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<document id="00E85F621F550C99123460EC86C3D4AC" ID-DOI="10.5281/zenodo.177118" ID-GBIF-Dataset="1513d696-cd96-4d1f-b841-2ceaae2d40f0" ID-ISSN="1175-5326" ID-Zenodo-Dep="177118" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459850580550" checkinUser="plazi" docAuthor="Hippa, Heikki" docDate="2007" docId="03DA87CCFFF0FFE502ABFC34BC01FDC1" docLanguage="en" docName="zt01502p044.pdf" docOrigin="Zootaxa 1502" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Manota subspathula Hippa, 2007, sp. n." docType="treatment" docVersion="8" lastPageNumber="23" masterDocId="FFE3FFB4FFE4FFF3023CFF86B91EFFDD" masterDocTitle="The genus Manota Williston (Diptera: Mycetophilidae) in Melanesia and Oceania" masterLastPageNumber="44" masterPageNumber="1" pageNumber="21" updateTime="1698225258820" updateUser="plazi">
<mods:mods id="DD20B567CE36693CAE24CD4224A3A734" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C805F5B07D2DBB93BA373BF505A92231">
<mods:title id="A178D951C22224680FF9D0F3CBA37DF6">The genus Manota Williston (Diptera: Mycetophilidae) in Melanesia and Oceania</mods:title>
</mods:titleInfo>
<mods:name id="30AEAA6364BBC717F6617DD50AD3E51C" type="personal">
<mods:role id="5B5A34A9F3F6C04647C78B8C0BDB6DA0">
<mods:roleTerm id="1C352D67BC999714E82A0AB84E80525D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6408C866D2278A230E72EFDFAF754268">Hippa, Heikki</mods:namePart>
</mods:name>
<mods:typeOfResource id="99882095A74D241BC0AF54EE52C60A17">text</mods:typeOfResource>
<mods:relatedItem id="DDAAD2BEA5DF958A289355BD5BA7CFA5" type="host">
<mods:titleInfo id="933BA92E51DC6C471297418309BF9BC5">
<mods:title id="E9E37F66C62762B4FF8C676D0B1CECA3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="55E69517C4C92B84AD9298D3B17AA004">
<mods:date id="EF3B6011280550E7E2AB97A7409597A4">2007</mods:date>
<mods:detail id="ABDA8CDF3C8D783D90670C2BBA9018FD" type="volume">
<mods:number id="4A58463770909DB31D63490D66C50FFF">1502</mods:number>
</mods:detail>
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<mods:start id="08145719EDEEF2154F9F10321256F860">1</mods:start>
<mods:end id="92F114F95F520B0F069755824A8F7AAE">44</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="855D73131C55BE81B51FB91B1B29D237">journal article</mods:classification>
<mods:identifier id="D52BE4802FA49606059CE76CC4C44C62" type="DOI">10.5281/zenodo.177118</mods:identifier>
<mods:identifier id="11F049289BF5E6CA48EDF94CC1CC633F" type="GBIF-Dataset">1513d696-cd96-4d1f-b841-2ceaae2d40f0</mods:identifier>
<mods:identifier id="A98F0BB2848C4047F5F83B856F11A420" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F2E8AFD7BCD97EF1F1ECF7545617D62D" type="Zenodo-Dep">177118</mods:identifier>
</mods:mods>
<treatment id="03DA87CCFFF0FFE502ABFC34BC01FDC1" ID-DOI="http://doi.org/10.5281/zenodo.6251820" ID-GBIF-Taxon="119362368" ID-Zenodo-Dep="6251820" LSID="urn:lsid:plazi:treatment:03DA87CCFFF0FFE502ABFC34BC01FDC1" httpUri="http://treatment.plazi.org/id/03DA87CCFFF0FFE502ABFC34BC01FDC1" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
<subSubSection id="C3696551FFF0FFE702ABFC34B829FC32" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BCC36DAFFF0FFE702ABFC34B8C6FC11" blockId="20.[151,472,946,1007]" box="[151,472,946,972]" pageId="20" pageNumber="21">
<heading id="D08481B6FFF0FFE702ABFC34B8C6FC11" bold="true" box="[151,472,946,972]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="B907EAC8FFF0FFE702ABFC34B8C6FC11" bold="true" box="[151,472,946,972]" pageId="20" pageNumber="21">
<taxonomicName id="4C734D59FFF0FFE702ABFC34B890FC11" ID-CoL="3XVYV" box="[151,398,946,972]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="subspathula" status="sp. nov.">
<emphasis id="B907EAC8FFF0FFE702ABFC34B890FC11" bold="true" box="[151,398,946,972]" italics="true" pageId="20" pageNumber="21">Manota subspathula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23457B3FFF0FFE703AAFC34B8C6FC11" box="[406,472,946,972]" pageId="20" pageNumber="21" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC36DAFFF0FFE702ABFC53B829FC32" blockId="20.[151,472,946,1007]" box="[151,311,981,1007]" pageId="20" pageNumber="21">
(
<figureCitation id="13482A5FFFF0FFE702A3FC53B9EAFC32" box="[159,244,981,1007]" captionStart="FIGURE 11" captionStartId="21.[151,255,1962,1986]" captionTargetBox="[384,1200,201,1925]" captionTargetId="figure@21.[363,1227,194,1930]" captionTargetPageId="21" captionText="FIGURE 11. Manota subspathula sp. n. (holotype): A. Hypopygium, dorsal view. B. Hypopygium, ventral view. C. Antennal flagellomere 4, lateral view. Scale for A and B 0.10 mm, for C 0.05 mm." httpUri="https://zenodo.org/record/177129/files/figure.png" pageId="20" pageNumber="21">Figs 11</figureCitation>
A–C)
</paragraph>
</subSubSection>
<subSubSection id="C3696551FFF0FFE702ABFBA3BBB3F882" pageId="20" pageNumber="21" type="description">
<paragraph id="8BCC36DAFFF0FFE702ABFBA3B8BDFBE3" blockId="20.[151,1437,1061,1887]" box="[151,419,1061,1087]" pageId="20" pageNumber="21">
A small-sized
<emphasis id="B907EAC8FFF0FFE7037CFBA3B8BDFBE3" box="[320,419,1061,1086]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C734D59FFF0FFE7037CFBA3B881FBE3" box="[320,415,1061,1086]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Manota</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BCC36DAFFF0FFE702FAFBCBB890F8EA" blockId="20.[151,1437,1061,1887]" pageId="20" pageNumber="21">
<collectingCountry id="F364764AFFF0FFE702FAFBCBB818FBBA" box="[198,262,1101,1127]" name="Malta" pageId="20" pageNumber="21">Male</collectingCountry>
.
<emphasis id="B907EAC8FFF0FFE7032DFBCBB886FBBA" bold="true" box="[273,408,1101,1127]" pageId="20" pageNumber="21">Coloration</emphasis>
. Pale brown, frons, vertex and occiput darker brown, mouthparts and legs slightly paler than thorax but apex of coxa, trochanter and base and apex of femur on legs 2 and 3 darker brown, wing unicolorous, hyaline, pale yellowish-brown, knob of haltere black.
<emphasis id="B907EAC8FFF0FFE70143FB1BBADCFB6A" bold="true" box="[895,962,1181,1207]" pageId="20" pageNumber="21">Head</emphasis>
. Antennal flagellomere 4,
<figureCitation id="13482A5FFFF0FFE706C6FB1BBC52FB6A" box="[1274,1356,1181,1207]" captionStart="FIGURE 11" captionStartId="21.[151,255,1962,1986]" captionTargetBox="[384,1200,201,1925]" captionTargetId="figure@21.[363,1227,194,1930]" captionTargetPageId="21" captionText="FIGURE 11. Manota subspathula sp. n. (holotype): A. Hypopygium, dorsal view. B. Hypopygium, ventral view. C. Antennal flagellomere 4, lateral view. Scale for A and B 0.10 mm, for C 0.05 mm." httpUri="https://zenodo.org/record/177129/files/figure.png" pageId="20" pageNumber="21">Fig. 11</figureCitation>
C. Palpomere 3 of maxillary palpus with apicomesial extension, with ca. 4 apically expanded curved sensilla; the two apical palpomeres absent on both sides of the single specimen.
<emphasis id="B907EAC8FFF0FFE70192FB6BBD15FADA" bold="true" box="[942,1035,1261,1287]" pageId="20" pageNumber="21">Thorax</emphasis>
. Pleural setosity pattern similar to
<figureCitation id="13482A5FFFF0FFE702ABFA93B9C4FAF2" box="[151,218,1301,1327]" captionStart="FIGURE 1" captionStartId="2.[151,255,1779,1803]" captionTargetBox="[267,1309,466,1740]" captionTargetId="figure@2.[264,1323,456,1887]" captionTargetPageId="2" captionText="FIGURE 1. Pleura, lateral view. A. Manota serawei sp. n. (holotype). B. M. explicans sp. n. (paratype). C. M. perissochaeta sp. n. (paratype). D. M. spathula sp. n. (holotype). E. M. bicuspis sp. n. (paratype). Scale 0.10 mm." httpUri="https://zenodo.org/record/177119/files/figure.png" pageId="20" pageNumber="21">Fig. 1</figureCitation>
D: anepisternum setose, with 24 setae; anterior basalare setose, with 6 setae; preepisternum 2 non-setose; laterotergite setose, with 26 setae; episternum 3 setose, with ca. 4 setae.
<emphasis id="B907EAC8FFF0FFE70626FABBBD40FA8A" bold="true" box="[1050,1118,1341,1367]" pageId="20" pageNumber="21">Wing</emphasis>
. Length
<quantity id="4C8B9B3FFFF0FFE706FAFABBBC3AFA8A" box="[1222,1316,1341,1367]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="mm" value="1.5">1.5 mm</quantity>
.
<emphasis id="B907EAC8FFF0FFE70713FABBB9C9FAA2" bold="true" pageId="20" pageNumber="21">Hypopygium</emphasis>
(
<figureCitation id="13482A5FFFF0FFE702DAFAE3B823FAA2" box="[230,317,1381,1407]" captionStart="FIGURE 11" captionStartId="21.[151,255,1962,1986]" captionTargetBox="[384,1200,201,1925]" captionTargetId="figure@21.[363,1227,194,1930]" captionTargetPageId="21" captionText="FIGURE 11. Manota subspathula sp. n. (holotype): A. Hypopygium, dorsal view. B. Hypopygium, ventral view. C. Antennal flagellomere 4, lateral view. Scale for A and B 0.10 mm, for C 0.05 mm." httpUri="https://zenodo.org/record/177129/files/figure.png" pageId="20" pageNumber="21">Figs 11</figureCitation>
A, B). Sternite 9 laterally united with gonocoxa except for the extreme posterior part, posterior margin deeply emarginated, extending to the level of the base of gonostylus, anterior margin shallowly notched, setosity similar to ventral setosity of gonocoxa. Parastylar lobe present and well exposed, with two or three setae directed obliquely posteriad, associated with weakly sclerotized lobes on its lateral side. Paraapodemal lobe not exposed in ventral view of hypopygium. The dorsal mesial margin of gonocoxa with a large lobe in the middle, bearing many blunt-ended setae at the margin and a fur-like substance on the dorsal surface, unlike other similar species. Number of juxtagonostylar setae one: it is a strong non-flattened megaseta arising from a basal body, which is slightly shorter than half the length of the megaseta. Gonostylus elongate, widening towards the slightly bilobed apex, with two long apical setae, with a row of dorsomesial setae diverging from the other rather weak setosity. Tegmen unusually shaped, with very strong lateral shoulders. Hypoproct long, extending posteriorly nearly as far as the apex of gonostylus, with ca. 20 scattered ventral setae. Cerci separate.
</paragraph>
<paragraph id="8BCC36DAFFF0FFE702FAF8C3BBB3F882" blockId="20.[151,1437,1061,1887]" box="[198,685,1861,1887]" pageId="20" pageNumber="21">Female and preimaginal stages unknown.</paragraph>
</subSubSection>
<caption id="DF0C6652FFF1FFE602ABF82CBD18F839" httpUri="https://zenodo.org/record/177129/files/figure.png" pageId="21" pageNumber="22" targetBox="[384,1200,201,1925]" targetPageId="21">
<paragraph id="8BCC36DAFFF1FFE602ABF82CBD18F839" blockId="21.[151,1435,1962,2020]" pageId="21" pageNumber="22">
<emphasis id="B907EAC8FFF1FFE602ABF82CB83EF81F" bold="true" box="[151,288,1962,1986]" pageId="21" pageNumber="22">FIGURE 11</emphasis>
.
<taxonomicName id="4C734D59FFF1FFE6030DF82CBB11F81C" box="[305,527,1962,1985]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="subspathula" status="sp. nov.">
<emphasis id="B907EAC8FFF1FFE6030DF82CBB11F81C" box="[305,527,1962,1985]" italics="true" pageId="21" pageNumber="22">Manota subspathula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23457B3FFF1FFE60024F82CBB49F81F" box="[536,599,1962,1986]" pageId="21" pageNumber="22" rank="species">
<emphasis id="B907EAC8FFF1FFE60024F82CBB49F81F" bold="true" box="[536,599,1962,1986]" pageId="21" pageNumber="22">sp. n.</emphasis>
</taxonomicNameLabel>
(holotype):
<emphasis id="B907EAC8FFF1FFE600DCF82CBBEDF81F" bold="true" box="[736,755,1962,1986]" pageId="21" pageNumber="22">A</emphasis>
. Hypopygium, dorsal view.
<emphasis id="B907EAC8FFF1FFE60611F82CBD58F81F" bold="true" box="[1069,1094,1962,1986]" pageId="21" pageNumber="22">B.</emphasis>
Hypopygium, ventral view.
<emphasis id="B907EAC8FFF1FFE607BEF82CBC85F81F" bold="true" box="[1410,1435,1962,1986]" pageId="21" pageNumber="22">C.</emphasis>
Antennal flagellomere 4, lateral view. Scale for A and B 0.10 mm, for C 0.05 mm.
</paragraph>
</caption>
<subSubSection id="C3696551FFF2FFE502FAFF11BC01FDC1" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BCC36DAFFF2FFE502FAFF11BD30FE17" blockId="22.[151,1436,151,540]" pageId="22" pageNumber="23">
<emphasis id="B907EAC8FFF2FFE502FAFF11B84CFF6C" bold="true" box="[198,338,151,177]" pageId="22" pageNumber="23">Discussion.</emphasis>
<taxonomicName id="4C734D59FFF2FFE50365FF1EBB11FF6C" box="[345,527,152,177]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="subspathula">
<emphasis id="B907EAC8FFF2FFE50365FF1EBB11FF6C" box="[345,527,152,177]" italics="true" pageId="22" pageNumber="23">M. subspathula</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C734D59FFF2FFE50095FF1EBA28FF6C" box="[681,822,152,177]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="spathula">
<emphasis id="B907EAC8FFF2FFE50095FF1EBA28FF6C" box="[681,822,152,177]" italics="true" pageId="22" pageNumber="23">M. spathula</emphasis>
</taxonomicName>
but can be distinguished e.g. by the following characters: 1) the posterior margin of sternite 9 is notched, not convex, 2) the large lobe at the middle of the dorsal mesial margin of gonocoxa has a fur- or velvet-like substance on it dorsal surface, bare in
<taxonomicName id="4C734D59FFF2FFE50680FF6EBC57FEDC" box="[1212,1353,232,257]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="spathula">
<emphasis id="B907EAC8FFF2FFE50680FF6EBC57FEDC" box="[1212,1353,232,257]" italics="true" pageId="22" pageNumber="23">M. spathula</emphasis>
</taxonomicName>
, 3) the juxtagonostylar megaseta is narrow, greatly expanded in
<taxonomicName id="4C734D59FFF2FFE50102FE96BAD2FEF4" box="[830,972,272,297]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="spathula">
<emphasis id="B907EAC8FFF2FFE50102FE96BAD2FEF4" box="[830,972,272,297]" italics="true" pageId="22" pageNumber="23">M. spathula</emphasis>
</taxonomicName>
, 4) the long apical setae of gonostylus are wide apart, with one at the apicolateral corner and the other at the apicomesial corner, in
<taxonomicName id="4C734D59FFF2FFE506DAFEBEBC85FE8C" box="[1254,1435,312,337]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="subspathula">
<emphasis id="B907EAC8FFF2FFE506DAFEBEBC85FE8C" box="[1254,1435,312,337]" italics="true" pageId="22" pageNumber="23">M. subspathula</emphasis>
</taxonomicName>
both the setae are close to each other at the apicolateral corner, and 5) the lateral shoulders of tegmen are much more strongly pronounced. Both species are similar to
<taxonomicName id="4C734D59FFF2FFE501B4FE0EBD19FE7C" box="[904,1031,392,417]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B907EAC8FFF2FFE501B4FE0EBD19FE7C" box="[904,1031,392,417]" italics="true" pageId="22" pageNumber="23">M. hirsuta</emphasis>
</taxonomicName>
and to many species outside the Melanesian and Oceanian region. For further discussion, see under
<taxonomicName id="4C734D59FFF2FFE50191FE36BD39FE14" box="[941,1063,432,457]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B907EAC8FFF2FFE50191FE36BD39FE14" box="[941,1063,432,457]" italics="true" pageId="22" pageNumber="23">M. hirsuta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCC36DAFFF2FFE502FAFE51BC01FDC1" blockId="22.[151,1436,151,540]" pageId="22" pageNumber="23">
<emphasis id="B907EAC8FFF2FFE502FAFE51B80AFE2C" bold="true" box="[198,276,471,497]" pageId="22" pageNumber="23">
<typeStatus id="54C88878FFF2FFE502FAFE51B813FE2C" box="[198,269,471,497]" pageId="22" pageNumber="23">Types</typeStatus>
.
</emphasis>
<typeStatus id="54C88878FFF2FFE50321FE5EB897FE2C" box="[285,393,472,497]" pageId="22" pageNumber="23" type="holotype">
<emphasis id="B907EAC8FFF2FFE50321FE5EB897FE2C" box="[285,393,472,497]" italics="true" pageId="22" pageNumber="23">Holotype</emphasis>
</typeStatus>
.
<collectingCountry id="F364764AFFF2FFE503ABFE5EB8C8FE2F" box="[407,470,472,498]" name="Malta" pageId="22" pageNumber="23">Male</collectingCountry>
,
<collectingCountry id="F364764AFFF2FFE503DEFE5EBBD9FE2F" box="[482,711,472,498]" name="Papua New Guinea" pageId="22" pageNumber="23">Papua New Guinea</collectingCountry>
, Madang province, Baitabag village, Kau Wildlife Area,
<quantity id="4C8B9B3FFFF2FFE5075DFE5EBC82FE2F" box="[1377,1436,472,498]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="m" value="50.0">50 m</quantity>
a. s. l.,
<geoCoordinate id="EE47501DFFF2FFE502D7FD84B859FDC1" box="[235,327,512,540]" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="925" value="-5.133333">S 5o 08’</geoCoordinate>
<geoCoordinate id="EE47501DFFF2FFE50373FD84B8D2FDC1" box="[335,460,512,540]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="925" value="145.76666">E 145o 46’</geoCoordinate>
, primary forest, Malaise trap 8–
<date id="FFCD101AFFF2FFE5017FFD84BAD5FDC1" box="[835,971,514,540]" pageId="22" pageNumber="23" value="1999-11-21">21.11.1999</date>
, Lukáš Č iżek leg. (in NRM).
</paragraph>
</subSubSection>
</treatment>
</document>
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