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<document id="E58336994EA2EA9880ABB442E6C37277" ID-DOI="10.5281/zenodo.3676640" ID-GBIF-Dataset="33b3b569-c80d-4a03-9bc3-aa651fa1578a" ID-ISSN="1942-1354" ID-Zenodo-Dep="3676640" ID-ZooBank="37A2F301-0637-41B1-8CE7-A17A54564AF9" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582199722738" checkinUser="felipe" docAuthor="Schnepp, Kyle E." docDate="2019" docId="03E98787FFF20A33FF53C716FBD7FBFA" docLanguage="en" docName="InsectaMundi.0742.1-6.pdf.imf" docOrigin="Insecta Mundi 742 (742)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Phyllophaga rugosa" docType="treatment" docVersion="8" lastPageNumber="2" masterDocId="FFD0FFFFFFF10A30FF93C634FFA8FFD3" masterDocTitle="Three new synonymies in Phyllophaga Harris, 1827 (Coleoptera: Scarabaeidae), with lectotype and neotype designations" masterLastPageNumber="6" masterPageNumber="1" pageNumber="2" updateTime="1698720329174" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="E43FCE6EF273DA3C92FE65E3FCD02A55">Three new synonymies in Phyllophaga Harris, 1827 (Coleoptera: Scarabaeidae), with lectotype and neotype designations</mods:title>
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<mods:namePart id="5528D33031CEC3EABED9101E5CD3317E">Schnepp, Kyle E.</mods:namePart>
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<treatment id="03E98787FFF20A33FF53C716FBD7FBFA" ID-DOI="http://doi.org/10.5281/zenodo.3681513" ID-GBIF-Taxon="162233556" ID-Zenodo-Dep="3681513" LSID="urn:lsid:plazi:treatment:03E98787FFF20A33FF53C716FBD7FBFA" httpUri="http://treatment.plazi.org/id/03E98787FFF20A33FF53C716FBD7FBFA" lastPageNumber="2" pageId="3" pageNumber="2">
<subSubSection id="C35A651AFFF20A33FF53C716FD76FEE9" box="[192,734,290,314]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="8BFF3691FFF20A33FF53C716FD76FEE9" blockId="3.[192,734,290,347]" box="[192,734,290,314]" pageId="3" pageNumber="2">
<heading id="D0B781FDFFF20A33FF53C716FD76FEE9" bold="true" box="[192,734,290,314]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
<taxonomicName id="4C404D12FFF20A33FF53C716FD76FEE9" authority="(Melsheimer, 1845)" baseAuthorityName="Melsheimer" baseAuthorityYear="1845" box="[192,734,290,314]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B934EA83FFF20A33FF53C716FD76FEE9" bold="true" box="[192,734,290,314]" pageId="3" pageNumber="2">
<emphasis id="B934EA83FFF20A33FF53C716FE7CFEE9" bold="true" box="[192,468,290,314]" italics="true" pageId="3" pageNumber="2">Phyllophaga rugosa</emphasis>
(
<bibRefCitation id="EFD14B60FFF20A33FE77C716FD7DFEE9" author="Melsheimer, F. E." box="[484,725,290,314]" pageId="3" pageNumber="2" pagination="134 - 160" refId="ref3453" refString="Melsheimer, F. E. 1845. Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 2: 134 - 160." type="journal article" year="1845">Melsheimer, 1845</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35A651AFFF20A33FF53C777FE33FE88" box="[192,411,323,347]" pageId="3" pageNumber="2" type="description">
<paragraph id="8BFF3691FFF20A33FF53C777FE33FE88" blockId="3.[192,734,290,347]" box="[192,411,323,347]" pageId="3" pageNumber="2">
<figureCitation id="137B2A14FFF20A33FF53C777FEFCFE88" box="[192,340,323,347]" captionStart="Figures 16" captionStartId="6.[192,287,1726,1748]" captionTargetBox="[369,1328,213,1651]" captionTargetId="figure@6.[369,1328,213,1651]" captionTargetPageId="6" captionText="Figures 16. Male genitalia of Phyllophaga type specimens. 1) Phyllophaga sociata, lectotype. 2) Phyllophaga knausii, lectotype. 3) Phyllophaga rugosa, neotype. 4) Phyllophaga chippewa, holotype. 5) Phyllophaga bipartita, lectotype. 6) Phyllophaga falta, holotype." figureDoi="http://doi.org/10.5281/zenodo.3676642" httpUri="https://zenodo.org/record/3676642/files/figure.png" pageId="3" pageNumber="2">Figures 34</figureCitation>
,
<figureCitation id="137B2A14FFF20A33FEF3C777FE33FE88" box="[352,411,323,347]" captionStart="Figures 712" captionStartId="7.[192,287,1663,1685]" captionTargetBox="[192,1445,213,1629]" captionTargetId="figure@7.[192,1445,213,1629]" captionTargetPageId="7" captionText="Figures 712. Specimen labels of Phyllophaga type specimens. 7) Phyllophaga sociata, lectotype. 8) Phyllophaga knausii, lectotype. 9) Phyllophaga rugosa, neotype. 10) Phyllophaga chippewa, holotype. 11) Phyllophaga bipartita, lectotype. 12) Phyllophaga falta, holotype." figureDoi="http://doi.org/10.5281/zenodo.3676644" httpUri="https://zenodo.org/record/3676644/files/figure.png" pageId="3" pageNumber="2">910</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35A651AFFF20A33FF53C75AFB9FFE08" pageId="3" pageNumber="2" type="reference_group">
<paragraph id="8BFF3691FFF20A33FF53C75AFBD9FE57" blockId="3.[192,1168,366,475]" box="[192,1137,366,388]" pageId="3" pageNumber="2">
<treatmentCitationGroup id="AB5011BFFFF20A33FF53C75AFBD9FE57" box="[192,1137,366,388]" pageId="3" pageNumber="2">
<taxonomicName id="4C404D12FFF20A33FF53C75AFD32FE57" authority="Melsheimer 1845: 140" authorityName="Melsheimer" authorityPageNumber="140" authorityYear="1845" box="[192,666,366,388]" class="Insecta" family="Melolonthidae" genus="Ancylonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B934EA83FFF20A33FF53C75AFE35FE57" box="[192,413,366,388]" italics="true" pageId="3" pageNumber="2">Ancylonycha rugosa</emphasis>
<treatmentCitation id="0AE11080FFF20A33FE37C75AFD32FE57" author="Melsheimer, F. E." box="[420,666,366,388]" page="140" pageId="3" pageNumber="2" year="1845">
<bibRefCitation id="EFD14B60FFF20A33FE37C75AFD32FE57" author="Melsheimer, F. E." box="[420,666,366,388]" pageId="3" pageNumber="2" pagination="134 - 160" refId="ref3453" refString="Melsheimer, F. E. 1845. Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 2: 134 - 160." type="journal article" year="1845">Melsheimer 1845: 140</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
<emphasis id="B934EA83FFF20A33FD37C75AFCA4FE50" bold="true" box="[676,780,366,387]" pageId="3" pageNumber="2">
<typeStatus id="54FB8833FFF20A33FD37C75AFCA4FE50" box="[676,780,366,387]" pageId="3" pageNumber="2" type="neotype">Neotype</typeStatus>
</emphasis>
: MCZ,
<emphasis id="B934EA83FFF20A33FCCDC75AFB8EFE57" bold="true" box="[862,1062,366,388]" pageId="3" pageNumber="2">here designated</emphasis>
, male.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFF3691FFF20A33FF53C7BFFB38FE73" blockId="3.[192,1168,366,475]" box="[192,1168,395,417]" pageId="3" pageNumber="2">
<treatmentCitationGroup id="AB5011BFFFF20A33FF53C7BFFB38FE73" box="[192,1168,395,417]" pageId="3" pageNumber="2">
<taxonomicName id="4C404D12FFF20A33FF53C7BFFD90FE72" authority="(Melsheimer)" baseAuthorityName="Melsheimer" baseAuthorityYear="1845" box="[192,568,395,417]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B934EA83FFF20A33FF53C7BFFE34FE72" box="[192,412,395,417]" italics="true" pageId="3" pageNumber="2">Phyllophaga rugosa</emphasis>
(Melsheimer)
</taxonomicName>
:
<treatmentCitation id="0AE11080FFF20A33FDD1C7BFFC19FE72" author="Haldeman, S. S. &amp; J. L. LeConte" box="[578,945,395,417]" page="59" pageId="3" pageNumber="2" year="1853">
<bibRefCitation id="EFD14B60FFF20A33FDD1C7BFFC19FE72" author="Haldeman, S. S. &amp; J. L. LeConte" box="[578,945,395,417]" pageId="3" pageNumber="2" refId="ref3161" refString="Haldeman, S. S., and J. L. LeConte. 1853. Catalogue of the described Coleoptera of the United States by Friedrich Ernst Melsheimer, revised. Smithsonian Institution; Washington, DC. 174 p." type="book" year="1853">Haldeman and LeConte 1853: 59</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE11080FFF20A33FC2FC7BFFB23FE73" author="Glasgow, R. D." box="[956,1163,395,417]" page="371" pageId="3" pageNumber="2" year="1916">
<bibRefCitation id="EFD14B60FFF20A33FC2FC7BFFB23FE73" author="Glasgow, R. D." box="[956,1163,395,417]" pageId="3" pageNumber="2" pagination="365 - 379" refId="ref3117" refString="Glasgow, R. D. 1916. Phyllophaga Harris (Lachnosterna Hope): A revision of the synonymy, and one new name. Bulletin of the Illinois State Laboratory of Natural History 11 (5): 365 - 379." type="journal article" year="1916">
<collectingRegion id="4984F873FFF20A33FC2FC7BFFBB2FE72" box="[956,1050,395,417]" country="United Kingdom" name="Glasgow City" pageId="3" pageNumber="2">Glasgow</collectingRegion>
1916: 371
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFF3691FFF20A33FF53C79CFC80FE6D" blockId="3.[192,1168,366,475]" box="[192,808,424,446]" pageId="3" pageNumber="2">
<treatmentCitationGroup id="AB5011BFFFF20A33FF53C79CFC80FE6D" box="[192,808,424,446]" pageId="3" pageNumber="2">
<taxonomicName id="4C404D12FFF20A33FF53C79CFDEEFE6D" authority="(Melsheimer)" baseAuthorityName="Melsheimer" box="[192,582,424,446]" class="Insecta" family="Scarabaeidae" genus="Lachnosterna" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B934EA83FFF20A33FF53C79CFE02FE6E" box="[192,426,424,445]" italics="true" pageId="3" pageNumber="2">Lachnosterna rugosa</emphasis>
(Melsheimer)
</taxonomicName>
:
<treatmentCitation id="0AE11080FFF20A33FDC3C79CFC8CFE6D" author="LeConte, J. L." box="[592,804,424,446]" page="252" pageId="3" pageNumber="2" year="1856">
<bibRefCitation id="EFD14B60FFF20A33FDC3C79CFC8CFE6D" author="LeConte, J. L." box="[592,804,424,446]" pageId="3" pageNumber="2" pagination="225 - 288" refId="ref3382" refString="LeConte, J. L. 1856. Synopsis of the Melolonthidae of the United States. Journal of the Academy of Natural Sciences of Philadelphia, Second Series 3: 225 - 288." type="book chapter" year="1856">LeConte 1856: 252</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFF3691FFF20A33FF53C7F1FB9FFE08" blockId="3.[192,1168,366,475]" box="[192,1079,453,475]" pageId="3" pageNumber="2">
<treatmentCitationGroup id="AB5011BFFFF20A33FF53C7F1FB9FFE08" box="[192,1079,453,475]" pageId="3" pageNumber="2">
<taxonomicName id="4C404D12FFF20A33FF53C7F1FD2FFE08" authority="Saylor 1939 b: 455" authorityName="Saylor" authorityPageNumber="455" authorityYear="1939" box="[192,647,453,475]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="chippewa">
<emphasis id="B934EA83FFF20A33FF53C7F1FE10FE09" box="[192,440,453,474]" italics="true" pageId="3" pageNumber="2">Phyllophaga chippewa</emphasis>
<treatmentCitation id="0AE11080FFF20A33FE2DC7F1FD2FFE08" author="Saylor, L. W." box="[446,647,453,475]" page="455" pageId="3" pageNumber="2" year="1939">
<bibRefCitation id="EFD14B60FFF20A33FE2DC7F1FD2FFE08" author="Saylor, L. W." box="[446,647,453,475]" pageId="3" pageNumber="2" pagination="452 - 461" refId="ref3747" refString="Saylor, L. W. 1939 b. Notes and descriptions of United States Scarab beetles. Journal of the Washington Academy of Sciences 29 (10): 452 - 461." type="journal article" year="1939">Saylor 1939b: 455</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
<emphasis id="B934EA83FFF20A33FD02C7F1FCE3FE09" bold="true" box="[657,843,453,474]" pageId="3" pageNumber="2">New synonymy</emphasis>
.
<typeStatus id="54FB8833FFF20A33FCCBC7F1FC3BFE09" box="[856,915,453,474]" pageId="3" pageNumber="2">Type</typeStatus>
: USNM, male.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35A651AFFF20A33FF7BC7C4FBD7FBFA" pageId="3" pageNumber="2" type="discussion">
<paragraph id="8BFF3691FFF20A33FF7BC7C4FA08FC5A" blockId="3.[192,1440,496,1065]" pageId="3" pageNumber="2">
<taxonomicName id="4C404D12FFF20A33FF7BC7C4FE77FDDB" authorityName="Melsheimer" authorityYear="1845" box="[232,479,496,520]" class="Insecta" family="Melolonthidae" genus="Ancylonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B934EA83FFF20A33FF7BC7C4FE77FDDB" box="[232,479,496,520]" italics="true" pageId="3" pageNumber="2">Ancylonycha rugosa</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFD14B60FFF20A33FD5AC7C5FC19FDDA" author="Melsheimer, F. E." box="[713,945,497,521]" pageId="3" pageNumber="2" pagination="134 - 160" refId="ref3453" refString="Melsheimer, F. E. 1845. Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 2: 134 - 160." type="journal article" year="1845">Melsheimer (1845)</bibRefCitation>
from
<collectingRegion id="4984F873FFF20A33FC6CC7C5FBC0FDDA" box="[1023,1128,497,521]" country="United States of America" name="Virginia" pageId="3" pageNumber="2">Virginia</collectingRegion>
.
<bibRefCitation id="EFD14B60FFF20A33FBE5C7C5FEA4FDFA" author="Haldeman, S. S. &amp; J. L. LeConte" pageId="3" pageNumber="2" refId="ref3161" refString="Haldeman, S. S., and J. L. LeConte. 1853. Catalogue of the described Coleoptera of the United States by Friedrich Ernst Melsheimer, revised. Smithsonian Institution; Washington, DC. 174 p." type="book" year="1853">Haldeman and LeConte (1853)</bibRefCitation>
included it in
<taxonomicName id="4C404D12FFF20A33FE2EC424FDFFFDFB" authorityName="Harris" authorityYear="1827" box="[445,599,528,552]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B934EA83FFF20A33FE2EC424FDFFFDFB" box="[445,599,528,552]" italics="true" pageId="3" pageNumber="2">Phyllophaga</emphasis>
</taxonomicName>
in their revision of Melsheimers catalogue.
<bibRefCitation id="EFD14B60FFF20A33FBE5C425FA98FDFA" author="LeConte, J. L." box="[1142,1328,529,553]" pageId="3" pageNumber="2" pagination="225 - 288" refId="ref3382" refString="LeConte, J. L. 1856. Synopsis of the Melolonthidae of the United States. Journal of the Academy of Natural Sciences of Philadelphia, Second Series 3: 225 - 288." type="book chapter" year="1856">LeConte (1856)</bibRefCitation>
used the generic name
<taxonomicName id="4C404D12FFF20A33FEF9C404FDB8FD9B" authorityName="Hope" authorityYear="1837" box="[362,528,560,584]" class="Insecta" family="Scarabaeidae" genus="Lachnosterna" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B934EA83FFF20A33FEF9C404FDB8FD9B" box="[362,528,560,584]" italics="true" pageId="3" pageNumber="2">Lachnosterna</emphasis>
</taxonomicName>
for this species and most other
<taxonomicName id="4C404D12FFF20A33FC00C404FB85FD9B" authorityName="Harris" authorityYear="1827" box="[915,1069,560,584]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B934EA83FFF20A33FC00C404FB85FD9B" box="[915,1069,560,584]" italics="true" pageId="3" pageNumber="2">Phyllophaga</emphasis>
</taxonomicName>
(sensu stricto).
<bibRefCitation id="EFD14B60FFF20A33FB7CC405FA08FD9A" author="Saylor, L. W." box="[1263,1440,561,585]" pageId="3" pageNumber="2" pagination="452 - 461" refId="ref3747" refString="Saylor, L. W. 1939 b. Notes and descriptions of United States Scarab beetles. Journal of the Washington Academy of Sciences 29 (10): 452 - 461." type="journal article" year="1939">Saylor (1939b)</bibRefCitation>
described
<taxonomicName id="4C404D12FFF20A33FEA8C464FDF8FDBB" authorityName="Saylor" authorityYear="1939" box="[315,592,592,616]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="chippewa">
<emphasis id="B934EA83FFF20A33FEA8C464FDF8FDBB" box="[315,592,592,616]" italics="true" pageId="3" pageNumber="2">Phyllophaga chippewa</emphasis>
</taxonomicName>
from a single male collected in Schley,
<collectingRegion id="4984F873FFF20A33FBA0C465FB1DFDBA" box="[1075,1205,593,617]" country="United States of America" name="Minnesota" pageId="3" pageNumber="2">Minnesota</collectingRegion>
(
<figureCitation id="137B2A14FFF20A33FB56C465FAB3FDBA" box="[1221,1307,593,617]" captionStart="Figures 712" captionStartId="7.[192,287,1663,1685]" captionTargetBox="[192,1445,213,1629]" captionTargetId="figure@7.[192,1445,213,1629]" captionTargetPageId="7" captionText="Figures 712. Specimen labels of Phyllophaga type specimens. 7) Phyllophaga sociata, lectotype. 8) Phyllophaga knausii, lectotype. 9) Phyllophaga rugosa, neotype. 10) Phyllophaga chippewa, holotype. 11) Phyllophaga bipartita, lectotype. 12) Phyllophaga falta, holotype." figureDoi="http://doi.org/10.5281/zenodo.3676644" httpUri="https://zenodo.org/record/3676644/files/figure.png" pageId="3" pageNumber="2">Fig. 10</figureCitation>
), and compared it with
<taxonomicName id="4C404D12FFF20A33FEF2C444FC1CFD5A" authority="(Schoenherr and Gyllenhal)" baseAuthorityName="Schoenherr and Gyllenhal" box="[353,948,624,649]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="knochii">
<emphasis id="B934EA83FFF20A33FEF2C444FDF3FD5B" box="[353,603,624,648]" italics="true" pageId="3" pageNumber="2">Phyllophaga knochii</emphasis>
(Schoenherr and Gyllenhal)
</taxonomicName>
. While
<taxonomicName id="4C404D12FFF20A33FB9FC445FB35FD5B" authorityName="Saylor" authorityYear="1939" box="[1036,1181,624,648]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="chippewa">
<emphasis id="B934EA83FFF20A33FB9FC445FB35FD5B" box="[1036,1181,624,648]" italics="true" pageId="3" pageNumber="2">P. chippewa</emphasis>
</taxonomicName>
does belong in Group IX of
<bibRefCitation id="EFD14B60FFF20A33FE93C4A5FE0CFD7A" author="Horn, G. H." box="[256,420,657,681]" pageId="3" pageNumber="2" pagination="209 - 296" refId="ref3306" refString="Horn, G. H. 1887. Revision of the species of Lachnosterna of America north of Mexico. Transactions of the American Entomological Society 14: 209 - 296." type="journal article" year="1887">Horns (1887)</bibRefCitation>
revision, as stated by
<bibRefCitation id="EFD14B60FFF20A33FD23C4A5FCCCFD7A" author="Saylor, L. W." box="[688,868,657,681]" pageId="3" pageNumber="2" pagination="452 - 461" refId="ref3747" refString="Saylor, L. W. 1939 b. Notes and descriptions of United States Scarab beetles. Journal of the Washington Academy of Sciences 29 (10): 452 - 461." type="journal article" year="1939">Saylor (1939b)</bibRefCitation>
, there are some significant differences between this species and
<taxonomicName id="4C404D12FFF20A33FE1AC485FDABFD1B" authority="Horn 1887" authorityName="Horn" authorityYear="1887" box="[393,515,688,712]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="knochii">
<emphasis id="B934EA83FFF20A33FE1AC485FDABFD1B" box="[393,515,688,712]" italics="true" pageId="3" pageNumber="2">P. knochii</emphasis>
</taxonomicName>
, including pronotal punctation and clypeal emargination. The parameres of
<taxonomicName id="4C404D12FFF20A33FF53C4E5FEFBFD3B" authorityName="Saylor" authorityYear="1939" box="[192,339,720,744]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="chippewa">
<emphasis id="B934EA83FFF20A33FF53C4E5FEFBFD3B" box="[192,339,720,744]" italics="true" pageId="3" pageNumber="2">P. chippewa</emphasis>
</taxonomicName>
are mostly developed, looking normally sclerotized, but the base of the genitalia (
<figureCitation id="137B2A14FFF20A33FAA5C4E5FAD4FD3A" box="[1334,1404,721,745]" captionStart="Figures 16" captionStartId="6.[192,287,1726,1748]" captionTargetBox="[369,1328,213,1651]" captionTargetId="figure@6.[369,1328,213,1651]" captionTargetPageId="6" captionText="Figures 16. Male genitalia of Phyllophaga type specimens. 1) Phyllophaga sociata, lectotype. 2) Phyllophaga knausii, lectotype. 3) Phyllophaga rugosa, neotype. 4) Phyllophaga chippewa, holotype. 5) Phyllophaga bipartita, lectotype. 6) Phyllophaga falta, holotype." figureDoi="http://doi.org/10.5281/zenodo.3676642" httpUri="https://zenodo.org/record/3676642/files/figure.png" pageId="3" pageNumber="2">Fig. 4</figureCitation>
) is clearly underdeveloped and misshapen. The genitalia appear to be a deformity of
<taxonomicName id="4C404D12FFF20A33FB3DC4C5FA89FCDB" authorityName="Melsheimer" authorityYear="1845" box="[1198,1313,753,776]" class="Insecta" family="Melolonthidae" genus="Ancylonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B934EA83FFF20A33FB3DC4C5FA89FCDB" box="[1198,1313,753,776]" italics="true" pageId="3" pageNumber="2">P. rugosa</emphasis>
</taxonomicName>
, as stated by
<bibRefCitation id="EFD14B60FFF20A33FF75C525FDE6FCFA" author="Luginbill, P. &amp; H. R. Painter" box="[230,590,785,809]" pageId="3" pageNumber="2" pagination="1 - 102" refId="ref3417" refString="Luginbill, P., and H. R. Painter. 1953. May beetles of the United States and Canada. United States Department of Agriculture Technical Bulletin 1060: 1 - 102." type="journal article" year="1953">Luginbill and Painter (1953)</bibRefCitation>
. After examination of the
<typeStatus id="54FB8833FFF20A33FC05C525FC66FCFA" box="[918,974,785,809]" pageId="3" pageNumber="2">type</typeStatus>
,
<taxonomicName id="4C404D12FFF20A33FC4FC525FBDAFCFB" authorityName="Saylor" authorityYear="1939" box="[988,1138,784,808]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="chippewa">
<emphasis id="B934EA83FFF20A33FC4FC525FBDAFCFB" box="[988,1138,784,808]" italics="true" pageId="3" pageNumber="2">P. chippewa</emphasis>
</taxonomicName>
is externally within the variation of
<taxonomicName id="4C404D12FFF20A33FEC7C505FE6EFC9B" authorityName="Melsheimer" authorityYear="1845" box="[340,454,817,840]" class="Insecta" family="Melolonthidae" genus="Ancylonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B934EA83FFF20A33FEC7C505FE6EFC9B" box="[340,454,817,840]" italics="true" pageId="3" pageNumber="2">P. rugosa</emphasis>
</taxonomicName>
and the collection locality is also within the range of
<taxonomicName id="4C404D12FFF20A33FBC3C505FA34FC9A" authority="(Pike et al. 1977)" baseAuthorityName="Pike" baseAuthorityYear="1977" box="[1104,1436,817,841]" class="Insecta" family="Melolonthidae" genus="Ancylonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B934EA83FFF20A33FBC3C505FB6AFC9B" box="[1104,1218,817,840]" italics="true" pageId="3" pageNumber="2">P. rugosa</emphasis>
(
<bibRefCitation id="EFD14B60FFF20A33FB41C505FA3CFC9A" author="Pike, K. S. &amp; R. L. Rivers &amp; Z. B. Mayo" box="[1234,1428,817,841]" pageId="3" pageNumber="2" pagination="1 - 13" refId="ref3487" refString="Pike, K. S., R. L. Rivers, and Z. B. Mayo. 1977. Geographical distribution of the known Phyllophaga and Cyclocephala species in the North Central States. University of Nebraska Agricultural Experiment Station Miscellaneous Publication 34: 1 - 13." type="journal article" year="1977">Pike et al. 1977</bibRefCitation>
)
</taxonomicName>
. Being the
<typeStatus id="54FB8833FFF20A33FEACC565FEDCFCBA" box="[319,372,849,873]" pageId="3" pageNumber="2">type</typeStatus>
of
<taxonomicName id="4C404D12FFF20A33FE09C565FD86FCBB" authorityName="Saylor" authorityYear="1939" box="[410,558,848,872]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="chippewa">
<emphasis id="B934EA83FFF20A33FE09C565FD86FCBB" box="[410,558,848,872]" italics="true" pageId="3" pageNumber="2">P. chippewa</emphasis>
</taxonomicName>
remains the only known specimen, I concur with
<bibRefCitation id="EFD14B60FFF20A33FB06C565FEA4FC5A" author="Luginbill, P. &amp; H. R. Painter" pageId="3" pageNumber="2" pagination="1 - 102" refId="ref3417" refString="Luginbill, P., and H. R. Painter. 1953. May beetles of the United States and Canada. United States Department of Agriculture Technical Bulletin 1060: 1 - 102." type="journal article" year="1953">Luginbill and Painter (1953)</bibRefCitation>
in considering it a deformed specimen of
<taxonomicName id="4C404D12FFF20A33FD69C545FCC3FC5B" authority="(Melsheimer, 1845)" baseAuthorityName="Melsheimer" baseAuthorityYear="1845" box="[762,875,881,904]" class="Insecta" family="Melolonthidae" genus="Ancylonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B934EA83FFF20A33FD69C545FCC3FC5B" box="[762,875,881,904]" italics="true" pageId="3" pageNumber="2">P. rugosa</emphasis>
</taxonomicName>
and here formally synonymize the two species.
</paragraph>
<paragraph id="8BFF3691FFF20A33FF7BC5A5FBD7FBFA" blockId="3.[192,1440,496,1065]" pageId="3" pageNumber="2">
A search for Melsheimers type of
<taxonomicName id="4C404D12FFF20A33FD13C5A5FD5AFC7B" authority="Melsheimer 1845: 140" authorityName="Melsheimer" authorityPageNumber="140" authorityYear="1845" box="[640,754,913,936]" class="Insecta" family="Melolonthidae" genus="Ancylonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B934EA83FFF20A33FD13C5A5FD3BFC7B" box="[640,659,913,936]" italics="true" pageId="3" pageNumber="2">A</emphasis>
.
<emphasis id="B934EA83FFF20A33FD33C5A5FD5AFC7B" box="[672,754,913,936]" italics="true" pageId="3" pageNumber="2">rugosa</emphasis>
</taxonomicName>
turned up no specimens and is presumed lost. A
<typeStatus id="54FB8833FFF20A33FAD3C5A5FA08FC7A" box="[1344,1440,913,937]" pageId="3" pageNumber="2" type="neotype">neotype</typeStatus>
of
<taxonomicName id="4C404D12FFF20A33FF4EC585FEF8FC1B" authority="(Melsheimer, 1845)" baseAuthorityName="Melsheimer" baseAuthorityYear="1845" box="[221,336,945,968]" class="Insecta" family="Melolonthidae" genus="Ancylonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B934EA83FFF20A33FF4EC585FF58FC1B" box="[221,240,945,968]" italics="true" pageId="3" pageNumber="2">A</emphasis>
.
<emphasis id="B934EA83FFF20A33FF6DC585FEF8FC1B" box="[254,336,945,968]" italics="true" pageId="3" pageNumber="2">rugosa</emphasis>
</taxonomicName>
is here designated to fix the name to a single specimen to allow comparison to other types. The specimen chosen is a dissected male (
<figureCitation id="137B2A14FFF20A33FD26C5E5FD50FC3A" box="[693,760,977,1001]" captionStart="Figures 16" captionStartId="6.[192,287,1726,1748]" captionTargetBox="[369,1328,213,1651]" captionTargetId="figure@6.[369,1328,213,1651]" captionTargetPageId="6" captionText="Figures 16. Male genitalia of Phyllophaga type specimens. 1) Phyllophaga sociata, lectotype. 2) Phyllophaga knausii, lectotype. 3) Phyllophaga rugosa, neotype. 4) Phyllophaga chippewa, holotype. 5) Phyllophaga bipartita, lectotype. 6) Phyllophaga falta, holotype." figureDoi="http://doi.org/10.5281/zenodo.3676642" httpUri="https://zenodo.org/record/3676642/files/figure.png" pageId="3" pageNumber="2">Fig. 3</figureCitation>
) from
<collectingRegion id="4984F873FFF20A33FCD4C5E5FBA9FC3A" box="[839,1025,977,1001]" country="United States of America" name="North Carolina" pageId="3" pageNumber="2">North Carolina</collectingRegion>
in the Horn Collection. Label data (
<figureCitation id="137B2A14FFF20A33FF5AC5C5FEB8FBDA" box="[201,272,1009,1033]" captionStart="Figures 712" captionStartId="7.[192,287,1663,1685]" captionTargetBox="[192,1445,213,1629]" captionTargetId="figure@7.[192,1445,213,1629]" captionTargetPageId="7" captionText="Figures 712. Specimen labels of Phyllophaga type specimens. 7) Phyllophaga sociata, lectotype. 8) Phyllophaga knausii, lectotype. 9) Phyllophaga rugosa, neotype. 10) Phyllophaga chippewa, holotype. 11) Phyllophaga bipartita, lectotype. 12) Phyllophaga falta, holotype." figureDoi="http://doi.org/10.5281/zenodo.3676644" httpUri="https://zenodo.org/record/3676644/files/figure.png" pageId="3" pageNumber="2">Fig. 9</figureCitation>
) are as follows: “N. C //
<specimenCount id="9D46FD18FFF20A33FDADC5C5FDFAFBDB" box="[574,594,1009,1032]" pageId="3" pageNumber="2" type="male"></specimenCount>
// HornColl/ H [handwritten] 5774 // MCZ-ENT 00711292 // [on red]
<typeStatus id="54FB8833FFF20A33FF53C225FEEDFBFA" box="[192,325,1041,1065]" pageId="3" pageNumber="2" type="neotype">NEOTYPE</typeStatus>
/
<taxonomicName id="4C404D12FFF20A33FEC6C224FDE1FBFB" authority="(Melsheimer, 1845)" baseAuthorityName="Melsheimer" baseAuthorityYear="1845" box="[341,585,1040,1064]" class="Insecta" family="Melolonthidae" genus="Ancylonycha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="B934EA83FFF20A33FEC6C224FDE1FBFB" box="[341,585,1040,1064]" italics="true" pageId="3" pageNumber="2">Ancylonycha rugosa</emphasis>
</taxonomicName>
/
<bibRefCitation id="EFD14B60FFF20A33FDCAC225FC9AFBFA" author="Melsheimer, F. E." box="[601,818,1041,1065]" pageId="3" pageNumber="2" pagination="134 - 160" refId="ref3453" refString="Melsheimer, F. E. 1845. Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia 2: 134 - 160." type="journal article" year="1845">Melsheimer, 1845</bibRefCitation>
/ Det: K. E. Schnepp 2018”.
</paragraph>
</subSubSection>
</treatment>
</document>