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<document id="A30CC3545397A74B3D3CB4C18DE61685" ID-DOI="10.26028/cybium/2020-444-004" ID-ISSN="2101-0315" ID-Zenodo-Dep="12711981" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720646236220" checkinUser="felipe" docAuthor="Keith, Philippe &amp; Mennesson, Marion I." docDate="2020" docId="03DA87DB5118D622FF3DFBA615959736" docLanguage="en" docName="Cybium.44.4.331-349.pdf" docOrigin="Cybium 44 (4)" docSource="http://dx.doi.org/10.26028/cybium/2020-444-004" docStyle="DocumentStyle:EAB95795642928CB986F0923DE58F0B7.1:Cybium.2019-.journal_article" docStyleId="EAB95795642928CB986F0923DE58F0B7" docStyleName="Cybium.2019-.journal_article" docStyleVersion="1" docTitle="Ophiocara aporos subsp. rigonis , Whitley 1938" docType="treatment" docVersion="2" lastPageNumber="344" masterDocId="FFE3FFA35114D62FFFB9FFDC11319044" masterDocTitle="Review of Giuris (Teleostei: Eleotridae) from Indo-Pacific islands, with description of three new species" masterLastPageNumber="349" masterPageNumber="331" pageNumber="343" updateTime="1720650532356" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E318E89AA30ADEFF1757B8D0840778FD">Review of Giuris (Teleostei: Eleotridae) from Indo-Pacific islands, with description of three new species</mods:title>
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<mods:namePart id="8A0DA458971F38565F063BA0AE010449">Keith, Philippe</mods:namePart>
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<mods:namePart id="12E924444611220BB7DEC21DB24EDF9D">Mennesson, Marion I.</mods:namePart>
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<mods:identifier id="99220E647E143AD11F527E0ADBCF9305" type="ISSN">2101-0315</mods:identifier>
<mods:identifier id="994ED4507B5DFDA18F3FBB8989F31B0D" type="Zenodo-Dep">12711981</mods:identifier>
</mods:mods>
<treatment id="03DA87DB5118D622FF3DFBA615959736" LSID="urn:lsid:plazi:treatment:03DA87DB5118D622FF3DFBA615959736" httpUri="http://treatment.plazi.org/id/03DA87DB5118D622FF3DFBA615959736" lastPageId="13" lastPageNumber="344" pageId="12" pageNumber="343">
<subSubSection id="C36965465118D623FF3DFBA6130594D6" box="[132,564,1146,1170]" pageId="12" pageNumber="343" type="nomenclature">
<paragraph id="8BCC36CD5118D623FF3DFBA6130594D6" blockId="12.[132,564,1146,1170]" box="[132,564,1146,1170]" pageId="12" pageNumber="343">
<heading id="D08481A15118D623FF3DFBA6130594D6" box="[132,564,1146,1170]" fontSize="9" level="2" pageId="12" pageNumber="343" reason="5">
<taxonomicName id="4C734D4E5118D623FF3DFBA6130594D6" authority="Whitley, 1938" authorityName=", Whitley" authorityYear="1938" box="[132,564,1146,1170]" family="Eleotridae" genus="Ophiocara" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="343" phylum="Chordata" rank="subSpecies" species="aporos" subSpecies="rigonis">
<emphasis id="B907EADF5118D623FF3DFBA610A894D6" box="[132,409,1146,1170]" italics="true" pageId="12" pageNumber="343">Ophiocara aporos rigonis</emphasis>
Whitley, 1938
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36965465118D623FF3DFB66104596B6" pageId="12" pageNumber="343" type="materials_examined">
<paragraph id="8BCC36CD5118D623FF3DFB66106D9496" blockId="12.[132,775,1210,1779]" box="[132,348,1210,1234]" pageId="12" pageNumber="343">
<emphasis id="B907EADF5118D623FF3DFB66106D9496" bold="true" box="[132,348,1210,1234]" pageId="12" pageNumber="343">Material examined</emphasis>
</paragraph>
<paragraph id="8BCC36CD5118D623FF13FB06108F9556" blockId="12.[132,775,1210,1779]" pageId="12" pageNumber="343">
<materialsCitation id="3B1B3C905118D623FF13FB0610889556" country="Australia" location="Lillesmere Lagoon" municipality="Burdekin River" pageId="12" pageNumber="343" specimenCount="1" stateProvince="Queensland" typeStatus="syntype">
<emphasis id="B907EADF5118D623FF13FB0611CF94B5" box="[170,254,1242,1265]" italics="true" pageId="12" pageNumber="343">
<typeStatus id="54C8886F5118D623FF13FB0611CF94B5" box="[170,254,1242,1265]" pageId="12" pageNumber="343" type="syntype">Syntype</typeStatus>
</emphasis>
. AMS A.17837,
<location id="8EAC60165118D623FE70FB0613A994B7" LSID="urn:lsid:plazi:treatment:03DA87DB5118D622FF3DFBA615959736:8EAC60165118D623FE70FB0613A994B7" box="[457,664,1242,1267]" country="Australia" municipality="Burdekin River" name="Lillesmere Lagoon" pageId="12" pageNumber="343" stateProvince="Queensland">Lillesmere Lagoon</location>
,
<collectingMunicipality id="6BA8ACB75118D623FD1BFB0611F09556" pageId="12" pageNumber="343">Burdekin River</collectingMunicipality>
,
<collectingRegion id="49B7F82F5118D623FF72FB26107D9556" box="[203,332,1274,1298]" country="Australia" name="Queensland" pageId="12" pageNumber="343">Queensland</collectingRegion>
,
<collectingCountry id="F364765D5118D623FEEDFB2610889556" box="[340,441,1274,1298]" name="Australia" pageId="12" pageNumber="343">Australia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BCC36CD5118D623FF13FAC6104596B6" blockId="12.[132,775,1210,1779]" pageId="12" pageNumber="343">
<emphasis id="B907EADF5118D623FF13FAC611C69576" box="[170,247,1306,1330]" italics="true" pageId="12" pageNumber="343">Others</emphasis>
.
<taxonomicName id="4C734D4E5118D623FEA6FAC612329576" authority=", Whitley, 1938" authorityName=", Whitley" authorityYear="1938" box="[287,771,1306,1330]" family="Eleotridae" genus="Ophiocara" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="343" phylum="Chordata" rank="subSpecies" species="aporos" subSpecies="rigonis">
<emphasis id="B907EADF5118D623FEA6FAC613659576" box="[287,596,1306,1330]" italics="true" pageId="12" pageNumber="343">Ophiocara aporos rigonis,</emphasis>
<materialsCitation id="3B1B3C905118D623FDE5FAC612329576" box="[604,771,1306,1330]" collectingDate="2015-11-09" collectingDateMax="2018-10-23" collectingDateMin="2015-11-09" collectionCode="MNHN" collectorName="Kolobangara Is. &amp; Keith &amp; Lord &amp; Boseto &amp; Marquet &amp; Amick" country="Papua New Guinea" county="Freshwater creek" location="Water Lily Hole" municipality="Rigo" pageId="12" pageNumber="343" specimenCode="MNHN 2019-0242, MNHN 2019-0245, MNHN 2019-0243, MNHN 2019-0244, MNHN 2016-229, MNHN 2019-0252" specimenCount="12" specimenCount-female="8" specimenCount-male="4" stateProvince="West New Britain" typeStatus="holotype">Whitley, 1938</materialsCitation>
</taxonomicName>
.
<typeStatus id="54C8886F5118D623FF3DFAE611C19516" box="[132,240,1338,1362]" pageId="12" pageNumber="343" type="holotype">Holotype</typeStatus>
: AMS IA.5785.
<emphasis id="B907EADF5118D623FE1EFAE7132E9516" box="[423,543,1339,1362]" italics="true" pageId="12" pageNumber="343">
<typeStatus id="54C8886F5118D623FE1EFAE7132A9516" box="[423,539,1339,1362]" pageId="12" pageNumber="343" type="paratype">Paratypes</typeStatus>
.
</emphasis>
AMS IA.5787 (1), IA.5789 (1).
<collectingCounty id="62AD4E415118D623FEA8FA8610F89536" box="[273,457,1370,1394]" pageId="12" pageNumber="343">Freshwater creek</collectingCounty>
near
<collectingMunicipality id="6BA8ACB75118D623FDBCFA86130D9536" box="[517,572,1370,1394]" pageId="12" pageNumber="343">Rigo</collectingMunicipality>
,
<collectingCountry id="F364765D5118D623FDFFFA87119095D6" name="Papua New Guinea" pageId="12" pageNumber="343">Papua New Guinea</collectingCountry>
.
<specimenCode id="DBD59EB65118D623FF17FAA710BA95D6" box="[174,395,1402,1426]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="343">MNHN 2019-0242</specimenCode>
, male (
<quantity id="4C8B9B285118D623FE59FAA6137995D6" box="[480,584,1402,1426]" metricMagnitude="-2" metricUnit="m" metricValue="9.23" pageId="12" pageNumber="343" unit="mm" value="92.3">92.3 mm</quantity>
SL),
<location id="8EAC60165118D623FD32FAA7118D95F6" LSID="urn:lsid:plazi:treatment:03DA87DB5118D622FF3DFBA615959736:8EAC60165118D623FD32FAA7118D95F6" country="Papua New Guinea" county="Freshwater creek" municipality="Rigo" name="Water Lily Hole" pageId="12" pageNumber="343" stateProvince="West New Britain">Water Lily Hole</location>
,
<location id="8EAC60165118D623FF7EFA4711C295F6" LSID="urn:lsid:plazi:treatment:03DA87DB5118D622FF3DFBA615959736:8EAC60165118D623FF7EFA4711C295F6" box="[199,243,1435,1458]" country="Papua New Guinea" county="Freshwater creek" municipality="Rigo" name="Nut" pageId="12" pageNumber="343" stateProvince="West New Britain">Nut</location>
,
<collectingRegion id="49B7F82F5118D623FF47FA4710B695F6" box="[254,391,1434,1458]" country="Papua New Guinea" name="West New Britain" pageId="12" pageNumber="343">New Britain</collectingRegion>
,
<collectingCountry id="F364765D5118D623FE2BFA47135995F6" box="[402,616,1434,1458]" name="Papua New Guinea" pageId="12" pageNumber="343">Papua New Guinea</collectingCountry>
,
<date id="FFCD100D5118D623FDCBFA46123395F6" box="[626,770,1434,1458]" pageId="12" pageNumber="343" value="2018-10-23">
<collectingDate id="EF89E9E55118D623FDCBFA46123395F6" box="[626,770,1434,1458]" pageId="12" pageNumber="343" value="2018-10-23">23 Oct. 2018</collectingDate>
</date>
,
<location id="8EAC60165118D623FF3DFA6611F59596" LSID="urn:lsid:plazi:treatment:03DA87DB5118D622FF3DFBA615959736:8EAC60165118D623FF3DFA6611F59596" box="[132,196,1466,1490]" country="Papua New Guinea" county="Freshwater creek" municipality="Rigo" name="Keith" pageId="12" pageNumber="343" stateProvince="West New Britain">Keith</location>
,
<location id="8EAC60165118D623FF76FA6610369596" LSID="urn:lsid:plazi:treatment:03DA87DB5118D622FF3DFBA615959736:8EAC60165118D623FF76FA6610369596" box="[207,263,1466,1490]" country="Papua New Guinea" county="Freshwater creek" municipality="Rigo" name="Lord" pageId="12" pageNumber="343" stateProvince="West New Britain">Lord</location>
,
<location id="8EAC60165118D623FEA8FA66106F9596" LSID="urn:lsid:plazi:treatment:03DA87DB5118D622FF3DFBA615959736:8EAC60165118D623FEA8FA66106F9596" box="[273,350,1466,1490]" country="Papua New Guinea" county="Freshwater creek" municipality="Rigo" name="Amick" pageId="12" pageNumber="343" stateProvince="West New Britain">Amick</location>
,
<location id="8EAC60165118D623FED0FA6610879596" LSID="urn:lsid:plazi:treatment:03DA87DB5118D622FF3DFBA615959736:8EAC60165118D623FED0FA6610879596" box="[361,438,1466,1490]" country="Papua New Guinea" county="Freshwater creek" municipality="Rigo" name="Causse" pageId="12" pageNumber="343" stateProvince="West New Britain">Causse</location>
<emphasis id="B907EADF5118D623FE04FA6710C59596" box="[445,500,1466,1490]" italics="true" pageId="12" pageNumber="343">et al.</emphasis>
coll.; tag 17730.
<specimenCode id="DBD59EB65118D623FD0DFA67103295B6" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="343">MNHN 2019-0245</specimenCode>
,
<specimenCount id="9D75FD445118D623FEB6FA0610B095B6" box="[271,385,1498,1522]" count="4" pageId="12" pageNumber="343" type="female">4 females</specimenCount>
, same data as MNHN 2019-0242; tags 6826, 6828, 6829, 6830.
<specimenCode id="DBD59EB65118D623FE69FA2713979656" box="[464,678,1530,1554]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="343">MNHN 2019-0243</specimenCode>
,
<specimenCount id="9D75FD445118D623FD08FA2612369656" box="[689,775,1530,1554]" count="2" pageId="12" pageNumber="343" type="male">2 males</specimenCount>
and
<specimenCount id="9D75FD445118D623FF08F9C6102A9676" box="[177,283,1562,1586]" count="2" pageId="12" pageNumber="343" type="female">2 females</specimenCount>
, same data as
<collectionCode id="ED62AE085118D623FE0BF9C713359676" box="[434,516,1563,1586]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="343">MNHN</collectionCode>
2019-0242; tags 17731, 17732, 17733 &amp;17734.
<specimenCode id="DBD59EB65118D623FE3BF9E713609616" box="[386,593,1594,1618]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="343">MNHN 2019-0244</specimenCode>
, male, [extracted from
<specimenCode id="DBD59EB65118D623FF06F98710B29636" box="[191,387,1626,1650]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="343">MNHN 2016-229</specimenCode>
],
<collectorName id="2686531B5118D623FE2DF98613719636" box="[404,576,1626,1650]" pageId="12" pageNumber="343">Kolobangara Is.</collectorName>
, Zamba riv.,
<collectingCountry id="F364765D5118D623FD74F986103A96D6" name="Solomon Islands" pageId="12" pageNumber="343">Solomon Islands</collectingCountry>
,
<date id="FFCD100D5118D623FEAFF9A610AF96D6" box="[278,414,1658,1682]" pageId="12" pageNumber="343" value="2015-11-09">
<collectingDate id="EF89E9E55118D623FEAFF9A610AF96D6" box="[278,414,1658,1682]" pageId="12" pageNumber="343" value="2015-11-09">9 Nov. 2015</collectingDate>
</date>
,
<collectorName id="2686531B5118D623FE13F9A610D896D6" box="[426,489,1658,1682]" pageId="12" pageNumber="343">Keith</collectorName>
,
<collectorName id="2686531B5118D623FE4DF9A6131D96D6" box="[500,556,1658,1682]" pageId="12" pageNumber="343">Lord</collectorName>
,
<collectorName id="2686531B5118D623FD8EF9A713B496D6" box="[567,645,1659,1682]" pageId="12" pageNumber="343">Boseto</collectorName>
,
<collectorName id="2686531B5118D623FD29F9A713DD96D6" box="[656,748,1659,1682]" pageId="12" pageNumber="343">Marquet</collectorName>
<emphasis id="B907EADF5118D623FD4AF9A7119096F6" italics="true" pageId="12" pageNumber="343">et al.</emphasis>
coll.
<specimenCode id="DBD59EB65118D623FF5AF947108D96F6" box="[227,444,1690,1714]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="343">MNHN 2019-0252</specimenCode>
,
<specimenCount id="9D75FD445118D623FE71F947132E96F6" box="[456,543,1690,1715]" count="2" pageId="12" pageNumber="343" type="male">2 males</specimenCount>
and
<specimenCount id="9D75FD445118D623FDEEF94713F696F6" box="[599,711,1690,1715]" count="2" pageId="12" pageNumber="343" type="female">2 females</specimenCount>
,
<collectingRegion id="49B7F82F5118D623FD6AF94711E59696" country="Papua New Guinea" name="West New Britain" pageId="12" pageNumber="343">New Britain</collectingRegion>
,
<collectingCountry id="F364765D5118D623FF66F96710879696" box="[223,438,1722,1746]" name="Papua New Guinea" pageId="12" pageNumber="343">Papua New Guinea</collectingCountry>
, 2015,
<collectorName id="2686531B5118D623FDBCF96613619696" box="[517,592,1722,1746]" pageId="12" pageNumber="343">Amick</collectorName>
coll.; tags 6049, 6050, 16801 &amp; 16802.
</paragraph>
</subSubSection>
<subSubSection id="C36965465118D623FF3DF8C6124C9413" pageId="12" pageNumber="343" type="diagnosis">
<paragraph id="8BCC36CD5118D623FF3DF8C611C39776" blockId="12.[132,776,1818,1907]" box="[132,242,1818,1842]" pageId="12" pageNumber="343">
<emphasis id="B907EADF5118D623FF3DF8C611C39776" bold="true" box="[132,242,1818,1842]" pageId="12" pageNumber="343">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BCC36CD5118D623FF13F8E6124C9413" blockId="12.[132,776,1818,1907]" lastBlockId="12.[813,1455,1020,1111]" pageId="12" pageNumber="343">The species generally has 15 pectorals rays, a medium body depth at anus (20-26% SL), 11-13 scales in transverse forward series and a specific colourful reddish pattern in mature male and a brownish poorly coloured pattern in female.</paragraph>
</subSubSection>
<subSubSection id="C36965465118D622FC94FB5C159292F5" lastPageId="13" lastPageNumber="344" pageId="12" pageNumber="343" type="description">
<paragraph id="8BCC36CD5118D623FC94FB5C128094DC" blockId="12.[813,1456,1152,1906]" box="[813,945,1152,1176]" pageId="12" pageNumber="343">
<emphasis id="B907EADF5118D623FC94FB5C128094DC" bold="true" box="[813,945,1152,1176]" pageId="12" pageNumber="343">Description</emphasis>
</paragraph>
<paragraph id="8BCC36CD5118D623FCEBFB7D152F949E" blockId="12.[813,1456,1152,1906]" pageId="12" pageNumber="343">The scale and ray counts are given in table III and morphometrics in table IV.</paragraph>
<paragraph id="8BCC36CD5118D623FCEBFB3F124B95C4" blockId="12.[813,1456,1152,1906]" pageId="12" pageNumber="343">
The body is more ovoid than elongated. The body depth at anus is 21-26% SL, at first dorsal fin 20-25% SL, and the caudal peduncle depth is 13-16% SL. Predorsal length 44-51% SL and preanal length 62-67% SL. Size: up to
<quantity id="4C8B9B285118D623FA2DFA9B127F95C4" metricMagnitude="-1" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="343" unit="cm" value="22.0">22 cm</quantity>
SL.
</paragraph>
<paragraph id="8BCC36CD5118D623FCEBFA55129C96CE" blockId="12.[813,1456,1152,1906]" pageId="12" pageNumber="343">The head 31-36% SL is depressed, the snout is convex. The anterior nostrils are short, not reaching upper margin of upper lip. The mouth and jaw length 9-11% SL are small. Inwardly curved teeth set on both jaws and larger on outer row. Posterior end of maxillary extending before or to below around anterior margin of eye. Lips with numerous internal short papillae. Eye diameter 4-7% SL and interorbital length 13-16% SL.</paragraph>
<paragraph id="8BCC36CD5118D622FCEBF94F13F491D5" blockId="12.[813,1456,1152,1906]" lastBlockId="13.[132,775,217,658]" lastPageId="13" lastPageNumber="344" pageId="12" pageNumber="343">
Dorsal fins VI-I,8 with no filamentous rays. The first dorsal fin is with second, third and fourth rays longer. Anal fin I,9 directly opposite to the second dorsal fin. The caudal fin is with 13 branched rays and its posterior margin is rounded. Pelvic fins separate, I,5. Pectoral fins mostly 15 (
<specimenCount id="9D75FD445118D623FA94F8C4149E9774" box="[1325,1455,1816,1840]" count="3" pageId="12" pageNumber="343" type="generic">3 specimens</specimenCount>
with 14), with the posterior margin rounded. Lateral scales 28-30, with ctenoid scales on flanks and caudal peduncle. Cycloid scales from snout to top of head and anterior part of D1, on operculum, on base of pectoral fins and on belly extending to anus. Scales of top of head and back bigger than those of belly. Scales in transverse back series 8-10, in transverse forward 11-13, in predorsal 15-18 and in zigzag 8-10. Scales around the eye 9-14 (ENO). 25 vertebrae.
</paragraph>
<paragraph id="8BCC36CD5119D622FF13FE45119691B6" blockId="13.[132,775,217,658]" pageId="13" pageNumber="344">Absence of oculoscapular canal and supratemporals and presence of a short preopercular canal with the pores N and O.</paragraph>
<paragraph id="8BCC36CD5119D622FF13FE2510B59276" blockId="13.[132,775,217,658]" pageId="13" pageNumber="344">
Cephalic sensory papillae system developed as described by
<bibRefCitation id="EFE24B3C5119D622FF1AFDC510B19276" author="AKIHITO &amp; HAYASHI M. &amp; YOSHINO T." box="[163,384,537,562]" pageId="13" pageNumber="344" pagination="236 - 289" refId="ref16136" refString="AKIHITO, HAYASHI M. &amp; YOSHINO T., 1988. - Suborder Gobioidei. In: The Fishes of the Japanese Archipelago, 2 nd Edit. (Masuda H., Amaoko K., Araga C., Uyeno T. &amp; Yoshino T., eds), pp. 236 - 289, 445. Tokai University Press: Tokyo." type="journal article" year="1988">
Akihito
<emphasis id="B907EADF5119D622FF42FDC610189275" box="[251,297,537,561]" italics="true" pageId="13" pageNumber="344">et al</emphasis>
. (1988)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BCC36CD5119D622FF13FDE613E992D6" blockId="13.[132,775,217,658]" pageId="13" pageNumber="344">Males with a rounded/triangular urogenital papilla with distal tip rounded. The females have bulbous urogenital papilla with fimbriate projections around distal opening.</paragraph>
<paragraph id="8BCC36CD5119D622FF3DFD6510B49295" blockId="13.[132,776,697,1170]" box="[132,389,697,721]" pageId="13" pageNumber="344">
<emphasis id="B907EADF5119D622FF3DFD6510B49295" bold="true" box="[132,389,697,721]" pageId="13" pageNumber="344">Colour in preservation</emphasis>
</paragraph>
<paragraph id="8BCC36CD5119D622FF13FD06105893F6" blockId="13.[132,776,697,1170]" pageId="13" pageNumber="344">
<emphasis id="B907EADF5119D622FF13FD06103892B5" box="[170,265,730,754]" italics="true" pageId="13" pageNumber="344">Males. </emphasis>
Background of body dark brown on the back. The flanks are greyish to brownish. Each scale underlined by a dark grey dot. Belly greyish as the isthmus. Top of head brownish, lateral part greyish with 3 dark brown stripes radiating from the eye to the cheeks and operculum. The first and second dorsal fins as the anal, pelvic, pectoral and caudal fins are brownish.
</paragraph>
<paragraph id="8BCC36CD5119D622FF13FC66105294D6" blockId="13.[132,776,697,1170]" pageId="13" pageNumber="344">
<emphasis id="B907EADF5119D622FF13FC66102F9396" box="[170,286,954,978]" italics="true" pageId="13" pageNumber="344">Females. </emphasis>
Background of body brownish to black on the back. Top of head brownish, lateral part greyish with 3 thin brown stripes radiating from the eye to the cheeks and operculum. Flanks greyish from pectoral base to hypural base. Belly whitish. The first and second dorsal fins as the anal fin are brownish. Pectoral and pelvic fins greyish to hyaline. Caudal fin brownish.
</paragraph>
<paragraph id="8BCC36CD5119D622FF3DFB66102B9496" blockId="13.[132,775,1210,1907]" box="[132,282,1210,1234]" pageId="13" pageNumber="344">
<emphasis id="B907EADF5119D622FF3DFB66102B9496" bold="true" box="[132,282,1210,1234]" pageId="13" pageNumber="344">Colour in life</emphasis>
</paragraph>
<paragraph id="8BCC36CD5119D622FF13FB0615B09115" blockId="13.[132,775,1210,1907]" lastBlockId="13.[813,1456,217,689]" pageId="13" pageNumber="344">
<emphasis id="B907EADF5119D622FF13FB06103A94B6" box="[170,267,1242,1266]" italics="true" pageId="13" pageNumber="344">Males. </emphasis>
Two patterns. (1) Background of body greyish on the back. A bluish or greenish tinge along midline from the pectoral fin base to the caudal peduncle. The flanks have several, orange to reddish dots from pectoral base to hypural base, giving a certain shine to the fish. Belly whitish. Top of head greyish, lateral part yellowish with 3 dark brown to black stripes radiating from the eye to the cheeks and operculum, the highest continuing on the pectoral base which is yellow. The first and second dorsal fins as the anal fin are greyish with a reddish stripe at the distal part. Pectoral fins greyish to hyaline. Pelvic fins greyish to hyaline with a red distal margin. Caudal fin brownish with reddish base (
<figureCitation id="13482A485119D622FF7AF986102B9636" box="[195,282,1626,1650]" captionStart="Figure 5" captionStartId="12.[813,877,845,867]" captionTargetBox="[813,1456,240,827]" captionTargetId="figure-716@12.[813,1456,215,827]" captionTargetPageId="12" captionText="Figure 5. A: Giuris aporocephalus, male; in MNHN 2019-0243 (tag 17734), New Britain, Papua New Guinea; LS 61.3 mm (Photo P. Keith). B: G. aporocephalus, male; in MNHN 2019-0252, New Britain, Papua New Guinea; LS 143 mm (Photo P. Amick). C: G. aporocephalus, female; in MNHN 2019-0252, New Britain, Papua New Guinea; LS 143 mm (Photo P.Amick)." figureDoi="http://doi.org/10.5281/zenodo.12711995" httpUri="https://zenodo.org/record/12711995/files/figure.png" pageId="13" pageNumber="344">Fig. 5A</figureCitation>
). (2) Background of body brownish to orange on the back but colourful on the flanks. Blue along midline from the pectoral fin base to the caudal peduncle. The flanks are orange to reddish and each scale is underlined by a dark red dot. Belly bright yellow or orange as the isthmus. Top of head brownish, lateral part orange to bright red with 3 dark red stripes radiating from the eye to the cheeks and operculum, the highest continuing on the pectoral base which is bright yellow. Lips orange. The first and second dorsal fins as the anal fin are dark red to dark brown with an orange stripe at the distal part and a lighter red one at the base. Pectoral fins brownish. Pelvic fins brown with a red distal margin. Caudal fin brown (
<figureCitation id="13482A485119D622FB98FEE515459115" box="[1057,1140,313,337]" captionStart="Figure 5" captionStartId="12.[813,877,845,867]" captionTargetBox="[813,1456,240,827]" captionTargetId="figure-716@12.[813,1456,215,827]" captionTargetPageId="12" captionText="Figure 5. A: Giuris aporocephalus, male; in MNHN 2019-0243 (tag 17734), New Britain, Papua New Guinea; LS 61.3 mm (Photo P. Keith). B: G. aporocephalus, male; in MNHN 2019-0252, New Britain, Papua New Guinea; LS 143 mm (Photo P. Amick). C: G. aporocephalus, female; in MNHN 2019-0252, New Britain, Papua New Guinea; LS 143 mm (Photo P.Amick)." figureDoi="http://doi.org/10.5281/zenodo.12711995" httpUri="https://zenodo.org/record/12711995/files/figure.png" pageId="13" pageNumber="344">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8BCC36CD5119D622FCEBFE85159292F5" blockId="13.[813,1456,217,689]" pageId="13" pageNumber="344">
<emphasis id="B907EADF5119D622FCEBFE8512819135" box="[850,944,345,369]" italics="true" pageId="13" pageNumber="344">Females</emphasis>
(
<figureCitation id="13482A485119D622FC78FE8515269135" box="[961,1047,345,369]" captionStart="Figure 5" captionStartId="12.[813,877,845,867]" captionTargetBox="[813,1456,240,827]" captionTargetId="figure-716@12.[813,1456,215,827]" captionTargetPageId="12" captionText="Figure 5. A: Giuris aporocephalus, male; in MNHN 2019-0243 (tag 17734), New Britain, Papua New Guinea; LS 61.3 mm (Photo P. Keith). B: G. aporocephalus, male; in MNHN 2019-0252, New Britain, Papua New Guinea; LS 143 mm (Photo P. Amick). C: G. aporocephalus, female; in MNHN 2019-0252, New Britain, Papua New Guinea; LS 143 mm (Photo P.Amick)." figureDoi="http://doi.org/10.5281/zenodo.12711995" httpUri="https://zenodo.org/record/12711995/files/figure.png" pageId="13" pageNumber="344">Fig. 5C</figureCitation>
)
<emphasis id="B907EADF5119D622FB9EFE8615059135" box="[1063,1076,346,369]" italics="true" pageId="13" pageNumber="344"></emphasis>
Background of body brownish to black on the back. Top of head brownish, lateral part greyish with a yellowish tinge. Three thin brown stripes radiating from the eye to the cheeks and operculum, the upper one continuing on the pectoral base, which is yellow with a black superior spot. Flanks greyish to yellow from pectoral base to hypural base; each scale is underlined by a greyish dot. Belly whitish. The first and second dorsal fins as the anal fin are brownish with a reddish stripe at the distal part. Pectoral fins greyish to hyaline. Pelvic fins greyish to hyaline with a red distal margin. Caudal fin brownish.
</paragraph>
</subSubSection>
<subSubSection id="C36965465119D622FC94FD0515F093D5" pageId="13" pageNumber="344" type="biology_ecology">
<paragraph id="8BCC36CD5119D622FC94FD0512B692B5" blockId="13.[813,1455,729,914]" box="[813,903,729,753]" pageId="13" pageNumber="344">
<emphasis id="B907EADF5119D622FC94FD0512B692B5" bold="true" box="[813,903,729,753]" pageId="13" pageNumber="344">Ecology</emphasis>
</paragraph>
<paragraph id="8BCC36CD5119D622FCEBFD2515F093D5" blockId="13.[813,1455,729,914]" pageId="13" pageNumber="344">
The species occurs mainly in the vegetation of ponds, lakes, swamps, and in the lower parts of rivers. It is omnivorous and feeds on filamentous algae, small crustaceans and aquatic insects. It was found in sympatry with
<taxonomicName id="4C734D4E5119D622FA82FC86149E9335" authorityName="Keith &amp; Mennesson" authorityYear="2020" box="[1339,1455,858,881]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="344" phylum="Chordata" rank="species" species="caussei">
<emphasis id="B907EADF5119D622FA82FC86149E9335" box="[1339,1455,858,881]" italics="true" pageId="13" pageNumber="344">G. caussei</emphasis>
</taxonomicName>
(present paper) and
<emphasis id="B907EADF5119D622FC46FCA615F093D5" box="[1023,1217,890,913]" italics="true" pageId="13" pageNumber="344">
<taxonomicName id="4C734D4E5119D622FC46FCA6158C93D5" baseAuthorityName="Valenciennes" baseAuthorityYear="1837" box="[1023,1213,890,913]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="344" phylum="Chordata" rank="species" species="margaritaceus">G. margaritaceus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36965465119D622FC94FC6515959736" pageId="13" pageNumber="344" type="distribution">
<paragraph id="8BCC36CD5119D622FC94FC6512899395" blockId="13.[813,1455,953,1041]" box="[813,952,953,977]" pageId="13" pageNumber="344">
<emphasis id="B907EADF5119D622FC94FC6512899395" bold="true" box="[813,952,953,977]" pageId="13" pageNumber="344">Distribution</emphasis>
</paragraph>
<paragraph id="8BCC36CD5119D622FCEBFC0615F59455" blockId="13.[813,1455,953,1041]" pageId="13" pageNumber="344">
<taxonomicName id="4C734D4E5119D622FCEBFC06157493B5" baseAuthorityName="Macleay" baseAuthorityYear="1884" box="[850,1093,985,1009]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="344" phylum="Chordata" rank="species" species="aporocephalus">
<emphasis id="B907EADF5119D622FCEBFC06157493B5" box="[850,1093,985,1009]" italics="true" pageId="13" pageNumber="344">Giuris aporocephalus</emphasis>
</taxonomicName>
is known from
<collectingCountry id="F364765D5119D622FB4FFC05145193B5" box="[1270,1376,985,1009]" name="Australia" pageId="13" pageNumber="344">Australia</collectingCountry>
,
<collectingCountry id="F364765D5119D622FAD5FC0612839455" name="Papua New Guinea" pageId="13" pageNumber="344">Papua New Guinea</collectingCountry>
and the
<collectingCountry id="F364765D5119D622FBB2FC2515F19455" box="[1035,1216,1017,1041]" name="Solomon Islands" pageId="13" pageNumber="344">Solomon Islands</collectingCountry>
.
</paragraph>
<paragraph id="8BCC36CD5119D622FC94FBE5128B9415" blockId="13.[813,1456,1081,1906]" box="[813,954,1081,1105]" pageId="13" pageNumber="344">
<emphasis id="B907EADF5119D622FC94FBE5128B9415" bold="true" box="[813,954,1081,1105]" pageId="13" pageNumber="344">Comparison</emphasis>
</paragraph>
<paragraph id="8BCC36CD5119D622FCEBFB8615959736" blockId="13.[813,1456,1081,1906]" pageId="13" pageNumber="344">
<taxonomicName id="4C734D4E5119D622FCEBFB86157E9436" baseAuthorityName="Macleay" baseAuthorityYear="1884" box="[850,1103,1114,1138]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="344" phylum="Chordata" rank="species" species="aporocephalus">
<emphasis id="B907EADF5119D622FCEBFB86157E9436" box="[850,1103,1114,1138]" italics="true" pageId="13" pageNumber="344">Giuris aporocephalus</emphasis>
</taxonomicName>
differs from the other species sequenced by having a significant percentage of divergence in
<emphasis id="B907EADF5119D622FCF1FB46124794F5" box="[840,886,1178,1201]" italics="true" pageId="13" pageNumber="344">COI</emphasis>
gene (2.5-11.3%) (Tab. II) and by the colourful pattern of the male. Moreover it differs from
<taxonomicName id="4C734D4E5119D622FAB9FB6614489496" authorityName="Keith &amp; Mennesson" authorityYear="2020" box="[1280,1401,1210,1234]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="344" phylum="Chordata" rank="species" species="yahayai">
<emphasis id="B907EADF5119D622FAB9FB6614489496" box="[1280,1401,1210,1234]" italics="true" pageId="13" pageNumber="344">G. yahayai</emphasis>
</taxonomicName>
by a combination of characters including fewer scales in transverse forward series (11-13
<emphasis id="B907EADF5119D622FBE1FB2615449555" box="[1112,1141,1274,1297]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
17-20), a smaller body depth at anus (20-25
<emphasis id="B907EADF5119D622FC69FAC612DF9575" box="[976,1006,1306,1329]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
26-36% SL) and a smaller body depth at first dorsal fin (21-25
<emphasis id="B907EADF5119D622FBACFAE715039516" box="[1045,1074,1339,1362]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
26-39% SL). It differs from
<taxonomicName id="4C734D4E5119D622FADCFAE6127C9536" authorityName="Keith, Mennesson, Lord &amp; Hubert" authorityYear="2020" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="344" phylum="Chordata" rank="species" species="viator">
<emphasis id="B907EADF5119D622FADCFAE6127C9536" italics="true" pageId="13" pageNumber="344">G. viator</emphasis>
</taxonomicName>
in having always 15 pectorals rays
<emphasis id="B907EADF5119D622FB64FA8715CA9536" box="[1245,1275,1371,1394]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
always 14 rays, fewer scales in transverse forward series (11-13
<emphasis id="B907EADF5119D622FAF8FAA7146E95D6" box="[1345,1375,1403,1426]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
14-15) and zigzag series (8-9
<emphasis id="B907EADF5119D622FBA6FA47150D95F6" box="[1055,1084,1435,1458]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
9-10). It differs from
<taxonomicName id="4C734D4E5119D622FA90FA4614A595F6" baseAuthorityName="Bleeker" baseAuthorityYear="1854" box="[1321,1428,1434,1458]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="344" phylum="Chordata" rank="species" species="tolsoni">
<emphasis id="B907EADF5119D622FA90FA4614A595F6" box="[1321,1428,1434,1458]" italics="true" pageId="13" pageNumber="344">G. tolsoni</emphasis>
</taxonomicName>
in having always 15 pectorals rays
<emphasis id="B907EADF5119D622FB20FA6715869596" box="[1177,1207,1467,1490]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
always 14 rays, and a greater interorbital length (13-16
<emphasis id="B907EADF5119D622FB23FA07158695B6" box="[1178,1207,1499,1522]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
9-12% SL), and fewer scales in transverse forward series (11-13
<emphasis id="B907EADF5119D622FB4FFA2714229656" box="[1270,1299,1531,1554]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
13-16). It differs from
<taxonomicName id="4C734D4E5119D622FC2EF9C615649676" baseAuthorityName="Valenciennes" baseAuthorityYear="1837" box="[919,1109,1562,1586]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="344" phylum="Chordata" rank="species" species="margaritaceus">
<emphasis id="B907EADF5119D622FC2EF9C615649676" box="[919,1109,1562,1586]" italics="true" pageId="13" pageNumber="344">G. margaritaceus</emphasis>
</taxonomicName>
in having fewer scales in transverse forward series (11-13
<emphasis id="B907EADF5119D622FBEFF9E715429616" box="[1110,1139,1595,1618]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
13-14) and zigzag series (8-9
<emphasis id="B907EADF5119D622FC94F987127B9636" box="[813,842,1627,1650]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
9-10), more scales around the eye (10-14
<emphasis id="B907EADF5119D622FAA2F98714089636" box="[1307,1337,1627,1650]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
8-11), and always 15 pectorals rays
<emphasis id="B907EADF5119D622FB8EF9A7156596D6" box="[1079,1108,1659,1682]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
14-15. It differs from
<taxonomicName id="4C734D4E5119D622FAF8F9A6127396F6" authorityName="Keith &amp; Mennesson" authorityYear="2020" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="344" phylum="Chordata" rank="species" species="charpini">
<emphasis id="B907EADF5119D622FAF8F9A6127396F6" italics="true" pageId="13" pageNumber="344">G. charpini</emphasis>
</taxonomicName>
in having always 15 pectorals rays
<emphasis id="B907EADF5119D622FB6AF94715C096F6" box="[1235,1265,1691,1714]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
13-14, and more scales in transverse forward series (11-13
<emphasis id="B907EADF5119D622FB4FF96714259696" box="[1270,1300,1723,1746]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
13-14). It differs from
<taxonomicName id="4C734D4E5119D622FC21F907153896B6" authorityName="Keith &amp; Mennesson" authorityYear="2020" box="[920,1033,1755,1778]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="344" phylum="Chordata" rank="species" species="caussei">
<emphasis id="B907EADF5119D622FC21F907153896B6" box="[920,1033,1755,1778]" italics="true" pageId="13" pageNumber="344">G. caussei</emphasis>
</taxonomicName>
in having a greater body depth at anus (21-25
<emphasis id="B907EADF5119D622FCC3F92712A69756" box="[890,919,1787,1810]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
16-21% SL). It differs from
<taxonomicName id="4C734D4E5119D622FB70F92714749756" baseAuthorityName="Sauvage" baseAuthorityYear="1880" box="[1225,1349,1786,1810]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="344" phylum="Chordata" rank="species" species="laglaizei">
<emphasis id="B907EADF5119D622FB70F92714749756" box="[1225,1349,1786,1810]" italics="true" pageId="13" pageNumber="344">G. laglaizei</emphasis>
</taxonomicName>
in having more scales around the eye (10-14
<emphasis id="B907EADF5119D622FB15F8C715FB9776" box="[1196,1226,1819,1842]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
6-8), fewer scales in transverse back series (8-10
<emphasis id="B907EADF5119D622FBDDF8E715B39716" box="[1124,1154,1851,1874]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
9-11) and a greater interorbital length (13-16
<emphasis id="B907EADF5119D622FC41F88715249736" box="[1016,1045,1883,1906]" italics="true" pageId="13" pageNumber="344">vs.</emphasis>
10-12% SL).
</paragraph>
</subSubSection>
</treatment>
</document>