213 lines
28 KiB
XML
213 lines
28 KiB
XML
<document id="3747A537EBD6734FF36A82D2A157CE41" ID-DOI="10.11646/zootaxa.3990.2.1" ID-GBIF-Dataset="2dd793f4-d2f7-46c2-abbf-9952384cb55b" ID-ISSN="1175-5326" ID-Zenodo-Dep="245442" ID-ZooBank="C3F0F9F4-2879-402C-8504-D12810018E81" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461267390392" checkinUser="plazi" docAuthor="Cai, Yan-Peng & Zhou, Hong-Zhang" docDate="2015" docId="03CC2578FFBC1446FF04F468FE59E5E7" docLanguage="en" docName="zt03990p196.pdf" docOrigin="Zootaxa 3990 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quedius (Raphirus) spissus Cai & Zhou, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="163" masterDocId="FFF55D00FFB5144AFF93F70FFFA8E62F" masterDocTitle="Taxonomy of the subgenus Quedius (Raphirus) Stephens (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China" masterLastPageNumber="196" masterPageNumber="151" pageNumber="160" updateTime="1698610850963" updateUser="plazi">
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<mods:titleInfo id="AB27C837DE3AAB73D5ACDABBD9F58EDC">
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<mods:title id="1CD2940603C0A4E64A61A8EF360F55D2">Taxonomy of the subgenus Quedius (Raphirus) Stephens (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China</mods:title>
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<mods:namePart id="160633895314E783A65CB0BD7057F6C1">Cai, Yan-Peng</mods:namePart>
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<mods:roleTerm id="E46194DAC341FC00BFDB4DC8C71EB377">Author</mods:roleTerm>
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<mods:namePart id="38650865D1701B7740DD9EB8C8C9BF63">Zhou, Hong-Zhang</mods:namePart>
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<mods:title id="CC51E6EE2AE1987E63F9051594BCF7E8">Zootaxa</mods:title>
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<mods:part id="7D3208FF86FA27435789414B31BB45FC">
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<mods:date id="33A99CA13011CBBEE2EC3E94DA92DC17">2015</mods:date>
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<mods:number id="AA1E5EBD1639B9181938BDFFA8046D3B">3990</mods:number>
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<mods:identifier id="474B3AEDA24BFC134386ED9F1DCC5745" type="DOI">10.11646/zootaxa.3990.2.1</mods:identifier>
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<mods:identifier id="436A8A307135A505603449EB2358F893" type="GBIF-Dataset">2dd793f4-d2f7-46c2-abbf-9952384cb55b</mods:identifier>
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<mods:identifier id="DE76A2E21EC4FB34B5246FCF2CDEE1A0" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03CC2578FFBC1446FF04F468FE59E5E7" ID-DOI="http://doi.org/10.5281/zenodo.6115639" ID-GBIF-Taxon="119638137" ID-Zenodo-Dep="6115639" LSID="urn:lsid:plazi:treatment:03CC2578FFBC1446FF04F468FE59E5E7" httpUri="http://treatment.plazi.org/id/03CC2578FFBC1446FF04F468FE59E5E7" lastPageId="12" lastPageNumber="163" pageId="9" pageNumber="160">
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<subSubSection id="C37FC7E5FFBC1443FF04F468FE81E58C" pageId="9" pageNumber="160" type="nomenclature">
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<paragraph id="8BDA946EFFBC1443FF04F468FDE0E5AD" blockId="9.[151,584,871,931]" box="[151,584,871,898]" pageId="9" pageNumber="160">
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<heading id="D0922302FFBC1443FF04F468FDE0E5AD" bold="true" box="[151,584,871,898]" fontSize="11" level="1" pageId="9" pageNumber="160" reason="1">
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<emphasis id="B911487CFFBC1443FF04F468FDE0E5AD" bold="true" box="[151,584,871,898]" pageId="9" pageNumber="160">
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<taxonomicName id="4C65EFEDFFBC1443FF04F468FE4BE5AE" ID-CoL="4R3B7" box="[151,483,871,898]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="spissus" status="sp. nov." subGenus="Raphirus">
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<emphasis id="B911487CFFBC1443FF04F468FF54E5AE" bold="true" box="[151,252,871,897]" italics="true" pageId="9" pageNumber="160">Quedius</emphasis>
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(
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<emphasis id="B911487CFFBC1443FE9EF468FED5E5AE" bold="true" box="[269,381,871,897]" italics="true" pageId="9" pageNumber="160">Raphirus</emphasis>
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)
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<emphasis id="B911487CFFBC1443FE1DF468FE4BE5AE" bold="true" box="[398,483,871,897]" italics="true" pageId="9" pageNumber="160">spissus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A222F507FFBC1443FE79F467FDE0E5AD" box="[490,584,872,898]" pageId="9" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDA946EFFBC1443FF04F485FE81E58C" blockId="9.[151,584,871,931]" box="[151,297,906,931]" pageId="9" pageNumber="160">
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(
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<figureCitation id="135E88EBFFBC1443FF0CF485FE88E58C" box="[159,288,906,931]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[172,1433,195,1855]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Raphirus) spissus sp. nov.: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="9" pageNumber="160">Figs. 4, 4-1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37FC7E5FFBC1443FF04F4DDFE5EE220" pageId="9" pageNumber="160" type="materials_examined">
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<paragraph id="8BDA946EFFBC1443FF04F4DDFE5EE220" blockId="9.[151,1437,978,1975]" pageId="9" pageNumber="160">
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<emphasis id="B911487CFFBC1443FF04F4DDFEEDE5C4" bold="true" box="[151,325,978,1003]" pageId="9" pageNumber="160">
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<typeStatus id="54DE2ACCFFBC1443FF04F4DDFF7AE5C4" box="[151,210,978,1003]" pageId="9" pageNumber="160">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54DE2ACCFFBC1443FEDFF4DDFE1CE5C4" box="[332,436,978,1003]" pageId="9" pageNumber="160" type="holotype">Holotype</typeStatus>
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♂,
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<emphasis id="B911487CFFBC1443FE4DF4DDFD04E5C4" bold="true" box="[478,684,978,1003]" pageId="9" pageNumber="160">
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<collectingCountry id="F372D4FEFFBC1443FE4DF4DDFD97E5C4" box="[478,575,978,1003]" name="China" pageId="9" pageNumber="160">CHINA</collectingCountry>
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: Sichuan
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</emphasis>
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Province: Songpan County, Huanglongsi Nature Reserve,
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<quantity id="4C9D398BFFBC1443FAD3F4DCFA3FE5C5" box="[1344,1431,979,1003]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="160" unit="m" value="4000.0">4000 m</quantity>
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,
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<date id="FFDBB2AEFFBC1443FF04F4F8FE8BE220" box="[151,291,1015,1039]" pageId="9" pageNumber="160" value="1990-07-03">3. VII. 1990</date>
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, coll. Huifu Wang.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37FC7E5FFBC1446FF54F315FE38E4AA" lastPageId="12" lastPageNumber="163" pageId="9" pageNumber="160" type="description">
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<paragraph id="8BDA946EFFBC1443FF54F315FB8CE254" blockId="9.[151,1437,978,1975]" pageId="9" pageNumber="160">
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<emphasis id="B911487CFFBC1443FF54F315FEF1E21C" bold="true" box="[199,345,1050,1075]" pageId="9" pageNumber="160">Description.</emphasis>
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Head black; pronotum, scutellum and elytra dark brown; abdomen entirely dark brown, slightly iridescent; pubescence of elytra and abdomen dark brown; antennae brown, labrum and mandibles dark brown, maxillary and labial palpi brown; legs brown, with mid and hind tibiae darkened.
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</paragraph>
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<paragraph id="8BDA946EFFBC1443FF54F388FC56E2EC" blockId="9.[151,1437,978,1975]" pageId="9" pageNumber="160">
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BL =
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<quantity id="4C9D398BFFBC1443FE99F388FECBE2B1" box="[266,355,1159,1183]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="160" unit="mm" value="4.4">4.4 mm</quantity>
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, BW =
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<quantity id="4C9D398BFFBC1443FE2EF388FDBEE2B1" box="[445,534,1159,1183]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="160" unit="mm" value="0.9">0.9 mm</quantity>
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, HL/PL/EL = 1.00: 1.42: 1.32, HW/PW/EW/AW = 1.00: 1.16: 1.27: 1.24, HW/ HL = 1.30, HEL/HTL = 6.24, PW/PL = 1.06, EW/EL = 1.25, ESL/EL = 0.48.
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</paragraph>
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<paragraph id="8BDA946EFFBC1443FF54F3C1FC81E028" blockId="9.[151,1437,978,1975]" pageId="9" pageNumber="160">
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Head (
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<figureCitation id="135E88EBFFBC1443FE87F3C1FEF1E2C8" box="[276,345,1230,1255]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[172,1433,195,1855]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Raphirus) spissus sp. nov.: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="9" pageNumber="160">Fig. 4</figureCitation>
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A) round, wider than long; eye very large and convex, nearly occupying entire lateral sides of head, in dorsal view tempora very short, markedly narrowed posteriad; no additional setiferous punctures between anterior frontal setiferous punctures; posterior frontal setiferous puncture situated before level of posteriomedial margin of eye, separated by distance about as long as diameter of puncture; temporal setiferous puncture touching posterior margin of eye, with several small setiferous punctures arranged in oblique curve behind and below it; one basal setiferous puncture situated about midway between posterior frontal setiferous puncture and nuchal constriction; head with fine and dense microsculpture of transverse and oblique waves. Antenna moderately long, with segment I longer than segment II or III, segment II and III nearly equal in length, segments IV–V, XI slightly longer than wide, segments VI–X slightly wider than long.
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</paragraph>
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<paragraph id="8BDA946EFFBC1443FF54F11CFBE3E0B8" blockId="9.[151,1437,978,1975]" pageId="9" pageNumber="160">
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Pronotum (
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<figureCitation id="135E88EBFFBC1443FED6F11DFE22E004" box="[325,394,1554,1579]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[172,1433,195,1855]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Raphirus) spissus sp. nov.: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="9" pageNumber="160">Fig. 4</figureCitation>
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B) slightly wider than long, slightly narrowed anteriad, posteriolateral and posterior margins continuously rounded, lateral margins not explanate; three setiferous punctures in each dorsal row and two setiferous punctures in each sublateral row, last sublateral row puncture situated before level of large lateral setiferous puncture; surface of pronotum with microsculpture similar to that of head.
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</paragraph>
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<paragraph id="8BDA946EFFBC1443FF54F1ADFEF5E0F1" blockId="9.[151,1437,978,1975]" pageId="9" pageNumber="160">
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Scutellum (
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<figureCitation id="135E88EBFFBC1443FEDFF1ADFE3BE094" box="[332,403,1698,1723]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[172,1433,195,1855]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Raphirus) spissus sp. nov.: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="9" pageNumber="160">Fig. 4</figureCitation>
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C) with several setiferous punctures, surface with very fine and dense microsculpture of transverse waves.
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</paragraph>
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<paragraph id="8BDA946EFFBC1443FF54F1E5FB01E164" blockId="9.[151,1437,978,1975]" pageId="9" pageNumber="160">
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Elytra (
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<figureCitation id="135E88EBFFBC1443FE8FF1E5FECAE12C" box="[284,354,1770,1795]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[172,1433,195,1855]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Raphirus) spissus sp. nov.: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="9" pageNumber="160">Fig. 4</figureCitation>
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C) short, wider than long, slightly broadened posteriad, each elytron with surface covered with dense setiferous punctures, transverse interspaces between punctures slightly smaller than diameters of punctures; surface between punctures without microsculpture. Wings extremely reduced, nonfunctional.
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</paragraph>
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<paragraph id="8BDA946EFFBC1443FF54F059FB90E199" blockId="9.[151,1437,978,1975]" pageId="9" pageNumber="160">Abdominal tergite II finely punctate and pubescent; setiferous punctures of other tergites distinctly finer and denser than those of elytra, becoming slightly sparser toward posterior margin of each tergite, and in general toward apex of abdomen; tergite VII without whitish apical seam of palisade setae.</paragraph>
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<caption id="DF1AC4E6FFBF1440FF04F05AFB60E1A5" httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="10" pageNumber="161" targetBox="[172,1433,195,1855]" targetPageId="10">
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<paragraph id="8BDA946EFFBF1440FF04F05AFB60E1A5" blockId="10.[151,1435,1877,1930]" pageId="10" pageNumber="161">
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<emphasis id="B911487CFFBF1440FF04F05AFEBBE145" bold="true" box="[151,275,1877,1899]" pageId="10" pageNumber="161">FIGURE 4.</emphasis>
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<taxonomicName id="4C65EFEDFFBF1440FE89F05AFD9AE144" box="[282,562,1877,1899]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="spissus" status="sp. nov." subGenus="Raphirus">
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<emphasis id="B911487CFFBF1440FE89F05AFEC6E144" box="[282,366,1877,1899]" italics="true" pageId="10" pageNumber="161">Quedius</emphasis>
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(
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<emphasis id="B911487CFFBF1440FEEEF05AFE72E144" box="[381,474,1877,1899]" italics="true" pageId="10" pageNumber="161">Raphirus</emphasis>
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)
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<emphasis id="B911487CFFBF1440FE79F059FD9AE144" box="[490,562,1878,1899]" italics="true" pageId="10" pageNumber="161">spissus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A222F507FFBF1440FDABF059FD21E144" box="[568,649,1878,1899]" pageId="10" pageNumber="161" rank="species">
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<emphasis id="B911487CFFBF1440FDABF059FD21E144" bold="true" box="[568,649,1878,1899]" pageId="10" pageNumber="161">sp. nov.</emphasis>
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</taxonomicNameLabel>
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: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses).
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</paragraph>
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</caption>
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<caption id="DF1AC4E6FFBE1441FF04F0AEFEEEE1DB" httpUri="https://zenodo.org/record/245448/files/figure.png" pageId="11" pageNumber="162" targetBox="[168,1419,193,1931]" targetPageId="11">
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<paragraph id="8BDA946EFFBE1441FF04F0AEFEEEE1DB" blockId="11.[151,1436,1953,2036]" pageId="11" pageNumber="162">
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<emphasis id="B911487CFFBE1441FF04F0AEFE80E199" bold="true" box="[151,296,1953,1975]" pageId="11" pageNumber="162">FIGURE 4-1.</emphasis>
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<taxonomicName id="4C65EFEDFFBE1441FEBCF0AEFDEEE198" box="[303,582,1953,1975]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="spissus" status="sp. nov." subGenus="Raphirus">
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<emphasis id="B911487CFFBE1441FEBCF0AEFE2BE198" box="[303,387,1953,1975]" italics="true" pageId="11" pageNumber="162">Quedius</emphasis>
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(
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<emphasis id="B911487CFFBE1441FE01F0AEFE47E198" box="[402,495,1953,1975]" italics="true" pageId="11" pageNumber="162">Raphirus</emphasis>
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)
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<emphasis id="B911487CFFBE1441FE6DF0ADFDEEE198" box="[510,582,1954,1975]" italics="true" pageId="11" pageNumber="162">spissus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A222F507FFBE1441FDDFF0AEFD35E199" box="[588,669,1953,1974]" pageId="11" pageNumber="162" rank="species">
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<emphasis id="B911487CFFBE1441FDDFF0AEFD35E199" bold="true" box="[588,669,1953,1974]" pageId="11" pageNumber="162">sp. nov.</emphasis>
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</taxonomicNameLabel>
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: A. male sternite VIII; B. male sternite IX; C. male tergite X; D. aedeagus, lateral view; E. aedeagus, parameral view; F. underside of paramere; G. apical portion of median lobe, parameral view. (Scale bars = 0.20 mm).
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</paragraph>
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</caption>
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<paragraph id="8BDA946EFFB91446FF54F798FC5EE44F" blockId="12.[151,1436,151,968]" pageId="12" pageNumber="163">
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Male with first four segments of foretarsus moderately dilated, sub-bilobed, each heavily covered with tenent setae ventrally, segment II slightly narrower than apex of tibia; tergite VIII with basal ridge complete, slightly arched backward in middle, with four long setae on each side; sternite VIII (
|
||
<figureCitation id="135E88EBFFB91446FC67F7D0FBE3E6D7" box="[1012,1099,223,248]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[172,1433,195,1855]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Raphirus) spissus sp. nov.: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="12" pageNumber="163">Fig. 4-1</figureCitation>
|
||
A) with basal ridge complete, slightly sinuate, with three long setae on each side, apical margin with a wide and moderately deep medioapical emargination, a small triangular area in front of the emargination impunctate; sternite IX (
|
||
<figureCitation id="135E88EBFFB91446FB35F628FAA8E76F" box="[1190,1280,295,320]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[172,1433,195,1855]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Raphirus) spissus sp. nov.: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="12" pageNumber="163">Fig. 4-1</figureCitation>
|
||
B) with basal portion long and narrow, apical margin complete, widely rounded; tergite X (
|
||
<figureCitation id="135E88EBFFB91446FC63F643FBEFE74B" box="[1008,1095,332,357]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[172,1433,195,1855]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Raphirus) spissus sp. nov.: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="12" pageNumber="163">Fig. 4-1</figureCitation>
|
||
C) with basal side broadly and deeply concave, apical margin rounded; aedeagus in lateral view (
|
||
<figureCitation id="135E88EBFFB91446FC0EF660FC50E7A7" box="[925,1016,367,392]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[172,1433,195,1855]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Raphirus) spissus sp. nov.: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="12" pageNumber="163">Fig. 4-1</figureCitation>
|
||
D) with apex of paramere not quite reaching that of median lobe, median lobe with short median carina a little below apex facing parameral side; aedeagus in parameral view (
|
||
<figureCitation id="135E88EBFFB91446FE4DF6B8FD9DE7FF" box="[478,565,439,464]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[172,1433,195,1855]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Raphirus) spissus sp. nov.: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="12" pageNumber="163">Fig. 4-1</figureCitation>
|
||
E) with paramere largely parallel-sided, entirely covering median lobe except for very apex, apical portion gradually narrowed into arcuate apex, median lobe quite narrow and elongate, apical portion narrowed into small and subacute apex (
|
||
<figureCitation id="135E88EBFFB91446FD24F6F0FCAEE437" box="[695,774,511,536]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[172,1433,195,1855]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Raphirus) spissus sp. nov.: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="12" pageNumber="163">Figs. 4</figureCitation>
|
||
E, 4-1G); apical portion of paramere with four apical setae at apical margin, and two similar setae on each lateral margin below apex, underside with about 12 sensory peg setae arranged in two short longitudinal rows along midline (
|
||
<figureCitation id="135E88EBFFB91446FCD2F548FC38E44F" box="[833,912,583,608]" captionStart="FIGURE 4" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[172,1433,195,1855]" captionTargetId="figure@10.[151,1436,193,1856]" captionTargetPageId="10" captionText="FIGURE 4. Quedius (Raphirus) spissus sp. nov.: A. head (0.5 mm); B. pronotum (0.5 mm); C. elytra (0.5 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245447/files/figure.png" pageId="12" pageNumber="163">Figs. 4</figureCitation>
|
||
D, 4-1F).
|
||
</paragraph>
|
||
<paragraph id="8BDA946EFFB91446FF54F563FE38E4AA" blockId="12.[151,1436,151,968]" box="[199,400,620,645]" pageId="12" pageNumber="163">Female unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37FC7E5FFB91446FF54F580FAB6E4E2" pageId="12" pageNumber="163" type="distribution">
|
||
<paragraph id="8BDA946EFFB91446FF54F580FAB6E4E2" blockId="12.[151,1436,151,968]" pageId="12" pageNumber="163">
|
||
<emphasis id="B911487CFFB91446FF54F580FEC8E487" bold="true" box="[199,352,655,680]" pageId="12" pageNumber="163">Distribution.</emphasis>
|
||
<taxonomicName id="4C65EFEDFFB91446FEF9F59FFD88E488" box="[362,544,656,680]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="spissus" status="sp. nov.">
|
||
<emphasis id="B911487CFFB91446FEF9F59FFD88E488" box="[362,544,656,680]" italics="true" pageId="12" pageNumber="163">Quedius spissus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A222F507FFB91446FDB9F580FD2EE487" box="[554,646,655,680]" pageId="12" pageNumber="163" rank="species">
|
||
<emphasis id="B911487CFFB91446FDB9F580FD2EE487" bold="true" box="[554,646,655,680]" pageId="12" pageNumber="163">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is at present known only from the
|
||
<typeStatus id="54DE2ACCFFB91446FBB1F59FFBFAE487" box="[1058,1106,656,680]" pageId="12" pageNumber="163">type</typeStatus>
|
||
locality in northern Sichuan Province: Huanglongsi Nature Reserve, at high altitude of
|
||
<quantity id="4C9D398BFFB91446FCBBF5BBFC28E4E3" box="[808,896,692,716]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="163" unit="m" value="4000.0">4000 m</quantity>
|
||
. The specimen was collected in July.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37FC7E5FFB91446FF54F5D8FEAAE550" pageId="12" pageNumber="163" type="diagnosis">
|
||
<paragraph id="8BDA946EFFB91446FF54F5D8FEAAE550" blockId="12.[151,1436,151,968]" pageId="12" pageNumber="163">
|
||
<emphasis id="B911487CFFB91446FF54F5D8FEE9E4DF" bold="true" box="[199,321,727,752]" pageId="12" pageNumber="163">Diagnosis.</emphasis>
|
||
This brachypterous species of the
|
||
<taxonomicName id="4C65EFEDFFB91446FD50F5D7FC9DE4DF" box="[707,821,728,752]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="muscicola">
|
||
<emphasis id="B911487CFFB91446FD50F5D7FC9DE4DF" box="[707,821,728,752]" italics="true" pageId="12" pageNumber="163">muscicola</emphasis>
|
||
</taxonomicName>
|
||
group was discovered in Sichuan Province of
|
||
<collectingCountry id="F372D4FEFFB91446FAA9F5D8FA29E4DF" box="[1338,1409,727,752]" name="China" pageId="12" pageNumber="163">China</collectingCountry>
|
||
. It is most similar to
|
||
<taxonomicName id="4C65EFEDFFB91446FECEF5F3FDC0E53B" authority="Smetana, 2011" authorityName="Smetana" authorityYear="2011" box="[349,616,764,788]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="chion">
|
||
<emphasis id="B911487CFFB91446FECEF5F3FE15E53B" box="[349,445,764,788]" italics="true" pageId="12" pageNumber="163">Q. chion</emphasis>
|
||
<bibRefCitation id="EFF4E99FFFB91446FE57F5F3FDC0E53B" author="Smetana" box="[452,616,764,788]" pageId="12" pageNumber="163" refString="Smetana, A. (2011) Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae, 51 (1), 145 - 156." type="journal article" year="2011">Smetana, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
discovered in Yunnan, by both having the paramere of the male aedeagus entirely covering median lobe except for very apex, but it can be distinguished from the latter by having the paramere wider, with fewer sensory peg setae arranged in shorter longitudinal rows, and median lobe of aedeagus narrower.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37FC7E5FFB91446FF54F483FE59E5E7" pageId="12" pageNumber="163" type="etymology">
|
||
<paragraph id="8BDA946EFFB91446FF54F483FE59E5E7" blockId="12.[151,1436,151,968]" pageId="12" pageNumber="163">
|
||
<emphasis id="B911487CFFB91446FF54F483FEE6E58A" bold="true" box="[199,334,908,933]" pageId="12" pageNumber="163">Etymology.</emphasis>
|
||
The specific name is from the Latin adjective
|
||
<taxonomicName id="4C65EFEDFFB91446FCF6F482FC1CE58B" box="[869,948,909,932]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="spissus">
|
||
<emphasis id="B911487CFFB91446FCF6F482FC1CE58B" box="[869,948,909,932]" italics="true" pageId="12" pageNumber="163">spissus</emphasis>
|
||
</taxonomicName>
|
||
(dense), referring to the dense pubescence on the elytra and the abdomen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |