1411,1354,1446]" box="[892,1411,1394,1414]" pageId="4" pageNumber="279">
<uri id="FFCE7A47FFE1FFA7FC83DC5EFA59FA1A" box="[892,1411,1394,1414]" pageId="4" pageNumber="279">
urn:lsid:zoobank. org:act:
<uuid id="FFF94C90FFE1FFA7FBD3DC5EFA59FA1A" box="[1068,1411,1394,1414]" pageId="4" pageNumber="279">63DF628A-E08F-4D9E-9518-B5C00599B8D8</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFE1FFA7FC83DCBEFC31FA3A" box="[892,1003,1426,1446]" pageId="4" pageNumber="279" type="description">
<paragraph id="8BE07645FFE1FFA7FC83DCBEFC31FA3A" blockId="4.[892,1411,1354,1446]" box="[892,1003,1426,1446]" pageId="4" pageNumber="279">
<figureCitation id="13646AC0FFE1FFA7FC83DCBEFC72FA3A" box="[892,936,1426,1446]" captionStart="Fig" captionStartId="31.[174,207,2052,2074]" captionTargetBox="[224,1371,498,2021]" captionTargetId="figure-49@31.[205,1390,491,2049]" captionTargetPageId="31" captionText="Fig. 2–6: Eleusis interrupta (2), E.humilis (3), E. nigerrima (4), E. boliviana (5), E. andina (6); aedeagus in lateral (A) and ventral (B)" figureDoi="http://doi.org/10.5281/zenodo.5742501" httpUri="https://zenodo.org/record/5742501/files/figure.png" pageId="4" pageNumber="279">Figs 5</figureCitation>
A-C,28C
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFE1FFA7FCA9DCECFC7DF9E4" pageId="4" pageNumber="279" type="materials_examined">
<paragraph id="8BE07645FFE1FFA7FCA9DCECFC7DF9E4" blockId="4.[854,1459,1471,1656]" pageId="4" pageNumber="279">
<emphasis id="B92BAA57FFE1FFA7FCA9DCECFC2EFA4B" bold="true" box="[854,1012,1471,1496]" pageId="4" pageNumber="279">Type material</emphasis>
:
<materialsCitation id="3B377C18FFE1FFA7FBF6DCECFA74F9A4" accessionNumber="G99027" collectingDate="1999-02-06" collectionCode="KNHM" collectorName="F. Genier" country="Bolivia" latitude="-17.141666" location="Villa Tunari Rd." longLatPrecision="13" longitude="-65.704834" pageId="4" pageNumber="279" specimenCount="1" stateProvince="Cochabamba" typeStatus="holotype">
male,
<typeStatus id="54E4C8E7FFE1FFA7FBAFDCECFB6CFA44" box="[1104,1206,1472,1496]" pageId="4" pageNumber="279" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34836D5FFE1FFA7FB38DCECFACFFA44" box="[1223,1301,1472,1496]" name="Bolivia" pageId="4" pageNumber="279">Bolivia</collectingCountry>
:
<collectingRegion id="499BB8A7FFE1FFA7FAD9DCECFC55FA6B" country="Bolivia" name="Cochabamba" pageId="4" pageNumber="279">Cochabamba Prov.</collectingRegion>
,
<collectingRegion id="499BB8A7FFE1FFA7FC5FDCCCFBF5FA64" box="[928,1071,1504,1528]" country="Bolivia" name="Cochabamba" pageId="4" pageNumber="279">Cochabamba</collectingRegion>
,
<locationDeviation id="9A526F67FFE1FFA7FBBFDCCCFADCFA64" box="[1088,1286,1504,1528]" location="Yungas" pageId="4" pageNumber="279">
<quantity id="4CA7DBA0FFE1FFA7FBBFDCCCFB55FA64" box="[1088,1167,1504,1528]" metricMagnitude="5" metricUnit="m" metricValue="1.09" pageId="4" pageNumber="279" unit="km" value="109.0">109 km</quantity>
E Yungas
</locationDeviation>
(Cochabamba-
<location id="8E80209EFFE1FFA7FCA9DF2CFBD9F984" LSID="urn:lsid:plazi:treatment:03F6C753FFE1FFA6FC83DC66FE1FF824:8E80209EFFE1FFA7FCA9DF2CFBD9F984" box="[854,1027,1536,1560]" country="Bolivia" latitude="-17.141666" longLatPrecision="13" longitude="-65.704834" name="Villa Tunari Rd." pageId="4" pageNumber="279" stateProvince="Cochabamba">Villa Tunari Rd.</location>
) (
<geoCoordinate id="EE6B1082FFE1FFA7FBE8DF2CFBA1F984" box="[1047,1147,1536,1560]" degrees="17" direction="south" minutes="8.50" orientation="latitude" pageId="4" pageNumber="279" precision="9" value="-17.141666">17°8.50'S</geoCoordinate>
,
<geoCoordinate id="EE6B1082FFE1FFA7FB79DF2CFADEF984" box="[1158,1284,1536,1560]" degrees="65" direction="west" minutes="42.29" orientation="longitude" pageId="4" pageNumber="279" precision="9" value="-65.704834">65°42.29'W</geoCoordinate>
), flight
<collectingMethod id="521E0E52FFE1FFA7FAABDF2CFA69F984" box="[1364,1459,1536,1560]" pageId="4" pageNumber="279">intercept</collectingMethod>
tap, 1.-
<date id="FFE15085FFE1FFA7FC67DF0CFC27F9A4" box="[920,1021,1568,1592]" pageId="4" pageNumber="279" value="1999-02-06">
<collectingDate id="EFA5A96DFFE1FFA7FC67DF0CFC27F9A4" box="[920,1021,1568,1592]" pageId="4" pageNumber="279" value="1999-02-06">6.2.1999</collectingDate>
</date>
, leg.
<collectorName id="26AA1393FFE1FFA7FBC9DF0CFB4DF9A4" box="[1078,1175,1568,1592]" pageId="4" pageNumber="279">F. Genier</collectorName>
, #BOL1G99027 (
<collectionCode id="ED4EEE80FFE1FFA7FAAFDF0CFA7DF9A4" box="[1360,1447,1568,1592]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="4" pageNumber="279">KNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B377C18FFE1FFA7FCA9DF6CFC79F9E4" accessionNumber="G99027" collectingDate="1999-02-06" collectionCode="KNHM, UIC" collectorName="F. Genier" country="Bolivia" latitude="-17.141666" location="Villa Tunari Rd." longLatPrecision="13" longitude="-65.704834" pageId="4" pageNumber="279" specimenCount="3" specimenCount-male="3" stateProvince="Cochabamba" typeStatus="paratype">
<typeStatus id="54E4C8E7FFE1FFA7FCA9DF6CFC1AF9C4" box="[854,960,1600,1624]" pageId="4" pageNumber="279" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D59BDCCFFE1FFA7FC37DF6CFBC2F9C4" box="[968,1048,1600,1624]" pageId="4" pageNumber="279" type="male">3 males</specimenCount>
with same data as holotype (2
<collectionCode id="ED4EEE80FFE1FFA7FAA4DF6CFA75F9CB" box="[1371,1455,1600,1623]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="4" pageNumber="279">KNHM</collectionCode>
, 1
<collectionCode id="ED4EEE80FFE1FFA7FC96DF4CFC41F9E4" box="[873,923,1632,1656]" pageId="4" pageNumber="279">UIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFE1FFA7FCA9DFB3FA6EF824" pageId="4" pageNumber="279" type="diagnosis">
<paragraph id="8BE07645FFE1FFA7FCA9DFB3FA6EF824" blockId="4.[854,1460,1695,1976]" pageId="4" pageNumber="279">
<emphasis id="B92BAA57FFE1FFA7FCA9DFB3FC19F92B" bold="true" box="[854,963,1695,1719]" pageId="4" pageNumber="279">Diagnosis</emphasis>
: Regarding the small size,
<taxonomicName id="4C5F0DC6FFE1FFA7FB09DF8CFAA8F924" authorityName="Irmler" authorityYear="2017" box="[1270,1394,1696,1720]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="boliviana">
<emphasis id="B92BAA57FFE1FFA7FB09DF8CFAA8F924" box="[1270,1394,1696,1720]" italics="true" pageId="4" pageNumber="279">E. boliviana</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4C5F0DC6FFE1FFA7FBD6DFECFB53F944" authorityName="Irmler" authorityYear="2017" box="[1065,1161,1728,1752]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="fauveli">
<emphasis id="B92BAA57FFE1FFA7FBD6DFECFB53F944" box="[1065,1161,1728,1752]" italics="true" pageId="4" pageNumber="279">E. fauveli</emphasis>
</taxonomicName>
.
<taxonomicName id="4C5F0DC6FFE1FFA7FB5FDFECFAC6F944" authorityName="Irmler" authorityYear="2017" box="[1184,1308,1728,1752]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="boliviana">
<emphasis id="B92BAA57FFE1FFA7FB5FDFECFAC6F944" box="[1184,1308,1728,1752]" italics="true" pageId="4" pageNumber="279">E. boliviana</emphasis>
</taxonomicName>
is distinctly darker than
<taxonomicName id="4C5F0DC6FFE1FFA7FC1DDFCCFB98F964" authorityName="Irmler" authorityYear="2017" box="[994,1090,1760,1784]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="fauveli">
<emphasis id="B92BAA57FFE1FFA7FC1DDFCCFB98F964" box="[994,1090,1760,1784]" italics="true" pageId="4" pageNumber="279">E. fauveli</emphasis>
</taxonomicName>
and in this respect, it resembles
<taxonomicName id="4C5F0DC6FFE1FFA7FCA9DE2CFC02F88B" authorityName="FAUVEL" authorityYear="1864" box="[854,984,1792,1815]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="nigerrima">
<emphasis id="B92BAA57FFE1FFA7FCA9DE2CFC02F88B" box="[854,984,1792,1815]" italics="true" pageId="4" pageNumber="279">E. nigerrima</emphasis>
</taxonomicName>
. In contrast to
<taxonomicName id="4C5F0DC6FFE1FFA7FB72DE2CFB37F884" authorityName="Irmler" authorityYear="2017" box="[1165,1261,1792,1816]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="fauveli">
<emphasis id="B92BAA57FFE1FFA7FB72DE2CFB37F884" box="[1165,1261,1792,1816]" italics="true" pageId="4" pageNumber="279">E. fauveli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F0DC6FFE1FFA7FAD5DE2CFA76F88B" authorityName="FAUVEL" authorityYear="1864" box="[1322,1452,1792,1815]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="nigerrima">
<emphasis id="B92BAA57FFE1FFA7FAD5DE2CFA76F88B" box="[1322,1452,1792,1815]" italics="true" pageId="4" pageNumber="279">E. nigerrima</emphasis>
</taxonomicName>
, the elytra are longer than wide and not quadrate or even wider than long. However, without dissection of the aedeagus, the separation from the other related species of the group is difficult. The long elongate prominence of the central aedeagal lobe clearly characterises the species.
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFE1FFA6FCA9DEF3FE3EF8C4" lastPageId="5" lastPageNumber="280" pageId="4" pageNumber="279" type="description">
<paragraph id="8BE07645FFE1FFA7FCA9DEF3FA80F7A4" blockId="4.[854,1459,2015,2104]" pageId="4" pageNumber="279">
<emphasis id="B92BAA57FFE1FFA7FCA9DEF3FC02F86B" bold="true" box="[854,984,2015,2039]" pageId="4" pageNumber="279">Description</emphasis>
: Length:
<quantity id="4CA7DBA0FFE1FFA7FBBDDECCFB4CF86B" box="[1090,1174,2016,2040]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="279" unit="mm" value="2.4">2.4 mm</quantity>
. Colouration: dark brown to blackish; elytra lighter brown with blackish posterior margin; legs and antennae light brown to yellow.
</paragraph>
<paragraph id="8BE07645FFE0FFA6FF51D82CFE4BFD04" blockId="5.[174,779,256,1880]" pageId="5" pageNumber="280">
Head:
<quantity id="4CA7DBA0FFE0FFA6FF09D82CFE8FFE8B" box="[246,341,256,280]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="280" unit="mm" value="0.31">0.31 mm</quantity>
long,
<quantity id="4CA7DBA0FFE0FFA6FE62D82CFE21FE8B" box="[413,507,256,280]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="5" pageNumber="280" unit="mm" value="0.36">0.36 mm</quantity>
wide; eyes large, prominent; nearly twice as long as temples; temples smoothly narrowed to neck; preocular sides shortly parallel to base of antennae; anterior margin of clypeus sinuate; apex widely curved; supraocular groove reaching from anterior edge of eyes to neck; punctation moderately deep and irregularly sparse; interstices between punctures two to four times as wide as diameter of punctures; at each end of supraocular groove with large setiferous puncture; long setae yellow; microsculpture irregularly deep; on vertex deeper than on clypeus; striae elongate; on clypeus transverse; on vertex longitudinal; close to neck diagonal; surface slightly shiny.
</paragraph>
<paragraph id="8BE07645FFE0FFA6FF51DB8CFD5BFC0B" blockId="5.[174,779,256,1880]" pageId="5" pageNumber="280">Antennae slightly longer than head and pronotum combined; first antennomere thicker and longer than following antennomeres; second antennomere two third as long as first; third slightly shorter than second; following antennomeres increasing in width and decreasing in length; fourth antennomere slightly longer than wide; tenth antennomere quadrate; antennomeres four to eleven pubescent and with long apical setae.</paragraph>
<paragraph id="8BE07645FFE0FFA6FF51DA8CFE99FAC4" blockId="5.[174,779,256,1880]" pageId="5" pageNumber="280">
Pronotum:
<quantity id="4CA7DBA0FFE0FFA6FECCDA8CFE49FC2B" box="[307,403,928,952]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="280" unit="mm" value="0.28">0.28 mm</quantity>
long,
<quantity id="4CA7DBA0FFE0FFA6FE17DA8CFD92FC2B" box="[488,584,928,952]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="280" unit="mm" value="0.34">0.34 mm</quantity>
wide; widest at anterior angles; anterior angles smoothly curved; sides in anterior half nearly parallel; in posterior half greatly narrowed; posterior margin half as wide as at anterior angles; punctation similarly dense and deep as on vertex; more uniformly punctate than head; close to anterior angles with three larger setiferous punctures; long setae yellow; along lateral margin with few more setiferous punctures; lateral margin interrupted at middle; interruption marked by deep setiferous puncture; microsculpture similarly deep as on vertex; meshes elongate; on disc longitudinal; close to posterior angles diagonal; close to anterior angles nearly circular; surface slightly shiny.
</paragraph>
<paragraph id="8BE07645FFE0FFA6FF51DC4CFE0EF984" blockId="5.[174,779,256,1880]" pageId="5" pageNumber="280">
Elytra:
<quantity id="4CA7DBA0FFE0FFA6FEFEDC4CFEBAFAEB" box="[257,352,1376,1400]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="280" unit="mm" value="0.46">0.46 mm</quantity>
long,
<quantity id="4CA7DBA0FFE0FFA6FE50DC4CFDD4FAEB" box="[431,526,1376,1400]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="280" unit="mm" value="0.44">0.44 mm</quantity>
wide; widest close to posterior angles; humeral angles smoothly curved; punctation weaker than on pronotum, but as dense as on pronotum; one large setiferous puncture on central disc; microsculpture as deep as on pronotum; meshes longitudinal; surface slightly shiny.
</paragraph>
<paragraph id="8BE07645FFE0FFA6FF51DF0CFD94F9CB" blockId="5.[174,779,256,1880]" pageId="5" pageNumber="280">Abdomen densely punctate and pubescent; among short hairs several punctures with long setae.</paragraph>
<paragraph id="8BE07645FFE0FFA6FF51DF4CFE3EF8C4" blockId="5.[174,779,256,1880]" pageId="5" pageNumber="280">Aedeagus moderately broad; close to apex slightly broader than at base; apex moderately acute; with long central process reaching apex of central lobe; at base of process with several sensillae; parameres slightly shorter than central lobe; with moderately wide apical transparent lobe; at apex of transparent lobe with numerous short setae; one long seta at outer central position; at base of parameres with few sensillae.</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFE0FFA6FF51DEACFE1FF824" pageId="5" pageNumber="280" type="etymology">
<paragraph id="8BE07645FFE0FFA6FF51DEACFE1FF824" blockId="5.[174,778,1920,1976]" pageId="5" pageNumber="280">
<emphasis id="B92BAA57FFE0FFA6FF51DEACFEFFF804" bold="true" box="[174,293,1920,1944]" pageId="5" pageNumber="280">Etymology</emphasis>
: The species name derived from the country’s name, where it was found.
</paragraph>
</subSubSection>
</treatment>
</document>
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