treatments-xml/data/03/89/B7/0389B761B71CAD36FF2EC4AA1F1D3084.xml
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<document id="C605ED86592D116A37522F22EA03FF3B" ID-CLB-Dataset="45797" ID-DOI="10.5281/zenodo.206783" ID-GBIF-Dataset="3320067b-952f-49e0-b68c-7be870bcd2f6" ID-ISSN="1175-5326" ID-Zenodo-Dep="206783" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460419069571" checkinUser="plazi" docAuthor="Marques, Antonio C., Peña, Álvaro L., Miranda, Thaís P. &amp; Migotto, Alvaro E." docDate="2011" docId="0389B761B71CAD36FF2EC4AA1F1D3084" docLanguage="en" docName="zt03129p028.pdf" docOrigin="Zootaxa 3129" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Filellum nitidum Watson 2005" docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="FFB0CF19B712AD38FFB9C43D1B1F341D" masterDocTitle="Revision of the genus Filellum Hincks, 1868 (Lafoeidae, Leptothecata, Hydrozoa)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="15" updateTime="1698329484829" updateUser="ExternalLinkService">
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<mods:title id="C331AF5ECB5243837E08730927BCA80F">Revision of the genus Filellum Hincks, 1868 (Lafoeidae, Leptothecata, Hydrozoa)</mods:title>
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<mods:namePart id="A89D93F9349BD8CCFF2CF43E3C9A535C">Miranda, Thaís P.</mods:namePart>
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<mods:namePart id="1E5F28739DFAD3C9500B1E162F054CA7">Migotto, Alvaro E.</mods:namePart>
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<paragraph id="8B9F0677B71CAD36FF2EC4AA190B34AC" blockId="14.[151,532,151,177]" box="[151,532,151,177]" pageId="14" pageNumber="15">
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<taxonomicName id="4C207DF4B71CAD36FF2EC4AA190B34AC" ID-CoL="6J2VP" authority="Watson, 2005" authorityName="Watson" authorityYear="2005" box="[151,532,151,177]" class="Hydrozoa" family="Lafoeidae" genus="Filellum" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="nitidum">
<emphasis id="B954DA65B71CAD36FF2EC4AA190B34AC" bold="true" box="[151,532,151,177]" pageId="14" pageNumber="15">
<emphasis id="B954DA65B71CAD36FF2EC4AA1A7B34AC" bold="true" box="[151,356,151,177]" italics="true" pageId="14" pageNumber="15">Filellum nitidum</emphasis>
<bibRefCitation id="EFB17B86B71CAD36FED5C4AA190B34AC" author="Watson" box="[364,532,151,177]" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson, 2005</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8B9F0677B71CAD36FF2EC4E219BC34E9" blockId="14.[151,675,222,245]" box="[151,675,222,245]" pageId="14" pageNumber="15">
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<taxonomicName id="4C207DF4B71CAD36FF2EC4E21AE034E9" ID-CoL="6J2VP" authority="Watson, 2005: 504" authorityName="Watson" authorityPageNumber="504" authorityYear="2005" box="[151,511,222,245]" class="Hydrozoa" family="Lafoeidae" genus="Filellum" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="nitidum">
<emphasis id="B954DA65B71CAD36FF2EC4E21A2234E8" box="[151,317,223,245]" italics="true" pageId="14" pageNumber="15">Filellum nitidum</emphasis>
<treatmentCitation id="0A812066B71CAD36FEFAC4E21AE034E9" author="Watson" box="[323,511,222,244]" page="504" pageId="14" pageNumber="15" year="2005">
<bibRefCitation id="EFB17B86B71CAD36FEFAC4E21AD134E9" author="Watson" box="[323,462,222,244]" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson, 2005</bibRefCitation>
: 504
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505, figs. 3ce.
</treatmentCitationGroup>
</paragraph>
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<typeStatus id="549BB8D5B71CAD36FF2EC51C1BCD3524" box="[151,210,289,313]" pageId="14" pageNumber="15">Type</typeStatus>
series.
</emphasis>
Holotype—
<taxonomicName id="4C207DF4B71CAD36FE05C51C18303524" authority="Watson, 2005" authorityName="Watson" authorityYear="2005" box="[444,815,289,313]" class="Hydrozoa" family="Lafoeidae" genus="Filellum" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="nitidum">
<emphasis id="B954DA65B71CAD36FE05C51C19623524" box="[444,637,289,313]" italics="true" pageId="14" pageNumber="15">Filellum nitidum</emphasis>
<bibRefCitation id="EFB17B86B71CAD36FD35C51F18303524" author="Watson" box="[652,815,289,313]" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson, 2005</bibRefCitation>
</taxonomicName>
, fertile colony on stem of
<taxonomicName id="4C207DF4B71CAD36FBC7C51F1E0A3524" box="[1150,1301,290,313]" class="Hydrozoa" family="Aglaopheniidae" genus="Gymnangium" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="genus">
<emphasis id="B954DA65B71CAD36FBC7C51F1E0A3524" box="[1150,1301,290,313]" italics="true" pageId="14" pageNumber="15">Gymnangium</emphasis>
</taxonomicName>
sp. (
<collectionCode id="ED319EB2B71CAD36FAE0C51C1E833524" LSID="urn:lsid:biocol.org:col:34978" box="[1369,1436,289,313]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="14" pageNumber="15">NMV</collectionCode>
F
<date id="FF9E20B7B71CAD36FF1EC5781BE03540" box="[167,255,325,349]" pageId="14" pageNumber="15" value="50-10-16">101650</date>
). Paratype—
<taxonomicName id="4C207DF4B71CAD36FE29C57819EF3540" authority="Watson, 2005" authorityName="Watson" authorityYear="2005" box="[400,752,325,349]" class="Hydrozoa" family="Lafoeidae" genus="Filellum" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="nitidum">
<emphasis id="B954DA65B71CAD36FE29C57819533540" box="[400,588,325,349]" italics="true" pageId="14" pageNumber="15">Filellum nitidum</emphasis>
<bibRefCitation id="EFB17B86B71CAD36FDEAC57B19EF3540" author="Watson" box="[595,752,325,349]" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson, 2005</bibRefCitation>
</taxonomicName>
, fertile colony on basal stems of additional colonies of
<taxonomicName id="4C207DF4B71CAD36FAE7C57B1BE6359C" class="Hydrozoa" family="Aglaopheniidae" genus="Gymnangium" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="genus">
<emphasis id="B954DA65B71CAD36FAE7C57B1BE6359C" italics="true" pageId="14" pageNumber="15">Gymnangium</emphasis>
</taxonomicName>
sp. (
<collectionCode id="ED319EB2B71CAD36FE97C5541A6A359C" LSID="urn:lsid:biocol.org:col:34981" box="[302,373,361,385]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="14" pageNumber="15">WAM</collectionCode>
Z29955) (
<bibRefCitation id="EFB17B86B71CAD36FE53C55419A3359C" author="Watson" box="[490,700,361,385]" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson 2005: 504</bibRefCitation>
).
</paragraph>
<paragraph id="8B9F0677B71CAD36FF7EC5B01F9235B8" blockId="14.[151,1437,288,1177]" box="[199,1165,397,422]" pageId="14" pageNumber="15">
<emphasis id="B954DA65B71CAD36FF7EC5B01A7A35BB" bold="true" box="[199,357,397,422]" pageId="14" pageNumber="15">
<typeStatus id="549BB8D5B71CAD36FF7EC5B01A1D35B8" box="[199,258,397,421]" pageId="14" pageNumber="15">Type</typeStatus>
locality.
</emphasis>
From the
<typeStatus id="549BB8D5B71CAD36FE60C5B0192135B8" box="[473,574,397,421]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
,
<collectingCountry id="F33746E7B71CAD36FDF0C5B019AC35B8" box="[585,691,397,421]" name="Australia" pageId="14" pageNumber="15">Australia</collectingCountry>
, Middle
<collectingCountry id="F33746E7B71CAD36FCACC5B0184335B8" box="[789,860,397,421]" name="Iceland" pageId="14" pageNumber="15">Island</collectingCountry>
,
<quantity id="4CD8AB92B71CAD36FCD1C5B018BE35B8" box="[872,929,397,421]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="15" unit="m" value="25.0">25 m</quantity>
(
<bibRefCitation id="EFB17B86B71CAD36FC16C5B01F9F35B8" author="Watson" box="[943,1152,397,421]" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson 2005: 504</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A55FCB71CAD36FF7EC58D196B35D4" box="[199,628,432,457]" pageId="14" pageNumber="15" type="description">
<paragraph id="8B9F0677B71CAD36FF7EC58D196B35D4" blockId="14.[151,1437,288,1177]" box="[199,628,432,457]" pageId="14" pageNumber="15">
<emphasis id="B954DA65B71CAD36FF7EC58D1A4635D4" bold="true" box="[199,345,432,457]" pageId="14" pageNumber="15">Description.</emphasis>
See
<bibRefCitation id="EFB17B86B71CAD36FE37C58F197035D4" author="Watson" box="[398,623,433,457]" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson (2005: 504)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A55FCB71CAD36FF7EC5E81A5C360C" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8B9F0677B71CAD36FF7EC5E81A5C360C" blockId="14.[151,1437,288,1177]" pageId="14" pageNumber="15">
<emphasis id="B954DA65B71CAD36FF7EC5E81A7F35F3" bold="true" box="[199,352,469,494]" pageId="14" pageNumber="15">Distribution.</emphasis>
<taxonomicName id="4C207DF4B71CAD36FED2C5E8193535F0" box="[363,554,469,493]" class="Hydrozoa" family="Lafoeidae" genus="Filellum" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="nitidum">
<emphasis id="B954DA65B71CAD36FED2C5E8193535F0" box="[363,554,469,493]" italics="true" pageId="14" pageNumber="15">Filellum nitidum</emphasis>
</taxonomicName>
is only known from its original description, from
<collectingCountry id="F33746E7B71CAD36FB3AC5E81FF235F0" box="[1155,1261,469,493]" name="Australia" pageId="14" pageNumber="15">Australia</collectingCountry>
, Middle
<collectingCountry id="F33746E7B71CAD36FAEEC5E81E8435F0" box="[1367,1435,469,493]" name="Iceland" pageId="14" pageNumber="15">Island</collectingCountry>
(
<bibRefCitation id="EFB17B86B71CAD36FF27C5C41A28360C" author="Watson" box="[158,311,505,529]" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A55FCB71CAD36FF7EC6201F1D3084" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8B9F0677B71CAD36FF7EC6201F1D3084" blockId="14.[151,1437,288,1177]" pageId="14" pageNumber="15">
<emphasis id="B954DA65B71CAD36FF7EC6201A243628" bold="true" box="[199,315,541,565]" pageId="14" pageNumber="15">Remarks.</emphasis>
<taxonomicName id="4C207DF4B71CAD36FEFBC6201AE23628" box="[322,509,541,565]" class="Hydrozoa" family="Lafoeidae" genus="Filellum" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="nitidum">
<emphasis id="B954DA65B71CAD36FEFBC6201AE23628" box="[322,509,541,565]" italics="true" pageId="14" pageNumber="15">Filellum nitidum</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFB17B86B71CAD36FD75C62318B53628" author="Watson" box="[716,938,541,565]" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson (2005: 504)</bibRefCitation>
as bearing “Hydrothecae sessile […] adnate part of base fairly short, free part long, perisarc of abcauline wall faintly wrinkled; perisarc of free wall rather thick, marginal rim slightly everted and thickened” and “hydrothecae with several regenerations along length”. Surprisingly, in spite of the original description and the figure 3C, where wrinkled hydrothecae are shown,
<bibRefCitation id="EFB17B86B71CAD36FB41C6B71BD536D8" author="Watson" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson (2005: 505)</bibRefCitation>
also stated that the “absence of marked abcauline rugosity and absence of a distinctly everted rim” would make the trophosome of the species similar to that of
<taxonomicName id="4C207DF4B71CAD36FD55C6EF189136F4" box="[748,910,722,745]" class="Hydrozoa" family="Lafoeidae" genus="Filellum" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="antarcticum">
<emphasis id="B954DA65B71CAD36FD55C6EF189136F4" box="[748,910,722,745]" italics="true" pageId="14" pageNumber="15">F. antarcticum</emphasis>
</taxonomicName>
. This seems to be an inapt description, since
<taxonomicName id="4C207DF4B71CAD36FA3EC6EF1A023710" class="Hydrozoa" family="Lafoeidae" genus="Filellum" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="antarcticum">
<emphasis id="B954DA65B71CAD36FA3EC6EF1A023710" italics="true" pageId="14" pageNumber="15">F. antarcticum</emphasis>
</taxonomicName>
has distinctly marked striae on the upper side of the adnate portion of the hydrothecae (
<bibRefCitation id="EFB17B86B71CAD36FB53C6C81A1A372C" author="Pena" pageId="14" pageNumber="15" refString="Pena Cantero, A. L., Svoboda, A. &amp; Vervoort, W. (2004 b) Antarctic hydroids (Cnidaria, Hydrozoa) of the families Campanulinidae, Lafoeidae and Campanulariidae from recent Antarctic expeditions with R. V. Polarstern, with the description of a new species. Journal of Natural History, 38, 2269 - 2303." type="journal article" year="2004" yearSuffix="b">
Peña Cantero
<emphasis id="B954DA65B71CAD36FA31C6CB1BAB372C" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2004b
</bibRefCitation>
). On the other hand, as also remarked by
<bibRefCitation id="EFB17B86B71CAD36FD63C727189E372C" author="Watson" box="[730,897,793,817]" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson (2005)</bibRefCitation>
, both species have clearly different gonothecae. Other morphological features listed by
<bibRefCitation id="EFB17B86B71CAD36FDEAC70319E63748" author="Watson" box="[595,761,829,853]" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson (2005)</bibRefCitation>
do not distinguish this species from the others of the genus
<taxonomicName id="4C207DF4B71CAD36FF2EC75C1BE93764" box="[151,246,865,889]" class="Hydrozoa" family="Lafoeidae" genus="Filellum" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="genus">
<emphasis id="B954DA65B71CAD36FF2EC75C1BE93764" box="[151,246,865,889]" italics="true" pageId="14" pageNumber="15">Filellum</emphasis>
</taxonomicName>
. Therefore, we believe that reliable diagnosis of
<taxonomicName id="4C207DF4B71CAD36FCFEC75F18A23764" box="[839,957,865,889]" class="Hydrozoa" family="Lafoeidae" genus="Filellum" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="nitidum">
<emphasis id="B954DA65B71CAD36FCFEC75F18A23764" box="[839,957,865,889]" italics="true" pageId="14" pageNumber="15">F. nitidum</emphasis>
</taxonomicName>
must be based on coppiniae characters, described as “surface of coppinia a mosaic covered by a thin pellicle of tissue perforated by small openings flush with surface; hydrothecae protruding above surface together with large inverted funnel-shaped tubules, some tubules bifurcated, apices of others closed” and the sectional view of “numerous rather haphazardly arranged elongate gonothecae” (
<bibRefCitation id="EFB17B86B71CAD36FED1C7CC196C3014" author="Watson" box="[360,627,1009,1033]" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson 2005: 504505</bibRefCitation>
). The figure illustrating the coppinia of this species (
<bibRefCitation id="EFB17B86B71CAD36FB7EC7CC1A0C3030" author="Watson" pageId="14" pageNumber="15" refString="Watson, J. E. (2005) Hydroids of the Archipelago of the Recherche and Esperance, Western Australia: Annotated list, redescription of species and description of new species. In: Weills, F. E., Walker, D. I. &amp; Kendrick, G. A. (Eds), The marine flora and fauna of Esperance, Western Australia. Western Australian Museum, Perth, pp. 495 - 612." type="book chapter" year="2005">Watson 2005: 503, figs. 3DE</bibRefCitation>
) shows protective tubes among the gonothecae; this, together with the different shape of the gonothecae, make the species distinct from
<taxonomicName id="4C207DF4B71CAD36FD9CC00719DA304C" box="[549,709,1081,1105]" class="Hydrozoa" family="Lafoeidae" genus="Filellum" kingdom="Animalia" order="Leptothecata" pageId="14" pageNumber="15" phylum="Cnidaria" rank="species" species="magnificum">
<emphasis id="B954DA65B71CAD36FD9CC00719DA304C" box="[549,709,1081,1105]" italics="true" pageId="14" pageNumber="15">F. magnificum</emphasis>
</taxonomicName>
, for which the defensive tubes are mainly situated on the periphery of the mass of gonothecae, like a fence, arching over the gonothecae. There is no information on the cnidome of the species, and no further comparison concerning this character can be made.
</paragraph>
</subSubSection>
</treatment>
</document>