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<document id="4513ABB77AF67E9BD264CBF8AD63ABDE" ID-DOI="10.5281/zenodo.178756" ID-GBIF-Dataset="f9ea8775-9b2d-4364-b20a-4738aab6a54c" ID-ISSN="1175-5326" ID-Zenodo-Dep="178756" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459920987951" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2007" docId="03E987D1FFBEFFF9FF178C6F34BFFC51" docLanguage="en" docName="zt01604p020.pdf" docOrigin="Zootaxa 1604" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Harpalus kaznakovi subsp. kaznakovi Kataev &amp; Wrase 1997" docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="FFD0FFA9FFB5FFF4FF8088153517FFD3" masterDocTitle="Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="12" updateTime="1698229235843" updateUser="plazi">
<mods:mods id="99020E56FA031ACEEE3E6888CF2D685E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7A309FBB45169CF95315F7F02F3AA1A0">
<mods:title id="C96DA3C6DC80DB8CA74FF37A0515FF90">Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini)</mods:title>
</mods:titleInfo>
<mods:name id="9ECF6A0D7410E646333B8163FAC47B0D" type="personal">
<mods:role id="20295A1D3EF17CC085C07605036445B2">
<mods:roleTerm id="6BAE0CB1DF24EB244A166D6CA147D9C4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="99CC1FD542386684293F3A8E95E93FB9">Kataev, Boris M.</mods:namePart>
</mods:name>
<mods:name id="62A46462803B0C736F3D9D8435430528" type="personal">
<mods:role id="64B1BB52C503BBA971402E927D425727">
<mods:roleTerm id="637579C7673515FE5F44AFD5A16F3041">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="165572DA93C01DF040D583DCF21290A7">Liang, Hongbin</mods:namePart>
</mods:name>
<mods:typeOfResource id="48ECD6745FAD0C7F59FCDB93E55C4545">text</mods:typeOfResource>
<mods:relatedItem id="1F7AECDAA5722E241A8C4A3B905DD8C7" type="host">
<mods:titleInfo id="4B4C56ACCE2FD010073FD2560CD3E7CE">
<mods:title id="DC26E16C8A85891BC1702AC4202137D8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="251D30E989E673F66F56EF7AF6E79EA1">
<mods:date id="DC717617729642964C11AD8FF021FDF1">2007</mods:date>
<mods:detail id="37402C25B7B96CDE13EE3E8441B6562D" type="volume">
<mods:number id="43305CE76E271F0C56AAB302B7E1381A">1604</mods:number>
</mods:detail>
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<mods:start id="57660FB3D1500D524155099535DAFF95">1</mods:start>
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</mods:extent>
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<mods:classification id="A598C0B9DC61A12AA7564923FEE27450">journal article</mods:classification>
<mods:identifier id="FF40F6A8032CBA6B4B10AB15543B3BDC" type="DOI">10.5281/zenodo.178756</mods:identifier>
<mods:identifier id="4DE9D9750F758FB97C755CBC33C17378" type="GBIF-Dataset">f9ea8775-9b2d-4364-b20a-4738aab6a54c</mods:identifier>
<mods:identifier id="88C6E0BC67D83CF13CBAD472635971A7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5C0F20B9CDFB6EDA7D7663F712035B31" type="Zenodo-Dep">178756</mods:identifier>
</mods:mods>
<treatment id="03E987D1FFBEFFF9FF178C6F34BFFC51" ID-DOI="http://doi.org/10.5281/zenodo.6250242" ID-GBIF-Taxon="119373714" ID-Zenodo-Dep="6250242" LSID="urn:lsid:plazi:treatment:03E987D1FFBEFFF9FF178C6F34BFFC51" httpUri="http://treatment.plazi.org/id/03E987D1FFBEFFF9FF178C6F34BFFC51" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
<subSubSection id="C35A654CFFBEFFFFFF178C6F3492FB64" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BFF36C7FFBEFFFFFF178C6F3639FB47" blockId="11.[151,814,1146,1207]" box="[151,814,1146,1172]" pageId="11" pageNumber="12">
<heading id="D0B781ABFFBEFFFFFF178C6F3639FB47" bold="true" box="[151,814,1146,1172]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="4C404D44FFBEFFFFFF178C6F3639FB47" ID-CoL="8MBXS" authority="Kataev &amp; Wrase, 1997" authorityName="Kataev &amp; Wrase" authorityYear="1997" box="[151,814,1146,1172]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subSpecies" species="kaznakovi" subSpecies="kaznakovi">
<emphasis id="B934EAD5FFBEFFFFFF178C6F3639FB47" bold="true" box="[151,814,1146,1172]" pageId="11" pageNumber="12">
<emphasis id="B934EAD5FFBEFFFFFF178C6F371BFB47" bold="true" box="[151,524,1146,1172]" italics="true" pageId="11" pageNumber="12">Harpalus kaznakovi kaznakovi</emphasis>
<bibRefCitation id="EFD14B36FFBEFFFFFD938C6F3639FB47" author="Kataev" box="[531,814,1146,1172]" pageId="11" pageNumber="20" refString="Kataev, B. M., Wrase, D. W. (1997) New taxa of the genus Harpalus Latr. from China and Turkey (Coleoptera, Carabidae). Linzer biologische Beitrage, 29 (2), 991 - 1014." type="journal article" year="1997">Kataev &amp; Wrase, 1997</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFF36C7FFBEFFFFFF178C883492FB64" blockId="11.[151,814,1146,1207]" box="[151,389,1181,1207]" pageId="11" pageNumber="12">
(
<figureCitation id="137B2A42FFBEFFFFFF1F8C883431FB64" box="[159,294,1181,1207]" captionStart="FIGURES 25 – 29" captionStartId="12.[151,269,1777,1801]" captionTargetBox="[153,1389,532,1776]" captionTargetId="figure@12.[151,1436,532,1843]" captionTargetPageId="12" captionText="FIGURES 25 – 29. Harpalus kaznakovi. 25 – 27, H. k. kaznakovi (25, 27, Sichuan, Garzê Co.; 26, Sichuan, Dêgê Co.); 28 – 29, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 25 – 26, 28, habitus, 27 – 29, left metepisternum. Scales = 1.0 mm (A: Figs 25 – 26, 28; B: 27, 29)." httpUri="https://zenodo.org/record/178761/files/figure.png" pageId="11" pageNumber="12">Figs 25–27</figureCitation>
,
<figureCitation id="137B2A42FFBEFFFFFEB28C883469FB64" box="[306,382,1181,1207]" captionStart="FIGURES 30 – 44" captionStartId="14.[151,269,1898,1922]" captionTargetBox="[186,1385,219,1897]" captionTargetId="figure@14.[151,1436,212,1938]" captionTargetPageId="14" captionText="FIGURES 30 – 44. Harpalus kaznakovi, median lobe of aedeagus (32 – 33, 38, 41, 44, terminal lamella). 30 – 41, H. k. kaznakovi (30 – 38, Sichuan, Garzê Co.; 39 – 41, Sichuan, Dêgê Co.). 42 – 44, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 30, 32 – 34, 36, 38 – 39, 41 – 42, 44, dorsal aspect; 31, 35, 37, 40, 43, lateral aspect. Scales: A = 0.5 mm (Figs 32 – 33, 38, 41, 44), B = 1.0 mm (Figs 30 – 31, 34 – 37, 39 – 40, 42 – 43)." httpUri="https://zenodo.org/record/178762/files/figure.png" pageId="11" pageNumber="12">30–41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFBEFFFFFF178CF83466FAF7" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="8BFF36C7FFBEFFFFFF178CF83466FAF7" blockId="11.[151,1435,1261,1316]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB5011E9FFBEFFFFFF178CF83466FAF7" pageId="11" pageNumber="12">
<taxonomicName id="4C404D44FFBEFFFFFF178CF837ACFAD7" ID-CoL="3JSM9" authority="Kataev &amp; Wrase, 1997: 996" authorityName="Kataev &amp; Wrase" authorityPageNumber="996" authorityYear="1997" box="[151,699,1261,1284]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kaznakovi">
<emphasis id="B934EAD5FFBEFFFFFF178CF83465FAD7" box="[151,370,1261,1284]" italics="true" pageId="11" pageNumber="12">Harpalus kaznakovi</emphasis>
<treatmentCitation id="0AE110D6FFBEFFFFFEFC8CF837ACFAD7" author="Kataev" box="[380,699,1261,1284]" page="996" pageId="11" pageNumber="12" year="1997">
<bibRefCitation id="EFD14B36FFBEFFFFFEFC8CF83796FAD7" author="Kataev" box="[380,641,1261,1284]" pageId="11" pageNumber="20" refString="Kataev, B. M., Wrase, D. W. (1997) New taxa of the genus Harpalus Latr. from China and Turkey (Coleoptera, Carabidae). Linzer biologische Beitrage, 29 (2), 991 - 1014." type="journal article" year="1997">Kataev &amp; Wrase, 1997</bibRefCitation>
: 996
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54FB8865FFBEFFFFFD4A8CF837E8FAD7" box="[714,767,1261,1284]" pageId="11" pageNumber="12">Type</typeStatus>
locality: road Luhuo–Sertar, pass
<quantity id="4CB89B22FFBEFFFFFB038CF831DCFAD7" box="[1155,1227,1261,1284]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="km" value="35.0">35 km</quantity>
NNE Luhuo, NW Sichuan,
<collectingCountry id="F3577657FFBEFFFFFEAA8D18347AFAF7" box="[298,365,1293,1316]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFBEFFFFFF178D403176F8A4" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BFF36C7FFBEFFFFFF178D4031D0FABC" blockId="11.[151,1436,1365,2031]" box="[151,1223,1365,1391]" pageId="11" pageNumber="12">
<emphasis id="B934EAD5FFBEFFFFFF178D403494FABC" bold="true" box="[151,387,1365,1391]" pageId="11" pageNumber="12">Material examined</emphasis>
. In addition to the
<typeStatus id="54FB8865FFBEFFFFFDE38D403781FABC" box="[611,662,1365,1391]" pageId="11" pageNumber="12">type</typeStatus>
series the following specimens were examined:
</paragraph>
<paragraph id="8BFF36C7FFBEFFFFFF478D683176F8A4" blockId="11.[151,1436,1365,2031]" pageId="11" pageNumber="12">
<collectingCountry id="F3577657FFBEFFFFFF478D683406FA44" box="[199,273,1405,1431]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
. SICHUAN: 1 ɗ, same data as
<typeStatus id="54FB8865FFBEFFFFFD258D683607FA44" box="[677,784,1405,1431]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
: NW Sichuan, road Luhuo–Sertar, pass
<quantity id="4CB89B22FFBEFFFFFA848D683044FA44" box="[1284,1363,1405,1431]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="km" value="35.0">35 km</quantity>
NNE Luhuo, alpine region,
<quantity id="4CB89B22FFBEFFFFFE268DB03758FA6C" box="[422,591,1445,1471]" metricMagnitude="3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="11" pageNumber="12" unit="m" value="3750.0" valueMax="4000.0" valueMin="3500.0">3500–4000 m</quantity>
,
<date id="FFFE1007FFBEFFFFFDDE8DB037E6FA6C" box="[606,753,1445,1471]" pageId="11" pageNumber="12" value="1994-07-27">27.VII.1994</date>
, L. Kalab leg. (cFCCH); 2 ɗ, 2 Ψ, Garzê Co.,
<quantity id="4CB89B22FFBEFFFFFAB88DB03080FA6C" box="[1336,1431,1445,1471]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="11" pageNumber="12" unit="m" value="3400.0">3400 m</quantity>
,
<date id="FFFE1007FFBEFFFFFF178DD83408FA34" box="[151,287,1485,1511]" pageId="11" pageNumber="12" value="1983-06-29">29.VI.1983</date>
, Chen Yuanging &amp; Zhang Xuezhong leg. (
<collectionCode id="ED51AE02FFBEFFFFFC948DD8365FFA34" box="[788,840,1485,1511]" pageId="11" pageNumber="12">IZB</collectionCode>
); 1 ɗ, same but
<quantity id="4CB89B22FFBEFFFFFB868DD83175FA34" box="[1030,1122,1485,1511]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="11" pageNumber="12" unit="m" value="3300.0">3300 m</quantity>
,
<date id="FFFE1007FFBEFFFFFBED8DD831E2FA34" box="[1133,1269,1485,1511]" pageId="11" pageNumber="12" value="1983-06-30">30.VI.1983</date>
, Chen Yuanging leg. (
<collectionCode id="ED51AE02FFBEFFFFFE818DE03422F9DC" box="[257,309,1525,1551]" pageId="11" pageNumber="12">IZB</collectionCode>
); 2 ɗ, 2 Ψ, same but
<quantity id="4CB89B22FFBEFFFFFDB48DE03787F9DC" box="[564,656,1525,1551]" metricMagnitude="3" metricUnit="m" metricValue="3.65" pageId="11" pageNumber="12" unit="m" value="3650.0">3650 m</quantity>
,
<date id="FFFE1007FFBEFFFFFD1D8DE03639F9DC" box="[669,814,1525,1551]" pageId="11" pageNumber="12" value="1983-07-11">11.VII.1983</date>
, Wang Shuyong (
<specimenCount id="9D46FD4EFFBEFFFFFB818DE03182F9DC" box="[1025,1173,1525,1551]" pageId="11" pageNumber="12" type="generic">2 specimens</specimenCount>
) &amp; Chen Yuanqing (
<specimenCount id="9D46FD4EFFBEFFFFFA0C8DE03404F9E4" pageId="11" pageNumber="12" type="generic">2 specimens</specimenCount>
) leg. (
<collectionCode id="ED51AE02FFBEFFFFFEDD8E083486F9E4" box="[349,401,1565,1591]" pageId="11" pageNumber="12">IZB</collectionCode>
); 2 ɗ, same but
<quantity id="4CB89B22FFBEFFFFFDCE8E0837BEF9E4" box="[590,681,1565,1591]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="11" pageNumber="12" unit="m" value="3600.0">3600 m</quantity>
,
<date id="FFFE1007FFBEFFFFFD358E083620F9E4" box="[693,823,1565,1591]" pageId="11" pageNumber="12" value="1983-07-01">1.VII.1983</date>
, Cheng Yuanqing &amp; Wang Shuyong leg. (
<collectionCode id="ED51AE02FFBEFFFFFAA28E083041F9E4" box="[1314,1366,1565,1591]" pageId="11" pageNumber="12">IZB</collectionCode>
); 3 ɗ, 1 Ψ, Wenchuan,
<quantity id="4CB89B22FFBEFFFFFED78E5034FAF98C" box="[343,493,1605,1631]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="11" pageNumber="12" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900–1000 m</quantity>
,
<date id="FFFE1007FFBEFFFFFE788E503793F98C" box="[504,644,1605,1631]" pageId="11" pageNumber="12" value="1983-08-01">1.VIII.1983</date>
, Zhang Xuezhong leg. (
<collectionCode id="ED51AE02FFBEFFFFFC1B8E5036D8F98C" box="[923,975,1605,1631]" pageId="11" pageNumber="12">IZB</collectionCode>
); 1 ɗ, W Sichuan,
<quantity id="4CB89B22FFBEFFFFFB2A8E5031E4F98C" box="[1194,1267,1605,1631]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="km" value="15.0">15 km</quantity>
NE Qianning,
<geoCoordinate id="EE745000FFBEFFFFFF178E7835E0F954" box="[151,247,1645,1671]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="925" value="30.583334">30°35'N</geoCoordinate>
<geoCoordinate id="EE745000FFBEFFFFFF7F8E783479F954" box="[255,366,1645,1671]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="101.683334">101°41'E</geoCoordinate>
,
<date id="FFFE1007FFBEFFFFFEFB8E78371BF954" box="[379,524,1645,1671]" pageId="11" pageNumber="12" value="1998-07-11">11.VII.1998</date>
, M. Bocak leg. (cWR); 1 ɗ, 1 Ψ, Kangding,
<quantity id="4CB89B22FFBEFFFFFB9D8E78316EF954" box="[1053,1145,1645,1671]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="11" pageNumber="12" unit="m" value="3700.0">3700 m</quantity>
,
<date id="FFFE1007FFBEFFFFFB058E783080F954" box="[1157,1431,1645,1671]" pageId="11" pageNumber="12" value="1982-08-30" valueMax="1982-09-07" valueMin="1982-08-30">30.VIII. and 7.IX.1982</date>
, Wang Shuyong leg. (
<collectionCode id="ED51AE02FFBEFFFFFE128E8034D1F97C" box="[402,454,1685,1711]" pageId="11" pageNumber="12">IZB</collectionCode>
); 4 ɗ, 2 Ψ, Barkam Co.,
<quantity id="4CB89B22FFBEFFFFFD768E803643F97C" box="[758,852,1685,1711]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="12" unit="m" value="2500.0">2500 m</quantity>
,
<date id="FFFE1007FFBEFFFFFCE18E8036EAF97C" box="[865,1021,1685,1711]" pageId="11" pageNumber="12" value="1983-08-20">20.VIII.1983</date>
, Wang Ruqi leg. (
<collectionCode id="ED51AE02FFBEFFFFFB578E80301CF97C" box="[1239,1291,1685,1711]" pageId="11" pageNumber="12">IZB</collectionCode>
); 1 ɗ, Dêgê Co.,
<quantity id="4CB89B22FFBEFFFFFF508EA8343AF904" box="[208,301,1725,1751]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="11" pageNumber="12" unit="m" value="3900.0">3900 m</quantity>
,
<date id="FFFE1007FFBEFFFFFEB98EA834ABF904" box="[313,444,1725,1751]" pageId="11" pageNumber="12" value="1983-07-06">6.VII.1983</date>
, Wang Shuyong leg. (
<collectionCode id="ED51AE02FFBEFFFFFD418EA837E2F904" box="[705,757,1725,1751]" pageId="11" pageNumber="12">IZB</collectionCode>
); 1 Ψ, Dêgê Co., Maniganggo,
<quantity id="4CB89B22FFBEFFFFFBE58EA831D5F904" box="[1125,1218,1725,1751]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="11" pageNumber="12" unit="m" value="3900.0">3900 m</quantity>
,
<date id="FFFE1007FFBEFFFFFB4E8EA83046F904" box="[1230,1361,1725,1751]" pageId="11" pageNumber="12" value="1983-07-08">8.VII.1983</date>
, Chen Yuanqing leg. (
<collectionCode id="ED51AE02FFBEFFFFFECA8EF03469F92C" box="[330,382,1765,1791]" pageId="11" pageNumber="12">IZB</collectionCode>
); 2 Ψ, Dêgê Co.,
<quantity id="4CB89B22FFBEFFFFFDC88EF037B4F92C" box="[584,675,1765,1791]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="12" unit="m" value="3800.0">3800 m</quantity>
,
<date id="FFFE1007FFBEFFFFFD2F8EF03624F92C" box="[687,819,1765,1791]" pageId="11" pageNumber="12" value="1983-07-09">9.VII.1983</date>
, Wang Shuyong leg. (
<collectionCode id="ED51AE02FFBEFFFFFBB58EF0317EF92C" box="[1077,1129,1765,1791]" pageId="11" pageNumber="12">IZB</collectionCode>
); 2 ɗ, 2 Ψ, same but
<quantity id="4CB89B22FFBEFFFFFAE18EF035A6F8F4" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="11" pageNumber="12" unit="m" value="3900.0">3900 m</quantity>
,
<date id="FFFE1007FFBEFFFFFF3D8F183458F8F4" box="[189,335,1805,1831]" pageId="11" pageNumber="12" value="1983-07-10">10.VII.1983</date>
, Zhang Xuezhong leg. (
<collectionCode id="ED51AE02FFBEFFFFFDEA8F183789F8F4" box="[618,670,1805,1831]" pageId="11" pageNumber="12">IZB</collectionCode>
); 4 ɗ, 1 Ψ, same but
<quantity id="4CB89B22FFBEFFFFFC178F1836E4F8F4" box="[919,1011,1805,1831]" metricMagnitude="3" metricUnit="m" metricValue="4.2" pageId="11" pageNumber="12" unit="m" value="4200.0">4200 m</quantity>
,
<date id="FFFE1007FFBEFFFFFC7F8F183186F8F4" box="[1023,1169,1805,1831]" pageId="11" pageNumber="12" value="1983-07-10">10.VII.1983</date>
, Niu Chunlai (
<specimenCount id="9D46FD4EFFBEFFFFFAC08F1835C0F89C" pageId="11" pageNumber="12" type="generic">3 specimens</specimenCount>
) and Chai Huaicheng (
<specimenCount id="9D46FD4EFFBEFFFFFE688F20376CF89C" box="[488,635,1845,1871]" pageId="11" pageNumber="12" type="generic">2 specimens</specimenCount>
) leg. (
<collectionCode id="ED51AE02FFBEFFFFFD478F2037ECF89C" box="[711,763,1845,1871]" pageId="11" pageNumber="12">IZB</collectionCode>
); 1 Ψ, same but
<quantity id="4CB89B22FFBEFFFFFC3C8F20310EF89C" box="[956,1049,1845,1871]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="12" unit="m" value="4000.0">4000 m</quantity>
,
<date id="FFFE1007FFBEFFFFFBA58F2031B0F89C" box="[1061,1191,1845,1871]" pageId="11" pageNumber="12" value="1983-07-08">8.VII.1983</date>
, Wang Shuyong leg. (
<collectionCode id="ED51AE02FFBEFFFFFF208F4835C2F8A4" box="[160,213,1885,1911]" pageId="11" pageNumber="12">IZB</collectionCode>
).
<collectionCode id="ED51AE02FFBEFFFFFF688F483429F8A4" box="[232,318,1885,1911]" pageId="11" pageNumber="12">TIBET</collectionCode>
: 1 Ψ, Jomda Co.,
<quantity id="4CB89B22FFBEFFFFFD8E8F48377DF8A4" box="[526,618,1885,1911]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="11" pageNumber="12" unit="m" value="3400.0">3400 m</quantity>
,
<date id="FFFE1007FFBEFFFFFDF58F483611F8A4" box="[629,774,1885,1911]" pageId="11" pageNumber="12" value="1976-07-27">27.VII.1976</date>
, Zhang Xuezhong leg. (
<collectionCode id="ED51AE02FFBEFFFFFBA08F483143F8A4" box="[1056,1108,1885,1911]" pageId="11" pageNumber="12">IZB</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFBEFFF9FF478F9036E2FCD9" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="description">
<paragraph id="8BFF36C7FFBEFFF8FF478F9031A5F84F" blockId="11.[151,1436,1365,2031]" lastBlockId="12.[151,1436,1921,2028]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="B934EAD5FFBEFFFFFF478F90344FF84C" bold="true" box="[199,344,1925,1951]" pageId="11" pageNumber="12">Description</emphasis>
. Habitus:
<figureCitation id="137B2A42FFBEFFFFFE518F90374FF84C" box="[465,600,1925,1951]" captionStart="FIGURES 25 – 29" captionStartId="12.[151,269,1777,1801]" captionTargetBox="[153,1389,532,1776]" captionTargetId="figure@12.[151,1436,532,1843]" captionTargetPageId="12" captionText="FIGURES 25 – 29. Harpalus kaznakovi. 25 – 27, H. k. kaznakovi (25, 27, Sichuan, Garzê Co.; 26, Sichuan, Dêgê Co.); 28 – 29, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 25 – 26, 28, habitus, 27 – 29, left metepisternum. Scales = 1.0 mm (A: Figs 25 – 26, 28; B: 27, 29)." httpUri="https://zenodo.org/record/178761/files/figure.png" pageId="11" pageNumber="12">Figs 25–26</figureCitation>
. Metallic tinge on dorsum variable, sometimes recognizable only along margins of pronotum and elytra. Body size very variable: body length
<quantity id="4CB89B22FFBEFFFFFC678FB8316EF814" box="[999,1145,1965,1991]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="8.4" metricValueMin="5.9" pageId="11" pageNumber="12" unit="mm" value="7.15" valueMax="8.4" valueMin="5.9">5.9–8.4 mm</quantity>
. Head moderate or relatively large (in male WHmax/WPmax = 0.58–0.72 and WHmin/WPmax = 0.53–0.61), with convex or almost flat eyes (WHmax/WHmin = 1.04–1.25). Pronotum variable in proportions and shape, 1.44–1.59 as wide as long, with sides rounded or straight, converging posteriad in basal half, sometimes sinuate; basal angles obtuse, somewhat sharp or rounded at apices; basal foveae small, variable in shape, usually elongate and not deep; base of pronotum finely punctate or almost smooth, only with few fine punctures in basal foveae. Elytra relatively short or elongate, 1.26–1.42 times as long as wide, 2.14–2.43 times as long and 1.09–1.21 times as wide as pronotum; parascutellar pore present or absent; each third interval with or without one discal pore. Metepisterna (
<figureCitation id="137B2A42FFB9FFF8FEBF899D3483FE71" box="[319,404,392,418]" captionStart="FIGURES 25 – 29" captionStartId="12.[151,269,1777,1801]" captionTargetBox="[153,1389,532,1776]" captionTargetId="figure@12.[151,1436,532,1843]" captionTargetPageId="12" captionText="FIGURES 25 – 29. Harpalus kaznakovi. 25 – 27, H. k. kaznakovi (25, 27, Sichuan, Garzê Co.; 26, Sichuan, Dêgê Co.); 28 – 29, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 25 – 26, 28, habitus, 27 – 29, left metepisternum. Scales = 1.0 mm (A: Figs 25 – 26, 28; B: 27, 29)." httpUri="https://zenodo.org/record/178761/files/figure.png" pageId="12" pageNumber="13">Fig. 27</figureCitation>
) wider than long, weakly narrowed posteriad. Metacoxae usually without additional setigerous pores, occasionally with a small such pore medially. Median lobe of aedeagus (
<figureCitation id="137B2A42FFB9FFF8FB3389A535F6FE21" captionStart="FIGURES 30 – 44" captionStartId="14.[151,269,1898,1922]" captionTargetBox="[186,1385,219,1897]" captionTargetId="figure@14.[151,1436,212,1938]" captionTargetPageId="14" captionText="FIGURES 30 – 44. Harpalus kaznakovi, median lobe of aedeagus (32 – 33, 38, 41, 44, terminal lamella). 30 – 41, H. k. kaznakovi (30 – 38, Sichuan, Garzê Co.; 39 – 41, Sichuan, Dêgê Co.). 42 – 44, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 30, 32 – 34, 36, 38 – 39, 41 – 42, 44, dorsal aspect; 31, 35, 37, 40, 43, lateral aspect. Scales: A = 0.5 mm (Figs 32 – 33, 38, 41, 44), B = 1.0 mm (Figs 30 – 31, 34 – 37, 39 – 40, 42 – 43)." httpUri="https://zenodo.org/record/178762/files/figure.png" pageId="12" pageNumber="13">Figs 30–31, 34–37, 39–40</figureCitation>
) more or less strongly bent after basal bulb; terminal lamella (
<figureCitation id="137B2A42FFB9FFF8FC4689CD31B0FE21" box="[966,1191,472,498]" captionStart="FIGURES 30 – 44" captionStartId="14.[151,269,1898,1922]" captionTargetBox="[186,1385,219,1897]" captionTargetId="figure@14.[151,1436,212,1938]" captionTargetPageId="14" captionText="FIGURES 30 – 44. Harpalus kaznakovi, median lobe of aedeagus (32 – 33, 38, 41, 44, terminal lamella). 30 – 41, H. k. kaznakovi (30 – 38, Sichuan, Garzê Co.; 39 – 41, Sichuan, Dêgê Co.). 42 – 44, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 30, 32 – 34, 36, 38 – 39, 41 – 42, 44, dorsal aspect; 31, 35, 37, 40, 43, lateral aspect. Scales: A = 0.5 mm (Figs 32 – 33, 38, 41, 44), B = 1.0 mm (Figs 30 – 31, 34 – 37, 39 – 40, 42 – 43)." httpUri="https://zenodo.org/record/178762/files/figure.png" pageId="12" pageNumber="13">Figs 32–33, 38, 41</figureCitation>
) comparatively long
<emphasis id="B934EAD5FFB9FFF8FF478F943477F848" bold="true" box="[199,352,1921,1947]" pageId="12" pageNumber="13">Distribution</emphasis>
. Known from north-western Sichuan and eastern Tibet (Jomba County).
</paragraph>
<caption id="DF3F664FFFB9FFF8FF178EE434F6F89D" httpUri="https://zenodo.org/record/178761/files/figure.png" pageId="12" pageNumber="13" targetBox="[153,1389,532,1776]" targetPageId="12">
<paragraph id="8BFF36C7FFB9FFF8FF178EE434F6F89D" blockId="12.[151,1436,1777,1870]" pageId="12" pageNumber="13">
<emphasis id="B934EAD5FFB9FFF8FF178EE4344AF8DA" bold="true" box="[151,349,1777,1801]" pageId="12" pageNumber="13">FIGURES 25–29.</emphasis>
<emphasis id="B934EAD5FFB9FFF8FEE38EE73757F8DA" box="[355,576,1778,1801]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C404D44FFB9FFF8FEE38EE7372BF8DA" box="[355,572,1778,1801]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kaznakovi">Harpalus kaznakovi</taxonomicName>
.
</emphasis>
25–27,
<taxonomicName id="4C404D44FFB9FFF8FD148EE73620F8DA" box="[660,823,1778,1801]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="kaznakovi" subSpecies="kaznakovi">
<emphasis id="B934EAD5FFB9FFF8FD148EE73620F8DA" box="[660,823,1778,1801]" italics="true" pageId="12" pageNumber="13">H. k. kaznakovi</emphasis>
</taxonomicName>
(25, 27, Sichuan, Garzê Co.; 26, Sichuan, Dêgê Co.); 28– 29,
<taxonomicName id="4C404D44FFB9FFF8FF3F8F01345AF8F8" box="[191,333,1812,1835]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subSpecies" species="kaznakovi" status="subsp. nov." subSpecies="lilliputa">
<emphasis id="B934EAD5FFB9FFF8FF3F8F01345AF8F8" box="[191,333,1812,1835]" italics="true" pageId="12" pageNumber="13">H. k. lilliputa</emphasis>
</taxonomicName>
<emphasis id="B934EAD5FFB9FFF8FED48F01348CF8FF" bold="true" box="[340,411,1812,1836]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A20757AEFFB9FFF8FED48F01348CF8FF" box="[340,411,1812,1836]" pageId="12" pageNumber="13" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
(Tibet, Zogang Co., holotype). 25–26, 28, habitus, 27–29, left metepisternum. Scales = 1.0 mm (A: Figs 25–26, 28; B: 27, 29).
</paragraph>
</caption>
<paragraph id="8BFF36C7FFB9FFF9FF478FBC36E2FCD9" blockId="12.[151,1436,1921,2028]" lastBlockId="13.[151,1437,152,898]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="B934EAD5FFB9FFF8FF478FBC342AF810" bold="true" box="[199,317,1961,1987]" pageId="12" pageNumber="13">Variation</emphasis>
. The nominotypical subspecies demonstrates geographical variation in several characters, mainly in body size and shape of eyes and pronotum, and apparently represents a complex of several local forms which however seem to be not sharply separated from each other. At least based on the scarce available material delimitation of these forms involves big difficulties. In Sichuan, the specimens from Garzê County, Wenchuang and Kangding are similar to those of the
<typeStatus id="54FB8865FFB8FFF9FC9788FD365DFED1" box="[791,842,232,258]" pageId="13" pageNumber="14">type</typeStatus>
series; they are, on average, rather large (
<quantity id="4CB89B22FFB8FFF9FAC288FD35D0FEF9" metricMagnitude="-3" metricUnit="m" metricValue="7.1499999999999995" metricValueMax="8.2" metricValueMin="6.1" pageId="13" pageNumber="14" unit="mm" value="7.1499999999999995" valueMax="8.2" valueMin="6.1">6.1–8.2 mm</quantity>
) and robust but eyes are only scarcely convex, almost flat (
<figureCitation id="137B2A42FFB8FFF9FC0D890536F2FEF9" box="[909,997,272,298]" captionStart="FIGURES 25 – 29" captionStartId="12.[151,269,1777,1801]" captionTargetBox="[153,1389,532,1776]" captionTargetId="figure@12.[151,1436,532,1843]" captionTargetPageId="12" captionText="FIGURES 25 – 29. Harpalus kaznakovi. 25 – 27, H. k. kaznakovi (25, 27, Sichuan, Garzê Co.; 26, Sichuan, Dêgê Co.); 28 – 29, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 25 – 26, 28, habitus, 27 – 29, left metepisternum. Scales = 1.0 mm (A: Figs 25 – 26, 28; B: 27, 29)." httpUri="https://zenodo.org/record/178761/files/figure.png" pageId="13" pageNumber="14">Fig. 25</figureCitation>
; WHmax/WHmin = 1.04–1.10). The median lobe of aedeagus of these specimens is usually almost straight from dorsal view or only slightly curved to right (
<figureCitation id="137B2A42FFB8FFF9FED589753787FEA9" box="[341,656,352,378]" captionStart="FIGURES 30 – 44" captionStartId="14.[151,269,1898,1922]" captionTargetBox="[186,1385,219,1897]" captionTargetId="figure@14.[151,1436,212,1938]" captionTargetPageId="14" captionText="FIGURES 30 – 44. Harpalus kaznakovi, median lobe of aedeagus (32 – 33, 38, 41, 44, terminal lamella). 30 – 41, H. k. kaznakovi (30 – 38, Sichuan, Garzê Co.; 39 – 41, Sichuan, Dêgê Co.). 42 – 44, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 30, 32 – 34, 36, 38 – 39, 41 – 42, 44, dorsal aspect; 31, 35, 37, 40, 43, lateral aspect. Scales: A = 0.5 mm (Figs 32 – 33, 38, 41, 44), B = 1.0 mm (Figs 30 – 31, 34 – 37, 39 – 40, 42 – 43)." httpUri="https://zenodo.org/record/178762/files/figure.png" pageId="13" pageNumber="14">Figs 30–31, 34–35, 36–37</figureCitation>
); in several specimens from Garzê County the median lobe is narrower and less strongly arcuate (
<figureCitation id="137B2A42FFB8FFF9FD9B899D37B4FE71" box="[539,675,392,418]" captionStart="FIGURES 30 – 44" captionStartId="14.[151,269,1898,1922]" captionTargetBox="[186,1385,219,1897]" captionTargetId="figure@14.[151,1436,212,1938]" captionTargetPageId="14" captionText="FIGURES 30 – 44. Harpalus kaznakovi, median lobe of aedeagus (32 – 33, 38, 41, 44, terminal lamella). 30 – 41, H. k. kaznakovi (30 – 38, Sichuan, Garzê Co.; 39 – 41, Sichuan, Dêgê Co.). 42 – 44, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 30, 32 – 34, 36, 38 – 39, 41 – 42, 44, dorsal aspect; 31, 35, 37, 40, 43, lateral aspect. Scales: A = 0.5 mm (Figs 32 – 33, 38, 41, 44), B = 1.0 mm (Figs 30 – 31, 34 – 37, 39 – 40, 42 – 43)." httpUri="https://zenodo.org/record/178762/files/figure.png" pageId="13" pageNumber="14">Figs 34–35</figureCitation>
) than that in most other specimens. The pronotum of the specimens from Garzê County, Wenchuang and Kangding is usually with sides rounded or straight, converging posteriad in basal half, sometimes sinuate (more markedly so in the specimens from Wenchuan), and basal angles obtuse and rounded at apices. In one male examined from the environs of Qianning, eyes are scarcely convex but pronotal sides clearly sinuate basally and basal angles nearly right and rather sharp at apices. By contrast, in the specimens from Barkam and Dêgê counties (Sichuan) as well as Jomba County (Tibet), body is, on average, smaller (5.9–7.0 mm) and more slender and eyes are more convex (
<figureCitation id="137B2A42FFB8FFF9FBDE8A6D31B9FD41" box="[1118,1198,632,658]" captionStart="FIGURES 25 – 29" captionStartId="12.[151,269,1777,1801]" captionTargetBox="[153,1389,532,1776]" captionTargetId="figure@12.[151,1436,532,1843]" captionTargetPageId="12" captionText="FIGURES 25 – 29. Harpalus kaznakovi. 25 – 27, H. k. kaznakovi (25, 27, Sichuan, Garzê Co.; 26, Sichuan, Dêgê Co.); 28 – 29, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 25 – 26, 28, habitus, 27 – 29, left metepisternum. Scales = 1.0 mm (A: Figs 25 – 26, 28; B: 27, 29)." httpUri="https://zenodo.org/record/178761/files/figure.png" pageId="13" pageNumber="14">Fig.26</figureCitation>
; WHmax/WHmin = 1.11–1.25) than those in the most specimens from Garzê, Wenchuang and Kangding counties. The median lobe of aedeagus of these specimens is markedly arcuate and with apical portion more strongly curved to right (
<figureCitation id="137B2A42FFB8FFF9FF1F8AE5343EFCD9" box="[159,297,752,778]" captionStart="FIGURES 30 – 44" captionStartId="14.[151,269,1898,1922]" captionTargetBox="[186,1385,219,1897]" captionTargetId="figure@14.[151,1436,212,1938]" captionTargetPageId="14" captionText="FIGURES 30 – 44. Harpalus kaznakovi, median lobe of aedeagus (32 – 33, 38, 41, 44, terminal lamella). 30 – 41, H. k. kaznakovi (30 – 38, Sichuan, Garzê Co.; 39 – 41, Sichuan, Dêgê Co.). 42 – 44, H. k. lilliputa ssp. n. (Tibet, Zogang Co., holotype). 30, 32 – 34, 36, 38 – 39, 41 – 42, 44, dorsal aspect; 31, 35, 37, 40, 43, lateral aspect. Scales: A = 0.5 mm (Figs 32 – 33, 38, 41, 44), B = 1.0 mm (Figs 30 – 31, 34 – 37, 39 – 40, 42 – 43)." httpUri="https://zenodo.org/record/178762/files/figure.png" pageId="13" pageNumber="14">Figs 39–40</figureCitation>
). The armature of the internal sac in all forms is very similar.
</paragraph>
</subSubSection>
<subSubSection id="C35A654CFFB8FFF9FF478B0234BFFC51" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BFF36C7FFB8FFF9FF478B0234BFFC51" blockId="13.[151,1437,152,898]" pageId="13" pageNumber="14">
<emphasis id="B934EAD5FFB8FFF9FF478B02342EFCE2" bold="true" box="[199,313,791,817]" pageId="13" pageNumber="14">Remarks</emphasis>
. Although the nominotypical subspecies is highly variable, it may be recognized by the short and wide metepisterna, the terminal lamella of the median lobe of aedeagus nearly parallel–sided and the body larger and more robust.
</paragraph>
</subSubSection>
</treatment>
</document>
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