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<mods:title id="14F8FD9FF05D5CAE429C777EB0307FC8">Taxonomy of the subgenus Quedius (Raphirus) Stephens (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China</mods:title>
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<mods:namePart id="CFAD2AA02CB2878DBA8CC94BB315745A">Cai, Yan-Peng</mods:namePart>
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<treatment id="03CC2578FF90146FFF04F798FD79E312" ID-DOI="http://doi.org/10.5281/zenodo.6115665" ID-GBIF-Taxon="119638127" ID-Zenodo-Dep="6115665" LSID="urn:lsid:plazi:treatment:03CC2578FF90146FFF04F798FD79E312" httpUri="http://treatment.plazi.org/id/03CC2578FF90146FFF04F798FD79E312" lastPageNumber="188" pageId="37" pageNumber="188">
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<paragraph id="8BDA946EFF90146FFF04F798FD65E69D" blockId="37.[151,717,151,211]" box="[151,717,151,178]" pageId="37" pageNumber="188">
<heading id="D0922302FF90146FFF04F798FD65E69D" bold="true" box="[151,717,151,178]" fontSize="11" level="1" pageId="37" pageNumber="188" reason="1">
<taxonomicName id="4C65EFEDFF90146FFF04F798FD65E69D" ID-CoL="4R37P" authority="Cameron, 1932" authorityName="Cameron" authorityYear="1932" box="[151,717,151,178]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="species" species="muscicola" subGenus="Raphirus">
<emphasis id="B911487CFF90146FFF04F798FD65E69D" bold="true" box="[151,717,151,178]" pageId="37" pageNumber="188">
<emphasis id="B911487CFF90146FFF04F798FF54E69E" bold="true" box="[151,252,151,177]" italics="true" pageId="37" pageNumber="188">Quedius</emphasis>
(
<emphasis id="B911487CFF90146FFE9EF798FED5E69E" bold="true" box="[269,381,151,177]" italics="true" pageId="37" pageNumber="188">Raphirus</emphasis>
)
<emphasis id="B911487CFF90146FFE1DF798FDA0E69E" bold="true" box="[398,520,151,177]" italics="true" pageId="37" pageNumber="188">muscicola</emphasis>
<bibRefCitation id="EFF4E99FFF90146FFD83F797FD65E69D" author="Cameron" box="[528,717,152,178]" pageId="37" pageNumber="188" refString="Cameron, M. (1932) The fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. 3. Taylor and Francis, London, 443 pp." type="book" year="1932">
<collectingCountry id="F372D4FEFF90146FFD83F797FD2EE69D" box="[528,646,152,178]" name="Cameroon" pageId="37" pageNumber="188">Cameron</collectingCountry>
, 1932
</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDA946EFF90146FFF04F7B5FEEDE6FC" blockId="37.[151,717,151,211]" box="[151,325,186,211]" pageId="37" pageNumber="188">
(
<figureCitation id="135E88EBFF90146FFF0CF7B5FE94E6FC" box="[159,316,186,211]" captionStart="FIGURE 15" captionStartId="38.[151,250,1877,1899]" captionTargetBox="[182,1420,193,1855]" captionTargetId="figure@38.[162,1424,193,1856]" captionTargetPageId="38" captionText="FIGURE 15. Quedius (Raphirus) muscicola Cameron, 1932: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245468/files/figure.png" pageId="37" pageNumber="188">Figs. 15, 15-1</figureCitation>
)
</paragraph>
<paragraph id="8BDA946EFF90146FFF04F60DFE81E7BC" blockId="37.[151,1436,258,587]" pageId="37" pageNumber="188">
<bibRefCitation id="EFF4E99FFF90146FFF04F60DFE91E737" author="Cameron" box="[151,313,258,280]" pageId="37" pageNumber="188" refString="Cameron, M. (1932) The fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. 3. Taylor and Francis, London, 443 pp." type="book" year="1932">
<collectingCountry id="F372D4FEFF90146FFF04F60DFF5EE737" box="[151,246,258,280]" name="Cameroon" pageId="37" pageNumber="188">Cameron</collectingCountry>
, 1932
</bibRefCitation>
: 295 (
<taxonomicName id="4C65EFEDFF90146FFEEEF60DFE79E737" box="[381,465,258,280]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B911487CFF90146FFEEEF60DFE79E737" box="[381,465,258,280]" italics="true" pageId="37" pageNumber="188">Quedius</emphasis>
</taxonomicName>
; subgenus
<taxonomicName id="4C65EFEDFF90146FFDDAF60DFD0EE737" box="[585,678,258,280]" class="Insecta" family="Staphylinidae" genus="Raphirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B911487CFF90146FFDDAF60DFD0EE737" box="[585,678,258,280]" italics="true" pageId="37" pageNumber="188">Raphirus</emphasis>
</taxonomicName>
;
<typeStatus id="54DE2ACCFF90146FFD26F60CFD4FE737" box="[693,743,259,280]" pageId="37" pageNumber="188">Type</typeStatus>
locality:
<collectingCountry id="F372D4FEFF90146FFCDEF60DFC2BE737" box="[845,899,258,280]" name="India" pageId="37" pageNumber="188">India</collectingCountry>
, Ghum district);
<bibRefCitation id="EFF4E99FFF90146FFBA4F60DFB5AE737" author="Scheerpeltz" box="[1079,1266,258,280]" pageId="37" pageNumber="188" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (ed.), Coleopterorum Catalogus. Junk, Berlin, pp. 989 - 1500." type="book chapter" year="1933">Scheerpeltz, 1933</bibRefCitation>
: 1450 (catalog);
<bibRefCitation id="EFF4E99FFF90146FFF54F62EFE2AE718" author="Scheerpeltz" box="[199,386,289,311]" pageId="37" pageNumber="188" refString="Scheerpeltz, O. (1965) Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except Megalopsidiinae et Steninae). Arkiv for Zoologi, Uppsala, 17, 93 - 371." type="journal article" year="1965">Scheerpeltz, 1965</bibRefCitation>
: 112 (
<collectingCountry id="F372D4FEFF90146FFE50F62EFD84E718" box="[451,556,289,311]" name="Myanmar" pageId="37" pageNumber="188">Myanmar</collectingCountry>
);
<bibRefCitation id="EFF4E99FFF90146FFDACF62EFD73E718" author="Smetana" box="[575,731,289,311]" pageId="37" pageNumber="188" refString="Smetana, A. (1988) Revision of the tribes Quediini and Atanygnathini. Part II. The Himalayan region (Coleoptera: Staphylinidae). Quaestiones Entomologicae, 24 (2), 163 - 464." type="journal article" year="1988">Smetana, 1988</bibRefCitation>
: 253 (
<typeStatus id="54DE2ACCFF90146FFC8EF62EFCD4E718" box="[797,892,289,311]" pageId="37" pageNumber="188" type="lectotype">lectotype</typeStatus>
designation; redescription;
<collectingCountry id="F372D4FEFF90146FFB0FF62EFB7AE718" box="[1180,1234,289,311]" name="India" pageId="37" pageNumber="188">India</collectingCountry>
,
<collectingCountry id="F372D4FEFF90146FFB73F62DFAECE718" box="[1248,1348,290,311]" name="Myanmar" pageId="37" pageNumber="188">Myanmar</collectingCountry>
,
<collectingCountry id="F372D4FEFF90146FFAC3F62EFA39E718" box="[1360,1425,289,311]" name="Nepal" pageId="37" pageNumber="188">Nepal</collectingCountry>
);
<bibRefCitation id="EFF4E99FFF90146FFF54F64FFECAE779" author="Smetana" box="[199,354,320,342]" pageId="37" pageNumber="188" refString="Smetana, A. (1992) Revision of the tribes Quediini and Atanygnathini: Part II. The Himalayan region: Supplement 2. (Coleoptera: Staphylinidae: Staphylininae). Stuttgarter Beitrage zur Naturkunde Serie A (Biologie), 487, 1 - 11." type="journal article" year="1992">Smetana, 1992</bibRefCitation>
: 5 (
<collectingCountry id="F372D4FEFF90146FFE1AF64FFE63E779" box="[393,459,320,342]" name="Nepal" pageId="37" pageNumber="188">Nepal</collectingCountry>
);
<bibRefCitation id="EFF4E99FFF90146FFE4DF64FFDD2E779" author="Smetana" box="[478,634,320,342]" pageId="37" pageNumber="188" refString="Smetana, A. (1997) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 8. Quediini collected by S. Ueno and Y. Watanabe in Yunnan. Elytra, 25 (1), 129 - 134." type="journal article" year="1997">Smetana, 1997</bibRefCitation>
: 131 (
<collectingCountry id="F372D4FEFF90146FFD29F64FFD50E779" box="[698,760,320,342]" name="China" pageId="37" pageNumber="188">China</collectingCountry>
(Yunnan));
<bibRefCitation id="EFF4E99FFF90146FFCE6F64EFBA3E779" author="Herman" box="[885,1035,320,342]" pageId="37" pageNumber="188" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3) Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina), Xantholinini. Staphylinidae incertae sedis. Fossils, Protactinae. Bulletin of the American Museum of Natural History, 265, 3021 - 3839." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3214 (world catalog);
<bibRefCitation id="EFF4E99FFF90146FFB6EF64FFA31E779" author="Smetana" box="[1277,1433,320,342]" pageId="37" pageNumber="188" refString="Smetana, A. (2004) Family STAPHYLINIDAE Latreille, 1802. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Apollo Books, Stenstrup, pp. 656 - 670." type="book chapter" year="2004">Smetana, 2004</bibRefCitation>
: 667 (catalog for Palaearctic region);
<bibRefCitation id="EFF4E99FFF90146FFDC4F651FCA9E75B" author="Smetana" box="[599,769,350,372]" pageId="37" pageNumber="188" refString="Smetana, A. (2012 a) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 42. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 10. Zootaxa, 3156, 43 - 68." type="journal article" year="2012" yearSuffix="a">Smetana, 2012a</bibRefCitation>
: 44 (redescription;
<collectingCountry id="F372D4FEFF90146FFC45F651FBBAE75B" box="[982,1042,350,372]" name="China" pageId="37" pageNumber="188">China</collectingCountry>
(Guizhou, Hubei, Shaanxi, Sichuan, Yunnan)).
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FF90146FFF04F693FD70E464" pageId="37" pageNumber="188" type="reference_group">
<paragraph id="8BDA946EFF90146FFF04F693FD70E7C0" blockId="37.[151,1436,258,587]" pageId="37" pageNumber="188">
<treatmentCitationGroup id="AB75B340FF90146FFF04F693FD70E7C0" pageId="37" pageNumber="188">
<taxonomicName id="4C65EFEDFF90146FFF04F693FD00E79D" authority="Cameron, 1932: 297" authorityName="Cameron" authorityPageNumber="297" authorityYear="1932" box="[151,680,412,434]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="species" species="dohertyi" subGenus="Raphirus">
<emphasis id="B911487CFF90146FFF04F693FF43E79D" box="[151,235,412,434]" italics="true" pageId="37" pageNumber="188">Quedius</emphasis>
(
<emphasis id="B911487CFF90146FFF6CF693FEF4E79D" box="[255,348,412,434]" italics="true" pageId="37" pageNumber="188">Raphirus</emphasis>
)
<emphasis id="B911487CFF90146FFEFCF693FE6BE79D" box="[367,451,412,434]" italics="true" pageId="37" pageNumber="188">dohertyi</emphasis>
<treatmentCitation id="0AC4B27FFF90146FFE5CF693FD00E79D" author="Cameron" box="[463,680,412,434]" page="297" pageId="37" pageNumber="188" year="1932">
<bibRefCitation id="EFF4E99FFF90146FFE5CF693FDDCE79D" author="Cameron" box="[463,628,412,434]" pageId="37" pageNumber="188" refString="Cameron, M. (1932) The fauna of British India including Ceylon and Burma. Coleoptera. Staphylinidae. Vol. 3. Taylor and Francis, London, 443 pp." type="book" year="1932">
<collectingCountry id="F372D4FEFF90146FFE5CF693FD86E79D" box="[463,558,412,434]" name="Cameroon" pageId="37" pageNumber="188">Cameron</collectingCountry>
, 1932
</bibRefCitation>
: 297
</treatmentCitation>
</taxonomicName>
(
<taxonomicName id="4C65EFEDFF90146FFD2FF693FCB8E79D" ID-CoL="8VS2M" box="[700,784,412,434]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B911487CFF90146FFD2FF693FCB8E79D" box="[700,784,412,434]" italics="true" pageId="37" pageNumber="188">Quedius</emphasis>
</taxonomicName>
; subgenus
<taxonomicName id="4C65EFEDFF90146FFC1EF693FC42E79D" box="[909,1002,412,434]" class="Insecta" family="Staphylinidae" genus="Raphirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B911487CFF90146FFC1EF693FC42E79D" box="[909,1002,412,434]" italics="true" pageId="37" pageNumber="188">Raphirus</emphasis>
</taxonomicName>
;
<typeStatus id="54DE2ACCFF90146FFC6FF692FB86E79D" box="[1020,1070,413,434]" pageId="37" pageNumber="188">Type</typeStatus>
locality:
<collectingCountry id="F372D4FEFF90146FFB0BF692FB54E79D" box="[1176,1276,413,434]" name="Myanmar" pageId="37" pageNumber="188">Myanmar</collectingCountry>
, Ruby Mines);
<treatmentCitation id="0AC4B27FFF90146FFF54F6B5FE6AE7FF" author="Scheerpeltz" box="[199,450,442,464]" page="1438" pageId="37" pageNumber="188" year="1933">
<bibRefCitation id="EFF4E99FFF90146FFF54F6B5FE2AE7FF" author="Scheerpeltz" box="[199,386,442,464]" pageId="37" pageNumber="188" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (ed.), Coleopterorum Catalogus. Junk, Berlin, pp. 989 - 1500." type="book chapter" year="1933">Scheerpeltz, 1933</bibRefCitation>
: 1438
</treatmentCitation>
(catalog);
<treatmentCitation id="0AC4B27FFF90146FFDA7F6B5FCABE7FF" author="Smetana" box="[564,771,442,464]" page="253" pageId="37" pageNumber="188" year="1988">
<bibRefCitation id="EFF4E99FFF90146FFDA7F6B5FD79E7FF" author="Smetana" box="[564,721,442,464]" pageId="37" pageNumber="188" refString="Smetana, A. (1988) Revision of the tribes Quediini and Atanygnathini. Part II. The Himalayan region (Coleoptera: Staphylinidae). Quaestiones Entomologicae, 24 (2), 163 - 464." type="journal article" year="1988">Smetana, 1988</bibRefCitation>
: 253
</treatmentCitation>
(synonym of
<taxonomicName id="4C65EFEDFF90146FFC0AF6B5FBA8E7FF" ID-CoL="4R37P" box="[921,1024,442,464]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="species" species="muscicola">
<emphasis id="B911487CFF90146FFC0AF6B5FBA8E7FF" box="[921,1024,442,464]" italics="true" pageId="37" pageNumber="188">muscicola</emphasis>
</taxonomicName>
);
<treatmentCitation id="0AC4B27FFF90146FFB8BF6B4FB46E7FF" author="Herman" box="[1048,1262,442,464]" page="3214" pageId="37" pageNumber="188" year="2001">
<bibRefCitation id="EFF4E99FFF90146FFB8BF6B4FB07E7FF" author="Herman" box="[1048,1199,442,464]" pageId="37" pageNumber="188" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3) Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina), Xantholinini. Staphylinidae incertae sedis. Fossils, Protactinae. Bulletin of the American Museum of Natural History, 265, 3021 - 3839." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3214
</treatmentCitation>
(world catalog);
<treatmentCitation id="0AC4B27FFF90146FFF54F6D6FE39E7C0" author="Smetana" box="[199,401,473,495]" page="667" pageId="37" pageNumber="188" year="2004">
<bibRefCitation id="EFF4E99FFF90146FFF54F6D6FEC9E7C0" author="Smetana" box="[199,353,473,495]" pageId="37" pageNumber="188" refString="Smetana, A. (2004) Family STAPHYLINIDAE Latreille, 1802. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Apollo Books, Stenstrup, pp. 656 - 670." type="book chapter" year="2004">Smetana, 2004</bibRefCitation>
: 667
</treatmentCitation>
(catalog for Palaearctic region).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDA946EFF90146FFF04F6F7FD70E464" blockId="37.[151,1436,258,587]" pageId="37" pageNumber="188">
<treatmentCitationGroup id="AB75B340FF90146FFF04F6F7FD70E464" pageId="37" pageNumber="188">
<taxonomicName id="4C65EFEDFF90146FFF04F6F7FD17E421" authority="Cameron, 1944: 14" authorityName="Cameron" authorityPageNumber="14" authorityYear="1944" box="[151,703,504,526]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="species" species="heterogaster" subGenus="Raphirus">
<emphasis id="B911487CFF90146FFF04F6F7FF43E421" box="[151,235,504,526]" italics="true" pageId="37" pageNumber="188">Quedius</emphasis>
(
<emphasis id="B911487CFF90146FFF6EF6F7FEF2E421" box="[253,346,504,526]" italics="true" pageId="37" pageNumber="188">Raphirus</emphasis>
)
<emphasis id="B911487CFF90146FFEFFF6F7FE43E421" box="[364,491,504,526]" italics="true" pageId="37" pageNumber="188">heterogaster</emphasis>
<treatmentCitation id="0AC4B27FFF90146FFE66F6F7FD17E421" author="Cameron" box="[501,703,504,526]" page="14" pageId="37" pageNumber="188" year="1944">
<bibRefCitation id="EFF4E99FFF90146FFE66F6F7FD30E421" author="Cameron" box="[501,664,504,526]" pageId="37" pageNumber="188" refString="Cameron, M. (1944) Descriptions of new Staphylinidae (Coleoptera). The Proceedings of the Royal Entomological Society of London (B), 13, 11 - 15, 49 - 52." type="journal article" year="1944">
<collectingCountry id="F372D4FEFF90146FFE66F6F7FDFCE421" box="[501,596,504,526]" name="Cameroon" pageId="37" pageNumber="188">Cameron</collectingCountry>
, 1944
</bibRefCitation>
: 14
</treatmentCitation>
</taxonomicName>
(
<taxonomicName id="4C65EFEDFF90146FFD41F6F7FC8EE421" ID-CoL="8VS2M" box="[722,806,504,526]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B911487CFF90146FFD41F6F7FC8EE421" box="[722,806,504,526]" italics="true" pageId="37" pageNumber="188">Quedius</emphasis>
</taxonomicName>
; subgenus
<taxonomicName id="4C65EFEDFF90146FFC0CF6F7FC54E421" box="[927,1020,504,526]" class="Insecta" family="Staphylinidae" genus="Raphirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B911487CFF90146FFC0CF6F7FC54E421" box="[927,1020,504,526]" italics="true" pageId="37" pageNumber="188">Raphirus</emphasis>
</taxonomicName>
;
<typeStatus id="54DE2ACCFF90146FFB9EF6F6FB97E421" box="[1037,1087,505,526]" pageId="37" pageNumber="188">Type</typeStatus>
locality:
<collectingCountry id="F372D4FEFF90146FFB35F6F7FB74E421" box="[1190,1244,504,526]" name="India" pageId="37" pageNumber="188">India</collectingCountry>
, Chakrata district, Kanasar);
<treatmentCitation id="0AC4B27FFF90146FFEA7F519FDAFE403" author="Smetana" box="[308,519,534,556]" page="253" pageId="37" pageNumber="188" year="1988">
<bibRefCitation id="EFF4E99FFF90146FFEA7F519FE7BE403" author="Smetana" box="[308,467,534,556]" pageId="37" pageNumber="188" refString="Smetana, A. (1988) Revision of the tribes Quediini and Atanygnathini. Part II. The Himalayan region (Coleoptera: Staphylinidae). Quaestiones Entomologicae, 24 (2), 163 - 464." type="journal article" year="1988">Smetana, 1988</bibRefCitation>
: 253
</treatmentCitation>
(
<typeStatus id="54DE2ACCFF90146FFD8BF519FDDEE403" box="[536,630,534,556]" pageId="37" pageNumber="188" type="lectotype">lectotype</typeStatus>
designation; synonym of
<taxonomicName id="4C65EFEDFF90146FFC03F519FC5EE403" ID-CoL="4R37P" box="[912,1014,534,556]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="species" species="muscicola">
<emphasis id="B911487CFF90146FFC03F519FC5EE403" box="[912,1014,534,556]" italics="true" pageId="37" pageNumber="188">muscicola</emphasis>
</taxonomicName>
);
<treatmentCitation id="0AC4B27FFF90146FFB82F518FB43E403" author="Herman" box="[1041,1259,534,556]" page="3215" pageId="37" pageNumber="188" year="2001">
<bibRefCitation id="EFF4E99FFF90146FFB82F518FB01E403" author="Herman" box="[1041,1193,534,556]" pageId="37" pageNumber="188" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3) Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina), Xantholinini. Staphylinidae incertae sedis. Fossils, Protactinae. Bulletin of the American Museum of Natural History, 265, 3021 - 3839." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3215
</treatmentCitation>
(world catalog);
<treatmentCitation id="0AC4B27FFF90146FFF54F53AFE39E464" author="Smetana" box="[199,401,565,587]" page="667" pageId="37" pageNumber="188" year="2004">
<bibRefCitation id="EFF4E99FFF90146FFF54F53AFEC9E464" author="Smetana" box="[199,353,565,587]" pageId="37" pageNumber="188" refString="Smetana, A. (2004) Family STAPHYLINIDAE Latreille, 1802. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Apollo Books, Stenstrup, pp. 656 - 670." type="book chapter" year="2004">Smetana, 2004</bibRefCitation>
: 667
</treatmentCitation>
(catalog for Palaearctic region).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FF90146FFF04F578FA39E24A" pageId="37" pageNumber="188" type="materials_examined">
<paragraph id="8BDA946EFF90146FFF04F578FA39E24A" blockId="37.[151,1437,631,1341]" pageId="37" pageNumber="188">
<emphasis id="B911487CFF90146FFF04F578FDFDE4BF" bold="true" box="[151,597,631,656]" pageId="37" pageNumber="188">
Material examined.
<collectingCountry id="F372D4FEFF90146FFE15F578FE4FE4BF" box="[390,487,631,656]" name="China" pageId="37" pageNumber="188">CHINA</collectingCountry>
: Sichuan
</emphasis>
Province:
<specimenCount id="9D635FE7FF90146FFD5EF577FD51E4A0" box="[717,761,632,656]" pageId="37" pageNumber="188" type="generic">1 ♂</specimenCount>
, Wan County, Wangerbao Nature Reserve,
<date id="FFDBB2AEFF90146FFB4EF577FAC8E4BF" box="[1245,1376,632,656]" pageId="37" pageNumber="188" value="1994-05-28">28. V. 1994</date>
, coll. Xingke Yang;
<specimenCount id="9D635FE7FF90146FFEAAF593FECDE49B" box="[313,357,668,692]" pageId="37" pageNumber="188" type="generic">1 ♂</specimenCount>
, Dujiangyan City, Qingcheng Mountain,
<date id="FFDBB2AEFF90146FFCA5F593FC11E49B" box="[822,953,668,692]" pageId="37" pageNumber="188" value="1997-06-04">4. VI. 1997</date>
, coll. Haisheng Zhou; 4 ♂♂, 9 ♀, Baoxing County, Fengtongzhai Nature Reserve,
<date id="FFDBB2AEFF90146FFDF9F5B0FCBDE4F7" box="[618,789,703,728]" pageId="37" pageNumber="188" value="1997-06-06" valueMax="1997-06-07" valueMin="1997-06-06">67. VI. 1997</date>
, coll. Haisheng Zhou;
<specimenCount id="9D635FE7FF90146FFBBFF5CFFBF7E4F8" box="[1068,1119,704,728]" pageId="37" pageNumber="188" type="generic">1 ♂</specimenCount>
, Baoxing County, Yongfu, Pujigou,
<quantity id="4C9D398BFF90146FFF6FF5EBFEFCE4D3" box="[252,340,740,764]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="37" pageNumber="188" unit="m" value="2300.0">2300 m</quantity>
,
<date id="FFDBB2AEFF90146FFEF3F5EBFD85E4D3" box="[352,557,740,764]" pageId="37" pageNumber="188" value="2003-08-11" valueMax="2003-08-14" valueMin="2003-08-11">1114. VIII. 2003</date>
, coll. Xiaodong Yu;
<specimenCount id="9D635FE7FF90146FFC86F5EBFCE9E4D3" box="[789,833,740,764]" pageId="37" pageNumber="188" type="generic">1 ♂</specimenCount>
, Wolong National Nature Reserve, Wulidun,
<quantity id="4C9D398BFF90146FFAD3F5EBFA3FE4D3" box="[1344,1431,740,765]" metricMagnitude="3" metricUnit="m" metricValue="2.46" pageId="37" pageNumber="188" unit="m" value="2460.0">2460 m</quantity>
,
<date id="FFDBB2AEFF90146FFF04F407FEE4E50F" box="[151,332,776,800]" pageId="37" pageNumber="188" value="2004-08-05" valueMax="2004-08-08" valueMin="2004-08-05">58. VIII. 2004</date>
, coll. Xiaodong Yu; 1 ♀, Wolong National Nature Reserve, Wulidun,
<quantity id="4C9D398BFF90146FFBE0F407FB63E530" box="[1139,1227,776,800]" metricMagnitude="3" metricUnit="m" metricValue="2.27" pageId="37" pageNumber="188" unit="m" value="2270.0">2270 m</quantity>
,
<date id="FFDBB2AEFF90146FFB49F407FA30E50F" box="[1242,1432,776,800]" pageId="37" pageNumber="188" value="2004-09-22" valueMax="2004-09-25" valueMin="2004-09-22">2225. IX. 2004</date>
, coll. Xiaodong Yu;
<specimenCount id="9D635FE7FF90146FFEEEF423FE03E56B" box="[381,427,812,836]" pageId="37" pageNumber="188" type="generic">1 ♂</specimenCount>
, 1 ♀, Dujiangyan City, Qingcheng Mountain,
<date id="FFDBB2AEFF90146FFC5EF423FBDFE56B" box="[973,1143,812,836]" pageId="37" pageNumber="188" value="2005-08-21">21. VIII. 2005</date>
, coll. Yejun Zhang; 1 ♀, Dujiangyan City, Qingcheng Mountain,
<date id="FFDBB2AEFF90146FFDCBF45FFD52E547" box="[600,762,847,872]" pageId="37" pageNumber="188" value="2005-08-23">23. VIII. 2005</date>
, coll. Xiaoyan Li;
<specimenCount id="9D635FE7FF90146FFC5AF45FFC5CE548" box="[969,1012,848,872]" pageId="37" pageNumber="188" type="generic">1 ♂</specimenCount>
, Dujiangyan City, Longchi Mountain,
<date id="FFDBB2AEFF90146FFF04F47BFE94E5A2" box="[151,316,884,909]" pageId="37" pageNumber="188" value="2005-08-26">26. VIII. 2005</date>
, coll. Caiyun Zhao;
<specimenCount id="9D635FE7FF90146FFDB0F47BFDF8E5A3" box="[547,592,884,908]" pageId="37" pageNumber="188" type="generic">1 ♂</specimenCount>
, 3 ♀♀, Dujiangyan City, Longchi Mountain,
<quantity id="4C9D398BFF90146FFBC6F47BFB05E5A3" box="[1109,1197,884,908]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="37" pageNumber="188" unit="m" value="1800.0">1800 m</quantity>
,
<date id="FFDBB2AEFF90146FFB29F47BFAF7E5A2" box="[1210,1375,884,909]" pageId="37" pageNumber="188" value="2005-08-26">26. VIII. 2005</date>
, coll. Xiaoyan Li;
<specimenCount id="9D635FE7FF90146FFEB5F497FEF9E580" box="[294,337,920,944]" pageId="37" pageNumber="188" type="generic">1 ♂</specimenCount>
, Emeishan City, Emei Mountain,
<date id="FFDBB2AEFF90146FFD40F497FCD0E59F" box="[723,888,920,944]" pageId="37" pageNumber="188" value="2005-08-28">28. VIII. 2005</date>
, coll. Caiyun Zhao; 2 ♀♀, Emeishan City, Emei Mountain,
<date id="FFDBB2AEFF90146FFE81F4B3FE1DE5FB" box="[274,437,956,980]" pageId="37" pageNumber="188" value="2005-08-28">28. VIII. 2005</date>
, coll. Yejun Zhang; 2 ♀♀, Emeishan City, Emei Mountain,
<quantity id="4C9D398BFF90146FFBCBF4B3FB18E5FC" box="[1112,1200,956,980]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="37" pageNumber="188" unit="m" value="1200.0">1200 m</quantity>
,
<date id="FFDBB2AEFF90146FFB2FF4B3FAF7E5FB" box="[1212,1375,956,980]" pageId="37" pageNumber="188" value="2005-08-29">29. VIII. 2005</date>
, coll. Xiaoyan Li; 1 ♀, Baoxing County, Fengtongzhai Nature Reserve, Dashuigou,
<quantity id="4C9D398BFF90146FFBC0F4EFFB1AE5D8" box="[1107,1202,992,1016]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="188" unit="m" value="1500.0">1500 m</quantity>
,
<date id="FFDBB2AEFF90146FFB55F4EFFAF0E5D7" box="[1222,1368,992,1016]" pageId="37" pageNumber="188" value="2005-09-02">2. IX. 2005</date>
, coll. Hongzhang Zhou and Caiyun Zhao;
<specimenCount id="9D635FE7FF90146FFDA0F30BFDC8E233" box="[563,608,1028,1052]" pageId="37" pageNumber="188" type="generic">1 ♂</specimenCount>
, 2 ♀♀, Baoxing County, Fengtongzhai Nature Reserve, Dashuigou,
<quantity id="4C9D398BFF90146FFAF7F30BFF07E210" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="188" unit="m" value="1500.0">1500 m</quantity>
,
<date id="FFDBB2AEFF90146FFF28F327FE96E26F" box="[187,318,1063,1088]" pageId="37" pageNumber="188" value="2005-09-03">3. IX. 2005</date>
, coll. Caiyun Zhao; 4 ♂♂, 2 ♀♀, Baoxing County, Fengtongzhai Nature Reserve, Dashuigou,
<quantity id="4C9D398BFF90146FFAF7F328FF07E24B" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="188" unit="m" value="1500.0">1500 m</quantity>
,
<date id="FFDBB2AEFF90146FFF28F343FEF1E24A" box="[187,345,1100,1125]" pageId="37" pageNumber="188" value="2005-09-02" valueMax="2005-09-03" valueMin="2005-09-02">23. IX. 2005</date>
, coll. Xiaoyan Li;
<specimenCount id="9D635FE7FF90146FFDB9F343FDFEE24B" box="[554,598,1100,1124]" pageId="37" pageNumber="188" type="generic">1 ♂</specimenCount>
, 1 ♀, Lushan County, Longmen Village,
<date id="FFDBB2AEFF90146FFBB0F343FB02E24A" box="[1059,1194,1100,1125]" pageId="37" pageNumber="188" value="2005-10-15">15. X. 2005</date>
, coll. Haifeng Chen.
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FF90146FFF54F360FB1AE282" pageId="37" pageNumber="188" type="description">
<paragraph id="8BDA946EFF90146FFF54F360FB1AE282" blockId="37.[151,1437,631,1341]" pageId="37" pageNumber="188">
<emphasis id="B911487CFF90146FFF54F360FED5E2A7" bold="true" box="[199,381,1135,1160]" pageId="37" pageNumber="188">Measurements.</emphasis>
BL =
<quantity id="4C9D398BFF90146FFE5AF360FD8AE2A8" box="[457,546,1135,1160]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="37" pageNumber="188" unit="mm" value="6.9">6.9 mm</quantity>
, BW =
<quantity id="4C9D398BFF90146FFDEFF37FFD7DE2A8" box="[636,725,1136,1160]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="188" unit="mm" value="1.4">1.4 mm</quantity>
, HL/PL/EL = 1.00: 1.10: 1.30, HW/PW/EW/AW = 1.00: 1.11: 1.42: 1.27, HW/HL = 1.02, HEL/HTL = 8.96, PW/PL = 1.03, EW/EL = 1.11, ESL/EL = 0.55.
</paragraph>
</subSubSection>
<subSubSection id="C37FC7E5FF90146FFF54F3B8FD79E312" pageId="37" pageNumber="188" type="distribution">
<paragraph id="8BDA946EFF90146FFF54F3B8FD79E312" blockId="37.[151,1437,631,1341]" pageId="37" pageNumber="188">
<emphasis id="B911487CFF90146FFF54F3B8FEC8E2FF" bold="true" box="[199,352,1207,1232]" pageId="37" pageNumber="188">Distribution.</emphasis>
<taxonomicName id="4C65EFEDFF90146FFEFAF3B7FD1AE2FF" authority="Cameron" authorityName="Cameron" box="[361,690,1208,1232]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="188" phylum="Arthropoda" rank="species" species="muscicola">
<emphasis id="B911487CFF90146FFEFAF3B7FDEAE2FF" box="[361,578,1208,1232]" italics="true" pageId="37" pageNumber="188">Quedius muscicola</emphasis>
<collectingCountry id="F372D4FEFF90146FFDD8F3B7FD1AE2FF" box="[587,690,1208,1232]" name="Cameroon" pageId="37" pageNumber="188">Cameron</collectingCountry>
</taxonomicName>
is known from
<collectingCountry id="F372D4FEFF90146FFCFFF3B6FC73E2FE" box="[876,987,1209,1233]" name="Myanmar" pageId="37" pageNumber="188">Myanmar</collectingCountry>
, and from the mountain areas from the Himalayan range (
<collectingCountry id="F372D4FEFF90146FFEFFF3D3FE01E2DA" box="[364,425,1244,1269]" name="India" pageId="37" pageNumber="188">India</collectingCountry>
,
<collectingCountry id="F372D4FEFF90146FFE2AF3D3FE56E2DA" box="[441,510,1244,1269]" name="Nepal" pageId="37" pageNumber="188">Nepal</collectingCountry>
) to southwestern
<collectingCountry id="F372D4FEFF90146FFD5EF3D3FCB8E2DA" box="[717,784,1244,1269]" name="China" pageId="37" pageNumber="188">China</collectingCountry>
(Gansu, Guizhou, Hubei, Shaanxi, Sichuan and Yunnan Province), at altitudes from
<quantity id="4C9D398BFF90146FFE5CF3F0FDBEE337" box="[463,534,1279,1304]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="37" pageNumber="188" unit="m" value="650.0">650 m</quantity>
to
<quantity id="4C9D398BFF90146FFDAAF20FFD39E337" box="[569,657,1280,1304]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="37" pageNumber="188" unit="m" value="2800.0">2800 m</quantity>
. Many of the examined specimens were collected in bushes, stacks of withered grass, leaf litter, or at the edges of creeks.
</paragraph>
</subSubSection>
</treatment>
</document>