treatments-xml/data/03/8F/E7/038FE731FFF176401E9EABBEFD2DFE86.xml

<document id="CDF57D363AA0902F5F68612D13FC7791" ID-CLB-Dataset="20790" ID-DOI="10.11646/zootaxa.5094.2.6" ID-GBIF-Dataset="feaa3d37-8cf6-4dc1-b538-867602d85627" ID-ISSN="1175-5326" ID-Zenodo-Dep="5965287" ID-ZooBank="21350F24-5440-4BD8-BD2B-5FF1AB3EDA99" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643872924138" checkinUser="plazi" docAuthor="Muthukatturaja, Marimuthu &amp; Balasubramanian, Chellaiah" docDate="2022" docId="038FE731FFF176401E9EABBEFD2DFE86" docLanguage="en" docName="zootaxa.5094.2.6.pdf" docOrigin="Zootaxa 5094 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procloeon (Procloeon) kottagudiensis Balasubramanian and Muthukatturaja 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="328" masterDocId="FFB69F49FFF076471E09A95BFF8EFFC7" masterDocTitle="A new record and new species of Procloeon (Procloeon) Bengtsson, 1915 (Ephemeroptera: Baetidae) from the Western Ghats, peninsular India" masterLastPageNumber="330" masterPageNumber="321" pageNumber="322" updateTime="1699306419422" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0FB5DD6C58C757BD08903FB707F6C6DF">A new record and new species of Procloeon (Procloeon) Bengtsson, 1915 (Ephemeroptera: Baetidae) from the Western Ghats, peninsular India</mods:title>
</mods:titleInfo>
<mods:name id="D49F7345CA9AC4D1F58AAFC844BAF684" type="personal">
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<mods:namePart id="F1054C1725399F86330CAA520E3DF07D">Muthukatturaja, Marimuthu</mods:namePart>
<mods:nameIdentifier id="399A8FC81F956C9A4D8BDF4E2D9330B2" type="ORCID">0000-0001-5796-0211</mods:nameIdentifier>
<mods:affiliation id="4F9CDD1ED9B7F8C2575D5769FAFD09AB">Department of Zoology, Thiagarajar College, Madurai- 625009, Tamil Nadu, India. &amp; drmkraja 83 @ gmail. com; https: // orcid. org / 0000 - 0001 - 5796 - 0211</mods:affiliation>
<mods:nameIdentifier id="AFDD624245287124AC9752F0DA263291" type="email">drmkraja83@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C36259B994CFB4D4BFB893FAAAF86B61">Balasubramanian, Chellaiah</mods:namePart>
<mods:nameIdentifier id="53FD340438BBFB865713345EC386E7A1" type="ORCID">0000-0001-8563-1677</mods:nameIdentifier>
<mods:affiliation id="46C5C3C5B059ABFCC14490861E24BE29">Department of Zoology, Thiagarajar College, Madurai- 625009, Tamil Nadu, India. &amp; ulab 62 @ yahoo. com; https: // orcid. org / 0000 - 0001 - 8563 - 1677</mods:affiliation>
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<treatment id="038FE731FFF176401E9EABBEFD2DFE86" ID-DOI="http://doi.org/10.5281/zenodo.6301628" ID-GBIF-Taxon="193277074" ID-Zenodo-Dep="6301628" LSID="urn:lsid:plazi:treatment:038FE731FFF176401E9EABBEFD2DFE86" httpUri="http://treatment.plazi.org/id/038FE731FFF176401E9EABBEFD2DFE86" lastPageId="7" lastPageNumber="328" pageId="1" pageNumber="322">
<subSubSection id="C33C05ACFFF176461E9EABBEFBDEFCC7" box="[151,1104,741,768]" pageId="1" pageNumber="322" type="nomenclature">
<paragraph id="8B995627FFF176461E9EABBEFBDEFCC7" blockId="1.[151,1104,741,805]" box="[151,1104,741,768]" pageId="1" pageNumber="322">
<heading id="D0D1E14BFFF176461E9EABBEFBDEFCC7" bold="true" box="[151,1104,741,768]" fontSize="11" level="1" pageId="1" pageNumber="322" reason="1">
<emphasis id="B9528A35FFF176461E9EABBEFBDEFCC7" bold="true" box="[151,1104,741,768]" pageId="1" pageNumber="322">
<taxonomicName id="4C262DA4FFF176461E9EABBEFC65FD38" authority="Balasubramanian and Muthukatturaja" authorityName="Balasubramanian and Muthukatturaja" authorityYear="2022" box="[151,1003,741,768]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="species" species="kottagudiensis" status="sp. nov." subGenus="Procloeon">
<emphasis id="B9528A35FFF176461E9EABBEFE9DFCC7" bold="true" box="[151,275,741,768]" italics="true" pageId="1" pageNumber="322">Procloeon</emphasis>
(
<emphasis id="B9528A35FFF176461F2DABBDFEB4FCC7" bold="true" box="[292,314,742,768]" italics="true" pageId="1" pageNumber="322">P.</emphasis>
) 
<emphasis id="B9528A35FFF176461F42ABBEFE77FCC7" bold="true" box="[331,505,741,768]" italics="true" pageId="1" pageNumber="322">kottagudiensis</emphasis>
Balasubramanian and Muthukatturaja
</taxonomicName>
<taxonomicNameLabel id="A261374EFFF176461DFBABBDFBDEFCC7" box="[1010,1104,742,768]" pageId="1" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33C05ACFFF176461E9EAA51FE94FCE2" box="[151,282,778,805]" pageId="1" pageNumber="322" type="description">
<paragraph id="8B995627FFF176461E9EAA51FE94FCE2" blockId="1.[151,1104,741,805]" box="[151,282,778,805]" pageId="1" pageNumber="322">
(
<figureCitation id="131D4AA2FFF176461E96AA51FE9FFCE2" box="[159,273,778,805]" captionStart-0="FIGURES 1–3" captionStart-1="FIGURES 4–9" captionStart-2="FIGURES 10–15" captionStart-3="FIGURES 16–23" captionStart-4="FIGURES 24–29" captionStart-5="FIGURES 30–36" captionStartId-0="2.[151,264,1405,1429]" captionStartId-1="3.[151,264,1367,1391]" captionStartId-2="4.[151,264,1477,1501]" captionStartId-3="5.[151,264,1516,1540]" captionStartId-4="6.[151,264,955,979]" captionStartId-5="6.[151,264,1971,1995]" captionTargetBox-0="[151,1436,345,1381]" captionTargetBox-1="[151,1436,181,1344]" captionTargetBox-2="[152,1436,182,1453]" captionTargetBox-3="[151,1436,181,1491]" captionTargetBox-4="[165,1423,181,931]" captionTargetBox-5="[187,1394,1044,1938]" captionTargetId-0="figure-104@2.[151,1436,345,1381]" captionTargetId-1="figure-17@3.[151,1436,181,1344]" captionTargetId-2="figure-20@4.[151,1436,181,1453]" captionTargetId-3="figure-17@5.[151,1436,181,1492]" captionTargetId-4="figure-20@6.[165,1423,181,931]" captionTargetId-5="figure-58@6.[180,1406,1040,1948]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="6" captionText-0="FIGURES 1–3. Procloeon (P.) kottagudiensis sp. nov., mature nymph. 1, dorsal view; 2, ventral view; 3, lateral view." captionText-1="FIGURES 4–9. Procloeon (P.) kottagudiensis sp. nov., mature nymph. Dorsal view: 4, labrum; 5, right mandible; 6, left mandible; 7, maxillae; 8, hypopharynx; 9, labium." captionText-2="FIGURES 10–15. Procloeon (P.) kottagudiensis sp. nov., mature nymph. Dorsal view: 10, foreleg; 11, tarsal claw; 12, gill I; 13, gill IV; 14, gill VII; 15, paraproct." captionText-3="FIGURES 16–23. Procloeon (P.) kottagudiensis sp. nov., mature nymph: 16, abdomen (dorsal view); 17, abdomen (ventral view); 18, abdominal segments VIII &amp; IX (dorsal view); 19, abdominal segment X (dorsal view); 20, abdominal tergum I; 21, abdominal tergum V; 22, paracercus, 23, cercus." captionText-4="FIGURES 24–29. Procloeon (P.) kottagudiensis sp. nov., imago: 24, male imago (dorsal view); 25, female imago (dorsal view); 26, abdomen (dorsal view); 27, abdomen (ventral view); 28, abdomen (lateral view); 29, thoracic sternum." captionText-5="FIGURES 30–36. Procloeon (P.) kottagudiensis sp. nov., male imago: 30, forewing; 31, hindwing; 32, hindwing enlarged; 33, male genitalia; 34, foreleg; 35, midleg; 36, hindleg." figureDoi-0="http://doi.org/10.5281/zenodo.5965291" figureDoi-1="http://doi.org/10.5281/zenodo.5965293" figureDoi-2="http://doi.org/10.5281/zenodo.5965297" figureDoi-3="http://doi.org/10.5281/zenodo.5965299" figureDoi-4="http://doi.org/10.5281/zenodo.5965303" figureDoi-5="http://doi.org/10.5281/zenodo.5965305" httpUri-0="https://zenodo.org/record/5965291/files/figure.png" httpUri-1="https://zenodo.org/record/5965293/files/figure.png" httpUri-2="https://zenodo.org/record/5965297/files/figure.png" httpUri-3="https://zenodo.org/record/5965299/files/figure.png" httpUri-4="https://zenodo.org/record/5965303/files/figure.png" httpUri-5="https://zenodo.org/record/5965305/files/figure.png" pageId="1" pageNumber="322">Figs 1–36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33C05ACFFF176461E9EAA12FDFBFB29" pageId="1" pageNumber="322" type="materials_examined">
<paragraph id="8B995627FFF176461E9EAA12FE2EFC35" blockId="1.[151,1437,840,2019]" pageId="1" pageNumber="322">
<emphasis id="B9528A35FFF176461E9EAA12FE00FCA5" bold="true" box="[151,398,840,867]" pageId="1" pageNumber="322">
Type material (
<collectionCode id="ED37CEE2FFF176461F59AA13FE0FFCA5" box="[336,385,840,866]" pageId="1" pageNumber="322">ZSI</collectionCode>
).
</emphasis>
<materialsCitation id="3B4E5C7AFFF176461F9EAA12FD18FC6D" ID-GBIF-Occurrence="3462299302" collectingDate="2021-02-30" collectorName=" Balasubramanian &amp; Muthukatturaja" country="India" county="Kottagudi" location="Theni District" municipality="Kuragani" pageId="1" pageNumber="322" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<emphasis id="B9528A35FFF176461F9EAA12FD80FCA4" bold="true" box="[407,526,841,867]" pageId="1" pageNumber="322">
<typeStatus id="549DE885FFF176461F9EAA12FD86FCA4" box="[407,520,841,867]" pageId="1" pageNumber="322" type="holotype">Holotype</typeStatus>
:
</emphasis>
male imago (
<collectorName id="26D333F1FFF176461CA5AA12FC85FCA4" box="[684,779,841,867]" pageId="1" pageNumber="322">Reg.No.</collectorName>
SRC/ ZSI-I/E 616) (reared from mature nymph), 
<collectingCountry id="F33116B7FFF176461B4FAA12FA16FCA4" box="[1350,1432,841,867]" name="India" pageId="1" pageNumber="322">INDIA</collectingCountry>
, 
<collectingCounty id="62F82EABFFF176461E9EAA37FE89FC41" box="[151,263,876,902]" pageId="1" pageNumber="322">Kottagudi</collectingCounty>
stream, 
<collectingMunicipality id="6BFDCC5DFFF176461F6CAA37FE5EFC41" box="[357,464,876,902]" pageId="1" pageNumber="322">Kuragani</collectingMunicipality>
, 
<location id="8EF900FCFFF176461FD3AA37FDF6FC41" LSID="urn:lsid:plazi:treatment:038FE731FFF176401E9EABBEFD2DFE86:8EF900FCFFF176461FD3AA37FDF6FC41" box="[474,632,876,902]" country="India" county="Kottagudi" municipality="Kuragani" name="Theni District" pageId="1" pageNumber="322" stateProvince="Tamil Nadu">Theni District</location>
, 
<collectingRegion id="49E298C5FFF176461C8BAA37FCCCFC40" box="[642,834,876,903]" country="India" name="Tamil Nadu" pageId="1" pageNumber="322">Tamil Nadu state</collectingRegion>
, 
<date id="FF9870E7FFF176461D45AA36FC46FC40" box="[844,968,877,903]" pageId="1" pageNumber="322" value="2021-02-30">
<collectingDate id="EFDC890FFFF176461D45AA36FC46FC40" box="[844,968,877,903]" pageId="1" pageNumber="322" value="2021-02-30">
30-II- 
<quantity id="4CDEFBC2FFF176461D87AA36FC46FC40" box="[910,968,877,903]" metricMagnitude="6" metricUnit="m" metricValue="2.0218" pageId="1" pageNumber="322" unit="m" value="2021800.0">2021</quantity>
</collectingDate>
</date>
, 800 m a.s.l, (10°04’576” N, 77°15’1100” E), colls. 
<collectorName id="26D333F1FFF176461F08AACBFE4DFC6D" box="[257,451,912,938]" pageId="1" pageNumber="322">Balasubramanian</collectorName>
&amp; 
<collectorName id="26D333F1FFF176461FEFAACBFD18FC6D" box="[486,662,912,938]" pageId="1" pageNumber="322">Muthukatturaja</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B4E5C7AFFF176461CA9AACAFE15FC35" ID-GBIF-Occurrence="3462299301" collectingDate="2021-02-30" collectorName=" Balasubramanian &amp; Muthukatturaja" country="India" county="Kottagudi" location="Theni District" municipality="Kuragani" pageId="1" pageNumber="322" specimenCount="2" specimenCount-female="2" stateProvince="Tamil Nadu" typeStatus="paratype">
<emphasis id="B9528A35FFF176461CA9AACAFCAFFC6C" bold="true" box="[672,801,913,939]" pageId="1" pageNumber="322">
<typeStatus id="549DE885FFF176461CA9AACAFC92FC6C" box="[672,796,913,939]" pageId="1" pageNumber="322" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D209DAEFFF176461D2EAACAFC06FC6D" box="[807,904,912,939]" pageId="1" pageNumber="322" type="female">2 female</specimenCount>
imagoes (reared from mature nymph) (
<collectorName id="26D333F1FFF176461B37AACAFA13FC6C" box="[1342,1437,913,939]" pageId="1" pageNumber="322">Reg.No.</collectorName>
SRC/ ZSI-I/E 616) and 2 mature nymphs (
<collectorName id="26D333F1FFF176461C79AAEEFD41FC08" box="[624,719,949,975]" pageId="1" pageNumber="322">Reg.No.</collectorName>
SRC/ ZSI-I/E 617), same data as holotype, colls. 
<collectorName id="26D333F1FFF176461B08AAEFFF49FC35" pageId="1" pageNumber="322">Balasubramanian</collectorName>
&amp; 
<collectorName id="26D333F1FFF176461EE2AA83FE15FC35" box="[235,411,984,1010]" pageId="1" pageNumber="322">Muthukatturaja</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8B995627FFF176461ECEAAA7FDFBFB29" blockId="1.[151,1437,840,2019]" pageId="1" pageNumber="322">
<emphasis id="B9528A35FFF176461ECEAAA7FDEBFBD1" bold="true" box="[199,613,1020,1046]" pageId="1" pageNumber="322">
Other material examined (
<collectionCode id="ED37CEE2FFF176461FF6AAA7FDD6FBD1" box="[511,600,1020,1046]" pageId="1" pageNumber="322">ZMTC</collectionCode>
).
</emphasis>
<specimenCount id="9D209DAEFFF176461C62AAA7FD3AFBD1" box="[619,692,1020,1047]" pageId="1" pageNumber="322" type="male">5 male</specimenCount>
(Reg.No. ZMTC-79) and 
<materialsCitation id="3B4E5C7AFFF176461DDEAAA7FF57FB98" ID-GBIF-Occurrence="3462299305" collectionCode="II-" collectorName="Reg. No." country="India" county="Kottagudi" location="Theni District" municipality="Kuragani" pageId="1" pageNumber="322" specimenCount="5" specimenCount-female="5" stateProvince="Tamil Nadu">
<specimenCount id="9D209DAEFFF176461DDEAAA7FBB9FBD1" box="[983,1079,1020,1047]" pageId="1" pageNumber="322" type="female">5 female</specimenCount>
imagoes (
<collectorName id="26D333F1FFF176461AA1AAA6FA89FBD0" box="[1192,1287,1021,1047]" pageId="1" pageNumber="322">Reg.No.</collectorName>
ZMTC-80),, and 10 mature nymphs (
<collectorName id="26D333F1FFF176461FAAAD7AFD8CFBFC" box="[419,514,1057,1083]" pageId="1" pageNumber="322">Reg.No.</collectorName>
ZMTC-81), 
<collectingCountry id="F33116B7FFF176461C98AD7AFD6DFBFC" box="[657,739,1057,1083]" name="India" pageId="1" pageNumber="322">INDIA</collectingCountry>
, 
<collectingCounty id="62F82EABFFF176461CE4AD7BFCD3FBFD" box="[749,861,1056,1082]" pageId="1" pageNumber="322">Kottagudi</collectingCounty>
stream, 
<collectingMunicipality id="6BFDCC5DFFF176461DB2AD7BFBA8FBFD" box="[955,1062,1056,1082]" pageId="1" pageNumber="322">Kuragani</collectingMunicipality>
, 
<location id="8EF900FCFFF176461A39AD7BFB40FBFD" LSID="urn:lsid:plazi:treatment:038FE731FFF176401E9EABBEFD2DFE86:8EF900FCFFF176461A39AD7BFB40FBFD" box="[1072,1230,1056,1082]" country="India" county="Kottagudi" municipality="Kuragani" name="Theni District" pageId="1" pageNumber="322" stateProvince="Tamil Nadu">Theni District</location>
, 
<collectingRegion id="49E298C5FFF176461AD1AD7BFA16FBFC" box="[1240,1432,1056,1083]" country="India" name="Tamil Nadu" pageId="1" pageNumber="322">Tamil Nadu state</collectingRegion>
, 
<date id="FF9870E7FFF176461E9EAD1EFF57FB98" box="[151,217,1093,1119]" pageId="1" pageNumber="322" value="2021-02-30">
30- 
<collectionCode id="ED37CEE2FFF176461EB5AD1EFF57FB98" box="[188,217,1093,1119]" pageId="1" pageNumber="322">II-</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B4E5C7AFFF176461ED0AD1EFBC1FB44" ID-GBIF-Occurrence="3462299303" country="India" elevation="2021800" location="Idukki District" municipality="Bison valley" pageId="1" pageNumber="322" specimenCount="1" stateProvince="Kerala">
<quantity id="4CDEFBC2FFF176461ED0AD1EFEE2FB98" box="[217,364,1093,1119]" metricMagnitude="6" metricUnit="m" metricValue="2.0218" pageId="1" pageNumber="322" unit="m" value="2021800.0">
<elevation id="000BB114FFF176461ED0AD1EFEE2FB98" box="[217,364,1093,1119]" metricMagnitude="6" metricUnit="m" metricValue="2.0218" pageId="1" pageNumber="322" unit="m" value="2021800.0">2021, 800 m</elevation>
</quantity>
a.s.l, (10°04’576” N, 77°15’1100” E), colls. 
<collectorName id="26D333F1FFF176461D74AD1FFBB1FB99" box="[893,1087,1092,1118]" pageId="1" pageNumber="322">Balasubramanian</collectorName>
&amp; 
<collectorName id="26D333F1FFF176461A63AD1FFA91FB99" box="[1130,1311,1092,1118]" pageId="1" pageNumber="322">Muthukatturaja.</collectorName>
15 mature nymphs (
<collectorName id="26D333F1FFF176461EF6AD32FED0FB44" box="[255,350,1129,1155]" pageId="1" pageNumber="322">Reg.No.</collectorName>
ZMTC-82) from 
<collectingMunicipality id="6BFDCC5DFFF176461C2FAD33FD3DFB45" box="[550,691,1128,1154]" pageId="1" pageNumber="322">Bison valley</collectingMunicipality>
, 
<location id="8EF900FCFFF176461CB6AD33FCE8FB45" LSID="urn:lsid:plazi:treatment:038FE731FFF176401E9EABBEFD2DFE86:8EF900FCFFF176461CB6AD33FCE8FB45" box="[703,870,1128,1154]" country="India" municipality="Bison valley" name="Idukki District" pageId="1" pageNumber="322" stateProvince="Kerala">Idukki District</location>
, 
<collectingRegion id="49E298C5FFF176461D7BAD33FC77FB44" box="[882,1017,1128,1155]" country="India" name="Kerala" pageId="1" pageNumber="322">Kerala state</collectingRegion>
, 
<date id="FF9870E7FFF176461A0CAD32FBC1FB44" box="[1029,1103,1129,1155]" pageId="1" pageNumber="322" value="2021-04-17">17-IV-</date>
</materialsCitation>
<materialsCitation id="3B4E5C7AFFF176461A46AD32FCBAFB0C" ID-GBIF-Occurrence="3462299304" collectionCode="III-" country="India" county="Periyavarai" elevation="2021882" latitude="10.004167" location="Idukki district" longLatPrecision="21" longitude="77.111664" municipality="Munnar" pageId="1" pageNumber="322" specimenCount="1" stateProvince="Kerala">
<quantity id="4CDEFBC2FFF176461A46AD32FB52FB44" box="[1103,1244,1129,1155]" metricMagnitude="6" metricUnit="m" metricValue="2.021882" pageId="1" pageNumber="322" unit="m" value="2021882.0">
<elevation id="000BB114FFF176461A46AD32FB52FB44" box="[1103,1244,1129,1155]" metricMagnitude="6" metricUnit="m" metricValue="2.021882" pageId="1" pageNumber="322" unit="m" value="2021882.0">2021, 882 m</elevation>
</quantity>
a.s.l, (
<geoCoordinate id="EE1230E0FFF176461B2EAD33FF20FB60" degrees="10" direction="north" minutes="00" orientation="latitude" pageId="1" pageNumber="322" precision="15" seconds="15" value="10.004167">10°00’15” N</geoCoordinate>
, 
<geoCoordinate id="EE1230E0FFF176461EB3ADD7FEC4FB60" box="[186,330,1164,1191]" degrees="77" direction="east" minutes="06" orientation="longitude" pageId="1" pageNumber="322" precision="15" seconds="42" value="77.111664">77°06’42” E</geoCoordinate>
), colls. 
<collectorName id="26D333F1FFF176461FABADD7FDEAFB61" box="[418,612,1164,1190]" pageId="1" pageNumber="322">Balasubramanian</collectorName>
&amp; 
<collectorName id="26D333F1FFF176461C80ADD7FCB3FB61" box="[649,829,1164,1190]" pageId="1" pageNumber="322">Muthukatturaja.</collectorName>
40 mature nymphs (
<collectorName id="26D333F1FFF176461A2DADD6FB0DFB60" box="[1060,1155,1165,1191]" pageId="1" pageNumber="322">Reg.No.</collectorName>
ZMTC-83) from 
<collectingCounty id="62F82EABFFF176461B42ADD7FF41FB0D" pageId="1" pageNumber="322">Periyavarai</collectingCounty>
stream (
<collectingMunicipality id="6BFDCC5DFFF176461F3BADEAFE1EFB0C" box="[306,400,1201,1227]" pageId="1" pageNumber="322">Munnar</collectingMunicipality>
), 
<location id="8EF900FCFFF176461FACADEBFDC9FB0D" LSID="urn:lsid:plazi:treatment:038FE731FFF176401E9EABBEFD2DFE86:8EF900FCFFF176461FACADEBFDC9FB0D" box="[421,583,1200,1226]" country="India" county="Periyavarai" latitude="10.004167" longLatPrecision="21" longitude="77.111664" municipality="Munnar" name="Idukki district" pageId="1" pageNumber="322" stateProvince="Kerala">Idukki district</location>
, 
<collectingRegion id="49E298C5FFF176461C5AADEBFD52FB0C" box="[595,732,1200,1227]" country="India" name="Kerala" pageId="1" pageNumber="322">Kerala state</collectingRegion>
, 
<date id="FF9870E7FFF176461CE1ADEAFCBAFB0C" box="[744,820,1201,1227]" pageId="1" pageNumber="322" value="2021-03-29">
29- 
<collectionCode id="ED37CEE2FFF176461D04ADEAFCBAFB0C" box="[781,820,1201,1227]" pageId="1" pageNumber="322">III-</collectionCode>
</date>
</materialsCitation>
2021, 
<quantity id="4CDEFBC2FFF176461D73ADEBFC59FB0C" box="[890,983,1200,1227]" metricMagnitude="3" metricUnit="m" metricValue="1.501" pageId="1" pageNumber="322" unit="m" value="1501.0">1,501 m</quantity>
a.s.l, (10°10’9476” N, 77°05’7512” E), colls. Balasubramanian &amp; Muthukatturaja.
</paragraph>
</subSubSection>
<subSubSection id="C33C05ACFFF176461ECEADA3FD6CF9F5" pageId="1" pageNumber="322" type="diagnosis">
<paragraph id="8B995627FFF176461ECEADA3FB7FFAB9" blockId="1.[151,1437,840,2019]" pageId="1" pageNumber="322">
<emphasis id="B9528A35FFF176461ECEADA3FECCFAD5" bold="true" box="[199,322,1272,1298]" pageId="1" pageNumber="322">Diagnosis.</emphasis>
The larva of 
<taxonomicName id="4C262DA4FFF176461FEDADA3FCBAFAD5" authority="Muthukatturaja &amp; Balasubramanian, 2022" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2022" box="[484,820,1272,1299]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="species" species="kottagudiensis" status="sp. nov." subGenus="Procloeon">
<emphasis id="B9528A35FFF176461FEDADA3FDD6FAD5" box="[484,600,1272,1298]" italics="true" pageId="1" pageNumber="322">Procloeon</emphasis>
(
<emphasis id="B9528A35FFF176461C62ADA2FD0EFAD5" box="[619,640,1273,1298]" italics="true" pageId="1" pageNumber="322">P.</emphasis>
) 
<emphasis id="B9528A35FFF176461C9BADA3FCBAFAD5" box="[658,820,1272,1298]" italics="true" pageId="1" pageNumber="322">kottagudiensis</emphasis>
</taxonomicName>
<emphasis id="B9528A35FFF176461D37ADA2FC14FAD4" bold="true" box="[830,922,1273,1299]" pageId="1" pageNumber="322">
<taxonomicNameLabel id="A261374EFFF176461D37ADA2FC14FAD4" box="[830,922,1273,1299]" pageId="1" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from congeners by the combination of the following diagnostic characters: (i) segment III of labial palp truncate, with sparse stout setae and hair-like setae on dorsal surface; (ii) abdominal tergum VIII with ca. 4 lateral spines; (iii) posterior margin of tergum I with acute spines; (iv) gill VII single without process; (v) paraproct with 7–9 acute spines.
</paragraph>
<paragraph id="8B995627FFF176461ECEACD3FD6CF9F5" blockId="1.[151,1437,840,2019]" pageId="1" pageNumber="322">
The imago of 
<taxonomicName id="4C262DA4FFF176461F6EACD3FD3AFA65" authority="Muthukatturaja &amp; Balasubramanian, 2022" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2022" box="[359,692,1416,1443]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="322" phylum="Arthropoda" rank="species" species="kottagudiensis" status="sp. nov." subGenus="Procloeon">
<emphasis id="B9528A35FFF176461F6EACD3FE55FA65" box="[359,475,1416,1442]" italics="true" pageId="1" pageNumber="322">Procloeon</emphasis>
(
<emphasis id="B9528A35FFF176461FE5ACD2FD8FFA65" box="[492,513,1417,1442]" italics="true" pageId="1" pageNumber="322">P.</emphasis>
) 
<emphasis id="B9528A35FFF176461C1BACD3FD3AFA65" box="[530,692,1416,1442]" italics="true" pageId="1" pageNumber="322">kottagudiensis</emphasis>
</taxonomicName>
<emphasis id="B9528A35FFF176461CB2ACD2FC9BFA64" bold="true" box="[699,789,1417,1443]" pageId="1" pageNumber="322">
<taxonomicNameLabel id="A261374EFFF176461CB2ACD2FC9BFA64" box="[699,789,1417,1443]" pageId="1" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from congeners by the combination of the following diagnostic characters: (i) stigmatic area of forewing milky transparent with 8 cross-veins; (ii) longitudinal veins with single crossvein except areas of R and MA with two cross-veins, Sc with three cross-veins; (iii) hind wings approximately five times longer than wide; (iv) costal projection of hind wing located at 1/2 distance from the base, and the longitudinal veins not forked.
</paragraph>
</subSubSection>
<subSubSection id="C33C05ACFFF176421ECEAF67FAE1F8AB" lastPageId="5" lastPageNumber="326" pageId="1" pageNumber="322" type="description">
<paragraph id="8B995627FFF176461ECEAF67FCB7F8CD" blockId="1.[151,1437,840,2019]" pageId="1" pageNumber="322">
<emphasis id="B9528A35FFF176461ECEAF67FD93F990" bold="true" box="[199,541,1596,1623]" pageId="1" pageNumber="322">
<emphasis id="B9528A35FFF176461ECEAF67FEDAF991" bold="true" box="[199,340,1596,1622]" italics="true" pageId="1" pageNumber="322">Description.</emphasis>
Mature Nymph
</emphasis>
(
<figureCitation id="131D4AA2FFF176461C38AF67FD2CF990" box="[561,674,1596,1623]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1405,1429]" captionTargetBox="[151,1436,345,1381]" captionTargetId="figure-104@2.[151,1436,345,1381]" captionTargetPageId="2" captionText="FIGURES 1–3. Procloeon (P.) kottagudiensis sp. nov., mature nymph. 1, dorsal view; 2, ventral view; 3, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5965291" httpUri="https://zenodo.org/record/5965291/files/figure.png" pageId="1" pageNumber="322">Figs. 1–3</figureCitation>
): Body length (excluding cerci) 
<quantity id="4CDEFBC2FFF176461A2AAF67FB20F990" box="[1059,1198,1596,1623]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="1" pageNumber="322" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">8.5–9.5 mm</quantity>
in male and female nymphs; cerci 
<quantity id="4CDEFBC2FFF176461F37AF3BFE47F9BC" box="[318,457,1632,1659]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="1" pageNumber="322" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5–2.8 mm</quantity>
; median terminal filaments 
<quantity id="4CDEFBC2FFF176461D0CAF3BFC1EF9BC" box="[773,912,1632,1659]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="1" pageNumber="322" unit="mm" value="2.3" valueMax="2.5" valueMin="2.1">2.1–2.5 mm</quantity>
; head width 1.0 mm; head length 
<quantity id="4CDEFBC2FFF176461B1BAF3AFAE2F9BC" box="[1298,1388,1633,1659]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="322" unit="mm" value="0.7">0.7 mm</quantity>
; antenna length 
<quantity id="4CDEFBC2FFF176461F22AFDFFE1DF958" box="[299,403,1668,1695]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="322" unit="mm" value="1.75">1.75 mm</quantity>
; labrum width 
<quantity id="4CDEFBC2FFF176461C49AFDEFD14F958" box="[576,666,1669,1695]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="322" unit="mm" value="0.3">0.3 mm</quantity>
; labrum length 
<quantity id="4CDEFBC2FFF176461D44AFDFFC38F958" box="[845,950,1668,1695]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="322" unit="mm" value="0.25">0.25 mm</quantity>
; compound eye length 
<quantity id="4CDEFBC2FFF176461AB6AFDEFA97F958" box="[1215,1305,1669,1695]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="322" unit="mm" value="0.3">0.3 mm</quantity>
; compound eye width 
<quantity id="4CDEFBC2FFF176461F06AFF3FEF6F904" box="[271,376,1704,1731]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="322" unit="mm" value="0.25">0.25 mm</quantity>
; distance between compound eye 
<quantity id="4CDEFBC2FFF176461CF4AFF2FCD9F904" box="[765,855,1705,1731]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="322" unit="mm" value="0.3">0.3 mm</quantity>
; length of fore femur/ tibia/ tarsi 1.3/ 0.8/ 
<quantity id="4CDEFBC2FFF176461B34AFF2FA16F904" box="[1341,1432,1705,1731]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="322" unit="mm" value="0.8">0.8 mm</quantity>
; length of middle femur/ tibia/ tarsi 1.3/ 0.7/ 
<quantity id="4CDEFBC2FFF176461C72AF96FD5DF920" box="[635,723,1741,1767]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="322" unit="mm" value="0.7">0.7 mm</quantity>
; length of hind femur/ tibia/ tarsi 1.5/ 0.7/ 
<quantity id="4CDEFBC2FFF176461AACAF96FB72F920" box="[1189,1276,1741,1767]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="322" unit="mm" value="0.8">0.8 mm</quantity>
. General body coloration light yellow to brown with dark brown markings.
</paragraph>
<paragraph id="8B995627FFF176451ECEAE4FFBADFE86" blockId="1.[151,1437,840,2019]" lastBlockId="2.[151,1436,151,321]" lastPageId="2" lastPageNumber="323" pageId="1" pageNumber="322">
Head: coloration light brown; vertex with paired dark brown stripes medially.Antennae pale yellow, longer than width of head; labrum (
<figureCitation id="131D4AA2FFF176461F95AE63FE50F894" box="[412,478,1848,1875]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1367,1391]" captionTargetBox="[151,1436,181,1344]" captionTargetId="figure-17@3.[151,1436,181,1344]" captionTargetPageId="3" captionText="FIGURES 4–9. Procloeon (P.) kottagudiensis sp. nov., mature nymph. Dorsal view: 4, labrum; 5, right mandible; 6, left mandible; 7, maxillae; 8, hypopharynx; 9, labium." figureDoi="http://doi.org/10.5281/zenodo.5965293" httpUri="https://zenodo.org/record/5965293/files/figure.png" pageId="1" pageNumber="322">Fig. 4</figureCitation>
) with submarginal stout setae laterally; margin of labrum with median W-shaped incision; anterior margin with row of long feathered setae and long fringed setae; right mandible (
<figureCitation id="131D4AA2FFF176461AADAE07FB6BF8B0" box="[1188,1253,1884,1911]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1367,1391]" captionTargetBox="[151,1436,181,1344]" captionTargetId="figure-17@3.[151,1436,181,1344]" captionTargetPageId="3" captionText="FIGURES 4–9. Procloeon (P.) kottagudiensis sp. nov., mature nymph. Dorsal view: 4, labrum; 5, right mandible; 6, left mandible; 7, maxillae; 8, hypopharynx; 9, labium." figureDoi="http://doi.org/10.5281/zenodo.5965293" httpUri="https://zenodo.org/record/5965293/files/figure.png" pageId="1" pageNumber="322">Fig. 5</figureCitation>
) with more than 7 blunt denticles on incisors; 8 denticles on prostheca and cluster of hair-like setae between prostheca and mola; left mandible (
<figureCitation id="131D4AA2FFF176461F06AEFFFEDCF878" box="[271,338,1956,1983]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1367,1391]" captionTargetBox="[151,1436,181,1344]" captionTargetId="figure-17@3.[151,1436,181,1344]" captionTargetPageId="3" captionText="FIGURES 4–9. Procloeon (P.) kottagudiensis sp. nov., mature nymph. Dorsal view: 4, labrum; 5, right mandible; 6, left mandible; 7, maxillae; 8, hypopharynx; 9, labium." figureDoi="http://doi.org/10.5281/zenodo.5965293" httpUri="https://zenodo.org/record/5965293/files/figure.png" pageId="1" pageNumber="322">Fig. 6</figureCitation>
) with 9 blunt denticles on incisors, 8 denticles on prostheca and cluster of hair-like setae between prostheca and mola; apex of maxilla (
<figureCitation id="131D4AA2FFF176461C34AE93FD0EF824" box="[573,640,1992,2019]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1367,1391]" captionTargetBox="[151,1436,181,1344]" captionTargetId="figure-17@3.[151,1436,181,1344]" captionTargetPageId="3" captionText="FIGURES 4–9. Procloeon (P.) kottagudiensis sp. nov., mature nymph. Dorsal view: 4, labrum; 5, right mandible; 6, left mandible; 7, maxillae; 8, hypopharynx; 9, labium." figureDoi="http://doi.org/10.5281/zenodo.5965293" httpUri="https://zenodo.org/record/5965293/files/figure.png" pageId="1" pageNumber="322">Fig. 7</figureCitation>
) with 3 prominent dentisetae and numerous stout monopectinate setae; galea-lacinia with 2–3 long hair-like setae at surface; palp 3-segmented; segment III ca. 0.8×length of segment I, conical, and with scattered hair-like setae; lingua and superlingua of hypopharynx (
<figureCitation id="131D4AA2FFF276451A27A9E0FBE1FF12" box="[1070,1135,187,213]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1367,1391]" captionTargetBox="[151,1436,181,1344]" captionTargetId="figure-17@3.[151,1436,181,1344]" captionTargetPageId="3" captionText="FIGURES 4–9. Procloeon (P.) kottagudiensis sp. nov., mature nymph. Dorsal view: 4, labrum; 5, right mandible; 6, left mandible; 7, maxillae; 8, hypopharynx; 9, labium." figureDoi="http://doi.org/10.5281/zenodo.5965293" httpUri="https://zenodo.org/record/5965293/files/figure.png" pageId="2" pageNumber="323">Fig. 8</figureCitation>
) with hair-like setae on anterior margin; labium (
<figureCitation id="131D4AA2FFF276451F9DA984FE59FF3E" box="[404,471,223,249]" captionStart="FIGURES 4–9" captionStartId="3.[151,264,1367,1391]" captionTargetBox="[151,1436,181,1344]" captionTargetId="figure-17@3.[151,1436,181,1344]" captionTargetPageId="3" captionText="FIGURES 4–9. Procloeon (P.) kottagudiensis sp. nov., mature nymph. Dorsal view: 4, labrum; 5, right mandible; 6, left mandible; 7, maxillae; 8, hypopharynx; 9, labium." figureDoi="http://doi.org/10.5281/zenodo.5965293" httpUri="https://zenodo.org/record/5965293/files/figure.png" pageId="2" pageNumber="323">Fig. 9</figureCitation>
) with V-shaped glossae slightly narrower than paraglossae, with row of stout marginal setae; paraglossae with row of stout marginal setae and few stout setae on dorsal surface; labial palp 3-segmented; segment III truncate, with sparse stout setae and hair-like setae on dorsal surface.
</paragraph>
<caption id="DF5906AFFFF276451E9EAC26FAC8FA52" ID-DOI="http://doi.org/10.5281/zenodo.5965291" ID-Zenodo-Dep="5965291" box="[151,1350,1405,1429]" httpUri="https://zenodo.org/record/5965291/files/figure.png" pageId="2" pageNumber="323" startId="2.[151,264,1405,1429]" targetBox="[151,1436,345,1381]" targetPageId="2">
<paragraph id="8B995627FFF276451E9EAC26FAC8FA52" blockId="2.[151,1350,1405,1429]" box="[151,1350,1405,1429]" pageId="2" pageNumber="323">
<emphasis id="B9528A35FFF276451E9EAC26FEB4FA52" bold="true" box="[151,314,1405,1429]" pageId="2" pageNumber="323">FIGURES 1–3.</emphasis>
<taxonomicName id="4C262DA4FFF276451F48AC26FDE5FA52" authority="Muthukatturaja &amp; Balasubramanian, 2022" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2022" box="[321,619,1405,1429]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="323" phylum="Arthropoda" rank="species" species="kottagudiensis" status="sp. nov." subGenus="Procloeon">
<emphasis id="B9528A35FFF276451F48AC26FE24FA52" box="[321,426,1405,1429]" italics="true" pageId="2" pageNumber="323">Procloeon</emphasis>
(
<emphasis id="B9528A35FFF276451FB1AC25FE42FA52" box="[440,460,1406,1429]" italics="true" pageId="2" pageNumber="323">P.</emphasis>
) 
<emphasis id="B9528A35FFF276451FD0AC26FDE5FA52" box="[473,619,1405,1429]" italics="true" pageId="2" pageNumber="323">kottagudiensis</emphasis>
</taxonomicName>
<emphasis id="B9528A35FFF276451C7BAC26FD47FA52" bold="true" box="[626,713,1405,1429]" pageId="2" pageNumber="323">
<taxonomicNameLabel id="A261374EFFF276451C7BAC26FD4BFA52" box="[626,709,1405,1429]" pageId="2" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
mature nymph. 1, dorsal view; 2, ventral view; 3, lateral view.
</paragraph>
</caption>
<paragraph id="8B995627FFF276451ECEAC98FAC5F9AA" blockId="2.[151,1437,1475,2042]" pageId="2" pageNumber="323">
Thorax: terga yellowish-brown; sterna white with dark brown markings near the base of legs. 
<emphasis id="B9528A35FFF276451AFDAC9FFAA7FA1A" box="[1268,1321,1476,1501]" italics="true" pageId="2" pageNumber="323">Legs</emphasis>
(
<figureCitation id="131D4AA2FFF276451B33AC98FA01FA1A" box="[1338,1423,1475,1501]" captionStart="FIGURES 10–15" captionStartId="4.[151,264,1477,1501]" captionTargetBox="[152,1436,182,1453]" captionTargetId="figure-20@4.[151,1436,181,1453]" captionTargetPageId="4" captionText="FIGURES 10–15. Procloeon (P.) kottagudiensis sp. nov., mature nymph. Dorsal view: 10, foreleg; 11, tarsal claw; 12, gill I; 13, gill IV; 14, gill VII; 15, paraproct." figureDoi="http://doi.org/10.5281/zenodo.5965297" httpUri="https://zenodo.org/record/5965297/files/figure.png" pageId="2" pageNumber="323">Fig. 10</figureCitation>
): light yellow; femur with 2 rows of stout monopectinate setae on inner margin and without stout apical setae; outer margin of femur with few hair-like setae; tibiae light brown, with 2 rows of stout setae on inner margin; tarsi light brown, with scattered hair-like setae dorsally and 2 rows of stout setae on inner margin; claws (
<figureCitation id="131D4AA2FFF276451ABFAF74FA8BF98E" box="[1206,1285,1583,1609]" captionStart="FIGURES 10–15" captionStartId="4.[151,264,1477,1501]" captionTargetBox="[152,1436,182,1453]" captionTargetId="figure-20@4.[151,1436,181,1453]" captionTargetPageId="4" captionText="FIGURES 10–15. Procloeon (P.) kottagudiensis sp. nov., mature nymph. Dorsal view: 10, foreleg; 11, tarsal claw; 12, gill I; 13, gill IV; 14, gill VII; 15, paraproct." figureDoi="http://doi.org/10.5281/zenodo.5965297" httpUri="https://zenodo.org/record/5965297/files/figure.png" pageId="2" pageNumber="323">Fig. 11</figureCitation>
) light brown, relatively long, ca. 0.37×length of tarsi, and with two rows of minute denticles progressively larger distally.
</paragraph>
<paragraph id="8B995627FFF276451ECEAF2CFB3DF83E" blockId="2.[151,1437,1475,2042]" pageId="2" pageNumber="323">
Abdomen: tergum I (
<figureCitation id="131D4AA2FFF276451FB2AF2CFD81F956" box="[443,527,1655,1681]" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure-17@5.[151,1436,181,1492]" captionTargetPageId="5" captionText="FIGURES 16–23. Procloeon (P.) kottagudiensis sp. nov., mature nymph: 16, abdomen (dorsal view); 17, abdomen (ventral view); 18, abdominal segments VIII &amp; IX (dorsal view); 19, abdominal segment X (dorsal view); 20, abdominal tergum I; 21, abdominal tergum V; 22, paracercus, 23, cercus." figureDoi="http://doi.org/10.5281/zenodo.5965299" httpUri="https://zenodo.org/record/5965299/files/figure.png" pageId="2" pageNumber="323">Fig. 20</figureCitation>
) light yellow with irregular brown markings; terga II, III and V (
<figureCitation id="131D4AA2FFF276451B0CAF2CFAD7F956" box="[1285,1369,1655,1681]" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure-17@5.[151,1436,181,1492]" captionTargetPageId="5" captionText="FIGURES 16–23. Procloeon (P.) kottagudiensis sp. nov., mature nymph: 16, abdomen (dorsal view); 17, abdomen (ventral view); 18, abdominal segments VIII &amp; IX (dorsal view); 19, abdominal segment X (dorsal view); 20, abdominal tergum I; 21, abdominal tergum V; 22, paracercus, 23, cercus." figureDoi="http://doi.org/10.5281/zenodo.5965299" httpUri="https://zenodo.org/record/5965299/files/figure.png" pageId="2" pageNumber="323">Fig. 21</figureCitation>
) with single triangular dark brown spot medially and irregular brown markings laterally; terga IV, VII and VIII somewhat pale; tergum VI pale brown to dark brown, with single paired distinct oblique streaks; terga (
<figureCitation id="131D4AA2FFF276451ABCAFE4FA89F91D" box="[1205,1287,1727,1754]" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure-17@5.[151,1436,181,1492]" captionTargetPageId="5" captionText="FIGURES 16–23. Procloeon (P.) kottagudiensis sp. nov., mature nymph: 16, abdomen (dorsal view); 17, abdomen (ventral view); 18, abdominal segments VIII &amp; IX (dorsal view); 19, abdominal segment X (dorsal view); 20, abdominal tergum I; 21, abdominal tergum V; 22, paracercus, 23, cercus." figureDoi="http://doi.org/10.5281/zenodo.5965299" httpUri="https://zenodo.org/record/5965299/files/figure.png" pageId="2" pageNumber="323">Fig. 16</figureCitation>
) with row of alternatively arranged long and small spines on posterior margin; tergum I with acute teeth on posterior margin (
<figureCitation id="131D4AA2FFF276451B66AFB8FF39F8E6" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure-17@5.[151,1436,181,1492]" captionTargetPageId="5" captionText="FIGURES 16–23. Procloeon (P.) kottagudiensis sp. nov., mature nymph: 16, abdomen (dorsal view); 17, abdomen (ventral view); 18, abdominal segments VIII &amp; IX (dorsal view); 19, abdominal segment X (dorsal view); 20, abdominal tergum I; 21, abdominal tergum V; 22, paracercus, 23, cercus." figureDoi="http://doi.org/10.5281/zenodo.5965299" httpUri="https://zenodo.org/record/5965299/files/figure.png" pageId="2" pageNumber="323">Fig. 20</figureCitation>
); posterior margin of tergum X straight (
<figureCitation id="131D4AA2FFF276451C73AE5CFD42F8E6" box="[634,716,1799,1825]" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure-17@5.[151,1436,181,1492]" captionTargetPageId="5" captionText="FIGURES 16–23. Procloeon (P.) kottagudiensis sp. nov., mature nymph: 16, abdomen (dorsal view); 17, abdomen (ventral view); 18, abdominal segments VIII &amp; IX (dorsal view); 19, abdominal segment X (dorsal view); 20, abdominal tergum I; 21, abdominal tergum V; 22, paracercus, 23, cercus." figureDoi="http://doi.org/10.5281/zenodo.5965299" httpUri="https://zenodo.org/record/5965299/files/figure.png" pageId="2" pageNumber="323">Fig. 19</figureCitation>
), with row of 11–13 acute spines; terga VIII and IX with ca. four and eight lateral spines respectively (
<figureCitation id="131D4AA2FFF276451C33AE70FD00F882" box="[570,654,1835,1861]" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure-17@5.[151,1436,181,1492]" captionTargetPageId="5" captionText="FIGURES 16–23. Procloeon (P.) kottagudiensis sp. nov., mature nymph: 16, abdomen (dorsal view); 17, abdomen (ventral view); 18, abdominal segments VIII &amp; IX (dorsal view); 19, abdominal segment X (dorsal view); 20, abdominal tergum I; 21, abdominal tergum V; 22, paracercus, 23, cercus." figureDoi="http://doi.org/10.5281/zenodo.5965299" httpUri="https://zenodo.org/record/5965299/files/figure.png" pageId="2" pageNumber="323">Fig. 18</figureCitation>
); sterna white with brown stripes laterally (
<figureCitation id="131D4AA2FFF276451A70AE70FB43F882" box="[1145,1229,1835,1861]" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure-17@5.[151,1436,181,1492]" captionTargetPageId="5" captionText="FIGURES 16–23. Procloeon (P.) kottagudiensis sp. nov., mature nymph: 16, abdomen (dorsal view); 17, abdomen (ventral view); 18, abdominal segments VIII &amp; IX (dorsal view); 19, abdominal segment X (dorsal view); 20, abdominal tergum I; 21, abdominal tergum V; 22, paracercus, 23, cercus." figureDoi="http://doi.org/10.5281/zenodo.5965299" httpUri="https://zenodo.org/record/5965299/files/figure.png" pageId="2" pageNumber="323">Fig. 17</figureCitation>
); sternum IX with dark brown stripe along lateral and anterior margins; paraprocts (
<figureCitation id="131D4AA2FFF276451D67AE14FC31F8AD" box="[878,959,1871,1898]" captionStart="FIGURES 10–15" captionStartId="4.[151,264,1477,1501]" captionTargetBox="[152,1436,182,1453]" captionTargetId="figure-20@4.[151,1436,181,1453]" captionTargetPageId="4" captionText="FIGURES 10–15. Procloeon (P.) kottagudiensis sp. nov., mature nymph. Dorsal view: 10, foreleg; 11, tarsal claw; 12, gill I; 13, gill IV; 14, gill VII; 15, paraproct." figureDoi="http://doi.org/10.5281/zenodo.5965297" httpUri="https://zenodo.org/record/5965297/files/figure.png" pageId="2" pageNumber="323">Fig. 15</figureCitation>
) with 7–9 acute spines on distal half; 
<emphasis id="B9528A35FFF276451B6FAE14FA12F8AE" box="[1382,1436,1871,1897]" italics="true" pageId="2" pageNumber="323">Gills</emphasis>
(
<figureCitation id="131D4AA2FFF276451E96AE28FE91F84A" box="[159,287,1907,1933]" captionStart="FIGURES 10–15" captionStartId="4.[151,264,1477,1501]" captionTargetBox="[152,1436,182,1453]" captionTargetId="figure-20@4.[151,1436,181,1453]" captionTargetPageId="4" captionText="FIGURES 10–15. Procloeon (P.) kottagudiensis sp. nov., mature nymph. Dorsal view: 10, foreleg; 11, tarsal claw; 12, gill I; 13, gill IV; 14, gill VII; 15, paraproct." figureDoi="http://doi.org/10.5281/zenodo.5965297" httpUri="https://zenodo.org/record/5965297/files/figure.png" pageId="2" pageNumber="323">Figs 12–14</figureCitation>
) pale brown with well-developed dark brown tracheae; gills I–VI bilamellate; dorsal lamella small; gill VII unilamellate. 
<emphasis id="B9528A35FFF276451F52AECCFD94F876" box="[347,538,1943,1969]" italics="true" pageId="2" pageNumber="323">Caudal filaments</emphasis>
: ca. 0.35×length of the body; ca. 1/2 basal part light yellow to brown and remaining distal part with dark brown bands; segments 4–6 narrow; paracercus with a row of long hair-like setae on inner and outer margins (
<figureCitation id="131D4AA2FFF276451F79AE84FE4BF83E" box="[368,453,2015,2041]" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure-17@5.[151,1436,181,1492]" captionTargetPageId="5" captionText="FIGURES 16–23. Procloeon (P.) kottagudiensis sp. nov., mature nymph: 16, abdomen (dorsal view); 17, abdomen (ventral view); 18, abdominal segments VIII &amp; IX (dorsal view); 19, abdominal segment X (dorsal view); 20, abdominal tergum I; 21, abdominal tergum V; 22, paracercus, 23, cercus." figureDoi="http://doi.org/10.5281/zenodo.5965299" httpUri="https://zenodo.org/record/5965299/files/figure.png" pageId="2" pageNumber="323">Fig. 22</figureCitation>
); cerci with a row of long hair-like setae on inner margin (
<figureCitation id="131D4AA2FFF276451A5BAE84FB28F83E" box="[1106,1190,2015,2041]" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1516,1540]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure-17@5.[151,1436,181,1492]" captionTargetPageId="5" captionText="FIGURES 16–23. Procloeon (P.) kottagudiensis sp. nov., mature nymph: 16, abdomen (dorsal view); 17, abdomen (ventral view); 18, abdominal segments VIII &amp; IX (dorsal view); 19, abdominal segment X (dorsal view); 20, abdominal tergum I; 21, abdominal tergum V; 22, paracercus, 23, cercus." figureDoi="http://doi.org/10.5281/zenodo.5965299" httpUri="https://zenodo.org/record/5965299/files/figure.png" pageId="2" pageNumber="323">Fig. 23</figureCitation>
).
</paragraph>
<caption id="DF5906AFFFF376441E9EAC0CFDE7FA53" ID-DOI="http://doi.org/10.5281/zenodo.5965293" ID-Zenodo-Dep="5965293" httpUri="https://zenodo.org/record/5965293/files/figure.png" pageId="3" pageNumber="324" startId="3.[151,264,1367,1391]" targetBox="[151,1436,181,1344]" targetPageId="3">
<paragraph id="8B995627FFF376441E9EAC0CFDE7FA53" blockId="3.[151,1436,1367,1428]" pageId="3" pageNumber="324">
<emphasis id="B9528A35FFF376441E9EAC0CFEB5FAA8" bold="true" box="[151,315,1367,1391]" pageId="3" pageNumber="324">FIGURES 4–9.</emphasis>
<taxonomicName id="4C262DA4FFF376441F48AC03FDE3FAB7" authority="Muthukatturaja &amp; Balasubramanian, 2022" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2022" box="[321,621,1368,1392]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="324" phylum="Arthropoda" rank="species" species="kottagudiensis" status="sp. nov." subGenus="Procloeon">
<emphasis id="B9528A35FFF376441F48AC03FE24FAB7" box="[321,426,1368,1392]" italics="true" pageId="3" pageNumber="324">Procloeon</emphasis>
(
<emphasis id="B9528A35FFF376441FB0AC03FE43FAA8" box="[441,461,1368,1391]" italics="true" pageId="3" pageNumber="324">P.</emphasis>
) 
<emphasis id="B9528A35FFF376441FD2AC03FDE3FAB7" box="[475,621,1368,1392]" italics="true" pageId="3" pageNumber="324">kottagudiensis</emphasis>
</taxonomicName>
<emphasis id="B9528A35FFF376441C7DAC03FD45FAB7" bold="true" box="[628,715,1368,1392]" pageId="3" pageNumber="324">
<taxonomicNameLabel id="A261374EFFF376441C7DAC03FD49FAB7" box="[628,711,1368,1392]" pageId="3" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
mature nymph. Dorsal view: 4, labrum; 5, right mandible; 6, left mandible; 7, maxillae; 8, hypopharynx; 9, labium.
</paragraph>
</caption>
<paragraph id="8B995627FFF376441ECEAC99FF49F957" blockId="3.[151,1437,1474,2004]" pageId="3" pageNumber="324">
<emphasis id="B9528A35FFF376441ECEAC99FED6FA1B" bold="true" box="[199,344,1474,1500]" pageId="3" pageNumber="324">Male Imago</emphasis>
(
<figureCitation id="131D4AA2FFF376441F60AC99FE31FA1B" box="[361,447,1474,1500]" captionStart="FIGURES 24–29" captionStartId="6.[151,264,955,979]" captionTargetBox="[165,1423,181,931]" captionTargetId="figure-20@6.[165,1423,181,931]" captionTargetPageId="6" captionText="FIGURES 24–29. Procloeon (P.) kottagudiensis sp. nov., imago: 24, male imago (dorsal view); 25, female imago (dorsal view); 26, abdomen (dorsal view); 27, abdomen (ventral view); 28, abdomen (lateral view); 29, thoracic sternum." figureDoi="http://doi.org/10.5281/zenodo.5965303" httpUri="https://zenodo.org/record/5965303/files/figure.png" pageId="3" pageNumber="324">Fig. 24</figureCitation>
): Body length (excluding cerci) 
<quantity id="4CDEFBC2FFF376441D26AC99FC34FA1B" box="[815,954,1474,1501]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.3" metricValueMin="5.7" pageId="3" pageNumber="324" unit="mm" value="6.0" valueMax="6.3" valueMin="5.7">5.7–6.3 mm</quantity>
; cerci 
<quantity id="4CDEFBC2FFF376441A0FAC99FB22FA1B" box="[1030,1196,1474,1500]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" metricValueMax="1.21" metricValueMin="1.17" pageId="3" pageNumber="324" unit="mm" value="11.899999999999999" valueMax="12.1" valueMin="11.7">11.7–12.1 mm</quantity>
; head width 
<quantity id="4CDEFBC2FFF376441B37AC99FA16FA1B" box="[1342,1432,1474,1500]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="324" unit="mm" value="0.8">0.8 mm</quantity>
; head length 
<quantity id="4CDEFBC2FFF376441F16ACBDFEF9F9C7" box="[287,375,1510,1537]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="324" unit="mm" value="0.5">0.5 mm</quantity>
; antenna 1ength 
<quantity id="4CDEFBC2FFF376441C3BACBDFD04F9C7" box="[562,650,1510,1536]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="324" unit="mm" value="0.7">0.7 mm</quantity>
; compound eye length 
<quantity id="4CDEFBC2FFF376441D82ACBDFC6DF9C7" box="[907,995,1510,1536]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="324" unit="mm" value="0.4">0.4 mm</quantity>
; compound eye width 
<quantity id="4CDEFBC2FFF376441AD7ACBDFAB8F9C7" box="[1246,1334,1510,1536]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="324" unit="mm" value="0.4">0.4 mm</quantity>
; distance between compound eye 0.0 mm; length of fore femur/ tibia/ tarsi 1.2/ 1.5/ 
<quantity id="4CDEFBC2FFF376441DEFAF51FBCEF9E3" box="[998,1088,1546,1572]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="324" unit="mm" value="1.7">1.7 mm</quantity>
; length of middle femur/ tibia/ tarsi 1.1/ 0.8/ 
<quantity id="4CDEFBC2FFF376441F3AAF75FE02F98F" box="[307,396,1582,1608]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="324" unit="mm" value="0.8">0.8 mm</quantity>
; length of hind femur/ tibia/ tarsi 1.1/ 0.8/ 
<quantity id="4CDEFBC2FFF376441D65AF75FC4BF98F" box="[876,965,1582,1608]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="324" unit="mm" value="0.8">0.8 mm</quantity>
; forewing length 
<quantity id="4CDEFBC2FFF376441A82AF75FB6AF98F" box="[1163,1252,1582,1609]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="324" unit="mm" value="5.3">5.3 mm</quantity>
; forewing width 
<quantity id="4CDEFBC2FFF376441E9EAF09FF7EF9AB" box="[151,240,1618,1644]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="324" unit="mm" value="2.1">2.1 mm</quantity>
; hindwing length 
<quantity id="4CDEFBC2FFF376441FB0AF09FD9CF9AB" box="[441,530,1618,1644]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="324" unit="mm" value="0.8">0.8 mm</quantity>
; hindwing width 
<quantity id="4CDEFBC2FFF376441CDDAF09FCB5F9AB" box="[724,827,1618,1645]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="324" unit="mm" value="0.15">0.15 mm</quantity>
. General body coloration yellowish brown to pale yellow.
</paragraph>
<paragraph id="8B995627FFF376441ECEAFC1FACCF91F" blockId="3.[151,1437,1474,2004]" pageId="3" pageNumber="324">Head: vertex pale yellow with paired dark brown stripes medially; compound eyes turbinate; dorsal eyes pale brown; ventral eyes pale grey; ocelli pale yellow; scape and pedicel of antennae paler; flagellum pale grey.</paragraph>
<paragraph id="8B995627FFF376441ECEAFB9FDA2F813" blockId="3.[151,1437,1474,2004]" pageId="3" pageNumber="324">
Thorax: prothorax pale brown; mesothorax pale yellow, with single paired chestnut brown vertical stripes medially; sterna pale yellow (
<figureCitation id="131D4AA2FFF376441FA4AE5DFD8CF8E7" box="[429,514,1798,1824]" captionStart="FIGURES 24–29" captionStartId="6.[151,264,955,979]" captionTargetBox="[165,1423,181,931]" captionTargetId="figure-20@6.[165,1423,181,931]" captionTargetPageId="6" captionText="FIGURES 24–29. Procloeon (P.) kottagudiensis sp. nov., imago: 24, male imago (dorsal view); 25, female imago (dorsal view); 26, abdomen (dorsal view); 27, abdomen (ventral view); 28, abdomen (lateral view); 29, thoracic sternum." figureDoi="http://doi.org/10.5281/zenodo.5965303" httpUri="https://zenodo.org/record/5965303/files/figure.png" pageId="3" pageNumber="324">Fig. 29</figureCitation>
). 
<emphasis id="B9528A35FFF376441C1CAE5CFDC4F8E7" box="[533,586,1799,1824]" italics="true" pageId="3" pageNumber="324">Legs</emphasis>
(
<figureCitation id="131D4AA2FFF376441C53AE5DFD68F8E6" box="[602,742,1798,1825]" captionStart="FIGURES 30–36" captionStartId="6.[151,264,1971,1995]" captionTargetBox="[187,1394,1044,1938]" captionTargetId="figure-58@6.[180,1406,1040,1948]" captionTargetPageId="6" captionText="FIGURES 30–36. Procloeon (P.) kottagudiensis sp. nov., male imago: 30, forewing; 31, hindwing; 32, hindwing enlarged; 33, male genitalia; 34, foreleg; 35, midleg; 36, hindleg." figureDoi="http://doi.org/10.5281/zenodo.5965305" httpUri="https://zenodo.org/record/5965305/files/figure.png" pageId="3" pageNumber="324">Figs. 36–36</figureCitation>
): femur pale yellow, tibia and tarsi pale yellow to translucent; claws brown basally. 
<emphasis id="B9528A35FFF376441F87AE70FE55F883" box="[398,475,1835,1860]" italics="true" pageId="3" pageNumber="324">Wings:</emphasis>
forewings (
<figureCitation id="131D4AA2FFF376441C6CAE71FD37F883" box="[613,697,1834,1860]" captionStart="FIGURES 30–36" captionStartId="6.[151,264,1971,1995]" captionTargetBox="[187,1394,1044,1938]" captionTargetId="figure-58@6.[180,1406,1040,1948]" captionTargetPageId="6" captionText="FIGURES 30–36. Procloeon (P.) kottagudiensis sp. nov., male imago: 30, forewing; 31, hindwing; 32, hindwing enlarged; 33, male genitalia; 34, foreleg; 35, midleg; 36, hindleg." figureDoi="http://doi.org/10.5281/zenodo.5965305" httpUri="https://zenodo.org/record/5965305/files/figure.png" pageId="3" pageNumber="324">Fig. 30</figureCitation>
) transparent, membrane hyaline and veins darker; stigmatic area milky transparent with 8 cross-veins; longitudinal veins with single crossvein except areas of R and MA with two cross-veins, Sc with three cross-veins; hindwings (
<figureCitation id="131D4AA2FFF376441CD7AE29FC0CF84B" box="[734,898,1906,1932]" captionStart="FIGURES 30–36" captionStartId="6.[151,264,1971,1995]" captionTargetBox="[187,1394,1044,1938]" captionTargetId="figure-58@6.[180,1406,1040,1948]" captionTargetPageId="6" captionText="FIGURES 30–36. Procloeon (P.) kottagudiensis sp. nov., male imago: 30, forewing; 31, hindwing; 32, hindwing enlarged; 33, male genitalia; 34, foreleg; 35, midleg; 36, hindleg." figureDoi="http://doi.org/10.5281/zenodo.5965305" httpUri="https://zenodo.org/record/5965305/files/figure.png" pageId="3" pageNumber="324">Figs. 31 &amp; 32</figureCitation>
) narrow and slender, approximately five times longer than wide, distal margin rounded, and with two longitudinal veins; apex of costal projection acute, located at 1/2 of distance from base to margin.
</paragraph>
<caption id="DF5906AFFFF476431E9EAC9EFD99F9C6" ID-DOI="http://doi.org/10.5281/zenodo.5965297" ID-Zenodo-Dep="5965297" httpUri="https://zenodo.org/record/5965297/files/figure.png" pageId="4" pageNumber="325" startId="4.[151,264,1477,1501]" targetBox="[152,1436,182,1453]" targetPageId="4">
<paragraph id="8B995627FFF476431E9EAC9EFD99F9C6" blockId="4.[151,1436,1477,1538]" pageId="4" pageNumber="325">
<emphasis id="B9528A35FFF476431E9EAC9EFEDBFA19" bold="true" box="[151,341,1477,1502]" pageId="4" pageNumber="325">FIGURES 10–15.</emphasis>
<taxonomicName id="4C262DA4FFF476431F55AC9DFD07FA19" authority="Muthukatturaja &amp; Balasubramanian, 2022" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2022" box="[348,649,1478,1502]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="325" phylum="Arthropoda" rank="species" species="kottagudiensis" status="sp. nov." subGenus="Procloeon">
<emphasis id="B9528A35FFF476431F55AC9DFE4BFA19" box="[348,453,1478,1502]" italics="true" pageId="4" pageNumber="325">Procloeon</emphasis>
(
<emphasis id="B9528A35FFF476431FDCAC9CFE66FA19" box="[469,488,1479,1502]" italics="true" pageId="4" pageNumber="325">P.</emphasis>
) 
<emphasis id="B9528A35FFF476431FFEAC9DFD07FA19" box="[503,649,1478,1502]" italics="true" pageId="4" pageNumber="325">kottagudiensis</emphasis>
</taxonomicName>
<emphasis id="B9528A35FFF476431C98AC9DFD67FA19" bold="true" box="[657,745,1478,1502]" pageId="4" pageNumber="325">
<taxonomicNameLabel id="A261374EFFF476431C98AC9DFD6AFA19" box="[657,740,1478,1502]" pageId="4" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
mature nymph. Dorsal view: 10, foreleg; 11, tarsal claw; 12, gill I; 13, gill IV; 14, gill VII; 15, paraproct.
</paragraph>
</caption>
<paragraph id="8B995627FFF476431ECEAF6BFDA5F939" blockId="4.[151,1437,1584,2006]" pageId="4" pageNumber="325">
Abdomen (
<figureCitation id="131D4AA2FFF476431F41AF6BFE58F98C" box="[328,470,1584,1611]" captionStart="FIGURES 24–29" captionStartId="6.[151,264,955,979]" captionTargetBox="[165,1423,181,931]" captionTargetId="figure-20@6.[165,1423,181,931]" captionTargetPageId="6" captionText="FIGURES 24–29. Procloeon (P.) kottagudiensis sp. nov., imago: 24, male imago (dorsal view); 25, female imago (dorsal view); 26, abdomen (dorsal view); 27, abdomen (ventral view); 28, abdomen (lateral view); 29, thoracic sternum." figureDoi="http://doi.org/10.5281/zenodo.5965303" httpUri="https://zenodo.org/record/5965303/files/figure.png" pageId="4" pageNumber="325">Figs. 26–28</figureCitation>
): terga pale yellow to translucent; terga I–V and ½ of tergum VI translucent; terga II–X with a pale brown posterior marginal line; ½ of tergum VI and terga VII–X pale brown to pale yellow; sternal coloration as in tergum except posterior marginal line absent; pleural margins grayish in color. 
<emphasis id="B9528A35FFF476431AC8AF23FAA5F955" box="[1217,1323,1656,1682]" italics="true" pageId="4" pageNumber="325">Genitalia</emphasis>
(
<figureCitation id="131D4AA2FFF476431B32AF23FA01F955" box="[1339,1423,1656,1682]" captionStart="FIGURES 30–36" captionStartId="6.[151,264,1971,1995]" captionTargetBox="[187,1394,1044,1938]" captionTargetId="figure-58@6.[180,1406,1040,1948]" captionTargetPageId="6" captionText="FIGURES 30–36. Procloeon (P.) kottagudiensis sp. nov., male imago: 30, forewing; 31, hindwing; 32, hindwing enlarged; 33, male genitalia; 34, foreleg; 35, midleg; 36, hindleg." figureDoi="http://doi.org/10.5281/zenodo.5965305" httpUri="https://zenodo.org/record/5965305/files/figure.png" pageId="4" pageNumber="325">Fig. 33</figureCitation>
): forceps pale yellow to translucent, 3-segmented; segment II ca. 3×length of segment I, segment III ca. 0.6×length of segment I; posterior margin of penal plate broadly round with sparse thin setae on the surface. Cerci pale yellow to translucent, ca. 2×length of body.
</paragraph>
<paragraph id="8B995627FFF476431ECEAE53FE91F811" blockId="4.[151,1437,1584,2006]" pageId="4" pageNumber="325">
<emphasis id="B9528A35FFF476431ECEAE53FEFFF8E5" bold="true" box="[199,369,1800,1826]" pageId="4" pageNumber="325">Female Imago</emphasis>
(
<figureCitation id="131D4AA2FFF476431F89AE53FE5AF8E4" box="[384,468,1800,1827]" captionStart="FIGURES 24–29" captionStartId="6.[151,264,955,979]" captionTargetBox="[165,1423,181,931]" captionTargetId="figure-20@6.[165,1423,181,931]" captionTargetPageId="6" captionText="FIGURES 24–29. Procloeon (P.) kottagudiensis sp. nov., imago: 24, male imago (dorsal view); 25, female imago (dorsal view); 26, abdomen (dorsal view); 27, abdomen (ventral view); 28, abdomen (lateral view); 29, thoracic sternum." figureDoi="http://doi.org/10.5281/zenodo.5965303" httpUri="https://zenodo.org/record/5965303/files/figure.png" pageId="4" pageNumber="325">Fig. 25</figureCitation>
): Body length (excluding cerci) 5.5–6.0 mm; cerci 13.0– 
<quantity id="4CDEFBC2FFF476431A44AE53FB3DF8E4" box="[1101,1203,1800,1827]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="4" pageNumber="325" unit="mm" value="13.3">13.3 mm</quantity>
; head width 
<quantity id="4CDEFBC2FFF476431B36AE53FA16F8E4" box="[1343,1432,1800,1827]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="325" unit="mm" value="0.6">0.6 mm</quantity>
; head length 
<quantity id="4CDEFBC2FFF476431F28AE77FEF5F880" box="[289,379,1836,1863]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="325" unit="mm" value="0.4">0.4 mm</quantity>
; antennae 1ength 
<quantity id="4CDEFBC2FFF476431C4FAE77FD11F880" box="[582,671,1836,1863]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="325" unit="mm" value="0.9">0.9 mm</quantity>
; compound eye length 
<quantity id="4CDEFBC2FFF476431DACAE77FC71F880" box="[933,1023,1836,1863]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="325" unit="mm" value="0.3">0.3 mm</quantity>
; compound eye width 
<quantity id="4CDEFBC2FFF476431AF6AE77FAE8F880" box="[1279,1382,1836,1863]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="325" unit="mm" value="0.25">0.25 mm</quantity>
; distance between compound eye 
<quantity id="4CDEFBC2FFF476431FE4AE0BFDC9F8AC" box="[493,583,1872,1899]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="325" unit="mm" value="0.1">0.1 mm</quantity>
; length of fore femur/ tibia/ tarsi 1.1/ 0.8/ 
<quantity id="4CDEFBC2FFF476431A21AE0BFB0CF8AC" box="[1064,1154,1872,1899]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="325" unit="mm" value="0.8">0.8 mm</quantity>
; length of middle femur/ tibia/ tarsi 0.9/ 0.6/ 
<quantity id="4CDEFBC2FFF476431F7CAE2FFE41F848" box="[373,463,1908,1935]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="325" unit="mm" value="0.6">0.6 mm</quantity>
; length of hind femur/ tibia/ tarsi 0.9/ 0.6/ 
<quantity id="4CDEFBC2FFF476431DB8AE2FFB85F848" box="[945,1035,1908,1935]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="325" unit="mm" value="0.6">0.6 mm</quantity>
; forewing length 
<quantity id="4CDEFBC2FFF476431AD8AE2FFAA5F848" box="[1233,1323,1908,1935]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="4" pageNumber="325" unit="mm" value="5.3">5.3 mm</quantity>
; forewing width 
<quantity id="4CDEFBC2FFF476431EE9AEC3FEB5F874" box="[224,315,1944,1971]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="325" unit="mm" value="2.1">2.1 mm</quantity>
; hindwing length 
<quantity id="4CDEFBC2FFF476431C03AEC3FDEBF874" box="[522,613,1944,1971]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="325" unit="mm" value="0.8">0.8 mm</quantity>
; hindwing width 
<quantity id="4CDEFBC2FFF476431D27AEC3FC19F874" box="[814,919,1944,1971]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="325" unit="mm" value="0.15">0.15 mm</quantity>
. General body coloration yellowish brown to pale yellow.
</paragraph>
<caption id="DF5906AFFFF576421E9EACB7FD0EF98C" ID-DOI="http://doi.org/10.5281/zenodo.5965299" ID-Zenodo-Dep="5965299" httpUri="https://zenodo.org/record/5965299/files/figure.png" pageId="5" pageNumber="326" startId="5.[151,264,1516,1540]" targetBox="[151,1436,181,1491]" targetPageId="5">
<paragraph id="8B995627FFF576421E9EACB7FD0EF98C" blockId="5.[151,1436,1516,1612]" pageId="5" pageNumber="326">
<emphasis id="B9528A35FFF576421E9EACB7FED8F9C3" bold="true" box="[151,342,1516,1540]" pageId="5" pageNumber="326">FIGURES 16–23.</emphasis>
<taxonomicName id="4C262DA4FFF576421F57ACB7FD03F9C3" authority="Muthukatturaja &amp; Balasubramanian, 2022" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2022" box="[350,653,1516,1540]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="kottagudiensis" status="sp. nov." subGenus="Procloeon">
<emphasis id="B9528A35FFF576421F57ACB7FE49F9C3" box="[350,455,1516,1540]" italics="true" pageId="5" pageNumber="326">Procloeon</emphasis>
(
<emphasis id="B9528A35FFF576421FDEACB6FE65F9C3" box="[471,491,1517,1540]" italics="true" pageId="5" pageNumber="326">P.</emphasis>
) 
<emphasis id="B9528A35FFF576421FF2ACB7FD03F9C3" box="[507,653,1516,1540]" italics="true" pageId="5" pageNumber="326">kottagudiensis</emphasis>
</taxonomicName>
<emphasis id="B9528A35FFF576421C9CACB7FD61F9C3" bold="true" box="[661,751,1516,1540]" pageId="5" pageNumber="326">
<taxonomicNameLabel id="A261374EFFF576421C9CACB7FD64F9C3" box="[661,746,1516,1540]" pageId="5" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
mature nymph: 16, abdomen (dorsal view); 17, abdomen (ventral view); 18, abdominal segments VIII &amp; IX (dorsal view); 19, abdominal segment X (dorsal view); 20, abdominal tergum I; 21, abdominal tergum V; 22, paracercus, 23, cercus.
</paragraph>
</caption>
<paragraph id="8B995627FFF576421ECEAF21FC9DF97E" blockId="5.[151,1437,1658,2009]" pageId="5" pageNumber="326">Head: vertex pale yellow with paired dark brown stripes medially; compound eyes black; ocelli pale yellow; scape and pedicel of antennae paler; flagellum pale grey.</paragraph>
<paragraph id="8B995627FFF576421ECEAF99FAD2F8C7" blockId="5.[151,1437,1658,2009]" pageId="5" pageNumber="326">
Thorax: prothorax pale brown; mesothorax pale yellow, with single paired chestnut brown vertical stripes medially; sterna pale yellow. 
<emphasis id="B9528A35FFF576421FA1AFBCFE53F8C7" box="[424,477,1767,1792]" italics="true" pageId="5" pageNumber="326">Legs</emphasis>
: coloration as in male imago. 
<emphasis id="B9528A35FFF576421D3BAFBCFCF1F8C7" box="[818,895,1767,1792]" italics="true" pageId="5" pageNumber="326">Wings:</emphasis>
coloration and venation as in male imago.
</paragraph>
<paragraph id="8B995627FFF576421ECEAE51FAE1F8AB" blockId="5.[151,1437,1658,2009]" pageId="5" pageNumber="326">Abdomen: terga pale yellow to translucent; terga I–VI pale yellow; terga II–X with a pale brown posterior marginal line; tergum VII paler; terga VIII–X pale brown to pale yellow; sternal coloration as in terga except posterior marginal line absent; pleural margins grayish in color. Cerci pale yellow to translucent, ca. 2.2×length of body.</paragraph>
</subSubSection>
<subSubSection id="C33C05ACFFF576421ECEAE2DFD15F873" pageId="5" pageNumber="326" type="etymology">
<paragraph id="8B995627FFF576421ECEAE2DFD15F873" blockId="5.[151,1437,1658,2009]" pageId="5" pageNumber="326">
<emphasis id="B9528A35FFF576421ECEAE2DFEC0F857" bold="true" box="[199,334,1910,1936]" pageId="5" pageNumber="326">Etymology.</emphasis>
The new species 
<taxonomicName id="4C262DA4FFF576421C14AE2CFC83F857" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2022" box="[541,781,1910,1936]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="326" phylum="Arthropoda" rank="species" species="kottagudiensis" subGenus="Procloeon">
<emphasis id="B9528A35FFF576421C14AE2CFC83F857" box="[541,781,1910,1936]" italics="true" pageId="5" pageNumber="326">P. (P.) kottagudiensis</emphasis>
</taxonomicName>
is named after the 
<typeStatus id="549DE885FFF576421DE5AE2CFB92F856" box="[1004,1052,1911,1937]" pageId="5" pageNumber="326">type</typeStatus>
locality, Kottagudi stream, Theni District, where the new species was collected.
</paragraph>
</subSubSection>
<subSubSection id="C33C05ACFFF576401ECEAEE5FD2DFE86" lastPageId="7" lastPageNumber="328" pageId="5" pageNumber="326" type="distribution">
<paragraph id="8B995627FFF576421ECEAEE5FCAFF81F" blockId="5.[151,1437,1658,2009]" box="[199,801,1982,2009]" pageId="5" pageNumber="326">
<emphasis id="B9528A35FFF576421ECEAEE5FEEFF81F" bold="true" box="[199,353,1982,2008]" pageId="5" pageNumber="326">Distribution.</emphasis>
Cardamom Hills, Western Ghats, 
<collectingCountry id="F33116B7FFF576421CE9AEE5FC93F81F" box="[736,797,1982,2008]" name="India" pageId="5" pageNumber="326">India</collectingCountry>
.
</paragraph>
<caption id="DF5906AFFFF676411E9EAAE0FB41FC30" ID-DOI="http://doi.org/10.5281/zenodo.5965303" ID-Zenodo-Dep="5965303" httpUri="https://zenodo.org/record/5965303/files/figure.png" pageId="6" pageNumber="327" startId="6.[151,264,955,979]" targetBox="[165,1423,181,931]" targetPageId="6">
<paragraph id="8B995627FFF676411E9EAAE0FB41FC30" blockId="6.[151,1436,955,1016]" pageId="6" pageNumber="327">
<emphasis id="B9528A35FFF676411E9EAAE0FEDDFC14" bold="true" box="[151,339,955,979]" pageId="6" pageNumber="327">FIGURES 24–29.</emphasis>
<taxonomicName id="4C262DA4FFF676411F51AAE0FD0EFC14" authority="Muthukatturaja &amp; Balasubramanian, 2022" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2022" box="[344,640,955,980]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="kottagudiensis" status="sp. nov." subGenus="Procloeon">
<emphasis id="B9528A35FFF676411F51AAE0FE4FFC14" box="[344,449,955,979]" italics="true" pageId="6" pageNumber="327">Procloeon</emphasis>
(
<emphasis id="B9528A35FFF676411FC7AAE7FE6FFC14" box="[462,481,956,979]" italics="true" pageId="6" pageNumber="327">P.</emphasis>
) 
<emphasis id="B9528A35FFF676411FE7AAE0FD0EFC14" box="[494,640,955,979]" italics="true" pageId="6" pageNumber="327">kottagudiensis</emphasis>
</taxonomicName>
<emphasis id="B9528A35FFF676411C8CAAE7FD55FC13" bold="true" box="[645,731,956,980]" pageId="6" pageNumber="327">
<taxonomicNameLabel id="A261374EFFF676411C8CAAE7FD59FC13" box="[645,727,956,980]" pageId="6" pageNumber="327" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
imago: 24, male imago (dorsal view); 25, female imago (dorsal view); 26, abdomen (dorsal view); 27, abdomen (ventral view); 28, abdomen (lateral view); 29, thoracic sternum.
</paragraph>
</caption>
<caption id="DF5906AFFFF676411E9EAEE8FD2CF828" ID-DOI="http://doi.org/10.5281/zenodo.5965305" ID-Zenodo-Dep="5965305" httpUri="https://zenodo.org/record/5965305/files/figure.png" pageId="6" pageNumber="327" startId="6.[151,264,1971,1995]" targetBox="[187,1394,1044,1938]" targetPageId="6">
<paragraph id="8B995627FFF676411E9EAEE8FD2CF828" blockId="6.[151,1436,1971,2031]" pageId="6" pageNumber="327">
<emphasis id="B9528A35FFF676411E9EAEE8FEDAF80C" bold="true" box="[151,340,1971,1995]" pageId="6" pageNumber="327">FIGURES 30–36.</emphasis>
<taxonomicName id="4C262DA4FFF676411F53AEE8FD0AF80C" authority="Muthukatturaja &amp; Balasubramanian, 2022" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2022" box="[346,644,1971,1995]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="327" phylum="Arthropoda" rank="species" species="kottagudiensis" status="sp. nov." subGenus="Procloeon">
<emphasis id="B9528A35FFF676411F53AEE8FE4DF80C" box="[346,451,1971,1995]" italics="true" pageId="6" pageNumber="327">Procloeon</emphasis>
(
<emphasis id="B9528A35FFF676411FD8AEEFFE6BF80C" box="[465,485,1972,1995]" italics="true" pageId="6" pageNumber="327">P.</emphasis>
) 
<emphasis id="B9528A35FFF676411FFBAEE8FD0AF80C" box="[498,644,1971,1995]" italics="true" pageId="6" pageNumber="327">kottagudiensis</emphasis>
</taxonomicName>
<emphasis id="B9528A35FFF676411C82AEE8FD6CF80C" bold="true" box="[651,738,1971,1995]" pageId="6" pageNumber="327">
<taxonomicNameLabel id="A261374EFFF676411C82AEE8FD53F80C" box="[651,733,1971,1995]" pageId="6" pageNumber="327" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male imago: 30, forewing; 31, hindwing; 32, hindwing enlarged; 33, male genitalia; 34, foreleg; 35, midleg; 36, hindleg.
</paragraph>
</caption>
<paragraph id="8B995627FFF776401ECEA9CDFD2DFE86" blockId="7.[151,1436,150,321]" pageId="7" pageNumber="328">
<emphasis id="B9528A35FFF776401ECEA9CDFEA4FF77" bold="true" box="[199,298,150,176]" pageId="7" pageNumber="328">Habitat.</emphasis>
The larvae of 
<taxonomicName id="4C262DA4FFF776401FC5A9CCFC98FF76" authority="Muthukatturaja &amp; Balasubramanian, 2022" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2022" box="[460,790,151,177]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="kottagudiensis" status="sp. nov." subGenus="Procloeon">
<emphasis id="B9528A35FFF776401FC5A9CCFDCEFF76" box="[460,576,151,177]" italics="true" pageId="7" pageNumber="328">Procloeon</emphasis>
(
<emphasis id="B9528A35FFF776401C46A9CCFDEBFF77" box="[591,613,151,176]" italics="true" pageId="7" pageNumber="328">P.</emphasis>
) 
<emphasis id="B9528A35FFF776401C7DA9CCFC98FF76" box="[628,790,151,177]" italics="true" pageId="7" pageNumber="328">kottagudiensis</emphasis>
</taxonomicName>
<emphasis id="B9528A35FFF776401D15A9CCFCFBFF76" bold="true" box="[796,885,151,177]" pageId="7" pageNumber="328">
<taxonomicNameLabel id="A261374EFFF776401D15A9CCFCFBFF76" box="[796,885,151,177]" pageId="7" pageNumber="328" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were collected from the pool areas of slow-flowing permanent streams with abundant fallen leaves, marginal roots and macrophytes. Water and air temperatures of the collection site were 23.6 and 26.5ºC respectively (at the time of sampling). The current velocity was approximately 
<quantity id="4CDEFBC2FFF776401EE4A858FEEDFEDA" box="[237,355,259,285]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="7" pageNumber="328" unit="m" value="0.2" valueMax="0.3" valueMin="0.1">0.1–0.3 m</quantity>
/s. Water depth ranged 
<quantity id="4CDEFBC2FFF776401C65A859FD6FFEDA" box="[620,737,258,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="7" pageNumber="328" unit="cm" value="17.5" valueMax="25.0" valueMin="10.0">10–25 cm</quantity>
. The substratum was composed of mainly boulders, cobbles, pebbles and a significant level of sand and silt.
</paragraph>
</subSubSection>
</treatment>
</document>