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<document id="C86DAED151A86681F7725F02E2E0F33A" ID-DOI="10.11646/zootaxa.3919.1.6" ID-GBIF-Dataset="4c40552e-e778-4d8e-a6b7-9662436cadc2" ID-ISSN="1175-5326" ID-Zenodo-Dep="244587" ID-ZooBank="F711CA99-1B2C-4E18-9F4B-7521D38D2303" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460989857372" checkinUser="plazi" docAuthor="Smith, Ian M., Cook, David R. &amp; Gerecke, Reinhard" docDate="2015" docId="03DA87C4FF971778FF4C6845A8CFF998" docLanguage="en" docName="zt03919p156.pdf" docOrigin="Zootaxa 3919 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexaxonopsis Viets 1926" docType="treatment" docVersion="9" lastPageNumber="142" masterDocId="FFE3FFBCFF881767FFDB6A61AA1CFF8D" masterDocTitle="Revision of the status of some genus-level water mite taxa in the families Pionidae Thor, 1900, Aturidae Thor, 1900, and Nudomideopsidae Smith, 1990 (Acari: Hydrachnidiae)" masterLastPageNumber="156" masterPageNumber="111" pageNumber="142" updateTime="1698561557693" updateUser="plazi">
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<mods:title id="F3A2DA87A5D61C84233F7D9A70B4D377">Revision of the status of some genus-level water mite taxa in the families Pionidae Thor, 1900, Aturidae Thor, 1900, and Nudomideopsidae Smith, 1990 (Acari: Hydrachnidiae)</mods:title>
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<mods:namePart id="3DF9789E6D3FE02082C18BB000C0CFA6">Smith, Ian M.</mods:namePart>
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<mods:namePart id="C752C6A06A15139D39AF1C93F14F61BC">Cook, David R.</mods:namePart>
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<mods:namePart id="0C3AB22BDE393D57C249E9415B5F63A6">Gerecke, Reinhard</mods:namePart>
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<treatment id="03DA87C4FF971778FF4C6845A8CFF998" ID-DOI="http://doi.org/10.5281/zenodo.6109901" ID-GBIF-Taxon="119596533" ID-Zenodo-Dep="6109901" LSID="urn:lsid:plazi:treatment:03DA87C4FF971778FF4C6845A8CFF998" httpUri="http://treatment.plazi.org/id/03DA87C4FF971778FF4C6845A8CFF998" lastPageNumber="142" pageId="31" pageNumber="142">
<subSubSection id="C3696559FF971778FF4C6845A886FDB3" box="[151,666,548,574]" pageId="31" pageNumber="142" type="nomenclature">
<paragraph id="8BCC36D2FF971778FF4C6845A886FDB3" blockId="31.[151,666,548,574]" box="[151,666,548,574]" pageId="31" pageNumber="142">
<heading id="D08481BEFF971778FF4C6845A886FDB3" bold="true" box="[151,666,548,574]" fontSize="11" level="1" pageId="31" pageNumber="142" reason="1">
<emphasis id="B907EAC0FF971778FF4C6845A886FDB3" bold="true" box="[151,666,548,574]" pageId="31" pageNumber="142">
Genus
<taxonomicName id="4C734D51FF971778FF356845A834FDB3" authority="Viets, 1926" authorityName="Viets" authorityYear="1926" box="[238,552,548,574]" class="Arachnida" family="Aturidae" genus="Hexaxonopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FF356845AB85FDB3" bold="true" box="[238,409,548,574]" italics="true" pageId="31" pageNumber="142">Hexaxonopsis</emphasis>
Viets, 1926
</taxonomicName>
<emphasis id="B907EAC0FF971778FDF46845A886FDB3" bold="true" box="[559,666,548,574]" italics="true" pageId="31" pageNumber="142">stat. nov.</emphasis>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF971778FF4C680AA972FD51" pageId="31" pageNumber="142" type="reference_group">
<paragraph id="8BCC36D2FF971778FF4C680AA904FD12" blockId="31.[151,886,618,733]" pageId="31" pageNumber="142">
<taxonomicName id="4C734D51FF971778FF4C680AABBBFD0D" authority="(Hexaxonopsis)" baseAuthorityName="Hexaxonopsis" box="[151,423,618,640]" class="Arachnida" family="Axonopsidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FF4C680AAAE3FD0D" box="[151,255,619,640]" italics="true" pageId="31" pageNumber="142">Axonopsis</emphasis>
(
<emphasis id="B907EAC0FF971778FED6680AAB87FD0D" box="[269,411,619,640]" italics="true" pageId="31" pageNumber="142">Hexaxonopsis</emphasis>
)
</taxonomicName>
:
<bibRefCitation id="EFE24B23FF971778FE6A680BA829FD0D" author="Cook" box="[433,565,618,640]" pageId="31" pageNumber="142" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook, 1974a</bibRefCitation>
, pp. 335336, figs.
<date id="FFCD1012FF971778FD26680BA96EFD0D" box="[765,882,618,640]" pageId="31" pageNumber="142" value="1413" valueMax="1417">14131417</date>
.
<taxonomicName id="4C734D51FF971778FF4C68EBABCDFD12" authority="(Plesiobrachypoda)" baseAuthorityName="Plesiobrachypoda" box="[151,465,649,671]" class="Arachnida" family="Axonopsidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FF4C68EBAAE3FD12" box="[151,255,650,671]" italics="true" pageId="31" pageNumber="142">Axonopsis</emphasis>
(
<emphasis id="B907EAC0FF971778FED668E8ABD9FD12" box="[269,453,649,671]" italics="true" pageId="31" pageNumber="142">Plesiobrachypoda</emphasis>
)
</taxonomicName>
:
<bibRefCitation id="EFE24B23FF971778FE0768E8A87CFD12" author="Cook" box="[476,608,649,671]" pageId="31" pageNumber="142" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook, 1974a</bibRefCitation>
, p. 336, fig. 1420.
</paragraph>
<paragraph id="8BCC36D2FF971778FF4C68C8A8F3FD33" blockId="31.[151,886,618,733]" box="[151,751,680,702]" pageId="31" pageNumber="142">
<taxonomicName id="4C734D51FF971778FF4C68C8ABBBFD33" authority="(Hexaxonopsis)" baseAuthorityName="Hexaxonopsis" box="[151,423,680,702]" class="Arachnida" family="Axonopsidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FF4C68C8AAE3FD33" box="[151,255,681,702]" italics="true" pageId="31" pageNumber="142">Axonopsis</emphasis>
(
<emphasis id="B907EAC0FF971778FED668C8AB87FD33" box="[269,411,681,702]" italics="true" pageId="31" pageNumber="142">Hexaxonopsis</emphasis>
)
</taxonomicName>
:
<bibRefCitation id="EFE24B23FF971778FE6A68C9A843FD33" author="Viets" box="[433,607,680,702]" pageId="31" pageNumber="142" refString="Viets, K. O. (1987) Die Milben des Sußwassers (Hydrachnellae und Halacaridae [part.], Acari). 2. Katalog. Sonderbande des Naturwissenschaftlichen Vereins in Hamburg, 8, 1 - 1012." type="journal article" year="1987">K.O. Viets, 1987</bibRefCitation>
, pp. 190192.
</paragraph>
<paragraph id="8BCC36D2FF971778FF4C68A6A972FD51" blockId="31.[151,886,618,733]" box="[151,878,710,733]" pageId="31" pageNumber="142">
<taxonomicName id="4C734D51FF971778FF4C68A6ABBBFD51" authority="(Hexaxonopsis)" baseAuthorityName="Hexaxonopsis" box="[151,423,710,733]" class="Arachnida" family="Axonopsidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FF4C68A6AAE3FD51" box="[151,255,711,732]" italics="true" pageId="31" pageNumber="142">Axonopsis</emphasis>
(
<emphasis id="B907EAC0FF971778FED668A9AB87FD50" box="[269,411,712,733]" italics="true" pageId="31" pageNumber="142">Hexaxonopsis</emphasis>
)
</taxonomicName>
:
<bibRefCitation id="EFE24B23FF971778FE6A68A7A864FD51" author="Smit" box="[433,632,710,732]" pageId="31" pageNumber="142" refString="Smit, H. &amp; Pesic, V. (2010) New species of water mites from Oman, with some zoogeographical notes (Acari: Hydrachnidia). Acarologia, 50, 151 - 192. http: // dx. doi. org / 10.1051 / acarologia / 20101953" type="journal article" year="2010">Smit &amp; Pešić, 2010</bibRefCitation>
, pp. 4548, figs. 3235.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF971778FF4C6968AAD6FB5F" pageId="31" pageNumber="142" type="diagnosis">
<paragraph id="8BCC36D2FF971778FF4C6968ABFDFCAF" blockId="31.[151,1437,777,1558]" box="[151,481,777,802]" pageId="31" pageNumber="142">
<emphasis id="B907EAC0FF971778FF4C6968AB40FCAC" bold="true" box="[151,348,777,802]" pageId="31" pageNumber="142">
Diagnosis.
<emphasis id="B907EAC0FF971778FEC3696BAB40FCAC" bold="true" box="[280,348,778,801]" italics="true" pageId="31" pageNumber="142">Larva</emphasis>
</emphasis>
: Unknown.
</paragraph>
<paragraph id="8BCC36D2FF971778FF1C694DAAD6FB5F" blockId="31.[151,1437,777,1558]" pageId="31" pageNumber="142">
<emphasis id="B907EAC0FF971778FF1C694DAB0EFCC8" bold="true" box="[199,274,812,837]" italics="true" pageId="31" pageNumber="142">Adults</emphasis>
(modified from
<bibRefCitation id="EFE24B23FF971778FE02694CA875FCCB" author="Cook" box="[473,617,813,838]" pageId="31" pageNumber="142" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook 1974a</bibRefCitation>
,
<bibRefCitation id="EFE24B23FF971778FDA3694CA8DCFCCB" author="Cook" box="[632,704,813,838]" pageId="31" pageNumber="142" refString="Cook, D. R. (1974 b) North American species of the genus Axonopsis (Acarina: Aturidae: Axonopsinae). The Great Lakes Entomologist, 7, 55 - 79." type="journal article" year="1974" yearSuffix="b">1974b</bibRefCitation>
): Character states of
<taxonomicName id="4C734D51FF971778FC19694FAE29FCC8" box="[962,1077,814,837]" class="Arachnida" family="Axonopsidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FC19694FAE29FCC8" box="[962,1077,814,837]" italics="true" pageId="31" pageNumber="142">Axonopsis</emphasis>
</taxonomicName>
-like mites. Dorsal and ventral shields separate or incompletely fused anteriorly; lacking caudal development posteriorly. Dorsal furrow lacking glandularia. Dorsal shield with or without paired rows of spinose projections anterolaterally; bearing six pairs of glandularia none of which flank excretory pore. Ventral shield on each side with ridge originating at lateral end of suture line between third and fourth coxal plates extending anterolaterally to lateral edge of shield; first and second coxal plates produced anteriorly into recurved, pointed, hook-like projections; fourth coxal plate usually bearing two pairs (rarely three pairs) of glandularia in region between genital field and opening for insertion of fourth leg. Genital field bearing three pairs of acetabula. Suture lines between genital field and ventral shield clearly delineated. Gnathosoma with mouth opening subterminal in position; gnathosomal apodemes moderately long. Pedipalp tibia moderately long and slender, bearing a long, thick seta on a prominent projection and a sessile slender seta ventrally; tarsus shorter than tibia. Fourth leg with segments not flattened and telofemur not reduced in size.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF971778FF1C6EBCAB13FAEC" pageId="31" pageNumber="142" type="materials_examined">
<paragraph id="8BCC36D2FF971778FF1C6EBCA8AEFB7B" blockId="31.[151,1437,777,1558]" box="[199,690,1245,1270]" pageId="31" pageNumber="142">
<emphasis id="B907EAC0FF971778FF1C6EBCAB7EFB7B" bold="true" box="[199,354,1245,1270]" pageId="31" pageNumber="142">
<typeStatus id="54C88870FF971778FF1C6EBCAB1EFB7B" box="[199,258,1245,1270]" pageId="31" pageNumber="142">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C734D51FF971778FEB26EBFA8AEFB7B" authority="Viets." authorityName="Viets." box="[361,690,1245,1270]" class="Arachnida" family="Aturidae" genus="Hexaxonopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="species" species="violacea">
<emphasis id="B907EAC0FF971778FEB26EBFA870FB7B" box="[361,620,1245,1270]" italics="true" pageId="31" pageNumber="142">Hexaxonopsis violacea</emphasis>
Viets.
</taxonomicName>
</paragraph>
<paragraph id="8BCC36D2FF971778FF1C6F60AB13FAEC" blockId="31.[151,1437,777,1558]" pageId="31" pageNumber="142">
<emphasis id="B907EAC0FF971778FF1C6F60AB8BFA97" bold="true" box="[199,407,1281,1306]" pageId="31" pageNumber="142">Species included.</emphasis>
Numerous species listed by
<bibRefCitation id="EFE24B23FF971778FD316F63A9A6FA97" author="Viets" box="[746,954,1281,1306]" pageId="31" pageNumber="142" refString="Viets, K. O. (1987) Die Milben des Sußwassers (Hydrachnellae und Halacaridae [part.], Acari). 2. Katalog. Sonderbande des Naturwissenschaftlichen Vereins in Hamburg, 8, 1 - 1012." type="journal article" year="1987">K.O. Viets (1987)</bibRefCitation>
and
<bibRefCitation id="EFE24B23FF971778FC236F60AF4EFA97" box="[1016,1362,1281,1306]" pageId="31" pageNumber="142" refString="Pesic, V., Smit, H. &amp; Saboori, A. (2012) Water mites delineating the Oriental and Palearctic regions - the unique fauna of southern Iran, with description of one new genus, one new subgenus and 14 new species (Acari: Hydrachnidia). Zootaxa, 3330, 1 - 67." type="journal article">Pešić, Smit &amp; Saboori (2012)</bibRefCitation>
under
<taxonomicName id="4C734D51FF971778FF4C6F47ABDFFAB3" authority="(Hexaxonopsis)" baseAuthorityName="Hexaxonopsis" box="[151,451,1318,1342]" class="Arachnida" family="Axonopsidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FF4C6F47AB16FAB0" box="[151,266,1318,1341]" italics="true" pageId="31" pageNumber="142">Axonopsis</emphasis>
(
<emphasis id="B907EAC0FF971778FEC76F47ABA5FAB0" box="[284,441,1318,1341]" italics="true" pageId="31" pageNumber="142">Hexaxonopsis</emphasis>
)
</taxonomicName>
and by
<bibRefCitation id="EFE24B23FF971778FDFF6F47A8EDFAB3" author="Viets" box="[548,753,1317,1342]" pageId="31" pageNumber="142" refString="Viets, K. O. (1987) Die Milben des Sußwassers (Hydrachnellae und Halacaridae [part.], Acari). 2. Katalog. Sonderbande des Naturwissenschaftlichen Vereins in Hamburg, 8, 1 - 1012." type="journal article" year="1987">K.O. Viets (1987)</bibRefCitation>
under
<taxonomicName id="4C734D51FF971778FC9F6F47AEBCFAB3" authority="(Plesiobrachypoda)" baseAuthorityName="Plesiobrachypoda" box="[836,1184,1317,1342]" class="Arachnida" family="Axonopsidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FC9F6F47A9ABFAB0" box="[836,951,1318,1341]" italics="true" pageId="31" pageNumber="142">Axonopsis</emphasis>
(
<emphasis id="B907EAC0FF971778FC116F44AE8AFAB3" box="[970,1174,1317,1342]" italics="true" pageId="31" pageNumber="142">Plesiobrachypoda</emphasis>
)
</taxonomicName>
, and see below under subgenera.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF971778FF1C6F0CAF19FA0B" box="[199,1285,1389,1414]" pageId="31" pageNumber="142" type="distribution">
<paragraph id="8BCC36D2FF971778FF1C6F0CAF19FA0B" blockId="31.[151,1437,777,1558]" box="[199,1285,1389,1414]" pageId="31" pageNumber="142">
<emphasis id="B907EAC0FF971778FF1C6F0CAB7CFA0B" bold="true" box="[199,352,1389,1414]" pageId="31" pageNumber="142">Distribution.</emphasis>
Holarctic (Europe, Southeast Asia,
<collectingCountry id="F3647642FF971778FD2F6F0CA926FA0B" box="[756,826,1389,1414]" name="China" pageId="31" pageNumber="142">China</collectingCountry>
,
<collectingCountry id="F3647642FF971778FC9E6F0FA994FA08" box="[837,904,1390,1413]" name="Japan" pageId="31" pageNumber="142">Japan</collectingCountry>
,
<collectingCountry id="F3647642FF971778FC496F0FA9DEFA08" box="[914,962,1390,1413]" name="Iran" pageId="31" pageNumber="142">Iran</collectingCountry>
, North
<collectingCountry id="F3647642FF971778FBCD6F0CAE65FA0B" box="[1046,1145,1389,1414]" name="United States of America" pageId="31" pageNumber="142">America</collectingCountry>
) and Africa.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF971778FF1C6FF0A8CFF998" pageId="31" pageNumber="142" type="discussion">
<paragraph id="8BCC36D2FF971778FF1C6FF0A8CFF998" blockId="31.[151,1437,777,1558]" pageId="31" pageNumber="142">
<emphasis id="B907EAC0FF971778FF1C6FF0AB50FA27" bold="true" box="[199,332,1425,1450]" pageId="31" pageNumber="142">Discussion.</emphasis>
<bibRefCitation id="EFE24B23FF971778FE816FF0ABE9FA27" author="Cook" box="[346,501,1425,1450]" pageId="31" pageNumber="142" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook (1974a</bibRefCitation>
,
<bibRefCitation id="EFE24B23FF971778FDD36FF0A853FA27" author="Cook" box="[520,591,1425,1450]" pageId="31" pageNumber="142" refString="Cook, D. R. (1974 b) North American species of the genus Axonopsis (Acarina: Aturidae: Axonopsinae). The Great Lakes Entomologist, 7, 55 - 79." type="journal article" year="1974" yearSuffix="b">1974b</bibRefCitation>
) treated
<taxonomicName id="4C734D51FF971778FD1B6FF3A941FA24" box="[704,861,1426,1449]" class="Arachnida" family="Aturidae" genus="Hexaxonopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FD1B6FF3A941FA24" box="[704,861,1426,1449]" italics="true" pageId="31" pageNumber="142">Hexaxonopsis</emphasis>
</taxonomicName>
as a subgenus of
<taxonomicName id="4C734D51FF971778FB9C6FF3AEA6FA24" box="[1095,1210,1426,1449]" class="Arachnida" family="Axonopsidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FB9C6FF3AEA6FA24" box="[1095,1210,1426,1449]" italics="true" pageId="31" pageNumber="142">Axonopsis</emphasis>
</taxonomicName>
. Here we propose elevation of this taxon to full generic rank based on the unique combination of character states listed above. We also propose that the taxon
<taxonomicName id="4C734D51FF971778FE116FB8A93CFA7F" authority="Viets, 1942" authorityName="Viets" authorityYear="1942" box="[458,800,1497,1522]" class="Arachnida" family="Aturidae" genus="Plesiobrachypoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FE116FB8A88AFA7F" box="[458,662,1497,1522]" italics="true" pageId="31" pageNumber="142">Plesiobrachypoda</emphasis>
Viets, 1942
</taxonomicName>
, previously regarded as a subgenus of
<taxonomicName id="4C734D51FF971778FB0B6FBBAF5FFA7C" box="[1232,1347,1498,1521]" class="Arachnida" family="Axonopsidae" genus="Axonopsis" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FB0B6FBBAF5FFA7C" box="[1232,1347,1498,1521]" italics="true" pageId="31" pageNumber="142">Axonopsis</emphasis>
</taxonomicName>
, should now be considered as a subgenus of
<taxonomicName id="4C734D51FF971778FDF46F9FA8D0F998" box="[559,716,1534,1557]" class="Arachnida" family="Aturidae" genus="Hexaxonopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="31" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF971778FDF46F9FA8D0F998" box="[559,716,1534,1557]" italics="true" pageId="31" pageNumber="142">Hexaxonopsis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>