treatments-xml/data/03/CC/0E/03CC0E3528378252FDB7FF59FDB7FA8C.xml
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<document id="C72B36417D8BEB606250C5616221041C" ID-DOI="10.1206/3855.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5368577" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630260853840" checkinUser="carolina" docAuthor="Engel, Michael S., Ortega-Blanco, Jaime &amp; Azevedo, Celso O." docDate="2016" docId="03CC0E3528378252FDB7FF59FDB7FA8C" docLanguage="en" docName="N3855.pdf" docOrigin="American Museum Novitates 2016 (3855)" docSource="http://www.bioone.org/doi/10.1206/3855.1" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Holopsenella Engel &amp; Ortega-Blanco &amp; Azevedo 2016, new genus" docType="treatment" docVersion="4" lastPageNumber="7" masterDocId="FFF5764D28318254FFBBFFB8FFEBFFAF" masterDocTitle="A New Bethylid Wasp in Lebanese Early Cretaceous Amber (Hymenoptera: Chrysidoidea), with Comments on other Mesozoic Taxa" masterLastPageNumber="16" masterPageNumber="1" pageNumber="7" updateTime="1699201309944" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F155CEF55F63C9EB39949753569CA3DF">A New Bethylid Wasp in Lebanese Early Cretaceous Amber (Hymenoptera: Chrysidoidea), with Comments on other Mesozoic Taxa</mods:title>
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<mods:namePart id="5085409F7475391F7E76E97E023F7685">Engel, Michael S.</mods:namePart>
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<mods:namePart id="6B0B3CBCB3FC3FC9561A2C905275ABE4">Ortega-Blanco, Jaime</mods:namePart>
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<mods:namePart id="9714AC9DA4C68EC24B283C169AE26B76">Azevedo, Celso O.</mods:namePart>
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<mods:title id="871B232E33D3F83B5B23852B302C821B">American Museum Novitates</mods:title>
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<mods:date id="96A6622EC3661BAC78A2BFD3032CBC0C">2016</mods:date>
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<subSubSection id="C37FECA828378252FDB7FF59FCDFFF53" box="[524,820,225,252]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BDABF2328378252FDB7FF59FCDFFF53" blockId="6.[524,820,225,252]" box="[524,820,225,252]" pageId="6" pageNumber="7">
<heading id="D092084F28378252FDB7FF59FCDFFF53" box="[524,820,225,252]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
<taxonomicName id="4C65C4A028378252FDB7FF59FD43FF53" authority="Engel &amp; Ortega-Blanco &amp; Azevedo, 2016" authorityName="Engel &amp; Ortega-Blanco &amp; Azevedo" authorityYear="2016" box="[524,680,225,252]" class="Insecta" family="Bethylidae" genus="Holopsenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911633128378252FDB7FF59FD43FF53" bold="true" box="[524,680,225,252]" italics="true" pageId="6" pageNumber="7">Holopsenella</emphasis>
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,
<taxonomicNameLabel id="A222DE4A28378252FD0CFF5AFCDFFF53" box="[695,820,226,252]" pageId="6" pageNumber="7" rank="genus">new genus</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C37FECA828378252FF13FEA2FCEBFE9C" box="[168,768,281,308]" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BDABF2328378252FF13FEA2FCEBFE9C" blockId="6.[120,1225,281,1316]" box="[168,768,281,308]" pageId="6" pageNumber="7">
<materialsCitation id="3B0DB57E28378252FF13FEA2FD16FE9C" box="[168,765,281,308]" collectionCode="SPECIES" pageId="6" pageNumber="7" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DE018128378252FF13FEA2FF08FE9B" box="[168,227,282,308]" pageId="6" pageNumber="7">TYPE</typeStatus>
<collectionCode id="ED7427E628378252FF50FEA1FEACFE9C" box="[235,327,281,307]" pageId="6" pageNumber="7">SPECIES</collectionCode>
:
<taxonomicName id="4C65C4A028378252FEEDFEA1FD8BFE9C" authority="Engel &amp; Ortega-Blanco &amp; Azevedo, 2016" authorityName="Engel &amp; Ortega-Blanco &amp; Azevedo" authorityYear="2016" box="[342,608,281,307]" class="Insecta" family="Bethylidae" genus="Holopsenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="primotica" status="sp. nov.">
<emphasis id="B911633128378252FEEDFEA1FD8BFE9C" box="[342,608,281,307]" italics="true" pageId="6" pageNumber="7">Holopsenella primotica</emphasis>
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,
<taxonomicNameLabel id="A222DE4A28378252FDD5FEA2FD16FE9C" box="[622,765,281,308]" pageId="6" pageNumber="7" rank="species">new species</taxonomicNameLabel>
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.
</paragraph>
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<subSubSection id="C37FECA828378252FF13FE87FD7AFBB1" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BDABF2328378252FF13FE87FD7AFBB1" blockId="6.[120,1225,281,1316]" pageId="6" pageNumber="7">
DIAGNOSIS:
<emphasis id="B911633128378252FE81FE87FE70FEF6" bold="true" box="[314,411,319,345]" pageId="6" pageNumber="7">Female.</emphasis>
Head broad, posterior margin in dorsal view straight, with rounded posterolateral corners (figs. 1, 2); vertex flat; occipital carina present (fig. 2A), weak; space between and anterior to antennal toruli depressed; antennal flagellum with 10 flagellomeres; scape distinctly thickened compared to pedicel and flagellum; compound eye circular, separated from posterior head margin in dorsal view by half of eye length; ocelli present (fig. 2B); clypeus projecting anteriorly (fig. 2B), surface comparatively flat except weak longitudinal medial ridge, apical margin with prominent but shallow medial emargination (fig. 2B); mandible short and thick, with at least two sharp apical teeth (likely three, as edge of a third seems partially apparent on left mandible), mandibles overlapping in slightly less than their apical halves, mandibles obscured by projection of clypeus in dorsal view; maxillary palpus elongate; labial palpus short. Forewing with marginal cell closed (fig. 2C), distal part of Rs almost straight resulting in evenly tapering marginal cell across its length, apex of cell acutely rounded; discoidal cell somewhat quadrate; cu-a straight; pterostigma longer than wide, with r-rs arising near pterostigmal midlength, r-rs long and arching apically to meet Rs; m-cu straight, nearly parallel with 1M (fig. 2C), and about as long as 1Cu; subdiscoidal cell slightly wider than discoidal cell, broader posteriorly than anteriorly. Tibial spur formula 1-2-2; basitarsi longest tarsomeres, tarsomeres IIIV distinctly shorter than basitarsi and tarsomeres V. Metasoma not petiolate; first metasomal tergum with weak transverse carina running across angle between anterior-facing and dorsal-facing surfaces (fig. 2A), then curving laterally along lateral disc for short distance.
</paragraph>
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<subSubSection id="C37FECA828378252FF13FB92FE9AFBEC" box="[168,369,1065,1092]" pageId="6" pageNumber="7" type="description">
<paragraph id="8BDABF2328378252FF13FB92FE9AFBEC" blockId="6.[120,1225,281,1316]" box="[168,369,1065,1092]" pageId="6" pageNumber="7">
<emphasis id="B911633128378252FF13FB92FF07FBEB" bold="true" box="[168,236,1066,1092]" pageId="6" pageNumber="7">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C37FECA828378252FF13FBF7FDB7FA8C" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8BDABF2328378252FF13FBF7FDB7FA8C" blockId="6.[120,1225,281,1316]" pageId="6" pageNumber="7">
ETYMOLOGY: The new genus-group name is a combination of the Greek terms
<emphasis id="B911633128378252FB8EFBF7FB9BFBC6" box="[1077,1136,1103,1129]" italics="true" pageId="6" pageNumber="7">holos</emphasis>
, meaning, “complete,” and the diminutive form of
<emphasis id="B911633128378252FD30FBCDFD55FB21" box="[651,702,1141,1166]" italics="true" pageId="6" pageNumber="7">psen</emphasis>
, meaning “wasp” (the Latinized diminutive
<emphasis id="B911633128378252FFC3FB21FF38FB1C" box="[120,211,1177,1203]" italics="true" pageId="6" pageNumber="7">psenella</emphasis>
serves as the stem for two of the most primitive bethyline genera exhibiting the most complete venation found in the family—
<taxonomicName id="4C65C4A028378252FDE4FB07FCAFFB76" authority="Kieffer" authorityName="Kieffer" authorityYear="1911" box="[607,836,1215,1241]" class="Insecta" family="Bethylidae" genus="Lytopsenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B911633128378252FDE4FB07FD07FB76" box="[607,748,1215,1241]" italics="true" pageId="6" pageNumber="7">Lytopsenella</emphasis>
Kieffer
</taxonomicName>
and
<taxonomicName id="4C65C4A028378252FC38FB07FB63FB75" authority="Westwood" authorityName="Westwood" authorityYear="1874" box="[899,1160,1215,1242]" class="Insecta" family="Bethylidae" genus="Eupsenella" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B911633128378252FC38FB07FC16FB76" box="[899,1021,1215,1241]" italics="true" pageId="6" pageNumber="7">Eupsenella</emphasis>
Westwood
</taxonomicName>
). The name refers to the comparatively complete wing venation relative to other lineages of
<taxonomicName id="4C65C4A028378252FBD1FB5CFF4FFA8C" class="Insecta" family="Bethylidae" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Bethylidae</taxonomicName>
. The gender of the name is feminine.
</paragraph>
</subSubSection>
</treatment>
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