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<document id="6162BAB21E0586746A88B92D1BE965D3" ID-CLB-Dataset="45303" ID-DOI="10.5281/zenodo.280262" ID-GBIF-Dataset="6b35b42d-86fe-40e4-a68c-047ed37376bc" ID-ISSN="1175-5326" ID-Zenodo-Dep="280262" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460448753740" checkinUser="plazi" docAuthor="Sroka, Pavel &amp; Godunko, Roman J." docDate="2012" docId="03CCB42FFFD81174FF11FA39FD028C7C" docLanguage="en" docName="zt03222p045.pdf" docOrigin="Zootaxa 3222" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Electrogena gibedede Godunko &amp; Sroka, sp. nov." docType="treatment" docVersion="9" lastPageNumber="42" masterDocId="FFF5CC57FFD9117AFF86FFA0FFBA8848" masterDocTitle="Mayflies of the Caucasus Mountains. I. A new species of the genus Electrogena Zurwerra &amp; Tomka, 1985" masterLastPageNumber="45" masterPageNumber="28" pageNumber="29" updateTime="1698338020022" updateUser="ExternalLinkService">
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<mods:title id="043F3A24782B0A98FDC80BDE57B8AF8D">Mayflies of the Caucasus Mountains. I. A new species of the genus Electrogena Zurwerra &amp; Tomka, 1985</mods:title>
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<mods:name id="F0C302ABBE7F4B28B53D7CC4F4C97BE5" type="personal">
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<mods:namePart id="AC1A91BEEC72D9ACD4BEF81D23AF75A6">Sroka, Pavel</mods:namePart>
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<mods:namePart id="D69103959EBFB934A48F148EF584A2F6">Godunko, Roman J.</mods:namePart>
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<mods:date id="3C94F23E56B4180184FB477E74E9A1B8">2012</mods:date>
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<treatment id="03CCB42FFFD81174FF11FA39FD028C7C" ID-DOI="http://doi.org/10.5281/zenodo.6174577" ID-GBIF-Taxon="119509965" ID-Zenodo-Dep="6174577" LSID="urn:lsid:plazi:treatment:03CCB42FFFD81174FF11FA39FD028C7C" httpUri="http://treatment.plazi.org/id/03CCB42FFFD81174FF11FA39FD028C7C" lastPageId="14" lastPageNumber="42" pageId="1" pageNumber="29">
<subSubSection id="C37F56B2FFD8117BFF11FA39FE908D9C" pageId="1" pageNumber="29" type="nomenclature">
<paragraph id="8BDA0539FFD8117BFF11FA39FD4F8DFA" blockId="1.[151,757,1432,1492]" box="[151,757,1432,1459]" pageId="1" pageNumber="29">
<heading id="D092B255FFD8117BFF11FA39FD4F8DFA" bold="true" box="[151,757,1432,1459]" fontSize="11" level="1" pageId="1" pageNumber="29" reason="1">
<emphasis id="B911D92BFFD8117BFF11FA39FD4F8DFA" bold="true" box="[151,757,1432,1459]" pageId="1" pageNumber="29">
<taxonomicName id="4C657EBAFFD8117BFF11FA39FD308DFA" ID-CoL="396SQ" authority="Godunko &amp; Sroka" authorityName="Godunko &amp; Sroka" box="[151,650,1432,1459]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="29" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD8117BFF11FA39FE218DFB" bold="true" box="[151,411,1433,1459]" italics="true" pageId="1" pageNumber="29">Electrogena gibedede</emphasis>
Godunko &amp; Sroka
</taxonomicName>
,
<taxonomicNameLabel id="A2226450FFD8117BFD11FA38FD4F8DFA" box="[663,757,1432,1458]" pageId="1" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA0539FFD8117BFF11FA1CFE908D9C" blockId="1.[151,757,1432,1492]" box="[151,298,1468,1492]" pageId="1" pageNumber="29">
<figureCitation id="135E19BCFFD8117BFF11FA1CFE908D9C" box="[151,298,1468,1492]" captionStart-0="FIGURES 1 3" captionStart-1="FIGURE 4" captionStart-2="FIGURES 5 6" captionStart-3="FIGURES 7 11" captionStart-4="FIGURES 12 14" captionStart-5="FIGURES 15 20" captionStart-6="FIGURES 21 26" captionStart-7="FIGURES 27 30" captionStart-8="FIGURES 31 34" captionStartId-0="2.[151,264,1702,1725]" captionStartId-1="3.[151,250,1737,1760]" captionStartId-2="4.[151,264,1825,1848]" captionStartId-3="5.[151,263,1980,2003]" captionStartId-4="6.[151,264,1631,1654]" captionStartId-5="7.[151,264,1843,1866]" captionStartId-6="9.[151,264,1898,1921]" captionStartId-7="10.[151,264,1220,1243]" captionStartId-8="13.[151,264,1743,1766]" captionTargetBox-0="[329,1251,209,1652]" captionTargetBox-1="[259,1325,210,1699]" captionTargetBox-2="[409,1186,217,1770]" captionTargetBox-3="[165,1411,338,1946]" captionTargetBox-4="[267,1320,228,1604]" captionTargetBox-5="[178,1407,454,1744]" captionTargetBox-6="[172,1416,351,1853]" captionTargetBox-7="[170,1415,186,1184]" captionTargetBox-8="[185,1403,741,1717]" captionTargetId-0="figure@2.[320,1267,185,1687]" captionTargetId-1="figure@3.[229,1358,193,1716]" captionTargetId-2="figure@4.[379,1208,193,1786]" captionTargetId-3="figure@5.[151,1436,337,1959]" captionTargetId-4="figure@6.[256,1330,193,1610]" captionTargetId-5="figure@7.[151,1436,445,1822]" captionTargetId-6="figure@9.[151,1436,351,1861]" captionTargetId-7="figure@10.[151,1436,163,1199]" captionTargetId-8="figure@13.[172,1414,733,1722]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="13" captionText-0="FIGURES 1 3. Electrogena gibedede sp. nov., male imago, paratype: 1, head, lateral view; 2, abdominal segments, lateral view; 3, abdominal segments, dorsal view." captionText-1="FIGURE 4. Electrogena gibedede sp. nov., male imago, paratype: genitalia, ventral view." captionText-2="FIGURES 5 6. Electrogena gibedede sp. nov., male imago, paratype: 5, penis lobes, ventral view; 6, penis lobes, dorsal view." captionText-3="FIGURES 7 11. Electrogena gibedede sp. nov., mature larva, paratype: 7, head, dorsal view; 8, hypopharynx, ventral view; 9, kinds of labrum shape, dorsal view; 10, variations of glossae shape, ventral view; 11, first segment of maxillary palp, dorsal view." captionText-4="FIGURES 12 14. Electrogena gibedede sp. nov., mature larva, paratype: 12, middle leg, dorsal view; 13, gill VII, dorsal view; 14, gill I, dorsal view." captionText-5="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." captionText-6="FIGURES 21 26. Electrogena gibedede sp. nov., male imago (21, 22) and mature larva (23 26): 21, penis lobes, ventral view; 22, genitalia, ventral view; 23, forefemora bristles, dorsal view; 24, forefemora bristles (detail), dorsal view; 25, hind femora bristles, dorsal view; 26, hind femora bristles (detail), dorsal view." captionText-7="FIGURES 27 30. Electrogena gibedede sp. nov., mature larva, paratype: 27, 28, posterior margin of abdominal tergites, dorsal view; 29, 30, tarsal claw, dorsal and ventral view." captionText-8="FIGURES 31 34. Electrogena gibedede sp. nov., egg: 31, general view; 32, chorionic surface with small rounded tubercles and granular ground matrix; 33, micropyle; 34, group of KCTs attachment structures." httpUri-0="https://zenodo.org/record/280263/files/figure.png" httpUri-1="https://zenodo.org/record/280264/files/figure.png" httpUri-2="https://zenodo.org/record/280265/files/figure.png" httpUri-3="https://zenodo.org/record/280266/files/figure.png" httpUri-4="https://zenodo.org/record/280267/files/figure.png" httpUri-5="https://zenodo.org/record/280268/files/figure.png" httpUri-6="https://zenodo.org/record/280269/files/figure.png" httpUri-7="https://zenodo.org/record/280270/files/figure.png" httpUri-8="https://zenodo.org/record/280271/files/figure.png" pageId="1" pageNumber="29">Figures 134</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37F56B2FFD81177FF11F9A3FC208FFB" lastPageId="13" lastPageNumber="41" pageId="1" pageNumber="29" type="description">
<paragraph id="8BDA0539FFD8117BFF11F9A3FD3F8E77" blockId="1.[151,1437,1539,2032]" pageId="1" pageNumber="29">
<emphasis id="B911D92BFFD8117BFF11F9A3FE928E54" bold="true" box="[151,296,1539,1564]" pageId="1" pageNumber="29">Description.</emphasis>
<emphasis id="B911D92BFFD8117BFEA9F9A4FE078E54" box="[303,445,1540,1564]" italics="true" pageId="1" pageNumber="29">Male imago.</emphasis>
Size: body length 7.58.0 mm, forewing length 8.09.0 mm, cerci length 20.024.0 mm (approximately 2.5 times longer than body).
</paragraph>
<paragraph id="8BDA0539FFD8117BFF4FF9ECFC318ECF" blockId="1.[151,1437,1539,2032]" pageId="1" pageNumber="29">
Head brown with paler clypeal part; compound eyes divided grayish dorsally and black ventrally (
<figureCitation id="135E19BCFFD8117BFA92F9ECFADB8E2C" box="[1300,1377,1612,1636]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1702,1725]" captionTargetBox="[329,1251,209,1652]" captionTargetId="figure@2.[320,1267,185,1687]" captionTargetPageId="2" captionText="FIGURES 1 3. Electrogena gibedede sp. nov., male imago, paratype: 1, head, lateral view; 2, abdominal segments, lateral view; 3, abdominal segments, dorsal view." httpUri="https://zenodo.org/record/280263/files/figure.png" pageId="1" pageNumber="29">Figs. 1</figureCitation>
,
<figureCitation id="135E19BCFFD8117BFAE9F9ECFA348E2C" box="[1391,1422,1612,1636]" captionStart="FIGURES 15 20" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[178,1407,454,1744]" captionTargetId="figure@7.[151,1436,445,1822]" captionTargetPageId="7" captionText="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="1" pageNumber="29">17</figureCitation>
). No stripes or bands on head. Ocelli black. Antennae pale brownish.
</paragraph>
<paragraph id="8BDA0539FFD8117BFF4FF934FE948EBC" blockId="1.[151,1437,1539,2032]" pageId="1" pageNumber="29">
Prothorax brown dorsally, paler in central part; ventrally dark brown (
<figureCitation id="135E19BCFFD8117BFC6FF934FB848EE4" box="[1001,1086,1684,1708]" captionStart="FIGURES 15 20" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[178,1407,454,1744]" captionTargetId="figure@7.[151,1436,445,1822]" captionTargetPageId="7" captionText="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="1" pageNumber="29">Fig. 17</figureCitation>
). Meso- and metahorax brown dorsally with light yellowish smudges; ventrally dark brown. Thorax laterally pale, yellowish with brownish sclerites (
<figureCitation id="135E19BCFFD8117BFF4BF97CFE9A8EBC" box="[205,288,1756,1780]" captionStart="FIGURES 15 20" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[178,1407,454,1744]" captionTargetId="figure@7.[151,1436,445,1822]" captionTargetPageId="7" captionText="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="1" pageNumber="29">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8BDA0539FFD8117BFF4FF8A1FD628FB8" blockId="1.[151,1437,1539,2032]" pageId="1" pageNumber="29">
Legs slender, all pairs with 5segmented tarsus with one sharp and one blunt claw. Forelegs generally darker than other legs. Forefemur proximally pale brown, with darker margins. Distinct dark brown transversal band approximately at 2/3 of femur length (
<figureCitation id="135E19BCFFD8117BFDD7F8E7FD1C8F17" box="[593,678,1863,1887]" captionStart="FIGURES 15 20" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[178,1407,454,1744]" captionTargetId="figure@7.[151,1436,445,1822]" captionTargetPageId="7" captionText="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="1" pageNumber="29">Fig. 17</figureCitation>
). Distal part of forefemur brownish. Dark brown transversal band also at distal end of femora, at articulation with tibia. Foretibia brown, darker than femur. Tarsal segments IIV brown, slightly paler than tibia. Last tarsal segment darker, brown. Tarsal claws brown. Middle and hind legs of same coloration as forelegs; femora pale brown, with darker brown spot situated approximately at 2/3 of femur length. Tibia and tarsi brownish, darker than femur.
</paragraph>
<caption id="DF1A55B1FFDB1178FF11F906FDFC8E93" httpUri="https://zenodo.org/record/280263/files/figure.png" pageId="2" pageNumber="30" targetBox="[329,1251,209,1652]" targetPageId="2">
<paragraph id="8BDA0539FFDB1178FF11F906FDFC8E93" blockId="2.[151,1436,1702,1755]" pageId="2" pageNumber="30">
<emphasis id="B911D92BFFDB1178FF11F906FE868EF4" bold="true" box="[151,316,1702,1725]" pageId="2" pageNumber="30">FIGURES 13.</emphasis>
<taxonomicName id="4C657EBAFFDB1178FEC3F907FD998EF5" box="[325,547,1703,1725]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="30" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFDB1178FEC3F907FD998EF5" box="[325,547,1703,1725]" italics="true" pageId="2" pageNumber="30">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFDB1178FDAAF907FDC58EF4" box="[556,639,1703,1724]" pageId="2" pageNumber="30" rank="species">
<emphasis id="B911D92BFFDB1178FDAAF907FDC58EF4" bold="true" box="[556,639,1703,1724]" pageId="2" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
, male imago, paratype: 1, head, lateral view; 2, abdominal segments, lateral view; 3, abdominal segments, dorsal view.
</paragraph>
</caption>
<paragraph id="8BDA0539FFDB1178FF4FF8A1FA358F75" blockId="2.[151,1436,1793,2033]" pageId="2" pageNumber="30">
Wings hyaline, transparent, with easily visible brown venation. Costal and subcostal field slightly milky colored (mainly in pterostigmatic area). Pterostigma with several cross veins. Wings with typical of
<taxonomicName id="4C657EBAFFDB1178FB16F886FAAC8F76" box="[1168,1302,1830,1854]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFDB1178FB16F886FAAC8F76" box="[1168,1302,1830,1854]" italics="true" pageId="2" pageNumber="30">Electrogena</emphasis>
</taxonomicName>
venations.
</paragraph>
<paragraph id="8BDA0539FFDB1178FF4FF8E9FD2F8FB9" blockId="2.[151,1436,1793,2033]" pageId="2" pageNumber="30">
Abdominal tergites laterally brown with whitish pattern, consisting of pale triangles visible on laterocaudal portions of each tergite. Additionally, pairs of elongated pale spots situated dorsally on tergites, to side of central thin longitudinal pale line (
<figureCitation id="135E19BCFFDB1178FE4BF831FD868FE1" box="[461,572,1937,1961]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1702,1725]" captionTargetBox="[329,1251,209,1652]" captionTargetId="figure@2.[320,1267,185,1687]" captionTargetPageId="2" captionText="FIGURES 1 3. Electrogena gibedede sp. nov., male imago, paratype: 1, head, lateral view; 2, abdominal segments, lateral view; 3, abdominal segments, dorsal view." httpUri="https://zenodo.org/record/280263/files/figure.png" pageId="2" pageNumber="30">Figs. 2, 3</figureCitation>
,
<figureCitation id="135E19BCFFDB1178FDCCF831FDD38FE1" box="[586,617,1937,1961]" captionStart="FIGURES 15 20" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[178,1407,454,1744]" captionTargetId="figure@7.[151,1436,445,1822]" captionTargetPageId="7" captionText="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="2" pageNumber="30">19</figureCitation>
). Sternites pale brownish, with very indistinct whitish pattern in central part, consisting of two oblique elongated spots situated frontally and two dots caudally. Distal part of sternite IX white. Neural ganglion visible in sternite VII.
</paragraph>
<caption id="DF1A55B1FFDA1179FF11F969FB938E97" box="[151,1065,1737,1760]" httpUri="https://zenodo.org/record/280264/files/figure.png" pageId="3" pageNumber="31" targetBox="[259,1325,210,1699]" targetPageId="3">
<paragraph id="8BDA0539FFDA1179FF11F969FB938E97" blockId="3.[151,1065,1737,1760]" box="[151,1065,1737,1760]" pageId="3" pageNumber="31">
<emphasis id="B911D92BFFDA1179FF11F969FEA98E96" bold="true" box="[151,275,1737,1760]" pageId="3" pageNumber="31">FIGURE 4.</emphasis>
<taxonomicName id="4C657EBAFFDA1179FE9FF969FE4F8E97" box="[281,501,1737,1759]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="31" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFDA1179FE9FF969FE4F8E97" box="[281,501,1737,1759]" italics="true" pageId="3" pageNumber="31">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFDA1179FE7AF96AFDF68E97" box="[508,588,1738,1759]" pageId="3" pageNumber="31" rank="species">
<emphasis id="B911D92BFFDA1179FE7AF96AFDF68E97" bold="true" box="[508,588,1738,1759]" pageId="3" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
, male imago, paratype: genitalia, ventral view.
</paragraph>
</caption>
<paragraph id="8BDA0539FFDA1179FF4FF8A6FA278F2E" blockId="3.[151,1437,1798,1894]" pageId="3" pageNumber="31">Cerci dark brown basally, towards distal end getting paler, with dark bands following individual articulations. Every second articulation more pronounced, accompanied by more distinct dark band than those between them. These differences are visible mainly in central part of cerci. All surface covered with dark hairs. Paracercus vestigial.</paragraph>
<caption id="DF1A55B1FFDD117EFF11F881FA278F70" box="[151,1437,1825,1848]" httpUri="https://zenodo.org/record/280265/files/figure.png" pageId="4" pageNumber="32" targetBox="[409,1186,217,1770]" targetPageId="4">
<paragraph id="8BDA0539FFDD117EFF11F881FA278F70" blockId="4.[151,1437,1825,1848]" box="[151,1437,1825,1848]" pageId="4" pageNumber="32">
<emphasis id="B911D92BFFDD117EFF11F881FE808F7F" bold="true" box="[151,314,1825,1848]" pageId="4" pageNumber="32">FIGURES 56.</emphasis>
<taxonomicName id="4C657EBAFFDD117EFEC6F882FDA68F70" box="[320,540,1826,1848]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="32" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFDD117EFEC6F882FDA68F70" box="[320,540,1826,1848]" italics="true" pageId="4" pageNumber="32">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFDD117EFDA5F882FDC98F7F" box="[547,627,1826,1847]" pageId="4" pageNumber="32" rank="species">
<emphasis id="B911D92BFFDD117EFDA5F882FDC98F7F" bold="true" box="[547,627,1826,1847]" pageId="4" pageNumber="32">sp. nov.</emphasis>
</taxonomicNameLabel>
, male imago, paratype: 5, penis lobes, ventral view; 6, penis lobes, dorsal view.
</paragraph>
</caption>
<paragraph id="8BDA0539FFDD117FFF4FF8FFFBDF889C" blockId="4.[151,1436,1887,2019]" lastBlockId="5.[151,1436,151,284]" lastPageId="5" lastPageNumber="33" pageId="4" pageNumber="32">
<emphasis id="B911D92BFFDD117EFF4FF8FFFE898F3F" box="[201,307,1887,1911]" italics="true" pageId="4" pageNumber="32">Genitalia</emphasis>
: Styliger plate with apparent rounded projections on posterior margin. Forceps brown, on the proximal portion darker. Surface of forceps with numerous tiny rounded projections on inner side (
<figureCitation id="135E19BCFFDD117EFB41F823FAB48FD3" box="[1223,1294,1923,1947]" captionStart="FIGURE 4" captionStartId="3.[151,250,1737,1760]" captionTargetBox="[259,1325,210,1699]" captionTargetId="figure@3.[229,1358,193,1716]" captionTargetPageId="3" captionText="FIGURE 4. Electrogena gibedede sp. nov., male imago, paratype: genitalia, ventral view." httpUri="https://zenodo.org/record/280264/files/figure.png" pageId="4" pageNumber="32">Fig. 4</figureCitation>
). Penis with contrasting brown pigmentation. Penis lobes somewhat squared basally with abrupt step at outer margin of basal parts of penis lobes. Lobes separated distally with wide interspace, U or Vshaped (
<figureCitation id="135E19BCFFDD117EFBD4F86BFB7A8FAB" box="[1106,1216,1995,2019]" captionStart="FIGURES 5 6" captionStartId="4.[151,264,1825,1848]" captionTargetBox="[409,1186,217,1770]" captionTargetId="figure@4.[379,1208,193,1786]" captionTargetPageId="4" captionText="FIGURES 5 6. Electrogena gibedede sp. nov., male imago, paratype: 5, penis lobes, ventral view; 6, penis lobes, dorsal view." httpUri="https://zenodo.org/record/280265/files/figure.png" pageId="4" pageNumber="32">Figs. 5, 6</figureCitation>
,
<figureCitation id="135E19BCFFDD117EFB4BF86BFAA28FAB" box="[1229,1304,1995,2019]" captionStart="FIGURES 21 26" captionStartId="9.[151,264,1898,1921]" captionTargetBox="[172,1416,351,1853]" captionTargetId="figure@9.[151,1436,351,1861]" captionTargetPageId="9" captionText="FIGURES 21 26. Electrogena gibedede sp. nov., male imago (21, 22) and mature larva (23 26): 21, penis lobes, ventral view; 22, genitalia, ventral view; 23, forefemora bristles, dorsal view; 24, forefemora bristles (detail), dorsal view; 25, hind femora bristles, dorsal view; 26, hind femora bristles (detail), dorsal view." httpUri="https://zenodo.org/record/280269/files/figure.png" pageId="4" pageNumber="32">21, 22</figureCitation>
). On dorsal side of penis lobes 12 larger teeth situated near lateral margin, may be accompanied by 12 smaller teeth, reaching length of approximately 1/3 of larger ones (
<figureCitation id="135E19BCFFDC117FFDF8FF1CFD78889C" box="[638,706,188,212]" captionStart="FIGURES 5 6" captionStartId="4.[151,264,1825,1848]" captionTargetBox="[409,1186,217,1770]" captionTargetId="figure@4.[379,1208,193,1786]" captionTargetPageId="4" captionText="FIGURES 5 6. Electrogena gibedede sp. nov., male imago, paratype: 5, penis lobes, ventral view; 6, penis lobes, dorsal view." httpUri="https://zenodo.org/record/280265/files/figure.png" pageId="5" pageNumber="33">Fig. 6</figureCitation>
). Titillators with small teeth apically.
</paragraph>
<paragraph id="8BDA0539FFDC117FFF4FFF40FDAB8954" blockId="5.[151,1436,151,284]" pageId="5" pageNumber="33">
<emphasis id="B911D92BFFDC117FFF4FFF40FEC988B0" box="[201,371,224,248]" italics="true" pageId="5" pageNumber="33">Female imago.</emphasis>
Size: body length
<quantity id="4C9DA8DCFFDC117FFDC8FF7FFD7F88BF" box="[590,709,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="5" pageNumber="33" unit="mm" value="8.25" valueMax="8.5" valueMin="8.0">88.5 mm</quantity>
, forewing length
<quantity id="4C9DA8DCFFDC117FFC09FF7FFBA988BF" box="[911,1043,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" metricValueMax="10.2" metricValueMin="9.0" pageId="5" pageNumber="33" unit="mm" value="9.6" valueMax="10.2" valueMin="9.0">910.2 mm</quantity>
, cerci length
<quantity id="4C9DA8DCFFDC117FFB36FF7FFA9188BF" box="[1200,1323,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="5" pageNumber="33" unit="mm" value="17.0" valueMax="18.0" valueMin="16.0">1618 mm</quantity>
(approximately 2 times longer than body).
</paragraph>
<caption id="DF1A55B1FFDC117FFF11F81CFA278FB9" httpUri="https://zenodo.org/record/280266/files/figure.png" pageId="5" pageNumber="33" targetBox="[165,1411,338,1946]" targetPageId="5">
<paragraph id="8BDA0539FFDC117FFF11F81CFA278FB9" blockId="5.[151,1437,1980,2034]" pageId="5" pageNumber="33">
<emphasis id="B911D92BFFDC117FFF11F81CFEFF8F9A" bold="true" box="[151,325,1980,2003]" pageId="5" pageNumber="33">FIGURES 711.</emphasis>
<taxonomicName id="4C657EBAFFDC117FFECDF81DFD9C8F9B" box="[331,550,1981,2003]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="33" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFDC117FFECDF81DFD9C8F9B" box="[331,550,1981,2003]" italics="true" pageId="5" pageNumber="33">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFDC117FFDABF81DFDC78F9A" box="[557,637,1981,2002]" pageId="5" pageNumber="33" rank="species">
<emphasis id="B911D92BFFDC117FFDABF81DFDC78F9A" bold="true" box="[557,637,1981,2002]" pageId="5" pageNumber="33">sp. nov.</emphasis>
</taxonomicNameLabel>
, mature larva, paratype: 7, head, dorsal view; 8, hypopharynx, ventral view; 9, kinds of labrum shape, dorsal view; 10, variations of glossae shape, ventral view; 11, first segment of maxillary palp, dorsal view.
</paragraph>
</caption>
<caption id="DF1A55B1FFDF117CFF11F9FFFECF8EDC" httpUri="https://zenodo.org/record/280267/files/figure.png" pageId="6" pageNumber="34" targetBox="[267,1320,228,1604]" targetPageId="6">
<paragraph id="8BDA0539FFDF117CFF11F9FFFECF8EDC" blockId="6.[151,1436,1631,1684]" pageId="6" pageNumber="34">
<emphasis id="B911D92BFFDF117CFF11F9FFFEE98E3D" bold="true" box="[151,339,1631,1654]" pageId="6" pageNumber="34">FIGURES 1214.</emphasis>
<taxonomicName id="4C657EBAFFDF117CFEDEF9C0FD8F8E3E" box="[344,565,1632,1654]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="34" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFDF117CFEDEF9C0FD8F8E3E" box="[344,565,1632,1654]" italics="true" pageId="6" pageNumber="34">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFDF117CFDBCF9C0FD308E3D" box="[570,650,1632,1653]" pageId="6" pageNumber="34" rank="species">
<emphasis id="B911D92BFFDF117CFDBCF9C0FD308E3D" bold="true" box="[570,650,1632,1653]" pageId="6" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
, mature larva, paratype: 12, middle leg, dorsal view; 13, gill VII, dorsal view; 14, gill I, dorsal view.
</paragraph>
</caption>
<paragraph id="8BDA0539FFDF117CFF4FF91CFB218E9C" blockId="6.[151,1437,1724,2037]" box="[201,1179,1724,1749]" pageId="6" pageNumber="34">
Head brown, compound eyes and ocelli black (
<figureCitation id="135E19BCFFDF117CFD51F91CFC908E9C" box="[727,810,1724,1748]" captionStart="FIGURES 15 20" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[178,1407,454,1744]" captionTargetId="figure@7.[151,1436,445,1822]" captionTargetPageId="7" captionText="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="6" pageNumber="34">Fig. 18</figureCitation>
). Antennae brown, paler basally.
</paragraph>
<paragraph id="8BDA0539FFDF117CFF4FF97FFC798F2C" blockId="6.[151,1437,1724,2037]" pageId="6" pageNumber="34">Thorax dorsally and ventrally dark brown. Lateral portions yellowish. Legs more robust than in male imago. Femora and tibiae of similar coloration as in male imago. Tarsi uniformly dark brown. In some specimens middle and hind tarsi paler centrally (segments III and IV). Wings hyaline, transparent, with well visible brown venation. Costal and subcostal field slightly milky (mainly in pterostigmatic area).</paragraph>
<paragraph id="8BDA0539FFDF117CFF4FF8CFFA3A8FE4" blockId="6.[151,1437,1724,2037]" pageId="6" pageNumber="34">
Color pattern of abdomen similar to those in male imago, slightly paler (
<figureCitation id="135E19BCFFDF117CFB82F8CFFBE08FCF" box="[1028,1114,1903,1927]" captionStart="FIGURES 15 20" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[178,1407,454,1744]" captionTargetId="figure@7.[151,1436,445,1822]" captionTargetPageId="7" captionText="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="6" pageNumber="34">Fig. 20</figureCitation>
). Neural ganglion visible on sternite VII. Subgenital plate widely rounded, reaching articulation of segment IX. Subanal plate distinctly bent.
</paragraph>
<paragraph id="8BDA0539FFDF117CFF4FF817FA278FBC" blockId="6.[151,1437,1724,2037]" pageId="6" pageNumber="34">Cerci dark brown basally. Towards distal end getting paler, with dark bands following individual articulations. Every second articulation more pronounced, accompanied by more distinct dark band than those between them.</paragraph>
<paragraph id="8BDA0539FFDE117DFF11FF37FD89889C" blockId="7.[151,1436,151,391]" pageId="7" pageNumber="35">These differences more distinct than in male imago and disappear only in distal third of cerci. All surface covered with dark hairs. Paracercus vestigial.</paragraph>
<paragraph id="8BDA0539FFDE117DFF41FF40FB0888BF" blockId="7.[151,1436,151,391]" box="[199,1202,223,248]" pageId="7" pageNumber="35">
<emphasis id="B911D92BFFDE117DFF41FF40FECE88B0" box="[199,372,224,248]" italics="true" pageId="7" pageNumber="35">Male subimago</emphasis>
Size: body length
<quantity id="4C9DA8DCFFDE117DFDC0FF7FFD3388BF" box="[582,649,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="35" unit="mm" value="8.0">8 mm</quantity>
, forewing length
<quantity id="4C9DA8DCFFDE117DFCCBFF7FFC1688BF" box="[845,940,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="7" pageNumber="35" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
, cerci length unknown.
</paragraph>
<paragraph id="8BDA0539FFDE117DFF4FFEA4FD978954" blockId="7.[151,1436,151,391]" box="[201,557,260,284]" pageId="7" pageNumber="35">Head and antennae dark brown.</paragraph>
<paragraph id="8BDA0539FFDE117DFF4FFE87FAA089CF" blockId="7.[151,1436,151,391]" pageId="7" pageNumber="35">Prothorax brownish. Mesothorax dorsally pale yellowish (especially in central part), with dark brown areas frontally, caudally and laterally. Lateral portions of mesothorax between wing insertion and coxae pale yellowish, with occasional darker sclerites. Ventral side of thorax mainly uniformly brownish or yellowish brown.</paragraph>
<caption id="DF1A55B1FFDE117DFF11F893FCC68FCF" httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="7" pageNumber="35" targetBox="[178,1407,454,1744]" targetPageId="7">
<paragraph id="8BDA0539FFDE117DFF11F893FCC68FCF" blockId="7.[151,1437,1843,1927]" pageId="7" pageNumber="35">
<emphasis id="B911D92BFFDE117DFF11F893FEE98F01" bold="true" box="[151,339,1843,1866]" pageId="7" pageNumber="35">FIGURES 1520.</emphasis>
<taxonomicName id="4C657EBAFFDE117DFEDEF894FD8E8F02" box="[344,564,1844,1866]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="35" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFDE117DFEDEF894FD8E8F02" box="[344,564,1844,1866]" italics="true" pageId="7" pageNumber="35">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFDE117DFDBCF894FD308F01" box="[570,650,1844,1865]" pageId="7" pageNumber="35" rank="species">
<emphasis id="B911D92BFFDE117DFDBCF894FD308F01" bold="true" box="[570,650,1844,1865]" pageId="7" pageNumber="35">sp. nov.</emphasis>
</taxonomicNameLabel>
, mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view.
</paragraph>
</caption>
<paragraph id="8BDA0539FFDE1172FF4FF80EFEE289CF" blockId="7.[151,1436,1966,2026]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="36" pageId="7" pageNumber="35">
Forefemora brown, with darker margins and two darker transversal bands distally (at approximately 2/3 of the femur length and near articulation with tibia). Foretibiae uniformly brown, slightly darker at articulations. Tarsi 5 segmented, with one sharp and one blunt claw. Tarsal segments IIV of the same color as tibia, segment V and claws darker. Middle and hind legs generally paler than forelegs. Middle femora yellowish brown, with brown patch at approximately 3/5 of femur length. Further area dark brown. Middle tibia of same color as middle femur, dark brown only at articulation with femur. Middle tarsus 5segmented, with one sharp and one blunt claw. Tarsal segments brownish, most distal segment and claws darker.
<collectingCountry id="F37245A9FFD11172FCC5FE87FCC18977" box="[835,891,295,319]" name="India" pageId="8" pageNumber="36">Hind</collectingCountry>
legs of same arrangement and color pattern as middle ones. Wings dull brownish with dark brown venation. No patches of darker coloration.
<collectingCountry id="F37245A9FFD11172FB54FEECFAB0892C" box="[1234,1290,332,356]" name="India" pageId="8" pageNumber="36">Hind</collectingCountry>
margin with row of tiny hairs.
</paragraph>
<paragraph id="8BDA0539FFD11172FF4FFE34FB368ACC" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="36">
Abdominal tergites IX brown with two hooked, approximately
<emphasis id="B911D92BFFD11172FC1BFE35FC1689E4" box="[925,940,405,428]" italics="true" pageId="8" pageNumber="36">L</emphasis>
shaped whitish spots situated laterally near the tergo-sternal suture. Longitudinal part of each
<emphasis id="B911D92BFFD11172FD40FE19FD6F8998" box="[710,725,441,464]" italics="true" pageId="8" pageNumber="36">L</emphasis>
shaped spot is pointing caudally. On tergites I, IX and X, these spots are not pronounced in some specimens.
<emphasis id="B911D92BFFD11172FD11FE7DFD1C89BC" box="[663,678,477,500]" italics="true" pageId="8" pageNumber="36">L</emphasis>
shaped spot may be accompanied at its caudal end by a single pale dot. Presence of further two longitudinally elongated whitish spots in central part of tergites IVX. These spots situated near fore margin of each tergite. Central part of tergites IIII paler, thus elongated spots are not apparent. Sternites IIVIII yellowish brown with white pattern in central part, consisting of two oblique elongated spots situated frontally and two white dots caudally. Sternites IX and X brownish, sternite I whitish.
</paragraph>
<paragraph id="8BDA0539FFD11172FF4FFD30FAA08B5C" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="36">
<emphasis id="B911D92BFFD11172FF4FFD30FE898AE0" box="[201,307,656,680]" italics="true" pageId="8" pageNumber="36">Genitalia</emphasis>
: Penis lobes yellowish, titillators dark brown. Otherwise penis lobes without any dark markings. Incision between penis lobes indistinct. Typical shape of penis (squared, with abrupt, almost square-angled step at outer margin of basal parts of penis lobes) already well apparent. Titillators with teeth. Penis lobes, styliger plate and forceps densely covered with short hairs. On surface of forceps tiny articulated spines also present.
</paragraph>
<paragraph id="8BDA0539FFD11172FF4FFCBFFAB08B7F" blockId="8.[151,1437,151,2013]" box="[201,1290,799,825]" pageId="8" pageNumber="36">Cerci dark brown, covered with short hairs. Individual articulations of segments with dark bands.</paragraph>
<paragraph id="8BDA0539FFD11172FF41FCE4FE8D8B37" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="36">
<emphasis id="B911D92BFFD11172FF41FCE4FE2C8B14" box="[199,406,836,860]" italics="true" pageId="8" pageNumber="36">Female subimago.</emphasis>
Size: body length
<quantity id="4C9DA8DCFFD11172FDE3FCE4FD128B14" box="[613,680,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="36" unit="mm" value="8.0">8 mm</quantity>
, forewing length
<quantity id="4C9DA8DCFFD11172FCECFCE4FC568B14" box="[874,1004,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="10.0" metricValueMin="9.5" pageId="8" pageNumber="36" unit="mm" value="9.75" valueMax="10.0" valueMin="9.5">9.510 mm</quantity>
, cerci length
<quantity id="4C9DA8DCFFD11172FB06FCE4FB778B14" box="[1152,1229,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="36" unit="mm" value="11.0">11 mm</quantity>
(approximately as long as body).
</paragraph>
<paragraph id="8BDA0539FFD11172FF4FFC2CFD978BEC" blockId="8.[151,1437,151,2013]" box="[201,557,908,932]" pageId="8" pageNumber="36">Head and antennae dark brown.</paragraph>
<paragraph id="8BDA0539FFD11172FF4FFC0FFD548BA4" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="36">Dorsal side of prothorax brown, whitish posteromedially. Mesothorax of same color pattern as in male imago. Metathorax pale brown, darker at base of hind wings.</paragraph>
<paragraph id="8BDA0539FFD11172FF4FFC59FBE98C7C" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="36">
Legs of same arrangement and color pattern as in male imago. Wings uniformly dull brownish with dark brown venation. No patches of darker coloration.
<collectingCountry id="F37245A9FFD11172FD42FBBCFD468C7C" box="[708,764,1052,1076]" name="India" pageId="8" pageNumber="36">Hind</collectingCountry>
margin with row of tiny hairs.
</paragraph>
<paragraph id="8BDA0539FFD11172FF4FFBE0FCAB8C10" blockId="8.[151,1437,151,2013]" box="[201,785,1088,1112]" pageId="8" pageNumber="36">Abdominal color pattern same as in male subimago.</paragraph>
<paragraph id="8BDA0539FFD11172FF4FFBC4FB718C34" blockId="8.[151,1437,151,2013]" box="[201,1227,1124,1148]" pageId="8" pageNumber="36">Cerci dark brown, evenly covered with short hairs. Darker bands at individual articulations.</paragraph>
<paragraph id="8BDA0539FFD11172FF4FFB29FEBB8CA0" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="36">
<emphasis id="B911D92BFFD11172FF4FFB29FED28CE8" box="[201,360,1160,1184]" italics="true" pageId="8" pageNumber="36">Mature larva.</emphasis>
Size: body length (slightly differs between males and females): in male
<quantity id="4C9DA8DCFFD11172FB41FB28FAEC8CE8" box="[1223,1366,1160,1184]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="6.9" metricValueMin="5.8" pageId="8" pageNumber="36" unit="mm" value="6.35" valueMax="6.9" valueMin="5.8">5.86.9 mm</quantity>
; cerci length
<quantity id="4C9DA8DCFFD11172FF65FB0CFED68C8C" box="[227,364,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="7.8999999999999995" metricValueMax="8.2" metricValueMin="7.6" pageId="8" pageNumber="36" unit="mm" value="7.8999999999999995" valueMax="8.2" valueMin="7.6">7.68.2 mm</quantity>
, paracercus length 8.69.0 mm; in female 7.08.0 mm, cerci length 10.0 mm, paracercus length 11.0 mm.
</paragraph>
<paragraph id="8BDA0539FFD11172FF4FFB54FE068D1C" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="36">
Head brownish with apparent light pattern, consisting of two spots near fore margin approximately in front of lateral ocelli. Further two light spots elongated transversally, situated near antennal bases (
<figureCitation id="135E19BCFFD11172FB03FAB8FB6E8D78" box="[1157,1236,1304,1328]" captionStart="FIGURES 7 11" captionStartId="5.[151,263,1980,2003]" captionTargetBox="[165,1411,338,1946]" captionTargetId="figure@5.[151,1436,337,1959]" captionTargetPageId="5" captionText="FIGURES 7 11. Electrogena gibedede sp. nov., mature larva, paratype: 7, head, dorsal view; 8, hypopharynx, ventral view; 9, kinds of labrum shape, dorsal view; 10, variations of glossae shape, ventral view; 11, first segment of maxillary palp, dorsal view." httpUri="https://zenodo.org/record/280266/files/figure.png" pageId="8" pageNumber="36">Figs. 7</figureCitation>
,
<figureCitation id="135E19BCFFD11172FB59FAB8FB448D78" box="[1247,1278,1304,1328]" captionStart="FIGURES 15 20" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[178,1407,454,1744]" captionTargetId="figure@7.[151,1436,445,1822]" captionTargetPageId="7" captionText="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="8" pageNumber="36">15</figureCitation>
). Lateral margins of head light colored.
</paragraph>
<paragraph id="8BDA0539FFD11172FF4FFAC0FB3E8D88" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="36">
Prothorax yellowish with dark brown pattern. Lateral margins whitish. Longitudinal pale line in middle, extending towards its ends. Dark smudges laterally from this line, interrupted by pale areas near fore and hind margins of prothorax. Meso- and metathorax yellowish with darker brown smudges (
<figureCitation id="135E19BCFFD11172FBA2FA08FBCD8D88" box="[1060,1143,1448,1472]" captionStart="FIGURES 15 20" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[178,1407,454,1744]" captionTargetId="figure@7.[151,1436,445,1822]" captionTargetPageId="7" captionText="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="8" pageNumber="36">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8BDA0539FFD11172FF4FFA6DFE338EA8" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="36">
Legs yellowish, with dark brown pattern. Femora with four distinct elongated dark spots dorsally; distal two spots partly fused near tibia insertion (
<figureCitation id="135E19BCFFD11172FDC4FA50FD2F8E40" box="[578,661,1520,1544]" captionStart="FIGURES 12 14" captionStartId="6.[151,264,1631,1654]" captionTargetBox="[267,1320,228,1604]" captionTargetId="figure@6.[256,1330,193,1610]" captionTargetPageId="6" captionText="FIGURES 12 14. Electrogena gibedede sp. nov., mature larva, paratype: 12, middle leg, dorsal view; 13, gill VII, dorsal view; 14, gill I, dorsal view." httpUri="https://zenodo.org/record/280267/files/figure.png" pageId="8" pageNumber="36">Fig. 12</figureCitation>
). Tibiae uniformly yellowish, sometimes with dark smudge in central part. Tarsi brownish, indistinctly darker apically. Shape of bristles on dorsal surface of femora in
<taxonomicName id="4C657EBAFFD11172FB52F9B4FF768E18" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="36" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD11172FB52F9B4FF768E18" italics="true" pageId="8" pageNumber="36">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD11172FF52F998FE948E18" box="[212,302,1592,1616]" pageId="8" pageNumber="36" rank="species">
<emphasis id="B911D92BFFD11172FF52F998FE948E18" bold="true" box="[212,302,1592,1616]" pageId="8" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
differs depending on leg pair and exact location of particular bristle on surface of respective femur. Bristles on forefemora unique among representatives of genus
<taxonomicName id="4C657EBAFFD11172FCEEF9FCFC4A8E3C" box="[872,1008,1628,1652]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD11172FCEEF9FCFC4A8E3C" box="[872,1008,1628,1652]" italics="true" pageId="8" pageNumber="36">Electrogena</emphasis>
</taxonomicName>
, rounded and widened distally (
<figureCitation id="135E19BCFFD11172FAE5F9FCFF588ED0" captionStart="FIGURES 21 26" captionStartId="9.[151,264,1898,1921]" captionTargetBox="[172,1416,351,1853]" captionTargetId="figure@9.[151,1436,351,1861]" captionTargetPageId="9" captionText="FIGURES 21 26. Electrogena gibedede sp. nov., male imago (21, 22) and mature larva (23 26): 21, penis lobes, ventral view; 22, genitalia, ventral view; 23, forefemora bristles, dorsal view; 24, forefemora bristles (detail), dorsal view; 25, hind femora bristles, dorsal view; 26, hind femora bristles (detail), dorsal view." httpUri="https://zenodo.org/record/280269/files/figure.png" pageId="8" pageNumber="36">Figs. 23, 24</figureCitation>
). Bristles of middle and hind femora more bluntly pointed or pointed, spine-like (
<figureCitation id="135E19BCFFD11172FBFAF920FABD8ED0" box="[1148,1287,1664,1688]" captionStart="FIGURES 21 26" captionStartId="9.[151,264,1898,1921]" captionTargetBox="[172,1416,351,1853]" captionTargetId="figure@9.[151,1436,351,1861]" captionTargetPageId="9" captionText="FIGURES 21 26. Electrogena gibedede sp. nov., male imago (21, 22) and mature larva (23 26): 21, penis lobes, ventral view; 22, genitalia, ventral view; 23, forefemora bristles, dorsal view; 24, forefemora bristles (detail), dorsal view; 25, hind femora bristles, dorsal view; 26, hind femora bristles (detail), dorsal view." httpUri="https://zenodo.org/record/280269/files/figure.png" pageId="8" pageNumber="36">Figs. 25, 26</figureCitation>
). Middle and hind tibiae with rows of hairs, approximately as long as tibia width.
<collectingCountry id="F37245A9FFD11172FC16F904FC728EF4" box="[912,968,1700,1724]" name="India" pageId="8" pageNumber="36">Hind</collectingCountry>
tibiae with rows of hairs accompanied by short pointed bristles.
</paragraph>
<paragraph id="8BDA0539FFD11173FF4FF94CFB108954" blockId="8.[151,1437,151,2013]" lastBlockId="9.[151,1436,151,284]" lastPageId="9" lastPageNumber="37" pageId="8" pageNumber="36">
Abdominal tergites with pronounced dark brown pattern and only several isolated spots of yellowish color. Each tergite with two light spots placed centrally and further two spots situated more posterolaterally, near hind margin of respective tergite. Moreover, another light smudge may appear posteromedially. This smudge may fuse with two light spots in central part of tergite and connect them together in some cases. Sometimes this light smudge extends even to further two posterolateral spots, forming band of lighter coloration along hind margin of tergite. Extension of posteromedial smudge is common mainly on tergites III, IVV and VIIIX. Therefore tergites III, VI and X appears darker than rest of abdomen (tergite X sometimes even completely dark, without any lighter pattern) (
<figureCitation id="135E19BCFFD01173FF5AFF37FE8B88E7" box="[220,305,151,175]" captionStart="FIGURES 15 20" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[178,1407,454,1744]" captionTargetId="figure@7.[151,1436,445,1822]" captionTargetPageId="7" captionText="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="9" pageNumber="37">Fig. 15</figureCitation>
). Isolated broad lighter areas present also laterally, near gill insertions. Abdominal sternites yellowish, well visible neural ganglion in segment VII; brown pattern of sternites mainly absent (
<figureCitation id="135E19BCFFD01173FB41FF1CFAA5889C" box="[1223,1311,188,212]" captionStart="FIGURES 15 20" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[178,1407,454,1744]" captionTargetId="figure@7.[151,1436,445,1822]" captionTargetPageId="7" captionText="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="9" pageNumber="37">Fig. 16</figureCitation>
). Posterior margin of tergites with the dense row of large spines pointed apically (
<figureCitation id="135E19BCFFD01173FC2DFF7FFB8988BF" box="[939,1075,223,247]" captionStart="FIGURES 27 30" captionStartId="10.[151,264,1220,1243]" captionTargetBox="[170,1415,186,1184]" captionTargetId="figure@10.[151,1436,163,1199]" captionTargetPageId="10" captionText="FIGURES 27 30. Electrogena gibedede sp. nov., mature larva, paratype: 27, 28, posterior margin of abdominal tergites, dorsal view; 29, 30, tarsal claw, dorsal and ventral view." httpUri="https://zenodo.org/record/280270/files/figure.png" pageId="9" pageNumber="37">Figs. 27, 28</figureCitation>
). Gills whitish with pale brownish tracheization; marginal areas transparent (
<figureCitation id="135E19BCFFD01173FD12FEA4FCA78954" box="[660,797,260,284]" captionStart="FIGURES 12 14" captionStartId="6.[151,264,1631,1654]" captionTargetBox="[267,1320,228,1604]" captionTargetId="figure@6.[256,1330,193,1610]" captionTargetPageId="6" captionText="FIGURES 12 14. Electrogena gibedede sp. nov., mature larva, paratype: 12, middle leg, dorsal view; 13, gill VII, dorsal view; 14, gill I, dorsal view." httpUri="https://zenodo.org/record/280267/files/figure.png" pageId="9" pageNumber="37">Figs. 13, 14</figureCitation>
). Cerci yellow to yellowish-brown.
</paragraph>
<caption id="DF1A55B1FFD01173FF11F8CAFC8D8FF6" httpUri="https://zenodo.org/record/280269/files/figure.png" pageId="9" pageNumber="37" targetBox="[172,1416,351,1853]" targetPageId="9">
<paragraph id="8BDA0539FFD01173FF11F8CAFC8D8FF6" blockId="9.[151,1436,1898,1982]" pageId="9" pageNumber="37">
<emphasis id="B911D92BFFD01173FF11F8CAFEE98FC8" bold="true" box="[151,339,1898,1921]" pageId="9" pageNumber="37">FIGURES 2126.</emphasis>
<taxonomicName id="4C657EBAFFD01173FEDFF8CBFD8E8FC9" box="[345,564,1899,1921]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="37" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD01173FEDFF8CBFD8E8FC9" box="[345,564,1899,1921]" italics="true" pageId="9" pageNumber="37">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD01173FDBDF8CBFD308FC8" box="[571,650,1899,1920]" pageId="9" pageNumber="37" rank="species">
<emphasis id="B911D92BFFD01173FDBDF8CBFD308FC8" bold="true" box="[571,650,1899,1920]" pageId="9" pageNumber="37">sp. nov.</emphasis>
</taxonomicNameLabel>
, male imago (21, 22) and mature larva (2326): 21, penis lobes, ventral view; 22, genitalia, ventral view; 23, forefemora bristles, dorsal view; 24, forefemora bristles (detail), dorsal view; 25, hind femora bristles, dorsal view; 26, hind femora bristles (detail), dorsal view.
</paragraph>
</caption>
<caption id="DF1A55B1FFD31170FF11FB64FD368CB1" httpUri="https://zenodo.org/record/280270/files/figure.png" pageId="10" pageNumber="38" targetBox="[170,1415,186,1184]" targetPageId="10">
<paragraph id="8BDA0539FFD31170FF11FB64FD368CB1" blockId="10.[151,1436,1220,1273]" pageId="10" pageNumber="38">
<emphasis id="B911D92BFFD31170FF11FB64FEE98C92" bold="true" box="[151,339,1220,1243]" pageId="10" pageNumber="38">FIGURES 2730.</emphasis>
<taxonomicName id="4C657EBAFFD31170FEDEFB65FD8E8C93" box="[344,564,1221,1243]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD31170FEDEFB65FD8E8C93" box="[344,564,1221,1243]" italics="true" pageId="10" pageNumber="38">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD31170FDBCFB65FD308C92" box="[570,650,1221,1242]" pageId="10" pageNumber="38" rank="species">
<emphasis id="B911D92BFFD31170FDBCFB65FD308C92" bold="true" box="[570,650,1221,1242]" pageId="10" pageNumber="38">sp. nov.</emphasis>
</taxonomicNameLabel>
, mature larva, paratype: 27, 28, posterior margin of abdominal tergites, dorsal view; 29, 30, tarsal claw, dorsal and ventral view.
</paragraph>
</caption>
<paragraph id="8BDA0539FFD31170FF41FA87FCD88E16" blockId="10.[151,1437,1318,2026]" pageId="10" pageNumber="38">
A set of standard larval diagnostic characteristics for identification of
<taxonomicName id="4C657EBAFFD31170FC65FA86FBD18D76" box="[995,1131,1318,1342]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD31170FC65FA86FBD18D76" box="[995,1131,1318,1342]" italics="true" pageId="10" pageNumber="38">Electrogena</emphasis>
</taxonomicName>
species is provided below. In the present study we provide states and values for 30 characteristics of
<taxonomicName id="4C657EBAFFD31170FC60FAEAFB618D2A" box="[998,1243,1354,1378]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD31170FC60FAEAFB618D2A" box="[998,1243,1354,1378]" italics="true" pageId="10" pageNumber="38">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD31170FB62FAEAFA858D2A" box="[1252,1343,1354,1378]" pageId="10" pageNumber="38" rank="species">
<emphasis id="B911D92BFFD31170FB62FAEAFA858D2A" bold="true" box="[1252,1343,1354,1378]" pageId="10" pageNumber="38">sp. nov.</emphasis>
</taxonomicNameLabel>
Several features treated in the present study had not previously been used for the genus
<taxonomicName id="4C657EBAFFD31170FBA5FACEFB118DCE" box="[1059,1195,1390,1414]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD31170FBA5FACEFB118DCE" box="[1059,1195,1390,1414]" italics="true" pageId="10" pageNumber="38">Electrogena</emphasis>
</taxonomicName>
, but Haybach (1999) used them for the differentiation of some species of the genus
<taxonomicName id="4C657EBAFFD31170FCC8FA32FBD88DE2" authority="Eaton, 1868" authorityName="Eaton" authorityYear="1868" box="[846,1122,1426,1450]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD31170FCC8FA32FC6E8DE2" box="[846,980,1426,1450]" italics="true" pageId="10" pageNumber="38">Ecdyonurus</emphasis>
Eaton, 1868
</taxonomicName>
(
<taxonomicName id="4C657EBAFFD31170FBF4FA33FB508DE2" box="[1138,1258,1427,1450]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="venosus">
<emphasis id="B911D92BFFD31170FBF4FA33FB508DE2" box="[1138,1258,1427,1450]" italics="true" pageId="10" pageNumber="38">E. venosus</emphasis>
</taxonomicName>
species-group). The set comprises namely four characteristics concerning setae on the maxillary palp (N_PLS, N_PLBas, N_VEN, N_LPH). Since the structure of the maxilla of the genus
<taxonomicName id="4C657EBAFFD31170FC8AFA7AFC2E8DBA" box="[780,916,1498,1522]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD31170FC8AFA7AFC2E8DBA" box="[780,916,1498,1522]" italics="true" pageId="10" pageNumber="38">Electrogena</emphasis>
</taxonomicName>
is very similar to that of representatives of the
<taxonomicName id="4C657EBAFFD31170FF11FA5FFEB48E5E" box="[151,270,1535,1558]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="venosus">
<emphasis id="B911D92BFFD31170FF11FA5FFEB48E5E" box="[151,270,1535,1558]" italics="true" pageId="10" pageNumber="38">E. venosus</emphasis>
</taxonomicName>
species-group, these characteristics are used for the genus
<taxonomicName id="4C657EBAFFD31170FC26FA5EFB928E5E" box="[928,1064,1534,1558]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD31170FC26FA5EFB928E5E" box="[928,1064,1534,1558]" italics="true" pageId="10" pageNumber="38">Electrogena</emphasis>
</taxonomicName>
here as well. One new characteristic (S_1GI) is added to the series of the standard diagnostic characteristics. It describes the shape of the first gill plate, which is unique in some
<taxonomicName id="4C657EBAFFD31170FE74F9E6FDC08E16" box="[498,634,1606,1630]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD31170FE74F9E6FDC08E16" box="[498,634,1606,1630]" italics="true" pageId="10" pageNumber="38">Electrogena</emphasis>
</taxonomicName>
species (see below).
</paragraph>
<paragraph id="8BDA0539FFD31170FF4FF9CAFC578E82" blockId="10.[151,1437,1318,2026]" pageId="10" pageNumber="38">
Quantitative characteristics were measured in
<specimenCount id="9D63CEB0FFD31170FD4CF9CAFCDB8ECA" box="[714,865,1642,1666]" pageId="10" pageNumber="38" type="generic">13 specimens</specimenCount>
. Most features states are compared with the related species (focusing mainly on the species from the closely situated geographical regions, e.g. Caucasus Mts., Crimean Peninsula and
<collectingCountry id="F37245A9FFD31170FE27F912FDB98E82" box="[417,515,1714,1738]" name="Turkey" pageId="10" pageNumber="38">Anatolia</collectingCountry>
) and respective differences are pointed out.
</paragraph>
<paragraph id="8BDA0539FFD31170FF4FF976FBE28FEA" blockId="10.[151,1437,1318,2026]" pageId="10" pageNumber="38">
Representatives of the genus
<taxonomicName id="4C657EBAFFD31170FD92F976FD268EA6" box="[532,668,1750,1774]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD31170FD92F976FD268EA6" box="[532,668,1750,1774]" italics="true" pageId="10" pageNumber="38">Electrogena</emphasis>
</taxonomicName>
with known larvae, mostly recorded from the Caucasus Mts. and/or adjacent areas, which were taken for comparison in the list of the standard diagnostic features below, encompass the following species:
<taxonomicName id="4C657EBAFFD31170FE13F8BFFD258F7E" authority="Eaton, 1883" authorityName="Eaton" authorityYear="1883" box="[405,671,1822,1846]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B911D92BFFD31170FE13F8BFFE408F7E" box="[405,506,1822,1846]" italics="true" pageId="10" pageNumber="38">E. affinis</emphasis>
(Eaton, 1883)
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD31170FD2DF8BFFB3C8F7E" authority="Braasch &amp; Kazanci, 1986" authorityName="Braasch &amp; Kazanci" authorityYear="1986" box="[683,1158,1822,1846]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD31170FD2DF8BFFCF38F7E" box="[683,841,1822,1846]" italics="true" pageId="10" pageNumber="38">E. antalyensis</emphasis>
(Braasch &amp;
<bibRefCitation id="EFF478C8FFD31170FC5FF8BFFBC58F7E" author="Kazanci" box="[985,1151,1822,1846]" pageId="10" pageNumber="38" refString="Kazanci, N. (1986) A new Ephemeroptera (Heptageniidae) species from Anatolia. Do g a Turk Biyoloji Dergisi, 10, 391 - 393." type="journal article" year="1986">Kazancı, 1986</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD31170FB15F8BFFF608F12" authority="Braasch, 1983" authorityName="Braasch" authorityYear="1983" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="armeniaca">
<emphasis id="B911D92BFFD31170FB15F8BFFA938F7E" box="[1171,1321,1823,1846]" italics="true" pageId="10" pageNumber="38">E. armeniaca</emphasis>
(Braasch, 1983)
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD31170FF61F8E3FDC78F12" authority="Braasch, 1978" authorityName="Braasch" authorityYear="1978" box="[231,637,1858,1882]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="azerbajdshanica">
<emphasis id="B911D92BFFD31170FF61F8E3FE058F12" box="[231,447,1858,1882]" italics="true" pageId="10" pageNumber="38">E. azerbajdshanica</emphasis>
(
<bibRefCitation id="EFF478C8FFD31170FE49F8E2FDCE8F12" author="Braasch" box="[463,628,1858,1882]" pageId="10" pageNumber="38" refString="Braasch, D. (1978) Neue Ecdyonurus - Arten (Ephemeroptera, Heptageniidae) aus dem Kaukasus und Transkaukasien (Sowjetunion). I. Entomologische Nachrichten und Berichte, 22 (10), 145 - 159." type="journal article" year="1978">Braasch, 1978</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD31170FD0FF8E3FC168F12" authority="Sowa, 1984" authorityName="Sowa" authorityYear="1984" box="[649,940,1858,1882]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="braaschi">
<emphasis id="B911D92BFFD31170FD0FF8E3FCB08F12" box="[649,778,1858,1882]" italics="true" pageId="10" pageNumber="38">E. braaschi</emphasis>
(
<bibRefCitation id="EFF478C8FFD31170FC9CF8E2FC1E8F12" author="Sowa" box="[794,932,1858,1882]" pageId="10" pageNumber="38" refString="Sowa, R. (1984) Two new species of Ecdyonurus Eaton of lateralis (Curt.) group (Ephemeroptera, Heptageniidae) from the Crimea and Western Caucasus. Acta Hydrobiologica, 25 / 26 (2), 181 - 188." type="journal article" year="1984">Sowa, 1984</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD31170FC3EF8E3FABE8F12" authority="Demoulin, 1973" authorityName="Demoulin" authorityYear="1973" box="[952,1284,1858,1882]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD31170FC3EF8E3FB8B8F12" box="[952,1073,1858,1882]" italics="true" pageId="10" pageNumber="38">E. galileae</emphasis>
(
<bibRefCitation id="EFF478C8FFD31170FBC7F8E2FB468F12" author="Demoulin" box="[1089,1276,1858,1882]" pageId="10" pageNumber="38" refString="Demoulin, G. (1973) Contribution a l`etude des Ephemeropteres d`Israel. Introduction et I. Heptageniidae. Bulletin de l'Institut royal des Sciences naturelles de Belgique, 49 (8), 1 - 19." type="journal article" year="1973">Demoulin, 1973</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD31170FA97F8E3FEF78F36" authority="Braasch, 1978" authorityName="Braasch" authorityYear="1978" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="kuraensis">
<emphasis id="B911D92BFFD31170FA97F8E3FA268F12" box="[1297,1436,1858,1882]" italics="true" pageId="10" pageNumber="38">E. kuraensis</emphasis>
(
<bibRefCitation id="EFF478C8FFD31170FF18F8C6FEFF8F36" author="Braasch" box="[158,325,1894,1918]" pageId="10" pageNumber="38" refString="Braasch, D. (1978) Neue Ecdyonurus - Arten (Ephemeroptera, Heptageniidae) aus dem Kaukasus und Transkaukasien (Sowjetunion). I. Entomologische Nachrichten und Berichte, 22 (10), 145 - 159." type="journal article" year="1978">Braasch, 1978</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD31170FEDFF8C7FDC58F36" authority="Curtis, 1834" authorityName="Curtis" authorityYear="1834" box="[345,639,1894,1918]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B911D92BFFD31170FEDFF8C7FE6C8F36" box="[345,470,1894,1918]" italics="true" pageId="10" pageNumber="38">E. lateralis</emphasis>
(Curtis, 1834)
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD31170FD0DF8C7FC7D8F36" authority="Braasch, 1983" authorityName="Braasch" authorityYear="1983" box="[651,967,1894,1918]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="malickyi">
<emphasis id="B911D92BFFD31170FD0DF8C7FCB08F36" box="[651,778,1894,1918]" italics="true" pageId="10" pageNumber="38">E. malickyi</emphasis>
(Braasch, 1983)
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD31170FC52F8C7FB458F36" authority="Kazanci, 1987" authorityName="Kazanci" authorityYear="1987" box="[980,1279,1894,1918]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="necatii">
<emphasis id="B911D92BFFD31170FC52F8C7FBFA8F36" box="[980,1088,1895,1918]" italics="true" pageId="10" pageNumber="38">E. necatii</emphasis>
(
<bibRefCitation id="EFF478C8FFD31170FBC9F8C6FB4D8F36" author="Kazanci" box="[1103,1271,1894,1918]" pageId="10" pageNumber="38" refString="Kazanci, N. (1987) Ecdyonurus necatii, a New Ephemeroptera (Heptageniidae) Species from Turkey. Aquatic Insects, 9 (1), 17 - 20." type="journal article" year="1987">Kazancı, 1987</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD31170FA8AF8C7FECE8FEA" authority="Braasch, 1980" authorityName="Braasch" authorityYear="1980" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="pseudaffinis">
<emphasis id="B911D92BFFD31170FA8AF8C7FF028FEA" italics="true" pageId="10" pageNumber="38">E. pseudaffinis</emphasis>
(Braasch, 1980)
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD31170FEF9F82BFD7F8FEA" authority="Braasch, 1978" authorityName="Braasch" authorityYear="1978" box="[383,709,1930,1954]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="squamata">
<emphasis id="B911D92BFFD31170FEF9F82BFDB18FEA" box="[383,523,1931,1954]" italics="true" pageId="10" pageNumber="38">E. squamata</emphasis>
(
<bibRefCitation id="EFF478C8FFD31170FD9CF82AFD078FEA" author="Braasch" box="[538,701,1930,1954]" pageId="10" pageNumber="38" refString="Braasch, D. (1978) Neue Ecdyonurus - Arten (Ephemeroptera, Heptageniidae) aus dem Kaukasus und Transkaukasien (Sowjetunion). I. Entomologische Nachrichten und Berichte, 22 (10), 145 - 159." type="journal article" year="1978">Braasch, 1978</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C657EBAFFD31170FD7AF82BFBEE8FEA" authority="Sowa, 1984" authorityName="Sowa" authorityYear="1984" box="[764,1108,1930,1954]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B911D92BFFD31170FD7AF82BFC098FEA" box="[764,947,1931,1954]" italics="true" pageId="10" pageNumber="38">E. zimmermanni</emphasis>
(
<bibRefCitation id="EFF478C8FFD31170FC44F82AFBF68FEA" author="Sowa" box="[962,1100,1930,1954]" pageId="10" pageNumber="38" refString="Sowa, R. (1984) Two new species of Ecdyonurus Eaton of lateralis (Curt.) group (Ephemeroptera, Heptageniidae) from the Crimea and Western Caucasus. Acta Hydrobiologica, 25 / 26 (2), 181 - 188." type="journal article" year="1984">Sowa, 1984</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BDA0539FFD31170FF4FF80FFA268FA2" blockId="10.[151,1437,1318,2026]" pageId="10" pageNumber="38">
Due to the general lack of knowledge of the species of the genus
<taxonomicName id="4C657EBAFFD31170FC2AF80EFB8E8F8E" box="[940,1076,1966,1990]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD31170FC2AF80EFB8E8F8E" box="[940,1076,1966,1990]" italics="true" pageId="10" pageNumber="38">Electrogena</emphasis>
</taxonomicName>
in the Caucasus Mts., ratio values of quantitative characteristics used in the present study for comparison with
<taxonomicName id="4C657EBAFFD31170FBB8F872FA8F8FA2" box="[1086,1333,2002,2026]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="38" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD31170FBB8F872FA8F8FA2" box="[1086,1333,2002,2026]" italics="true" pageId="10" pageNumber="38">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD31170FAB9F872FA268FA2" box="[1343,1436,2002,2026]" pageId="10" pageNumber="38" rank="species">
<emphasis id="B911D92BFFD31170FAB9F872FA268FA2" bold="true" box="[1343,1436,2002,2026]" pageId="10" pageNumber="38">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BDA0539FFD21171FF11FF38FB3188B0" blockId="11.[151,1436,151,357]" pageId="11" pageNumber="39">were in some cases derived from the original drawings. Thus, they can be taken only as a pointer to the most pronounced differences. Moreover, the states of maxillae structure are unknown in most Caucasian species; since descriptions of the species do not often contain any specification of maxillae arrangement.</paragraph>
<paragraph id="8BDA0539FFD21171FF4FFEA5FC308977" blockId="11.[151,1436,151,357]" pageId="11" pageNumber="39">
Recent redescriptions of two species (
<taxonomicName id="4C657EBAFFD21171FDF2FEA5FD548954" box="[628,750,260,284]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD21171FDF2FEA5FD548954" box="[628,750,260,284]" italics="true" pageId="11" pageNumber="39">E. galileae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C657EBAFFD21171FCA3FEA5FC798954" box="[805,963,260,284]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD21171FCA3FEA5FC798954" box="[805,963,260,284]" italics="true" pageId="11" pageNumber="39">E. antalyensis</emphasis>
</taxonomicName>
, see
<bibRefCitation id="EFF478C8FFD21171FC7BFEA4FAB38954" author="Belfiore" box="[1021,1289,260,284]" pageId="11" pageNumber="39" refString="Belfiore, C. &amp; Sartori, M. (1999) Redescription of Electrogena galileae (Demoulin, 1973) (Ephemeroptera, Heptageniidae). Revue Suisse de Zoologie, 106 (4), 1025 - 1034." type="journal article" year="1999">Belfiore &amp; Sartori 1999</bibRefCitation>
and
<bibRefCitation id="EFF478C8FFD21171FAC6FEA4FEAB8977" author="Belfiore" pageId="11" pageNumber="39" refString="Belfiore, C., Tanatmis, M. &amp; Kazanci, N. (2000) Taxonomy of Electrogena antalyensis (Kazanci &amp; Braasch, 1986) (Ephemeroptera, Heptageniidae). Aquatic Insects, 22 (4), 261 - 270." type="journal article" year="2000">
Belfiore
<emphasis id="B911D92BFFD21171FF11FE89FF758908" box="[151,207,296,320]" italics="true" pageId="11" pageNumber="39">et al.</emphasis>
2000
</bibRefCitation>
, respectively) are most useful for the comparative study.
</paragraph>
<paragraph id="8BDA0539FFD21171FF4FFEEDFBAD892C" blockId="11.[151,1436,151,357]" box="[201,1047,332,357]" pageId="11" pageNumber="39">Mean, range and variance are presented for each quantitative characteristic.</paragraph>
<paragraph id="8BDA0539FFD21171FF11FE2EFE6289ED" blockId="11.[151,1436,397,1524]" box="[151,472,397,421]" pageId="11" pageNumber="39">
<emphasis id="B911D92BFFD21171FF11FE2EFE6A89ED" box="[151,464,397,421]" italics="true" pageId="11" pageNumber="39">Quantitative characteristics</emphasis>
:
</paragraph>
<paragraph id="8BDA0539FFD21171FF11FE11FDBD8981" blockId="11.[151,1436,397,1524]" box="[151,519,433,457]" pageId="11" pageNumber="39">1. N_PLP 15.07, 1416, 0.46</paragraph>
<paragraph id="8BDA0539FFD21171FF41FE75FE268A7C" blockId="11.[151,1436,397,1524]" pageId="11" pageNumber="39">
The number of hairs near the fore margin of the first segment of maxillary palp (
<figureCitation id="135E19BCFFD21171FBF8FE75FB6889A5" box="[1150,1234,469,493]" captionStart="FIGURES 7 11" captionStartId="5.[151,263,1980,2003]" captionTargetBox="[165,1411,338,1946]" captionTargetId="figure@5.[151,1436,337,1959]" captionTargetPageId="5" captionText="FIGURES 7 11. Electrogena gibedede sp. nov., mature larva, paratype: 7, head, dorsal view; 8, hypopharynx, ventral view; 9, kinds of labrum shape, dorsal view; 10, variations of glossae shape, ventral view; 11, first segment of maxillary palp, dorsal view." httpUri="https://zenodo.org/record/280266/files/figure.png" pageId="11" pageNumber="39">Fig. 11</figureCitation>
). Non-overlaping range has
<taxonomicName id="4C657EBAFFD21171FE8EFE5AFE1F8A59" box="[264,421,505,529]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD21171FE8EFE5AFE1F8A59" box="[264,421,505,529]" italics="true" pageId="11" pageNumber="39">E. antalyensis</emphasis>
</taxonomicName>
(06). Range overlaps with
<taxonomicName id="4C657EBAFFD21171FD62FE5AFCE78A59" box="[740,861,505,529]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD21171FD62FE5AFCE78A59" box="[740,861,505,529]" italics="true" pageId="11" pageNumber="39">E. galileae</emphasis>
</taxonomicName>
(719) and
<taxonomicName id="4C657EBAFFD21171FC63FE5AFBD88A59" box="[997,1122,505,529]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B911D92BFFD21171FC63FE5AFBD88A59" box="[997,1122,505,529]" italics="true" pageId="11" pageNumber="39">E. lateralis</emphasis>
</taxonomicName>
(721). 2. N_PLH 0, 00, 0
</paragraph>
<paragraph id="8BDA0539FFD21171FF41FDE0FF5F8A33" blockId="11.[151,1436,397,1524]" pageId="11" pageNumber="39">
Long hairs on the hind margin of the first segment of maxillary palp missing. Such long hairs present only in
<emphasis id="B911D92BFFD21171FA02FDE0FF5F8A33" italics="true" pageId="11" pageNumber="39">
<taxonomicName id="4C657EBAFFD21171FA02FDE0FF5B8A33" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="affinis">E. affinis</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BDA0539FFD21171FF11FD26FE1B8AD6" blockId="11.[151,1436,397,1524]" box="[151,417,646,670]" pageId="11" pageNumber="39">3. N_OUT 0, 00, 0</paragraph>
<paragraph id="8BDA0539FFD21171FF41FD0AFDB68B64" blockId="11.[151,1436,397,1524]" pageId="11" pageNumber="39">
Bristles on the outer margin of galea-lacinia are always missing. This characteristic is shared with
<taxonomicName id="4C657EBAFFD21171FAA4FD0BFA268A8A" box="[1314,1436,682,706]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD21171FAA4FD0BFA268A8A" box="[1314,1436,682,706]" italics="true" pageId="11" pageNumber="39">E. galileae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C657EBAFFD21171FF40FD6FFEFE8AAE" box="[198,324,718,742]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="malickyi">
<emphasis id="B911D92BFFD21171FF40FD6FFEFE8AAE" box="[198,324,718,742]" italics="true" pageId="11" pageNumber="39">E. malickyi</emphasis>
</taxonomicName>
. In other representatives of the genus bristles are present at least in some specimens (
<taxonomicName id="4C657EBAFFD21171FB78FD6FFAD98AAE" box="[1278,1379,718,742]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B911D92BFFD21171FB78FD6FFAD98AAE" box="[1278,1379,718,742]" italics="true" pageId="11" pageNumber="39">E. affinis</emphasis>
</taxonomicName>
01;
<taxonomicName id="4C657EBAFFD21171FF11FD52FEAE8B41" box="[151,276,753,777]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B911D92BFFD21171FF11FD52FEAE8B41" box="[151,276,753,777]" italics="true" pageId="11" pageNumber="39">E. lateralis</emphasis>
</taxonomicName>
010). In
<taxonomicName id="4C657EBAFFD21171FE0EFD52FD9F8B41" box="[392,549,753,777]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD21171FE0EFD52FD9F8B41" box="[392,549,753,777]" italics="true" pageId="11" pageNumber="39">E. antalyensis</emphasis>
</taxonomicName>
, the number of bristles is particularly high (mean 6.524). 4. N_CBS 14.23, 1216, 1.10
</paragraph>
<paragraph id="8BDA0539FFD21171FF41FC97FEDE8BDF" blockId="11.[151,1436,397,1524]" pageId="11" pageNumber="39">
The number of comb-shaped bristles on the fore margin of galea-lacinia is low.
<taxonomicName id="4C657EBAFFD21171FBCCFC99FB5D8B18" box="[1098,1255,824,848]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD21171FBCCFC99FB5D8B18" box="[1098,1255,824,848]" italics="true" pageId="11" pageNumber="39">E. antalyensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C657EBAFFD21171FA99FC99FA268B18" box="[1311,1436,824,848]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B911D92BFFD21171FA99FC99FA268B18" box="[1311,1436,824,848]" italics="true" pageId="11" pageNumber="39">E. lateralis</emphasis>
</taxonomicName>
have a slightly higher number of bristles (means 15.93 and 17.30, respectively).
<taxonomicName id="4C657EBAFFD21171FBA9FCFCFB108B3B" box="[1071,1194,859,883]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD21171FBA9FCFCFB108B3B" box="[1071,1194,859,883]" italics="true" pageId="11" pageNumber="39">E. galileae</emphasis>
</taxonomicName>
is a non-overlapping species (1921.5).
</paragraph>
<paragraph id="8BDA0539FFD21171FF11FC02FE5E8BF2" blockId="11.[151,1436,397,1524]" box="[151,484,930,954]" pageId="11" pageNumber="39">5. N_TCB 7.42, 78, 0.20</paragraph>
<paragraph id="8BDA0539FFD21171FF41FC68FE008C6F" blockId="11.[151,1436,397,1524]" pageId="11" pageNumber="39">
The number of pointed teeth on the 5th comb-shaped bristle is low.
<taxonomicName id="4C657EBAFFD21171FC57FC69FBF78BA8" box="[977,1101,968,992]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD21171FC57FC69FBF78BA8" box="[977,1101,968,992]" italics="true" pageId="11" pageNumber="39">E. galileae</emphasis>
</taxonomicName>
has a non-overlapping range (1316).
<taxonomicName id="4C657EBAFFD21171FF7BFC4DFE208C4C" box="[253,410,1004,1028]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD21171FF7BFC4DFE208C4C" box="[253,410,1004,1028]" italics="true" pageId="11" pageNumber="39">E. antalyensis</emphasis>
</taxonomicName>
has an overlapping range, but a generally higher number of teeth (10.34).
<taxonomicName id="4C657EBAFFD21171FB55FC4DFAF58C4C" box="[1235,1359,1004,1028]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B911D92BFFD21171FB55FC4DFAF58C4C" box="[1235,1359,1004,1028]" italics="true" pageId="11" pageNumber="39">E. lateralis</emphasis>
</taxonomicName>
has an overlapping range (713).
</paragraph>
<paragraph id="8BDA0539FFD21171FF11FB93FE198C03" blockId="11.[151,1436,397,1524]" box="[151,419,1075,1099]" pageId="11" pageNumber="39">6. N_CLW 2, 22, 0</paragraph>
<paragraph id="8BDA0539FFD21171FF41FBF6FDA68C90" blockId="11.[151,1436,397,1524]" pageId="11" pageNumber="39">
The species always posses 2 teeth on the tarsal claws (
<figureCitation id="135E19BCFFD21171FCAAFBF6FC0C8C26" box="[812,950,1110,1134]" captionStart="FIGURES 27 30" captionStartId="10.[151,264,1220,1243]" captionTargetBox="[170,1415,186,1184]" captionTargetId="figure@10.[151,1436,163,1199]" captionTargetPageId="10" captionText="FIGURES 27 30. Electrogena gibedede sp. nov., mature larva, paratype: 27, 28, posterior margin of abdominal tergites, dorsal view; 29, 30, tarsal claw, dorsal and ventral view." httpUri="https://zenodo.org/record/280270/files/figure.png" pageId="11" pageNumber="39">Figs. 29, 30</figureCitation>
). This characteristic is shared by numerous
<taxonomicName id="4C657EBAFFD21171FF11FBDAFEA58CDA" box="[151,287,1146,1170]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD21171FF11FBDAFEA58CDA" box="[151,287,1146,1170]" italics="true" pageId="11" pageNumber="39">Electrogena</emphasis>
</taxonomicName>
species, only
<taxonomicName id="4C657EBAFFD21171FE3BFBDBFDE08CDA" box="[445,602,1146,1170]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD21171FE3BFBDBFDE08CDA" box="[445,602,1146,1170]" italics="true" pageId="11" pageNumber="39">E. antalyensis</emphasis>
</taxonomicName>
posses more teeth (
<quantity id="4C9DA8DCFFD21171FCBEFBDAFC3A8CDA" box="[824,896,1146,1170]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="2.286" metricValueMin="1.016" pageId="11" pageNumber="39" unit="in" value="6.5" valueMax="9.0" valueMin="4.0">49 in</quantity>
two rows). Some specimens of
<taxonomicName id="4C657EBAFFD21171FB61FBDBFADA8CDA" box="[1255,1376,1146,1170]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD21171FB61FBDBFADA8CDA" box="[1255,1376,1146,1170]" italics="true" pageId="11" pageNumber="39">E. galileae</emphasis>
</taxonomicName>
have up to 4 teeth (24). Invariably one tooth is present in
<taxonomicName id="4C657EBAFFD21171FD6AFB3EFCD28CFD" box="[748,872,1181,1205]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B911D92BFFD21171FD6AFB3EFCD28CFD" box="[748,872,1181,1205]" italics="true" pageId="11" pageNumber="39">E. lateralis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C657EBAFFD21171FC19FB3EFB918CFD" box="[927,1067,1182,1205]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="squamata">
<emphasis id="B911D92BFFD21171FC19FB3EFB918CFD" box="[927,1067,1182,1205]" italics="true" pageId="11" pageNumber="39">E. squamata</emphasis>
</taxonomicName>
. 7. N_BVF 30.08, 2534, 8.076
</paragraph>
<paragraph id="8BDA0539FFD21171FF41FB44FE3C8D2F" blockId="11.[151,1436,397,1524]" pageId="11" pageNumber="39">
The number of bristles on the ventral side of femora near the hind margin is high. Other species with a high number of bristles is
<taxonomicName id="4C657EBAFFD21171FE04FAA9FE418D68" box="[386,507,1288,1312]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD21171FE04FAA9FE418D68" box="[386,507,1288,1312]" italics="true" pageId="11" pageNumber="39">E. galileae</emphasis>
</taxonomicName>
(22.45). The maximum number of bristles shared by other species is 3. Bristles are short and pointed apically, which is a characteristic shared by all other species except for
<taxonomicName id="4C657EBAFFD21171FB03FA8CFB448D0B" box="[1157,1278,1323,1347]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD21171FB03FA8CFB448D0B" box="[1157,1278,1323,1347]" italics="true" pageId="11" pageNumber="39">E. galileae</emphasis>
</taxonomicName>
with the bristles rounded apically.
</paragraph>
<paragraph id="8BDA0539FFD21171FF11FAD2FE208DC2" blockId="11.[151,1436,397,1524]" box="[151,410,1394,1418]" pageId="11" pageNumber="39">8. N_HFF 0, 00, 0</paragraph>
<paragraph id="8BDA0539FFD21171FF41FA35FEF18DBC" blockId="11.[151,1436,397,1524]" pageId="11" pageNumber="39">
The number of long hairs (at least twice as long as neighboring bristles) on the fore margin of femora.
<taxonomicName id="4C657EBAFFD21171FAC7FA36FE8A8D99" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="galileae" status="sp. nov.">
<emphasis id="B911D92BFFD21171FAC7FA36FE8A8D99" italics="true" pageId="11" pageNumber="39">Electrogena galileae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD21171FEBEFA19FE298D99" box="[312,403,1465,1489]" pageId="11" pageNumber="39" rank="species">
<emphasis id="B911D92BFFD21171FEBEFA19FE298D99" bold="true" box="[312,403,1465,1489]" pageId="11" pageNumber="39">sp. nov.</emphasis>
</taxonomicNameLabel>
has three very long hairs near the base of femora.
<taxonomicName id="4C657EBAFFD21171FC5DFA1AFBFB8D99" box="[987,1089,1465,1489]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B911D92BFFD21171FC5DFA1AFBFB8D99" box="[987,1089,1465,1489]" italics="true" pageId="11" pageNumber="39">E. affinis</emphasis>
</taxonomicName>
has 858 such hairs along the femoral margin.
</paragraph>
<paragraph id="8BDA0539FFD21171FF41F9BEFD0E8E35" blockId="11.[151,1436,1565,2032]" pageId="11" pageNumber="39">
Four more meristic characteristics were measured for
<taxonomicName id="4C657EBAFFD21171FCC8F9BDFBF38E7D" box="[846,1097,1565,1589]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD21171FCC8F9BDFBF38E7D" box="[846,1097,1565,1589]" italics="true" pageId="11" pageNumber="39">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD21171FBD0F9BEFB0C8E7E" box="[1110,1206,1566,1590]" pageId="11" pageNumber="39" rank="species">
<emphasis id="B911D92BFFD21171FBD0F9BEFB0C8E7E" bold="true" box="[1110,1206,1566,1590]" pageId="11" pageNumber="39">sp. nov.</emphasis>
</taxonomicNameLabel>
The characteristics proved to be useful for the taxonomy of the related
<taxonomicName id="4C657EBAFFD21171FD69F9E2FCD38E11" box="[751,873,1602,1625]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="species" species="venosus">
<emphasis id="B911D92BFFD21171FD69F9E2FCD38E11" box="[751,873,1602,1625]" italics="true" pageId="11" pageNumber="39">E. venosus</emphasis>
</taxonomicName>
species-group (Haybach 1999) and are proposed here for the genus
<taxonomicName id="4C657EBAFFD21171FEE1F9C6FE558E36" box="[359,495,1638,1662]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD21171FEE1F9C6FE558E36" box="[359,495,1638,1662]" italics="true" pageId="11" pageNumber="39">Electrogena</emphasis>
</taxonomicName>
for the first time.
</paragraph>
<paragraph id="8BDA0539FFD21171FF11F929FD998EE9" blockId="11.[151,1436,1565,2032]" box="[151,547,1673,1697]" pageId="11" pageNumber="39">9. N_PLS 33.077, 2541, 14.80</paragraph>
<paragraph id="8BDA0539FFD21171FF41F90DFDBF8EA1" blockId="11.[151,1436,1565,2032]" pageId="11" pageNumber="39">The number of bristles on the hind margin of the first segment of the maxillary palp. 10. N_PLBas 7.63, 611, 4.59</paragraph>
<paragraph id="8BDA0539FFD21171FF41F959FC9A8F59" blockId="11.[151,1436,1565,2032]" box="[199,800,1785,1809]" pageId="11" pageNumber="39">The number of hairs at the base of the maxillary palp.</paragraph>
<paragraph id="8BDA0539FFD21171FF11F8BDFDA48F7D" blockId="11.[151,1436,1565,2032]" box="[151,542,1821,1845]" pageId="11" pageNumber="39">11. N_VEN 13.88, 1019, 10.01</paragraph>
<paragraph id="8BDA0539FFD21171FF41F8E5FCE58F15" blockId="11.[151,1436,1565,2032]" box="[199,863,1861,1885]" pageId="11" pageNumber="39">The number of hairs on the ventral surface of galea-lacinia.</paragraph>
<paragraph id="8BDA0539FFD21171FF11F8C8FDB68FC8" blockId="11.[151,1436,1565,2032]" box="[151,524,1896,1920]" pageId="11" pageNumber="39">12. N_LPH 20.05, 1724, 5.66</paragraph>
<paragraph id="8BDA0539FFD21171FF41F830FEE68FB8" blockId="11.[151,1436,1565,2032]" pageId="11" pageNumber="39">The number of bristles on the hind margin of the first segment of the labial palp. Several of these bristles form a separate group at the distal part of the hind margin, near the articulation of the second segment. This group contains 27 bristles.</paragraph>
<paragraph id="8BDA0539FFD51176FF11FF38FE7088E7" blockId="12.[151,1437,152,1677]" box="[151,458,152,176]" pageId="12" pageNumber="40">
<emphasis id="B911D92BFFD51176FF11FF38FE7888F8" box="[151,450,152,176]" italics="true" pageId="12" pageNumber="40">Qualitative characteristics</emphasis>
:
</paragraph>
<paragraph id="8BDA0539FFD51176FF11FF1CFDB889CE" blockId="12.[151,1437,152,1677]" pageId="12" pageNumber="40">
13. S_HLB The tips of hypopharyngeal superlinguae are covered with hairs shorter than those on the fore margin of superlinguae (
<figureCitation id="135E19BCFFD51176FED0FF7FFE2688BF" box="[342,412,223,247]" captionStart="FIGURES 7 11" captionStartId="5.[151,263,1980,2003]" captionTargetBox="[165,1411,338,1946]" captionTargetId="figure@5.[151,1436,337,1959]" captionTargetPageId="5" captionText="FIGURES 7 11. Electrogena gibedede sp. nov., mature larva, paratype: 7, head, dorsal view; 8, hypopharynx, ventral view; 9, kinds of labrum shape, dorsal view; 10, variations of glossae shape, ventral view; 11, first segment of maxillary palp, dorsal view." httpUri="https://zenodo.org/record/280266/files/figure.png" pageId="12" pageNumber="40">Fig. 8</figureCitation>
). Most of the hairs on the tips are no longer than 1/3 of the length of the fore hairs. However, several individual hairs approximately as long as 1/2 of the fore hairs may occur. This arrangement is unique and different from the species with very short hairs of the same length on the tips, in those cases not exceeding 1/4 of the fore hair length (
<taxonomicName id="4C657EBAFFD51176FE1CFEECFDAC892B" box="[410,534,331,355]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B911D92BFFD51176FE1CFEECFDAC892B" box="[410,534,331,355]" italics="true" pageId="12" pageNumber="40">E. lateralis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD51176FDA2FEECFD27892B" box="[548,669,331,355]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD51176FDA2FEECFD27892B" box="[548,669,331,355]" italics="true" pageId="12" pageNumber="40">E. galileae</emphasis>
</taxonomicName>
), or from all other species with hairs of the same length as fore hairs situated on the superlingual tips.
</paragraph>
<paragraph id="8BDA0539FFD51176FF11FE32FB0F8986" blockId="12.[151,1437,152,1677]" pageId="12" pageNumber="40">
14. S_GLO The shape of glossae quadrangular,
<taxonomicName id="4C657EBAFFD51176FD55FE32FCE389E2" box="[723,857,402,426]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD51176FD55FE32FCE389E2" box="[723,857,402,426]" italics="true" pageId="12" pageNumber="40">Ecdyonurus</emphasis>
</taxonomicName>
-like, in contrast to the subtriangular shape of glossae in
<taxonomicName id="4C657EBAFFD51176FF58FE17FEC18986" box="[222,379,438,462]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD51176FF58FE17FEC18986" box="[222,379,438,462]" italics="true" pageId="12" pageNumber="40">E. antalyensis</emphasis>
</taxonomicName>
(
<figureCitation id="135E19BCFFD51176FE0DFE16FE648986" box="[395,478,438,462]" captionStart="FIGURES 7 11" captionStartId="5.[151,263,1980,2003]" captionTargetBox="[165,1411,338,1946]" captionTargetId="figure@5.[151,1436,337,1959]" captionTargetPageId="5" captionText="FIGURES 7 11. Electrogena gibedede sp. nov., mature larva, paratype: 7, head, dorsal view; 8, hypopharynx, ventral view; 9, kinds of labrum shape, dorsal view; 10, variations of glossae shape, ventral view; 11, first segment of maxillary palp, dorsal view." httpUri="https://zenodo.org/record/280266/files/figure.png" pageId="12" pageNumber="40">Fig. 10</figureCitation>
). This characteristic is shared with all other
<taxonomicName id="4C657EBAFFD51176FC4BFE16FBEF8986" box="[973,1109,438,462]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD51176FC4BFE16FBEF8986" box="[973,1109,438,462]" italics="true" pageId="12" pageNumber="40">Electrogena</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="8BDA0539FFD51176FF11FE79FB378A5D" blockId="12.[151,1437,152,1677]" pageId="12" pageNumber="40">
15. S_PGL The paraglossae are widely rounded laterally. In another Caucasian species
<taxonomicName id="4C657EBAFFD51176FB06FE7BFA9C89BA" box="[1152,1318,474,498]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="pseudaffinis">
<emphasis id="B911D92BFFD51176FB06FE7BFA9C89BA" box="[1152,1318,474,498]" italics="true" pageId="12" pageNumber="40">E. pseudaffinis</emphasis>
</taxonomicName>
the lateral parts of paraglossae are almost straight, whereas in
<taxonomicName id="4C657EBAFFD51176FD5EFE5FFCE88A5E" box="[728,850,510,534]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD51176FD5EFE5FFCE88A5E" box="[728,850,510,534]" italics="true" pageId="12" pageNumber="40">E. galileae</emphasis>
</taxonomicName>
they are somewhat pointed.
</paragraph>
<paragraph id="8BDA0539FFD51176FF11FD81FB328A15" blockId="12.[151,1437,152,1677]" pageId="12" pageNumber="40">
16. S_PNT The hind corners of the pronotum are smoothly rounded, in contrast to the abrupt step present in related Caucasian species, especially in
<taxonomicName id="4C657EBAFFD51176FDD0FDE6FD598A15" box="[598,739,582,605]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="squamata">
<emphasis id="B911D92BFFD51176FDD0FDE6FD598A15" box="[598,739,582,605]" italics="true" pageId="12" pageNumber="40">E. squamata</emphasis>
</taxonomicName>
or some specimens of
<taxonomicName id="4C657EBAFFD51176FC62FDE6FB3B8A15" box="[996,1153,581,605]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD51176FC62FDE6FB3B8A15" box="[996,1153,581,605]" italics="true" pageId="12" pageNumber="40">E. antalyensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDA0539FFD51176FF11FDC8FE998AA4" blockId="12.[151,1437,152,1677]" pageId="12" pageNumber="40">
17. S_BFE The bristles on the upper surface of forefemora are spatulate, extended towards the apex and rounded apically. This is a rare arrangement among the genus, shared only with
<taxonomicName id="4C657EBAFFD51176FC3DFD2EFB8E8AED" box="[955,1076,653,677]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD51176FC3DFD2EFB8E8AED" box="[955,1076,653,677]" italics="true" pageId="12" pageNumber="40">E. galileae</emphasis>
</taxonomicName>
. Most
<taxonomicName id="4C657EBAFFD51176FB02FD2DFAB68AED" box="[1156,1292,653,677]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD51176FB02FD2DFAB68AED" box="[1156,1292,653,677]" italics="true" pageId="12" pageNumber="40">Electrogena</emphasis>
</taxonomicName>
species posses long and pointed or bluntly pointed bristles of forefemora. Such an arrangement is typical for all other Caucasian species.
</paragraph>
<paragraph id="8BDA0539FFD51176FF11FD58FC148B33" blockId="12.[151,1437,152,1677]" pageId="12" pageNumber="40">
18. S_BFF The bristles arranged in a row on the hind margin of femora are long, their length is approximately equal to 1/2 of the maximal femur width in all leg pairs. This characteristic state can be observed in almost all other
<taxonomicName id="4C657EBAFFD51176FF11FC9FFEA58B1F" box="[151,287,831,855]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD51176FF11FC9FFEA58B1F" box="[151,287,831,855]" italics="true" pageId="12" pageNumber="40">Electrogena</emphasis>
</taxonomicName>
species. A different arrangement with very short bristles may be found in
<taxonomicName id="4C657EBAFFD51176FBE6FCE0FB478B1F" box="[1120,1277,831,855]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD51176FBE6FCE0FB478B1F" box="[1120,1277,831,855]" italics="true" pageId="12" pageNumber="40">E. antalyensis</emphasis>
</taxonomicName>
. 19. S_TAR The dark band on tarsi is indistinct and situated apically.
</paragraph>
<paragraph id="8BDA0539FFD51176FF11FC26FEFA8D4C" blockId="12.[151,1437,152,1677]" pageId="12" pageNumber="40">
20. S_1GI The shape of the first gill is slightly curved, with convex, almost parallel margins, steeply converging and rounded apically, forming a “hump” on the fore margin (
<figureCitation id="135E19BCFFD51176FCBBFC0AFC2A8B8A" box="[829,912,938,962]" captionStart="FIGURES 12 14" captionStartId="6.[151,264,1631,1654]" captionTargetBox="[267,1320,228,1604]" captionTargetId="figure@6.[256,1330,193,1610]" captionTargetPageId="6" captionText="FIGURES 12 14. Electrogena gibedede sp. nov., mature larva, paratype: 12, middle leg, dorsal view; 13, gill VII, dorsal view; 14, gill I, dorsal view." httpUri="https://zenodo.org/record/280267/files/figure.png" pageId="12" pageNumber="40">Fig. 14</figureCitation>
). It differs from species with margins converging throughout the gill length (e. g.
<taxonomicName id="4C657EBAFFD51176FDA1FC6FFD458BAE" box="[551,767,974,998]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="azerbajdshanica">
<emphasis id="B911D92BFFD51176FDA1FC6FFD458BAE" box="[551,767,974,998]" italics="true" pageId="12" pageNumber="40">E. azerbajdshanica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD51176FC88FC6FFC0E8BAE" box="[782,948,974,998]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="pseudaffinis">
<emphasis id="B911D92BFFD51176FC88FC6FFC0E8BAE" box="[782,948,974,998]" italics="true" pageId="12" pageNumber="40">E. pseudaffinis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD51176FC45FC6FFBF78BAE" box="[963,1101,974,998]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="kuraensis">
<emphasis id="B911D92BFFD51176FC45FC6FFBF78BAE" box="[963,1101,974,998]" italics="true" pageId="12" pageNumber="40">E. kuraensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD51176FBDDFC6FFB528BAE" box="[1115,1256,975,998]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="squamata">
<emphasis id="B911D92BFFD51176FBDDFC6FFB528BAE" box="[1115,1256,975,998]" italics="true" pageId="12" pageNumber="40">E. squamata</emphasis>
</taxonomicName>
). It is similar to the gill shape of
<taxonomicName id="4C657EBAFFD51176FED6FC53FE748C42" box="[336,462,1010,1034]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="malickyi">
<emphasis id="B911D92BFFD51176FED6FC53FE748C42" box="[336,462,1010,1034]" italics="true" pageId="12" pageNumber="40">E. malickyi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C657EBAFFD51176FD82FC53FDD28C42" box="[516,616,1010,1034]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B911D92BFFD51176FD82FC53FDD28C42" box="[516,616,1010,1034]" italics="true" pageId="12" pageNumber="40">E. affinis</emphasis>
</taxonomicName>
. The convexity of the fore margin is less pronounced in some specimens. 21. S_7GI The shape of the seventh gill gradually narrows towards the apex (
<figureCitation id="135E19BCFFD51176FBABFBB5FB3B8C65" box="[1069,1153,1045,1069]" captionStart="FIGURES 12 14" captionStartId="6.[151,264,1631,1654]" captionTargetBox="[267,1320,228,1604]" captionTargetId="figure@6.[256,1330,193,1610]" captionTargetPageId="6" captionText="FIGURES 12 14. Electrogena gibedede sp. nov., mature larva, paratype: 12, middle leg, dorsal view; 13, gill VII, dorsal view; 14, gill I, dorsal view." httpUri="https://zenodo.org/record/280267/files/figure.png" pageId="12" pageNumber="40">Fig. 13</figureCitation>
), differring from species with a broad proximal part of the gill, suddenly narrowing at the apex; see
<taxonomicName id="4C657EBAFFD51176FC70FB9AFB158C19" box="[1014,1199,1082,1105]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B911D92BFFD51176FC70FB9AFB158C19" box="[1014,1199,1082,1105]" italics="true" pageId="12" pageNumber="40">E. zimmermanni</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFF478C8FFD51176FB51FB99FF058C3D" author="Sowa" pageId="12" pageNumber="40" refString="Sowa, R. (1984) Two new species of Ecdyonurus Eaton of lateralis (Curt.) group (Ephemeroptera, Heptageniidae) from the Crimea and Western Caucasus. Acta Hydrobiologica, 25 / 26 (2), 181 - 188." type="journal article" year="1984">Sowa (1984: Fig. 37)</bibRefCitation>
. The gill plate is rounded apically, with a distinct row of short hairs along the distal 1/3 of the fore margin. 22. S_PLB The bristles on the hind margin of the first segment of the maxillary palp are distributed in three irregular rows. The length of bristles is approximately 1/3 of the segment width. This is shared by all other
<taxonomicName id="4C657EBAFFD51176FAC7FB05FF748CA8" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD51176FAC7FB05FF748CA8" italics="true" pageId="12" pageNumber="40">Electrogena</emphasis>
</taxonomicName>
species except for two species with shorter bristles:
<taxonomicName id="4C657EBAFFD51176FCAFFB69FC198CA8" box="[809,931,1224,1248]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD51176FCAFFB69FC198CA8" box="[809,931,1224,1248]" italics="true" pageId="12" pageNumber="40">E. galileae</emphasis>
</taxonomicName>
(about 1/41/5) and about 1/
<quantity id="4C9DA8DCFFD51176FB76FB68FAA58CA8" box="[1264,1311,1224,1248]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="12" pageNumber="40" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C657EBAFFD51176FAA1FB69FE868D4C" authority="Hagen, 1864" authorityName="Hagen" authorityYear="1864" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="zebrata">
<emphasis id="B911D92BFFD51176FAA1FB69FA268CA8" box="[1319,1436,1224,1248]" italics="true" pageId="12" pageNumber="40">E. zebrata</emphasis>
(Hagen, 1864)
</taxonomicName>
.
</paragraph>
<paragraph id="8BDA0539FFD51176FF11FAB0FE738E0D" blockId="12.[151,1437,152,1677]" pageId="12" pageNumber="40">
23. S_LBB The paramedian bristles on the ventral surface of the labrum are arranged in a single row. Such an arrangement is present in most other species, with the exception of
<taxonomicName id="4C657EBAFFD51176FC00FA94FC458D03" box="[902,1023,1331,1355]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD51176FC00FA94FC458D03" box="[902,1023,1331,1355]" italics="true" pageId="12" pageNumber="40">E. galileae</emphasis>
</taxonomicName>
, which exhibits two rows of bristles. 24. S_HEM The head is brownish, with light but clear markings (
<figureCitation id="135E19BCFFD51176FC19FAF7FC5E8D27" box="[927,996,1367,1391]" captionStart="FIGURES 7 11" captionStartId="5.[151,263,1980,2003]" captionTargetBox="[165,1411,338,1946]" captionTargetId="figure@5.[151,1436,337,1959]" captionTargetPageId="5" captionText="FIGURES 7 11. Electrogena gibedede sp. nov., mature larva, paratype: 7, head, dorsal view; 8, hypopharynx, ventral view; 9, kinds of labrum shape, dorsal view; 10, variations of glossae shape, ventral view; 11, first segment of maxillary palp, dorsal view." httpUri="https://zenodo.org/record/280266/files/figure.png" pageId="12" pageNumber="40">Fig. 7</figureCitation>
). There are two light spots on the fore margin of the head capsule (distance between these spots approximately equal to distance between lateral ocelli). There are further areas of lighter coloration along the lateral margins, anterior to the antennal insertions and along the posterior margin. Presence of a narrow light stripe anterior to the median ocellum and dark brown spots in the area of the lateral ocelli and eyes. Among other representatives of the genus, similar light markings are present only in
<taxonomicName id="4C657EBAFFD51176FF32F9ABFEA28E6A" box="[180,280,1546,1570]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B911D92BFFD51176FF32F9ABFEA28E6A" box="[180,280,1546,1570]" italics="true" pageId="12" pageNumber="40">E. affinis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD51176FEA0F9ABFE258E6A" box="[294,415,1546,1570]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD51176FEA0F9ABFE258E6A" box="[294,415,1546,1570]" italics="true" pageId="12" pageNumber="40">E. galileae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C657EBAFFD51176FE50F9ABFDC68E6A" box="[470,636,1546,1570]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="pseudaffinis">
<emphasis id="B911D92BFFD51176FE50F9ABFDC68E6A" box="[470,636,1546,1570]" italics="true" pageId="12" pageNumber="40">E. pseudaffinis</emphasis>
</taxonomicName>
, but the arrangement is different in
<taxonomicName id="4C657EBAFFD51176FB88F9ABFBC98E6A" box="[1038,1139,1546,1570]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="B911D92BFFD51176FB88F9ABFBC98E6A" box="[1038,1139,1546,1570]" italics="true" pageId="12" pageNumber="40">E. affinis</emphasis>
</taxonomicName>
(light spot along posterior margin of head is missing).
</paragraph>
<paragraph id="8BDA0539FFD51176FF11F9F1FE748EC5" blockId="12.[151,1437,152,1677]" pageId="12" pageNumber="40">
25. S_HEA The shape of the head capsule is symmetrical and oval, without distinct narrowing in the fore part (contrary to
<taxonomicName id="4C657EBAFFD51176FEA7F9D6FE048EC5" box="[289,446,1653,1677]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD51176FEA7F9D6FE048EC5" box="[289,446,1653,1677]" italics="true" pageId="12" pageNumber="40">E. antalyensis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BDA0539FFD51176FF11F917FE338E86" blockId="12.[151,1436,1718,2030]" box="[151,393,1718,1742]" pageId="12" pageNumber="40">
<emphasis id="B911D92BFFD51176FF11F917FE3B8E86" box="[151,385,1718,1742]" italics="true" pageId="12" pageNumber="40">Ratio characteristics</emphasis>
:
</paragraph>
<paragraph id="8BDA0539FFD51176FF11F97AFD918EBA" blockId="12.[151,1436,1718,2030]" box="[151,555,1754,1778]" pageId="12" pageNumber="40">26. R_1GI 2.06, 1.892.14, 0.005</paragraph>
<paragraph id="8BDA0539FFD51176FF41F95EFDA78FCA" blockId="12.[151,1436,1718,2030]" pageId="12" pageNumber="40">
Length/width ratio of the first gill plate. It differs from that of species with a distinctly narrower first gill (
<taxonomicName id="4C657EBAFFD51176FA02F8A0FEBE8F73" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="squamata">
<emphasis id="B911D92BFFD51176FA02F8A0FEBE8F73" italics="true" pageId="12" pageNumber="40">E. squamata</emphasis>
</taxonomicName>
, the ratio 4.45 and
<taxonomicName id="4C657EBAFFD51176FE7EF884FD0A8F73" box="[504,688,1828,1851]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="zimmermanni">
<emphasis id="B911D92BFFD51176FE7EF884FD0A8F73" box="[504,688,1828,1851]" italics="true" pageId="12" pageNumber="40">E. zimmermanni</emphasis>
</taxonomicName>
, the ratio 2.67), and from that of species with a distinctly wider first gill (
<taxonomicName id="4C657EBAFFD51176FE87F8E8FED68F16" box="[257,364,1863,1887]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="necatii">
<emphasis id="B911D92BFFD51176FE87F8E8FED68F16" box="[257,364,1863,1887]" italics="true" pageId="12" pageNumber="40">E. necatii</emphasis>
</taxonomicName>
, the ratio 1.57;
<taxonomicName id="4C657EBAFFD51176FDB4F8E8FD758F16" box="[562,719,1862,1887]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD51176FDB4F8E8FD758F16" box="[562,719,1862,1887]" italics="true" pageId="12" pageNumber="40">E. antalyensis</emphasis>
</taxonomicName>
, the ratio mean 1.60 and
<taxonomicName id="4C657EBAFFD51176FB87F8E8FB2D8F16" box="[1025,1175,1863,1887]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="armeniaca">
<emphasis id="B911D92BFFD51176FB87F8E8FB2D8F16" box="[1025,1175,1863,1887]" italics="true" pageId="12" pageNumber="40">E. armeniaca</emphasis>
</taxonomicName>
, the ratio 1.78). 27. R_7GI 2.24, 2.02.55, 0.043
</paragraph>
<paragraph id="8BDA0539FFD51176FF41F82EFCFC8F82" blockId="12.[151,1436,1718,2030]" pageId="12" pageNumber="40">
Length/width ratio of seventh gill plate. It differs from that of species with a distinctly wider seventh gill (
<taxonomicName id="4C657EBAFFD51176FA02F830FEB58F83" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="armeniaca">
<emphasis id="B911D92BFFD51176FA02F830FEB58F83" italics="true" pageId="12" pageNumber="40">E. armeniaca</emphasis>
</taxonomicName>
, the ratio 1.83 and
<taxonomicName id="4C657EBAFFD51176FE7AF814FDC48F83" box="[508,638,1971,1995]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="braaschi">
<emphasis id="B911D92BFFD51176FE7AF814FDC48F83" box="[508,638,1971,1995]" italics="true" pageId="12" pageNumber="40">E. braaschi</emphasis>
</taxonomicName>
, the ratio 1.84).
</paragraph>
<paragraph id="8BDA0539FFD51176FF11F876FD8C8FA6" blockId="12.[151,1436,1718,2030]" box="[151,566,2006,2030]" pageId="12" pageNumber="40">28. R_LBR 4.72, 4.555.25, 0.049</paragraph>
<paragraph id="8BDA0539FFD41177FF41FF37FF548954" blockId="13.[151,1436,151,464]" pageId="13" pageNumber="41">
Total width of labrum/mean width of the lateral lobe of labrum. The value of this ratio is approximately shared by most species from the genus
<taxonomicName id="4C657EBAFFD41177FD8DFF1CFD29889C" box="[523,659,188,212]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD41177FD8DFF1CFD29889C" box="[523,659,188,212]" italics="true" pageId="13" pageNumber="41">Electrogena</emphasis>
</taxonomicName>
, occurring in the Caucasus Mts. and adjacent territories (
<figureCitation id="135E19BCFFD41177FAA0FF1CFAD6889C" box="[1318,1388,188,212]" captionStart="FIGURES 7 11" captionStartId="5.[151,263,1980,2003]" captionTargetBox="[165,1411,338,1946]" captionTargetId="figure@5.[151,1436,337,1959]" captionTargetPageId="5" captionText="FIGURES 7 11. Electrogena gibedede sp. nov., mature larva, paratype: 7, head, dorsal view; 8, hypopharynx, ventral view; 9, kinds of labrum shape, dorsal view; 10, variations of glossae shape, ventral view; 11, first segment of maxillary palp, dorsal view." httpUri="https://zenodo.org/record/280266/files/figure.png" pageId="13" pageNumber="41">Fig. 9</figureCitation>
).
<taxonomicName id="4C657EBAFFD41177FA02FF1DFF4B88B0" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="41" phylum="Arthropoda" rank="species" species="galileae">
<emphasis id="B911D92BFFD41177FA02FF1DFF4B88B0" italics="true" pageId="13" pageNumber="41">E. galileae</emphasis>
</taxonomicName>
(the ratio 3.86) and
<taxonomicName id="4C657EBAFFD41177FE7AFF41FD3A88B0" box="[508,640,224,248]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="41" phylum="Arthropoda" rank="species" species="braaschi">
<emphasis id="B911D92BFFD41177FE7AFF41FD3A88B0" box="[508,640,224,248]" italics="true" pageId="13" pageNumber="41">E. braaschi</emphasis>
</taxonomicName>
(the ratio 4) posses significantly more pronounced lateral lobes of labrum.
</paragraph>
<paragraph id="8BDA0539FFD41177FF11FE87FD838977" blockId="13.[151,1436,151,464]" box="[151,569,295,319]" pageId="13" pageNumber="41">29. R_GLA 3.83, 3.464.56, 0.124</paragraph>
<paragraph id="8BDA0539FFD41177FF41FEECFE0A89CF" blockId="13.[151,1436,151,464]" pageId="13" pageNumber="41">
Distance between glossae. This feature differs in species with an apparently narrower distance between glossae (
<taxonomicName id="4C657EBAFFD41177FF26FED1FEB189C0" box="[160,267,369,392]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="41" phylum="Arthropoda" rank="species" species="necatii">
<emphasis id="B911D92BFFD41177FF26FED1FEB189C0" box="[160,267,369,392]" italics="true" pageId="13" pageNumber="41">E. necatii</emphasis>
</taxonomicName>
, the ratio 6).
</paragraph>
<paragraph id="8BDA0539FFD41177FF11FE34FD8D89E4" blockId="13.[151,1436,151,464]" box="[151,567,404,428]" pageId="13" pageNumber="41">30. R_GLB 2.74, 2.553.06, 0.017</paragraph>
<paragraph id="8BDA0539FFD41177FF41FE17FB808987" blockId="13.[151,1436,151,464]" box="[199,1082,439,464]" pageId="13" pageNumber="41">
Width of glossae. Glossae are moderately wide among all
<taxonomicName id="4C657EBAFFD41177FCD5FE18FC618998" box="[851,987,440,464]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD41177FCD5FE18FC618998" box="[851,987,440,464]" italics="true" pageId="13" pageNumber="41">Electrogena</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="8BDA0539FFD41177FF41FDA1FBEA8AEF" blockId="13.[151,1436,511,679]" pageId="13" pageNumber="41">
<emphasis id="B911D92BFFD41177FF41FDA1FF4E8A50" box="[199,244,513,536]" italics="true" pageId="13" pageNumber="41">Egg</emphasis>
(
<figureCitation id="135E19BCFFD41177FE85FE5FFE338A5F" box="[259,393,511,535]" captionStart="FIGURES 31 34" captionStartId="13.[151,264,1743,1766]" captionTargetBox="[185,1403,741,1717]" captionTargetId="figure@13.[172,1414,733,1722]" captionTargetPageId="13" captionText="FIGURES 31 34. Electrogena gibedede sp. nov., egg: 31, general view; 32, chorionic surface with small rounded tubercles and granular ground matrix; 33, micropyle; 34, group of KCTs attachment structures." httpUri="https://zenodo.org/record/280271/files/figure.png" pageId="13" pageNumber="41">Figs. 3134</figureCitation>
). Size: length 110160 μm; width 7095 μm. Egg oval, slightly elongated (
<figureCitation id="135E19BCFFD41177FB57FE5FFA9E8A5F" box="[1233,1316,511,535]" captionStart="FIGURES 31 34" captionStartId="13.[151,264,1743,1766]" captionTargetBox="[185,1403,741,1717]" captionTargetId="figure@13.[172,1414,733,1722]" captionTargetPageId="13" captionText="FIGURES 31 34. Electrogena gibedede sp. nov., egg: 31, general view; 32, chorionic surface with small rounded tubercles and granular ground matrix; 33, micropyle; 34, group of KCTs attachment structures." httpUri="https://zenodo.org/record/280271/files/figure.png" pageId="13" pageNumber="41">Fig. 31</figureCitation>
). The chorionic surface only with rare KCTs attachment structures, which are assembled in small groups (1530 KCTs) in the subequatorial region or in the region of the pole cap (
<figureCitation id="135E19BCFFD41177FCA0FDE7FCC68A17" box="[806,892,583,607]" captionStart="FIGURES 31 34" captionStartId="13.[151,264,1743,1766]" captionTargetBox="[185,1403,741,1717]" captionTargetId="figure@13.[172,1414,733,1722]" captionTargetPageId="13" captionText="FIGURES 31 34. Electrogena gibedede sp. nov., egg: 31, general view; 32, chorionic surface with small rounded tubercles and granular ground matrix; 33, micropyle; 34, group of KCTs attachment structures." httpUri="https://zenodo.org/record/280271/files/figure.png" pageId="13" pageNumber="41">Fig. 34</figureCitation>
). The chorionic surface covered with numerous small rounded tubercles (1.21.8 μm) and delicate granules (granular ground matrix) (
<figureCitation id="135E19BCFFD41177FBEDFDCCFB4C8ACC" box="[1131,1270,620,644]" captionStart="FIGURES 31 34" captionStartId="13.[151,264,1743,1766]" captionTargetBox="[185,1403,741,1717]" captionTargetId="figure@13.[172,1414,733,1722]" captionTargetPageId="13" captionText="FIGURES 31 34. Electrogena gibedede sp. nov., egg: 31, general view; 32, chorionic surface with small rounded tubercles and granular ground matrix; 33, micropyle; 34, group of KCTs attachment structures." httpUri="https://zenodo.org/record/280271/files/figure.png" pageId="13" pageNumber="41">Figs. 32, 33</figureCitation>
). Two to three micropyles are visible in the subequatorial region; micropylar rim is absent (
<figureCitation id="135E19BCFFD41177FC69FD2FFBFE8AEF" box="[1007,1092,655,679]" captionStart="FIGURES 31 34" captionStartId="13.[151,264,1743,1766]" captionTargetBox="[185,1403,741,1717]" captionTargetId="figure@13.[172,1414,733,1722]" captionTargetPageId="13" captionText="FIGURES 31 34. Electrogena gibedede sp. nov., egg: 31, general view; 32, chorionic surface with small rounded tubercles and granular ground matrix; 33, micropyle; 34, group of KCTs attachment structures." httpUri="https://zenodo.org/record/280271/files/figure.png" pageId="13" pageNumber="41">Fig. 32</figureCitation>
).
</paragraph>
<caption id="DF1A55B1FFD41177FF11F96FFC4D8F4C" httpUri="https://zenodo.org/record/280271/files/figure.png" pageId="13" pageNumber="41" targetBox="[185,1403,741,1717]" targetPageId="13">
<paragraph id="8BDA0539FFD41177FF11F96FFC4D8F4C" blockId="13.[151,1436,1743,1796]" pageId="13" pageNumber="41">
<emphasis id="B911D92BFFD41177FF11F96FFEEF8EAC" bold="true" box="[151,341,1743,1766]" pageId="13" pageNumber="41">FIGURES 3134.</emphasis>
<taxonomicName id="4C657EBAFFD41177FEDBF96FFD818EAD" box="[349,571,1743,1765]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="41" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD41177FEDBF96FFD818EAD" box="[349,571,1743,1765]" italics="true" pageId="13" pageNumber="41">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD41177FDC5F970FD2F8EAD" box="[579,661,1744,1765]" pageId="13" pageNumber="41" rank="species">
<emphasis id="B911D92BFFD41177FDC5F970FD2F8EAD" bold="true" box="[579,661,1744,1765]" pageId="13" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
, egg: 31, general view; 32, chorionic surface with small rounded tubercles and granular ground matrix; 33, micropyle; 34, group of KCTs attachment structures.
</paragraph>
</caption>
<paragraph id="8BDA0539FFD41177FF41F88FFC208FFB" blockId="13.[151,1436,1839,1971]" pageId="13" pageNumber="41">
<bibRefCitation id="EFF478C8FFD41177FF41F88FFEC68F0F" author="Haybach" box="[199,380,1839,1863]" pageId="13" pageNumber="41" refString="Haybach, A. (2008) Egg structure morphology gives evidence for a cryptic subspecies of Electrogena lateralis (Curtis, 1834) in Southern Europe: Electrogena lateralis concii (Grandi, 1953) nov. stat. [Ephemeroptera, Heptageniidae]. Ephemera (2007), 9 (1), 1 - 7." type="journal article" year="2008">Haybach (2008)</bibRefCitation>
used the word “naked” for the description of the eggs with lacking attachment structures. The same structure of chorion is known in
<taxonomicName id="4C657EBAFFD41177FDC9F8F4FCCF8F23" authority="Grandi, 1953" authorityName="Grandi" authorityYear="1953" box="[591,885,1875,1899]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="41" phylum="Arthropoda" rank="species" species="gridelii">
<emphasis id="B911D92BFFD41177FDC9F8F4FD788F23" box="[591,706,1875,1899]" italics="true" pageId="13" pageNumber="41">E. gridelii</emphasis>
(Grandi, 1953)
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD41177FC04F8F4FB348F23" authority="Hagen, 1864" authorityName="Hagen" authorityYear="1864" box="[898,1166,1875,1899]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="41" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="B911D92BFFD41177FC04F8F4FC598F23" box="[898,995,1875,1899]" italics="true" pageId="13" pageNumber="41">E. fallax</emphasis>
(Hagen, 1864)
</taxonomicName>
and
<taxonomicName id="4C657EBAFFD41177FB4EF8F4FADC8F23" box="[1224,1382,1875,1899]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="41" phylum="Arthropoda" rank="species" species="antalyensis">
<emphasis id="B911D92BFFD41177FB4EF8F4FADC8F23" box="[1224,1382,1875,1899]" italics="true" pageId="13" pageNumber="41">E. antalyensis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFF478C8FFD41177FF11F8D7FEE08FC7" author="Gaino" box="[151,346,1911,1935]" pageId="13" pageNumber="41" refString="Gaino, E., Belfiore, C. &amp; Mazzini, M. (1987) Ootaxonomic investigation of the Italian Species of the genus Electrogena (Ephemeroptera, Heptageniidae). Bollettino di Zoologii, 54, 169 - 175." type="journal article" year="1987">
Gaino
<emphasis id="B911D92BFFD41177FF64F8D8FEE08FC7" box="[226,346,1911,1935]" italics="true" pageId="13" pageNumber="41">et al. 1987</emphasis>
</bibRefCitation>
;
<bibRefCitation id="EFF478C8FFD41177FEEEF8D7FDF98FC7" author="Belfiore" box="[360,579,1911,1935]" pageId="13" pageNumber="41" refString="Belfiore, C., Tanatmis, M. &amp; Kazanci, N. (2000) Taxonomy of Electrogena antalyensis (Kazanci &amp; Braasch, 1986) (Ephemeroptera, Heptageniidae). Aquatic Insects, 22 (4), 261 - 270." type="journal article" year="2000">Belfiore et al. 2000</bibRefCitation>
), in contrast to another species of
<taxonomicName id="4C657EBAFFD41177FC44F8D7FBF28FC7" box="[962,1096,1911,1935]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="41" phylum="Arthropoda" rank="genus">Electrogena</taxonomicName>
(e.g.
<taxonomicName id="4C657EBAFFD41177FB00F8D8FB4A8FC7" box="[1158,1264,1911,1935]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="41" phylum="Arthropoda" rank="species" species="affinis">E. affinis</taxonomicName>
; see Belfiore
<emphasis id="B911D92BFFD41177FA0EF8D8FF0E8FFB" italics="true" pageId="13" pageNumber="41">et al.</emphasis>
1999) with numerous adhesive elements on the chorionic surface.
</paragraph>
</subSubSection>
<subSubSection id="C37F56B2FFD71174FF41FF38FD028C7C" pageId="14" pageNumber="42" type="etymology">
<paragraph id="8BDA0539FFD71174FF41FF38FD6D88BF" blockId="14.[151,1437,151,1076]" pageId="14" pageNumber="42">
<emphasis id="B911D92BFFD71174FF41FF38FEF488F8" bold="true" box="[199,334,152,176]" pageId="14" pageNumber="42">Etymology.</emphasis>
The species epithet refers to the name of the Russian State Inspection of Road Safety (acronym GIBDD).The species is named in honor of those few honest officers of the Russian road police whom we have met during our travels while collecting the new species.
</paragraph>
<paragraph id="8BDA0539FFD71174FF41FEA3FC7189E4" blockId="14.[151,1437,151,1076]" pageId="14" pageNumber="42">
<emphasis id="B911D92BFFD71174FF41FEA3FE878954" bold="true" box="[199,317,259,284]" pageId="14" pageNumber="42">Affinities.</emphasis>
Comparison of
<taxonomicName id="4C657EBAFFD71174FE77FEA4FD5F8954" box="[497,741,260,284]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="42" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD71174FE77FEA4FD5F8954" box="[497,741,260,284]" italics="true" pageId="14" pageNumber="42">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD71174FD6AFEA4FCFF8954" box="[748,837,260,284]" pageId="14" pageNumber="42" rank="species">
<emphasis id="B911D92BFFD71174FD6AFEA4FCFF8954" bold="true" box="[748,837,260,284]" pageId="14" pageNumber="42">sp. nov.</emphasis>
</taxonomicNameLabel>
with the related Caucasian species is particularly difficult due to the very poor available descriptions of almost all possibly sympatric species (the values of standard diagnostic characteristics are mostly unknown and only some of them can be roughly estimated from the original drawings). Conclusions about the closeness of relationships between individual Caucasian species of
<taxonomicName id="4C657EBAFFD71174FA92FED0FA2689C0" box="[1300,1436,368,392]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD71174FA92FED0FA2689C0" box="[1300,1436,368,392]" italics="true" pageId="14" pageNumber="42">Electrogena</emphasis>
</taxonomicName>
based on morphological similarities can therefore be only very restricted.
</paragraph>
<paragraph id="8BDA0539FFD71174FF4FFE18FB418B5C" blockId="14.[151,1437,151,1076]" pageId="14" pageNumber="42">
Most morphological differences in the larval stage between
<taxonomicName id="4C657EBAFFD71174FCE5FE18FBEC8998" box="[867,1110,440,464]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="42" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD71174FCE5FE18FBEC8998" box="[867,1110,440,464]" italics="true" pageId="14" pageNumber="42">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD71174FBDAFE18FB0F8998" box="[1116,1205,440,464]" pageId="14" pageNumber="42" rank="species">
<emphasis id="B911D92BFFD71174FBDAFE18FB0F8998" bold="true" box="[1116,1205,440,464]" pageId="14" pageNumber="42">sp. nov.</emphasis>
</taxonomicNameLabel>
and other individual species of the genus
<taxonomicName id="4C657EBAFFD71174FE04FE7CFDB089BC" box="[386,522,476,500]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD71174FE04FE7CFDB089BC" box="[386,522,476,500]" italics="true" pageId="14" pageNumber="42">Electrogena</emphasis>
</taxonomicName>
occurring in the area of question have been mentioned above in the descriptions of the individual standard diagnostic characteristics. Nevertheless we can generalize that from all species of the genus with known larvae, occurring in the area of question,
<taxonomicName id="4C657EBAFFD71174FCD6FD84FBFC8A74" box="[848,1094,548,572]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="42" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD71174FCD6FD84FBFC8A74" box="[848,1094,548,572]" italics="true" pageId="14" pageNumber="42">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD71174FBD6FD84FB168A74" box="[1104,1196,548,572]" pageId="14" pageNumber="42" rank="species">
<emphasis id="B911D92BFFD71174FBD6FD84FB168A74" bold="true" box="[1104,1196,548,572]" pageId="14" pageNumber="42">sp. nov.</emphasis>
</taxonomicNameLabel>
can be easily distinguished according to the current knowledge mainly by the combination of characteristics, as follows: (i) shape and color pattern of the head capsule (
<figureCitation id="135E19BCFFD71174FD93FDCCFDE08ACC" box="[533,602,620,644]" captionStart="FIGURES 7 11" captionStartId="5.[151,263,1980,2003]" captionTargetBox="[165,1411,338,1946]" captionTargetId="figure@5.[151,1436,337,1959]" captionTargetPageId="5" captionText="FIGURES 7 11. Electrogena gibedede sp. nov., mature larva, paratype: 7, head, dorsal view; 8, hypopharynx, ventral view; 9, kinds of labrum shape, dorsal view; 10, variations of glossae shape, ventral view; 11, first segment of maxillary palp, dorsal view." httpUri="https://zenodo.org/record/280266/files/figure.png" pageId="14" pageNumber="42">Fig. 7</figureCitation>
); (ii) shape of paraglossae, rounded laterally; (iii) setation of superlinguae with long hairs on the tips (
<figureCitation id="135E19BCFFD71174FE4AFD2FFDAA8AEF" box="[460,528,655,679]" captionStart="FIGURES 7 11" captionStartId="5.[151,263,1980,2003]" captionTargetBox="[165,1411,338,1946]" captionTargetId="figure@5.[151,1436,337,1959]" captionTargetPageId="5" captionText="FIGURES 7 11. Electrogena gibedede sp. nov., mature larva, paratype: 7, head, dorsal view; 8, hypopharynx, ventral view; 9, kinds of labrum shape, dorsal view; 10, variations of glossae shape, ventral view; 11, first segment of maxillary palp, dorsal view." httpUri="https://zenodo.org/record/280266/files/figure.png" pageId="14" pageNumber="42">Fig. 8</figureCitation>
); (iv) joint between pronotum and mesonotum smooth, without abrupt step (
<figureCitation id="135E19BCFFD71174FAE9FD2FFF0F8A84" captionStart="FIGURES 15 20" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[178,1407,454,1744]" captionTargetId="figure@7.[151,1436,445,1822]" captionTargetPageId="7" captionText="FIGURES 15 20. Electrogena gibedede sp. nov., mature larva (15, 16), male (17, 19) and female (18, 20) imagines, paratypes: 15, details of body coloration, dorsal view; 16, details of abdominal sternites coloration, ventral view; 17, 18, head and thorax, lateral view; 19, 20, details of abdominal tergites coloration, lateral view." httpUri="https://zenodo.org/record/280268/files/figure.png" pageId="14" pageNumber="42">Fig. 15</figureCitation>
); (v) unique shape of bristles on upper surface of forefemora, widened towards the apex and rounded (
<figureCitation id="135E19BCFFD71174FABFFD14FF0D8AA7" captionStart="FIGURES 21 26" captionStartId="9.[151,264,1898,1921]" captionTargetBox="[172,1416,351,1853]" captionTargetId="figure@9.[151,1436,351,1861]" captionTargetPageId="9" captionText="FIGURES 21 26. Electrogena gibedede sp. nov., male imago (21, 22) and mature larva (23 26): 21, penis lobes, ventral view; 22, genitalia, ventral view; 23, forefemora bristles, dorsal view; 24, forefemora bristles (detail), dorsal view; 25, hind femora bristles, dorsal view; 26, hind femora bristles (detail), dorsal view." httpUri="https://zenodo.org/record/280269/files/figure.png" pageId="14" pageNumber="42">Figs. 23, 24</figureCitation>
); (vi) presence of two teeth on the tarsal claws (
<figureCitation id="135E19BCFFD71174FD5AFD77FCDD8AA7" box="[732,871,727,751]" captionStart="FIGURES 27 30" captionStartId="10.[151,264,1220,1243]" captionTargetBox="[170,1415,186,1184]" captionTargetId="figure@10.[151,1436,163,1199]" captionTargetPageId="10" captionText="FIGURES 27 30. Electrogena gibedede sp. nov., mature larva, paratype: 27, 28, posterior margin of abdominal tergites, dorsal view; 29, 30, tarsal claw, dorsal and ventral view." httpUri="https://zenodo.org/record/280270/files/figure.png" pageId="14" pageNumber="42">Figs. 29, 30</figureCitation>
); (vii) shape of gill I with subparallel margins for most of the gill length (
<figureCitation id="135E19BCFFD71174FE18FD5CFE4B8B5C" box="[414,497,764,788]" captionStart="FIGURES 12 14" captionStartId="6.[151,264,1631,1654]" captionTargetBox="[267,1320,228,1604]" captionTargetId="figure@6.[256,1330,193,1610]" captionTargetPageId="6" captionText="FIGURES 12 14. Electrogena gibedede sp. nov., mature larva, paratype: 12, middle leg, dorsal view; 13, gill VII, dorsal view; 14, gill I, dorsal view." httpUri="https://zenodo.org/record/280267/files/figure.png" pageId="14" pageNumber="42">Fig. 14</figureCitation>
); (viii) shape of gill VII, gradually narrowing towards apex (
<figureCitation id="135E19BCFFD71174FB1CFD5CFB578B5C" box="[1178,1261,764,788]" captionStart="FIGURES 12 14" captionStartId="6.[151,264,1631,1654]" captionTargetBox="[267,1320,228,1604]" captionTargetId="figure@6.[256,1330,193,1610]" captionTargetPageId="6" captionText="FIGURES 12 14. Electrogena gibedede sp. nov., mature larva, paratype: 12, middle leg, dorsal view; 13, gill VII, dorsal view; 14, gill I, dorsal view." httpUri="https://zenodo.org/record/280267/files/figure.png" pageId="14" pageNumber="42">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="8BDA0539FFD71174FF4FFC80FDB58BEC" blockId="14.[151,1437,151,1076]" pageId="14" pageNumber="42">
At the imaginal stage males of
<taxonomicName id="4C657EBAFFD71174FDABFC80FC988B70" box="[557,802,800,824]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="42" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD71174FDABFC80FC988B70" box="[557,802,800,824]" italics="true" pageId="14" pageNumber="42">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD71174FCADFC80FC3C8B70" box="[811,902,800,824]" pageId="14" pageNumber="42" rank="species">
<emphasis id="B911D92BFFD71174FCADFC80FC3C8B70" bold="true" box="[811,902,800,824]" pageId="14" pageNumber="42">sp. nov.</emphasis>
</taxonomicNameLabel>
exhibit a unique arrangement of genitalia with almost square angled proximal parts of penis lobes. These diagnostic characteristics distinctly differentiate the new species from all other representatives of the genus
<taxonomicName id="4C657EBAFFD71174FD5BFCC8FCDF8BC8" box="[733,869,872,896]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B911D92BFFD71174FD5BFCC8FCDF8BC8" box="[733,869,872,896]" italics="true" pageId="14" pageNumber="42">Electrogena</emphasis>
</taxonomicName>
described so far, the latter displaying penis lobes mostly rounded or sub-triangular.
</paragraph>
<paragraph id="8BDA0539FFD71174FF4FFC0FFD028C7C" blockId="14.[151,1437,151,1076]" pageId="14" pageNumber="42">
The coloration of the adult abdomen, consisting of a triangular pattern from the lateral point of view, is also characteristic for
<taxonomicName id="4C657EBAFFD71174FEDAFC74FDEA8BA4" box="[348,592,980,1004]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="42" phylum="Arthropoda" rank="species" species="gibedede" status="sp. nov.">
<emphasis id="B911D92BFFD71174FEDAFC74FDEA8BA4" box="[348,592,980,1004]" italics="true" pageId="14" pageNumber="42">Electrogena gibedede</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2226450FFD71174FDD1FC74FD0B8BA4" box="[599,689,980,1004]" pageId="14" pageNumber="42" rank="species">
<emphasis id="B911D92BFFD71174FDD1FC74FD0B8BA4" bold="true" box="[599,689,980,1004]" pageId="14" pageNumber="42">sp. nov.</emphasis>
</taxonomicNameLabel>
, shared only by
<taxonomicName id="4C657EBAFFD71174FCEAFC75FBBE8BA4" box="[876,1028,981,1004]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="42" phylum="Arthropoda" rank="species" species="armeniaca">
<emphasis id="B911D92BFFD71174FCEAFC75FBBE8BA4" box="[876,1028,981,1004]" italics="true" pageId="14" pageNumber="42">E. armeniaca</emphasis>
</taxonomicName>
,
<taxonomicName id="4C657EBAFFD71174FB94FC75FBD48BA4" box="[1042,1134,980,1004]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="42" phylum="Arthropoda" rank="species" species="ressli">
<emphasis id="B911D92BFFD71174FB94FC75FBD48BA4" box="[1042,1134,980,1004]" italics="true" pageId="14" pageNumber="42">E. ressli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C657EBAFFD71174FB22FC75FAB28BA4" box="[1188,1288,980,1004]" class="Insecta" family="Heptageniidae" genus="Electrogena" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="42" phylum="Arthropoda" rank="species" species="dirmil">
<emphasis id="B911D92BFFD71174FB22FC75FAB28BA4" box="[1188,1288,980,1004]" italics="true" pageId="14" pageNumber="42">E. dirmil</emphasis>
</taxonomicName>
. The characteristic coloration of abdominal tergites together with the absence of distinct coloration of the sternites can also be used in discriminating females and subimagines.
</paragraph>
</subSubSection>
</treatment>
</document>