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<mods:title id="672B25A7E1D700BC7BDE91AA2E541C61">Molecular phylogenetics, systematics and host-plant associations of the Bruchidius albosparsus (Fåhraeus) species group (Coleoptera, Chrysomelidae, Bruchinae) with the description of four new species</mods:title>
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<mods:namePart id="B7409988418216217CB9EDBCAC4BD16D">Ru, Bruno Le</mods:namePart>
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<treatment id="03DA87F3FFB6FFDB17D7FA23770FFE00" ID-DOI="http://doi.org/10.5281/zenodo.5629624" ID-GBIF-Taxon="119600264" ID-Zenodo-Dep="5629624" LSID="urn:lsid:plazi:treatment:03DA87F3FFB6FFDB17D7FA23770FFE00" httpUri="http://treatment.plazi.org/id/03DA87F3FFB6FFDB17D7FA23770FFE00" lastPageId="21" lastPageNumber="472" pageId="19" pageNumber="470">
<subSubSection id="C369656EFFB6FFDD17D7FA2374F4F9F7" box="[151,708,1513,1539]" pageId="19" pageNumber="470" type="nomenclature">
<paragraph id="8BCC36E5FFB6FFDD17D7FA2374F4F9F7" blockId="19.[151,708,1513,1539]" box="[151,708,1513,1539]" pageId="19" pageNumber="470">
<heading id="D0848189FFB6FFDD17D7FA2374F4F9F7" bold="true" box="[151,708,1513,1539]" fontSize="11" level="1" pageId="19" pageNumber="470" reason="1">
<taxonomicName id="4C734D66FFB6FFDD17D7FA2374F4F9F7" authority="Anton &amp; Delobel, 2003" authorityName="Anton &amp; Delobel" authorityYear="2003" box="[151,708,1513,1539]" class="Insecta" family="Chrysomelidae" genus="Bruchidius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="raddianae">
<emphasis id="B907EAF7FFB6FFDD17D7FA2374F4F9F7" bold="true" box="[151,708,1513,1539]" pageId="19" pageNumber="470">
<emphasis id="B907EAF7FFB6FFDD17D7FA2377AFF9F7" bold="true" box="[151,415,1513,1539]" italics="true" pageId="19" pageNumber="470">Bruchidius raddianae</emphasis>
<bibRefCitation id="EFE24B14FFB6FFDD16E7FA2374F4F9F7" author="Anton" box="[423,708,1513,1539]" pageId="19" pageNumber="470" refString="Anton, K. - W. &amp; Delobel, A. (2003) African species of the Bruchidius centromaculatus group with &quot; eyed &quot; female pygidium (Coleoptera: Bruchidae: Bruchinae). Genus - International Journal of Invertebrate Taxonomy, 14, 159 - 190." type="journal article" year="2003">Anton &amp; Delobel, 2003</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369656EFFB6FFDD17D7F9FA74A7F990" pageId="19" pageNumber="470" type="reference_group">
<paragraph id="8BCC36E5FFB6FFDD17D7F9FA74A7F990" blockId="19.[151,678,1584,1636]" pageId="19" pageNumber="470">
<treatmentCitationGroup id="AB6311CBFFB6FFDD17D7F9FA74A7F990" pageId="19" pageNumber="470">
<taxonomicName id="4C734D66FFB6FFDD17D7F9FA77CDF9B2" authority="Decelle, 1979" authorityName="Decelle" authorityYear="1979" box="[151,509,1584,1606]" class="Insecta" family="Chrysomelidae" genus="Bruchidius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="sahelicus">
<emphasis id="B907EAF7FFB6FFDD17D7F9FA775AF9B2" box="[151,362,1584,1606]" italics="true" pageId="19" pageNumber="470">Bruchidius sahelicus</emphasis>
<bibRefCitation id="EFE24B14FFB6FFDD1631F9FA77CDF9B2" author="Decelle" box="[369,509,1584,1606]" pageId="19" pageNumber="470" refString="Decelle, J. (1979) Insects of Saudi Arabia. Coleoptera: Fam. Bruchidae. Fauna of Saudi Arabia, 1, 318 - 330." type="journal article" year="1979">Decelle, 1979</bibRefCitation>
</taxonomicName>
(nomen nudum)
<taxonomicName id="4C734D66FFB6FFDD17D7F98474A7F990" authority="Anton &amp; Delobel, 2003:178" authorityName="Anton &amp; Delobel" authorityPageNumber="178" authorityYear="2003" box="[151,663,1614,1636]" class="Insecta" family="Chrysomelidae" genus="Bruchidius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="raddianae">
<emphasis id="B907EAF7FFB6FFDD17D7F9847744F990" box="[151,372,1614,1636]" italics="true" pageId="19" pageNumber="470">Bruchidius raddianae</emphasis>
<treatmentCitation id="0AD210F4FFB6FFDD163BF98574A7F990" author="Anton" box="[379,663,1614,1636]" page="178" pageId="19" pageNumber="470" year="2003">
<bibRefCitation id="EFE24B14FFB6FFDD163BF985745DF990" author="Anton" box="[379,621,1614,1636]" pageId="19" pageNumber="470" refString="Anton, K. - W. &amp; Delobel, A. (2003) African species of the Bruchidius centromaculatus group with &quot; eyed &quot; female pygidium (Coleoptera: Bruchidae: Bruchinae). Genus - International Journal of Invertebrate Taxonomy, 14, 159 - 190." type="journal article" year="2003">Anton &amp; Delobel, 2003</bibRefCitation>
:178
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C369656EFFB6FFDB17D7F95B7720FEE9" lastPageId="21" lastPageNumber="472" pageId="19" pageNumber="470" type="materials_examined">
<paragraph id="8BCC36E5FFB6FFDA17D7F95B75CCFE59" blockId="19.[151,1437,1681,2030]" lastBlockId="20.[151,1437,151,573]" lastPageId="20" lastPageNumber="471" pageId="19" pageNumber="470">
<emphasis id="B907EAF7FFB6FFDD17D7F95B77B4F95E" bold="true" box="[151,388,1681,1706]" pageId="19" pageNumber="470">Material examined.</emphasis>
<collectingCountry id="F3647675FFB6FFDD16CEF95877E5F95E" box="[398,469,1682,1706]" name="Egypt" pageId="19" pageNumber="470">Egypt</collectingCountry>
:
<specimenCount id="9D75FD6CFFB6FFDD16A4F9587438F95E" box="[484,520,1682,1706]" pageId="19" pageNumber="470" type="generic">1♂</specimenCount>
, 2♀, Ras Mohamed,
<date id="FFCD1025FFB6FFDD1444F95B754CF95E" box="[772,892,1681,1706]" pageId="19" pageNumber="470" value="2001-04-25">25.iv.2001</date>
, ex
<taxonomicName id="4C734D66FFB6FFDD14F1F95B72D0F95E" box="[945,1248,1681,1706]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="subSpecies" species="tortilis" subSpecies="raddiana">
<emphasis id="B907EAF7FFB6FFDD14F1F95B72D0F95E" box="[945,1248,1681,1706]" italics="true" pageId="19" pageNumber="470">Vachellia tortilis raddiana</emphasis>
</taxonomicName>
[1♀ 02301] (G. Fédière) [
<collectionCode id="ED62AE20FFB6FFDD1641F97C7764F93A" box="[257,340,1718,1742]" pageId="19" pageNumber="470">CBAD</collectionCode>
,
<collectionCode id="ED62AE20FFB6FFDD161FF97C779EF93A" box="[351,430,1718,1742]" pageId="19" pageNumber="470">CBGP</collectionCode>
];
<specimenCount id="9D75FD6CFFB6FFDD1680F97C77D5F93A" box="[448,485,1718,1742]" pageId="19" pageNumber="470" type="generic">2♂</specimenCount>
, Aïn Sukna,
<date id="FFCD1025FFB6FFDD1534F97F74DFF93A" box="[628,751,1717,1742]" pageId="19" pageNumber="470" value="2001-06-15">15.vi.2001</date>
, ex
<taxonomicName id="4C734D66FFB6FFDD145AF97C7582F93A" box="[794,946,1717,1742]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="subSpecies" species="tortilis" subSpecies="raddiana">
<emphasis id="B907EAF7FFB6FFDD145AF97C7582F93A" box="[794,946,1717,1742]" italics="true" pageId="19" pageNumber="470">V. t. raddiana</emphasis>
</taxonomicName>
[
<specimenCount id="9D75FD6CFFB6FFDD14FFF97C75D4F93A" box="[959,996,1718,1742]" pageId="19" pageNumber="470" type="generic">2♂</specimenCount>
0 3701, 00503] (G. Fédière) [
<collectionCode id="ED62AE20FFB6FFDD1266F97C734CF93A" box="[1318,1404,1718,1742]" pageId="19" pageNumber="470">CBAD</collectionCode>
];
<specimenCount id="9D75FD6CFFB6FFDD12CEF97C769EF905" pageId="19" pageNumber="470" type="generic">3 ♂</specimenCount>
, 3♀, Karhur Tall,
<date id="FFCD1025FFB6FFDD163DF91377D2F906" box="[381,482,1753,1778]" pageId="19" pageNumber="470" value="2003-02-07">7.ii.2003</date>
, ex
<taxonomicName id="4C734D66FFB6FFDD1551F91074E0F906" box="[529,720,1753,1778]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="species" species="ehrenbergiana">
<emphasis id="B907EAF7FFB6FFDD1551F91074E0F906" box="[529,720,1753,1778]" italics="true" pageId="19" pageNumber="470">V. ehrenbergiana</emphasis>
</taxonomicName>
[1♀ 02203] (G. Fédière) [
<collectionCode id="ED62AE20FFB6FFDD14B6F910727CF906" box="[1014,1100,1754,1778]" pageId="19" pageNumber="470">CBAD</collectionCode>
];
<specimenCount id="9D75FD6CFFB6FFDD131FF91072B4F906" box="[1119,1156,1754,1778]" pageId="19" pageNumber="470" type="generic">1♂</specimenCount>
, 1♀, Assuan,
<date id="FFCD1025FFB6FFDD1263F91373A8F906" box="[1315,1432,1753,1778]" pageId="19" pageNumber="470" value="2002-07-03">3.vii.2002</date>
, ex
<taxonomicName id="4C734D66FFB6FFDD17F8F9347799F8E2" authority="G. Fediere" authorityName="G. Fediere" box="[184,425,1789,1814]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="species" species="tortilis">
<emphasis id="B907EAF7FFB6FFDD17F8F934772EF8E2" box="[184,286,1789,1814]" italics="true" pageId="19" pageNumber="470">V. tortilis</emphasis>
(G. Fédière)
</taxonomicName>
[
<collectionCode id="ED62AE20FFB6FFDD16F7F934743DF8E2" box="[439,525,1790,1814]" pageId="19" pageNumber="470">CBAD</collectionCode>
].
<collectingCountry id="F3647675FFB6FFDD155EF9347460F8E1" box="[542,592,1790,1813]" name="Iran" pageId="19" pageNumber="470">Iran</collectingCountry>
:
<specimenCount id="9D75FD6CFFB6FFDD151AF934744EF8E2" box="[602,638,1790,1814]" pageId="19" pageNumber="470" type="generic">1♂</specimenCount>
, Bushehr, ex
<taxonomicName id="4C734D66FFB6FFDD1455F9347598F8E2" box="[789,936,1790,1814]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="species" species="farnesiana">
<emphasis id="B907EAF7FFB6FFDD1455F9347598F8E2" box="[789,936,1790,1814]" italics="true" pageId="19" pageNumber="470">V. farnesiana</emphasis>
</taxonomicName>
[
<specimenCount id="9D75FD6CFFB6FFDD14F7F93475ECF8E2" box="[951,988,1790,1814]" pageId="19" pageNumber="470" type="generic">1♂</specimenCount>
0 0 611, specimen GK335 used for
<collectionCode id="ED62AE20FFB6FFDD121FF93473ACF8E2" LSID="urn:lsid:biocol.org:col:15536" box="[1375,1436,1790,1814]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="19" pageNumber="470">DNA</collectionCode>
extraction] (L. Abbaszadeh) [
<collectionCode id="ED62AE20FFB6FFDD16A8F8E87409F8CE" box="[488,569,1826,1850]" pageId="19" pageNumber="470">CBGP</collectionCode>
];
<specimenCount id="9D75FD6CFFB6FFDD150EF8E87443F8CE" box="[590,627,1826,1850]" pageId="19" pageNumber="470" type="generic">1♂</specimenCount>
, 1♀, Bushehr,
<date id="FFCD1025FFB6FFDD1463F8E875A4F8CE" box="[803,916,1826,1850]" pageId="19" pageNumber="470" value="2012-05-30">30.v.2012</date>
, ex
<taxonomicName id="4C734D66FFB6FFDD1486F8EB726DF8CE" box="[966,1117,1825,1850]" class="Magnoliopsida" family="Achariaceae" genus="Hydnocarpus" kingdom="Plantae" order="Malpighiales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="genus">
<emphasis id="B907EAF7FFB6FFDD1486F8EB726DF8CE" box="[966,1117,1825,1850]" italics="true" pageId="19" pageNumber="470">Hydnocarpus</emphasis>
</taxonomicName>
sp. [
<specimenCount id="9D75FD6CFFB6FFDD13D9F8E8728DF8CE" box="[1177,1213,1826,1850]" pageId="19" pageNumber="470" type="generic">1♂</specimenCount>
05713] (N. Farrar) [
<collectionCode id="ED62AE20FFB6FFDD17DEF88C76C4F8AA" box="[158,244,1862,1886]" pageId="19" pageNumber="470">CBAD</collectionCode>
].
<collectingCountry id="F3647675FFB6FFDD164AF88F777DF8AA" box="[266,333,1861,1886]" name="Israel" pageId="19" pageNumber="470">Israel</collectingCountry>
:
<specimenCount id="9D75FD6CFFB6FFDD161CF88C77B0F8AA" box="[348,384,1862,1886]" pageId="19" pageNumber="470" type="generic">1♂</specimenCount>
, Negev,
<date id="FFCD1025FFB6FFDD16ACF88F7472F8AA" box="[492,578,1861,1886]" pageId="19" pageNumber="470" value="1996-06">vi.1996</date>
, ex
<taxonomicName id="4C734D66FFB6FFDD1539F88C752AF8AA" box="[633,794,1861,1886]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="subSpecies" species="tortilis" subSpecies="raddiana">
<emphasis id="B907EAF7FFB6FFDD1539F88C752AF8AA" box="[633,794,1861,1886]" italics="true" pageId="19" pageNumber="470">V. t. raddiana</emphasis>
</taxonomicName>
[specimen Bd11 used for
<collectionCode id="ED62AE20FFB6FFDD1319F88C72A6F8AA" LSID="urn:lsid:biocol.org:col:15536" box="[1113,1174,1862,1886]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="19" pageNumber="470">DNA</collectionCode>
extraction] (D. Ward) [
<collectionCode id="ED62AE20FFB6FFDD17DEF8A076DFF876" box="[158,239,1898,1922]" pageId="19" pageNumber="470">CBGP</collectionCode>
];
<specimenCount id="9D75FD6CFFB6FFDD1642F8A07717F876" box="[258,295,1898,1922]" pageId="19" pageNumber="470" type="generic">1♂</specimenCount>
, 1♀, Wadi Bitaron,
<date id="FFCD1025FFB6FFDD154BF8A37452F876" box="[523,610,1897,1922]" pageId="19" pageNumber="470" value="1998-06">vi.1998</date>
, ex
<taxonomicName id="4C734D66FFB6FFDD15D0F8A075B9F876" authority="K. Or." authorityName="K. Or." box="[656,905,1897,1922]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="subSpecies" species="tortilis" subSpecies="raddiana">
<emphasis id="B907EAF7FFB6FFDD15D0F8A07519F876" box="[656,809,1897,1922]" italics="true" pageId="19" pageNumber="470">V. t. raddiana</emphasis>
(K. Or.)
</taxonomicName>
[
<collectionCode id="ED62AE20FFB6FFDD14D8F8A075DEF876" box="[920,1006,1898,1922]" pageId="19" pageNumber="470">CBAD</collectionCode>
];
<specimenCount id="9D75FD6CFFB6FFDD1342F8A37216F876" box="[1026,1062,1897,1922]" pageId="19" pageNumber="470" type="generic">5♂</specimenCount>
, 5♀, Wadi Saif,
<date id="FFCD1025FFB6FFDD13A3F8A37372F876" box="[1251,1346,1897,1922]" pageId="19" pageNumber="470" value="1998-07">vii.1998</date>
, ex
<taxonomicName id="4C734D66FFB6FFDD1231F8A0775BF852" authority="K. Or." authorityName="K. Or." class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="subSpecies" species="tortilis" subSpecies="raddiana">
<emphasis id="B907EAF7FFB6FFDD1231F8A076CEF852" italics="true" pageId="19" pageNumber="470">V. t. raddiana</emphasis>
, (K. Or.)
</taxonomicName>
[
<collectionCode id="ED62AE20FFB6FFDD163DF84477E3F852" box="[381,467,1934,1958]" pageId="19" pageNumber="470">CBAD</collectionCode>
].
<collectingCountry id="F3647675FFB6FFDD16A8F8447462F851" box="[488,594,1934,1957]" name="Morocco" pageId="19" pageNumber="470">Morocco</collectingCountry>
: 1♀, Vallée du Agdz,
<date id="FFCD1025FFB6FFDD141BF84475F9F852" box="[859,969,1934,1958]" pageId="19" pageNumber="470" value="2003-05-24">24.v.2003</date>
(A. Jaeger) [
<collectionCode id="ED62AE20FFB6FFDD1326F844728CF852" box="[1126,1212,1934,1958]" pageId="19" pageNumber="470">CBAD</collectionCode>
].
<taxonomicName id="4C734D66FFB6FFDD1392F847731FF852" box="[1234,1327,1933,1958]" class="Magnoliopsida" family="Fabaceae" genus="Senegalia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="477" phylum="Tracheophyta" rank="species" species="senegal">
<collectingCountry id="F3647675FFB6FFDD1392F847731FF852" box="[1234,1327,1933,1958]" name="Senegal" pageId="19" pageNumber="470">Senegal</collectingCountry>
</taxonomicName>
:
<specimenCount id="9D75FD6CFFB6FFDD127DF8447352F852" box="[1341,1378,1934,1958]" pageId="19" pageNumber="470" type="generic">4♂</specimenCount>
, 2♀, Khatali,
<date id="FFCD1025FFB6FFDD17BBF87B7743F83E" box="[251,371,1969,1994]" pageId="19" pageNumber="470" value="1995-04-29">29.iv.1995</date>
, ex
<taxonomicName id="4C734D66FFB6FFDD16E8F8787477F83E" box="[424,583,1969,1994]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="subSpecies" species="tortilis" subSpecies="raddiana">
<emphasis id="B907EAF7FFB6FFDD16E8F8787477F83E" box="[424,583,1969,1994]" italics="true" pageId="19" pageNumber="470">V. t. raddiana</emphasis>
</taxonomicName>
[
<specimenCount id="9D75FD6CFFB6FFDD1519F878744DF83E" box="[601,637,1970,1994]" pageId="19" pageNumber="470" type="generic">4♂</specimenCount>
11295, 12602, 12702, 12802, 2♀ 11695, 12695] (A. &amp; H. Delobel) [
<collectionCode id="ED62AE20FFB6FFDD17DEF81C76C4F81A" box="[158,244,2006,2030]" pageId="19" pageNumber="470">CBAD</collectionCode>
]; Dahra,
<date id="FFCD1025FFB6FFDD1619F81C7796F81A" box="[345,422,2006,2030]" pageId="19" pageNumber="470" value="1998-05">v.1998</date>
, ex
<taxonomicName id="4C734D66FFB6FFDD1694F81C745DF81A" box="[468,621,2005,2030]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="470" phylum="Tracheophyta" rank="subSpecies" species="tortilis" subSpecies="raddiana">
<emphasis id="B907EAF7FFB6FFDD1694F81C745DF81A" box="[468,621,2005,2030]" italics="true" pageId="19" pageNumber="470">V. t. raddiana</emphasis>
</taxonomicName>
[2♀ 0 2700, 02800] (A. &amp; H. Delobel) [
<collectionCode id="ED62AE20FFB6FFDD136FF81C72B5F81A" box="[1071,1157,2006,2030]" pageId="19" pageNumber="470">CBAD</collectionCode>
];
<specimenCount id="9D75FD6CFFB6FFDD13D7F81C728CF81A" box="[1175,1212,2006,2030]" pageId="19" pageNumber="470" type="generic">1♂</specimenCount>
, 1♀, Boké Namadi,
<date id="FFCD1025FFB1FFDA17D7FF5D7733FF44" box="[151,259,151,176]" pageId="20" pageNumber="471" value="1999-01-30">30.i.1999</date>
, ex
<taxonomicName id="4C734D66FFB1FFDA167CFF5377D1FF44" box="[316,481,151,176]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="20" pageNumber="471" phylum="Tracheophyta" rank="subSpecies" species="tortilis" subSpecies="raddiana">
<emphasis id="B907EAF7FFB1FFDA167CFF5377D1FF44" box="[316,481,151,176]" italics="true" pageId="20" pageNumber="471">V. t. raddiana</emphasis>
</taxonomicName>
[specimen R11 used for
<collectionCode id="ED62AE20FFB1FFDA1457FF527564FF44" LSID="urn:lsid:biocol.org:col:15536" box="[791,852,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="20" pageNumber="471">DNA</collectionCode>
extraction] (M. Sembène) [
<collectionCode id="ED62AE20FFB1FFDA13E1FF5272C3FF44" box="[1185,1267,152,176]" pageId="20" pageNumber="471">CBGP</collectionCode>
]. United Arab Republic:
<specimenCount id="9D75FD6CFFB1FFDA1650FF767705FF20" box="[272,309,188,212]" pageId="20" pageNumber="471" type="generic">1♂</specimenCount>
, Wadi Safad,
<date id="FFCD1025FFB1FFDA16A2FF7674A5FF21" box="[482,661,188,213]" pageId="20" pageNumber="471" value="2006-01-31" valueMax="2006-02-21" valueMin="2006-01-31">31.i.21.ii.2006</date>
, light trap [
<specimenCount id="9D75FD6CFFB1FFDA1467FF76757BFF20" box="[807,843,188,212]" pageId="20" pageNumber="471" type="generic">1♂</specimenCount>
05308] (A. Van Harten) [
<collectionCode id="ED62AE20FFB1FFDA13C6FF7672ECFF20" box="[1158,1244,188,212]" pageId="20" pageNumber="471">CBAD</collectionCode>
];
<specimenCount id="9D75FD6CFFB1FFDA13B4FF767329FF20" box="[1268,1305,188,212]" pageId="20" pageNumber="471" type="generic">1♂</specimenCount>
, 1♀, Wadi Maidaq,
<date id="FFCD1025FFB1FFDA17BBFF15774BFF0C" box="[251,379,223,248]" pageId="20" pageNumber="471" value="2005-12-21">21.xii.2005</date>
<date id="FFCD1025FFB1FFDA16CAFF1577C7FF0C" box="[394,503,223,248]" pageId="20" pageNumber="471" value="2006-03-02">2.iii.2006</date>
, light trap (A. Van Harten) [
<collectionCode id="ED62AE20FFB1FFDA1473FF2A75B9FF0C" box="[819,905,224,248]" pageId="20" pageNumber="471">CBAD</collectionCode>
]; 2♀, same data,
<date id="FFCD1025FFB1FFDA130CFF1572D0FF0C" box="[1100,1248,223,248]" pageId="20" pageNumber="471" value="2006-01-14" valueMax="2006-01-25" valueMin="2006-01-14">1425.i.2006</date>
(A. Van Harten) [
<collectionCode id="ED62AE20FFB1FFDA17DEFECE76C4FEE8" box="[158,244,260,284]" pageId="20" pageNumber="471">CBAD</collectionCode>
];
<specimenCount id="9D75FD6CFFB1FFDA1648FECE771DFEE8" box="[264,301,260,284]" pageId="20" pageNumber="471" type="generic">1♂</specimenCount>
, 2♀, Al-Ajban,
<date id="FFCD1025FFB1FFDA16A9FECE74ABFEE9" box="[489,667,260,285]" pageId="20" pageNumber="471" value="2005-10-22" valueMax="2005-11-09" valueMin="2005-10-22">22.x.9.xi.2005</date>
, Malaise trap (A. Van Harten) [
<collectionCode id="ED62AE20FFB1FFDA1348FECE726EFEE8" box="[1032,1118,260,284]" pageId="20" pageNumber="471">CBAD</collectionCode>
];
<specimenCount id="9D75FD6CFFB1FFDA1333FECE72A7FEE8" box="[1139,1175,260,284]" pageId="20" pageNumber="471" type="generic">1♂</specimenCount>
, 1♀,
<date id="FFCD1025FFB1FFDA1399FECE73A7FEE9" box="[1241,1431,260,285]" pageId="20" pageNumber="471" value="2006-05-27" valueMax="2006-06-26" valueMin="2006-05-27">27.v.26.vi.2006</date>
, light trap (A. Van Harten) [
<collectionCode id="ED62AE20FFB1FFDA1688FEE2742EFEB4" box="[456,542,296,320]" pageId="20" pageNumber="471">CBAD</collectionCode>
];
<specimenCount id="9D75FD6CFFB1FFDA1571FEE27465FEB4" box="[561,597,296,320]" pageId="20" pageNumber="471" type="generic">1♂</specimenCount>
, 1♀, Khor-Al-Kwair,
<date id="FFCD1025FFB1FFDA140FFEED75CEFEB4" box="[847,1022,295,320]" pageId="20" pageNumber="471" value="2007-04-24" valueMax="2007-05-02" valueMin="2007-04-24">24.iv.2.v.2007</date>
, light trap (A. Van Harten) [
<collectionCode id="ED62AE20FFB1FFDA127BFEE273A1FEB4" box="[1339,1425,296,320]" pageId="20" pageNumber="471">CBAD</collectionCode>
];
<specimenCount id="9D75FD6CFFB1FFDA17D7FE86768BFE90" box="[151,187,332,356]" pageId="20" pageNumber="471" type="generic">1♂</specimenCount>
, Hatta,
<date id="FFCD1025FFB1FFDA1659FE867784FE91" box="[281,436,332,357]" pageId="20" pageNumber="471" value="2006-04-04" valueMax="2006-04-11" valueMin="2006-04-04">4.11.iv.2006</date>
, light trap (A. Van Harten) [
<collectionCode id="ED62AE20FFB1FFDA1442FE867568FE90" box="[770,856,332,356]" pageId="20" pageNumber="471">CBAD</collectionCode>
];
<specimenCount id="9D75FD6CFFB1FFDA142EFE8675A3FE90" box="[878,915,332,356]" pageId="20" pageNumber="471" type="generic">1♂</specimenCount>
, Mahafiz,
<date id="FFCD1025FFB1FFDA1352FE867284FE91" box="[1042,1204,332,357]" pageId="20" pageNumber="471" value="2006-04-19" valueMax="2006-04-26" valueMin="2006-04-19">1926.iv.2006</date>
, water trap (A. Van Harten) [
<collectionCode id="ED62AE20FFB1FFDA17BAFEBA7760FE7C" box="[250,336,368,392]" pageId="20" pageNumber="471">CBAD</collectionCode>
]. Other material as listed in
<bibRefCitation id="EFE24B14FFB1FFDA15CAFEBA7591FE7C" author="Anton" box="[650,929,367,392]" pageId="20" pageNumber="471" refString="Anton, K. - W. &amp; Delobel, A. (2003) African species of the Bruchidius centromaculatus group with &quot; eyed &quot; female pygidium (Coleoptera: Bruchidae: Bruchinae). Genus - International Journal of Invertebrate Taxonomy, 14, 159 - 190." type="journal article" year="2003">Anton &amp; Delobel (2003)</bibRefCitation>
, including specimens from
<collectingCountry id="F3647675FFB1FFDA1397FEA5731EFE7C" box="[1239,1326,367,392]" name="Algeria" pageId="20" pageNumber="471">Algeria</collectingCountry>
, Burkina- Faso,
<collectingCountry id="F3647675FFB1FFDA179AFE5E7726FE59" box="[218,278,404,429]" name="India" pageId="20" pageNumber="471">India</collectingCountry>
,
<collectingCountry id="F3647675FFB1FFDA1661FE5E775EFE59" box="[289,366,404,429]" name="Jordan" pageId="20" pageNumber="471">Jordan</collectingCountry>
, Lybia,
<collectingCountry id="F3647675FFB1FFDA1688FE5E7430FE59" box="[456,512,404,429]" name="Serbia" pageId="20" pageNumber="471">Mali</collectingCountry>
,
<collectingCountry id="F3647675FFB1FFDA154BFE5E747CFE59" box="[523,588,404,429]" name="Niger" pageId="20" pageNumber="471">Niger</collectingCountry>
,
<collectingCountry id="F3647675FFB1FFDA1517FE5E74AEFE58" box="[599,670,404,428]" name="Oman" pageId="20" pageNumber="471">Oman</collectingCountry>
,
<collectingCountry id="F3647675FFB1FFDA15EAFE5E752AFE59" box="[682,794,404,429]" name="Sri Lanka" pageId="20" pageNumber="471">Sri Lanka</collectingCountry>
,
<collectingCountry id="F3647675FFB1FFDA1465FE5E755BFE59" box="[805,875,404,429]" name="Sudan" pageId="20" pageNumber="471">Sudan</collectingCountry>
and
<collectingCountry id="F3647675FFB1FFDA14E2FE5E75C9FE59" box="[930,1017,404,429]" name="Tunisia" pageId="20" pageNumber="471">Tunisia</collectingCountry>
.
</paragraph>
<paragraph id="8BCC36E5FFB1FFDA1787FE7275AAFDC9" blockId="20.[151,1437,151,573]" pageId="20" pageNumber="471">
A small to medium-sized (
<quantity id="4C8B9B00FFB1FFDA16B6FE7274B1FE24" box="[502,641,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.7" metricValueMin="1.4" pageId="20" pageNumber="471" unit="mm" value="2.05" valueMax="2.7" valueMin="1.4">1.42.7 mm</quantity>
) species, from light yellowish to checkered with blackish and whitish spots, elytra with double-toothed protuberance at base of striae 34, and a sharp isolated tooth at stria 2. Other distinctive morphological traits: in male, a large circular spot of short erect setae in basal angle of first ventrite; last visible tergite of female with pair of small foveae with shiny margin.
</paragraph>
<caption id="DF0C666DFFB1FFDA17D7F8527328F85A" box="[151,1304,1944,1966]" httpUri="https://zenodo.org/record/244573/files/figure.png" pageId="20" pageNumber="471" targetBox="[462,1133,647,1898]" targetPageId="20">
<paragraph id="8BCC36E5FFB1FFDA17D7F8527328F85A" blockId="20.[151,1304,1944,1966]" box="[151,1304,1944,1966]" pageId="20" pageNumber="471">
<emphasis id="B907EAF7FFB1FFDA17D7F8527763F85A" bold="true" box="[151,339,1944,1966]" pageId="20" pageNumber="471">FIGURES 2526.</emphasis>
<taxonomicName id="4C734D66FFB1FFDA161AF8527418F85A" box="[346,552,1944,1966]" class="Insecta" family="Chrysomelidae" genus="Bruchidius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="uberatus">
<emphasis id="B907EAF7FFB1FFDA161AF8527418F85A" box="[346,552,1944,1966]" italics="true" pageId="20" pageNumber="471">Bruchidius uberatus</emphasis>
</taxonomicName>
(male): 25—median lobe (ventral view); 26—lateral lobes (ventral view).
</paragraph>
</caption>
<paragraph id="8BCC36E5FFB0FFDB1787FF5D7720FEE9" blockId="21.[151,1437,151,501]" pageId="21" pageNumber="472">
Genitalia [see
<figureCitation id="13482A60FFB0FFDB162DFF5D77C7FF44" box="[365,503,151,176]" captionStart-0="FIGURES 25 26" captionStart-1="FIGURE 27" captionStart-2="FIGURE 28" captionStartId-0="20.[151,264,1944,1966]" captionStartId-1="23.[151,250,1858,1880]" captionStartId-2="25.[151,250,1286,1308]" captionTargetBox-0="[462,1133,647,1898]" captionTargetBox-1="[206,1373,423,1829]" captionTargetBox-2="[171,1420,719,1244]" captionTargetId-0="figure@20.[434,1154,625,1923]" captionTargetId-1="figure@23.[194,1392,411,1836]" captionTargetId-2="figure@25.[150,1436,697,1264]" captionTargetPageId-0="20" captionTargetPageId-1="23" captionTargetPageId-2="25" captionText-0="FIGURES 25 26. Bruchidius uberatus (male): 25 — median lobe (ventral view); 26 — lateral lobes (ventral view)." captionText-1="FIGURE 27. Results of molecular phylogenetic analyses. On the left side the best-fit maximum likelihood (ML) tree is figured. Support values are only provided for nodes leading to species (BV &lt;50 % are not figured). On the right side the majority-rule consensus topology from Bayesian inference (BI) analyses is figured. Support values are only provided for nodes leading to species (PP &lt;50 % are not figured). Representatives of the Bruchidius albosparsus species group are highlighted using a red frame." captionText-2="FIGURE 28. Color variation in Bruchidius grandemaculatus adults (a, b—males; c, d—females)." httpUri-0="https://zenodo.org/record/244573/files/figure.png" httpUri-1="https://zenodo.org/record/244574/files/figure.png" httpUri-2="https://zenodo.org/record/244575/files/figure.png" pageId="21" pageNumber="472">Figs. 2528</figureCitation>
, p.
<quantity id="4C8B9B00FFB0FFDB1556FF527452FF44" box="[534,610,151,176]" metricMagnitude="0" metricUnit="m" metricValue="4.8006" pageId="21" pageNumber="472" unit="in" value="189.0">189 in</quantity>
<bibRefCitation id="EFE24B14FFB0FFDB152BFF52754AFF44" author="Anton" box="[619,890,151,176]" pageId="21" pageNumber="472" refString="Anton, K. - W. &amp; Delobel, A. (2003) African species of the Bruchidius centromaculatus group with &quot; eyed &quot; female pygidium (Coleoptera: Bruchidae: Bruchinae). Genus - International Journal of Invertebrate Taxonomy, 14, 159 - 190." type="journal article" year="2003">Anton &amp; Delobel 2003</bibRefCitation>
]: Internal sac entirely lined with small broad or slim based spines, and 36 large denticles with base as long as point; lateral lobes separated to about 80% their length; tegminal strut with very small carina. In female, vagina long, no dorsal sclerite at entrance of bursa copulatrix.
</paragraph>
</subSubSection>
<subSubSection id="C369656EFFB0FFDB1787FEED72C2FE7C" pageId="21" pageNumber="472" type="biology_ecology">
<paragraph id="8BCC36E5FFB0FFDB1787FEED72C2FE7C" blockId="21.[151,1437,151,501]" pageId="21" pageNumber="472">
<emphasis id="B907EAF7FFB0FFDB1787FEED7717FEB4" bold="true" box="[199,295,295,320]" pageId="21" pageNumber="472">Biology.</emphasis>
Reared from seeds of
<emphasis id="B907EAF7FFB0FFDB1563FEED7543FEB4" box="[547,883,295,320]" italics="true" pageId="21" pageNumber="472">
<taxonomicName id="4C734D66FFB0FFDB1563FEED753DFEB4" box="[547,781,295,320]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="species" species="farnesiana">Vachellia farnesiana</taxonomicName>
,
<taxonomicName id="4C734D66FFB0FFDB1458FEE3755EFEB4" box="[792,878,295,320]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="species" species="flava">V. flava</taxonomicName>
,
</emphasis>
<taxonomicName id="4C734D66FFB0FFDB1439FEE372D5FEB4" authority="(Zohary)" authorityName="(Zohary) Ragup. et al." baseAuthorityName="Zohary" box="[889,1253,295,320]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="subSpecies" species="gerrardii" subSpecies="negevensis">
<emphasis id="B907EAF7FFB0FFDB1439FEE3724AFEB4" box="[889,1146,295,320]" italics="true" pageId="21" pageNumber="472">V. gerrardii negevensis</emphasis>
(Zohary)
</taxonomicName>
Ragup. et al.,
<emphasis id="B907EAF7FFB0FFDB12C7FEE37591FE91" italics="true" pageId="21" pageNumber="472">
<taxonomicName id="4C734D66FFB0FFDB12C7FEE376EEFE91" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="species" species="hockii">V. hockii</taxonomicName>
,
<taxonomicName id="4C734D66FFB0FFDB17ABFE8777EBFE90" box="[235,475,332,357]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="subSpecies" species="nilotica" subSpecies="tomentosa">V. nilotica tomentosa</taxonomicName>
,
<taxonomicName id="4C734D66FFB0FFDB16A7FE877470FE91" box="[487,576,332,357]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="species" species="seyal">V. seyal</taxonomicName>
,
<taxonomicName id="4C734D66FFB0FFDB150DFE877487FE91" box="[589,695,332,357]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="species" species="tortilis">V. tortilis</taxonomicName>
,
<taxonomicName id="4C734D66FFB0FFDB1584FE8775ADFE91" box="[708,925,332,357]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="subSpecies" species="tortilis" subSpecies="raddiana">V. tortilis raddiana</taxonomicName>
,
</emphasis>
and an unidentified species of
<taxonomicName id="4C734D66FFB0FFDB1245FE867703FE7C" class="Magnoliopsida" family="Achariaceae" genus="Hydnocarpus" kingdom="Plantae" order="Malpighiales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="genus">
<emphasis id="B907EAF7FFB0FFDB1245FE8673ACFE91" box="[1285,1436,332,357]" italics="true" pageId="21" pageNumber="472">Hydnocarpus</emphasis>
(Achariaceae)
</taxonomicName>
.
<emphasis id="B907EAF7FFB0FFDB1602FEA57526FE7C" box="[322,790,367,392]" italics="true" pageId="21" pageNumber="472">
<taxonomicName id="4C734D66FFB0FFDB1602FEA57470FE7C" box="[322,576,367,392]" class="Magnoliopsida" family="Fabaceae" genus="Dichrostachys" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="species" species="cinerea">Dichrostachys cinerea</taxonomicName>
,
<taxonomicName id="4C734D66FFB0FFDB150AFEA57526FE7C" box="[586,790,367,392]" class="Magnoliopsida" family="Fabaceae" genus="Senegalia" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="species" species="senegal">
<collectingCountry id="F3647675FFB0FFDB150AFEA57488FE7C" box="[586,696,367,392]" name="Senegal" pageId="21" pageNumber="472">Senegalia</collectingCountry>
senegal
</taxonomicName>
</emphasis>
and
<taxonomicName id="4C734D66FFB0FFDB140CFEBB75EDFE7C" box="[844,989,367,392]" class="Magnoliopsida" family="Fabaceae" genus="Vachellia" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="472" phylum="Tracheophyta" rank="species" species="sieberiana">
<emphasis id="B907EAF7FFB0FFDB140CFEBB75EDFE7C" box="[844,989,367,392]" italics="true" pageId="21" pageNumber="472">V. sieberiana</emphasis>
</taxonomicName>
are other possible hosts.
</paragraph>
</subSubSection>
<subSubSection id="C369656EFFB0FFDB1787FE5E770FFE00" pageId="21" pageNumber="472" type="distribution">
<paragraph id="8BCC36E5FFB0FFDB1787FE5E770FFE00" blockId="21.[151,1437,151,501]" pageId="21" pageNumber="472">
<emphasis id="B907EAF7FFB0FFDB1787FE5E7750FE59" bold="true" box="[199,352,404,429]" pageId="21" pageNumber="472">Distribution.</emphasis>
<collectingCountry id="F3647675FFB0FFDB162BFE5E77F2FE59" box="[363,450,404,429]" name="Algeria" pageId="21" pageNumber="472">Algeria</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB1691FE5E745EFE58" box="[465,622,404,429]" name="Burkina Faso" pageId="21" pageNumber="472">Burkina Faso</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB153DFE5F74F3FE59" box="[637,707,405,429]" name="Egypt" pageId="21" pageNumber="472">Egypt</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB1592FE5E753EFE59" box="[722,782,404,429]" name="India" pageId="21" pageNumber="472">India</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB145DFE5F757DFE58" box="[797,845,405,428]" name="Iran" pageId="21" pageNumber="472">Iran</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB141CFE5E75ADFE59" box="[860,925,404,429]" name="Israel" pageId="21" pageNumber="472">Israel</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB14EDFE5E75CAFE59" box="[941,1018,404,429]" name="Jordan" pageId="21" pageNumber="472">Jordan</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB1349FE5E727DFE59" box="[1033,1101,404,429]" name="Libya" pageId="21" pageNumber="472">Libya</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB131CFE5E72A4FE59" box="[1116,1172,404,429]" name="Serbia" pageId="21" pageNumber="472">Mali</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB13E3FE5E7311FE59" box="[1187,1313,404,429]" name="Mauritania" pageId="21" pageNumber="472">Mauritania</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB1270FE5F73A8FE58" box="[1328,1432,405,428]" name="Morocco" pageId="21" pageNumber="472">Morocco</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB17D7FE7D76E9FE24" box="[151,217,439,464]" name="Niger" pageId="21" pageNumber="472">Niger</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB17A7FE72771EFE24" box="[231,302,440,464]" name="Oman" pageId="21" pageNumber="472">Oman</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB167DFE7D77E7FE24" box="[317,471,439,464]" name="Saudi Arabia" pageId="21" pageNumber="472">Saudi Arabia</collectingCountry>
,
<taxonomicName id="4C734D66FFB0FFDB16A6FE7D7471FE24" box="[486,577,439,464]" class="Magnoliopsida" family="Fabaceae" genus="Senegalia" kingdom="Plantae" order="Fabales" pageId="21" pageNumber="477" phylum="Tracheophyta" rank="species" species="senegal">
<collectingCountry id="F3647675FFB0FFDB16A6FE7D7471FE24" box="[486,577,439,464]" name="Senegal" pageId="21" pageNumber="472">Senegal</collectingCountry>
</taxonomicName>
,
<collectingCountry id="F3647675FFB0FFDB1511FE7D74F5FE24" box="[593,709,439,464]" name="Sri Lanka" pageId="21" pageNumber="472">Sri Lanka</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB1594FE7D752DFE24" box="[724,797,439,464]" name="Sudan" pageId="21" pageNumber="472">Sudan</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB146CFE7D75B2FE24" box="[812,898,439,464]" name="Tunisia" pageId="21" pageNumber="472">Tunisia</collectingCountry>
,
<collectingCountry id="F3647675FFB0FFDB14D1FE7D72A1FE24" box="[913,1169,439,464]" name="United Arab Emirates" pageId="21" pageNumber="472">United Arab Emirates</collectingCountry>
, and
<collectingCountry id="F3647675FFB0FFDB1393FE727311FE3B" box="[1235,1313,440,463]" name="Yemen" pageId="21" pageNumber="472">Yemen</collectingCountry>
(
<bibRefCitation id="EFE24B14FFB0FFDB1274FE727703FE00" author="Anton" pageId="21" pageNumber="472" refString="Anton, K. - W. &amp; Delobel, A. (2003) African species of the Bruchidius centromaculatus group with &quot; eyed &quot; female pygidium (Coleoptera: Bruchidae: Bruchinae). Genus - International Journal of Invertebrate Taxonomy, 14, 159 - 190." type="journal article" year="2003">Anton &amp; Delobel 2003</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>