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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1113.82172" ID-Pensoft-Pub="1313-2970-1113-1" ID-Pensoft-UUID="EB46BF265F2E51B3A83799886C5F084A" ModsDocID="1313-2970-1113-1" checkinTime="1658252710949" checkinUser="pensoft" docAuthor="Bribiesca-Contreras, Guadalupe, Dahlgren, Thomas G., Amon, Diva J., Cairns, Stephen, Drennan, Regan, Durden, Jennifer M., Eleaume, Marc P., Hosie, Andrew M., Kremenetskaia, Antonina, McQuaid, Kirsty, O'Hara, Timothy D., Rabone, Muriel, Simon-Lledo, Erik, Smith, Craig R., Watling, Les, Wiklund, Helena &amp; Glover, Adrian G." docDate="2022" docId="DFCD19FD1EBA5FD98FAFBD1DE8BC0981" docLanguage="en" docName="ZooKeys 1113: 1-110" docOrigin="ZooKeys 1113" docPubDate="2022-07-18" docSource="http://dx.doi.org/10.3897/zookeys.1113.82172" docTitle="Peniagone leander Pawson &amp; Foell 1986" docType="treatment" docVersion="1" id="EB46BF265F2E51B3A83799886C5F084A" lastPageNumber="1" masterDocId="EB46BF265F2E51B3A83799886C5F084A" masterDocTitle="Benthic megafauna of the western Clarion-Clipperton Zone, Pacific Ocean" masterLastPageNumber="110" masterPageNumber="1" pageNumber="1" updateTime="1658253014331" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Benthic megafauna of the western Clarion-Clipperton Zone, Pacific Ocean</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bribiesca-Contreras, Guadalupe</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8163-8724</mods:nameIdentifier>
<mods:affiliation>Life Sciences Department, Natural History Museum, London, UK</mods:affiliation>
<mods:nameIdentifier type="email">l.bribiesca-contreras@nhm.ac.uk</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dahlgren, Thomas G.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6854-2031</mods:nameIdentifier>
<mods:affiliation>Department of Marine Sciences, University of Gothenburg, Gothenburg, Sweden &amp; Norwegian Research Centre, NORCE, Bergen, Norway</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Amon, Diva J.</mods:namePart>
<mods:affiliation>SpeSeas, D'Abadie, Trinidad and Tobago</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cairns, Stephen</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7209-9271</mods:nameIdentifier>
<mods:affiliation>Department of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, D. C., USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Drennan, Regan</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0137-5464</mods:nameIdentifier>
<mods:affiliation>National Oceanography Centre, Southampton, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Durden, Jennifer M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6529-9109</mods:nameIdentifier>
<mods:affiliation>UMR ISYEB, Department Origines et Evolution, Museum national d'Histoire Naturelle, Paris, France</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Eleaume, Marc P.</mods:namePart>
<mods:affiliation>Collections &amp; Research, Western Australia Museum, Perth, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hosie, Andrew M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5683-662X</mods:nameIdentifier>
<mods:affiliation>Shirshov Institute of Oceanology, Russian Academy of Sciences, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kremenetskaia, Antonina</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8851-3318</mods:nameIdentifier>
<mods:affiliation>School of Biological and Marine Sciences, University of Plymouth, Plymouth, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>McQuaid, Kirsty</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0395-8332</mods:nameIdentifier>
<mods:affiliation>Museums Victoria, Melbourne, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>O'Hara, Timothy D.</mods:namePart>
<mods:affiliation>Department of Oceanography, University of Hawai'i at Manoa, Honolulu, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rabone, Muriel</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8351-2313</mods:nameIdentifier>
<mods:affiliation>National Oceanography Centre, Southampton, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Simon-Lledo, Erik</mods:namePart>
<mods:affiliation>UMR ISYEB, Department Origines et Evolution, Museum national d'Histoire Naturelle, Paris, France</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Craig R.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3976-0889</mods:nameIdentifier>
<mods:affiliation>School of Life Sciences, University of Hawai'i at Manoa, Honolulu, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Watling, Les</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6901-1168</mods:nameIdentifier>
<mods:affiliation>School of Life Sciences, University of Hawai'i at Manoa, Honolulu, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wiklund, Helena</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8252-3504</mods:nameIdentifier>
<mods:affiliation>Department of Marine Sciences, University of Gothenburg, Gothenburg, Sweden</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Glover, Adrian G.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9489-074X</mods:nameIdentifier>
<mods:affiliation>National Oceanography Centre, Southampton, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-07-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1113</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>110</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1113.82172</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1113.82172</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1113-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">EB46BF265F2E51B3A83799886C5F084A</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:DFCD19FD1EBA5FD98FAFBD1DE8BC0981" httpUri="http://treatment.plazi.org/id/DFCD19FD1EBA5FD98FAFBD1DE8BC0981" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="DFCD19FD-1EBA-5FD9-8FAF-BD1DE8BC0981" authority="Pawson &amp; Foell, 1986" authorityName="Pawson &amp; Foell" authorityYear="1986" class="Holothuroidea" family="Elpidiidae" genus="Peniagone" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Peniagone leander" order="Elasipodida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leander">Peniagone leander Pawson &amp; Foell, 1986</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Peniagone leander Pawson &amp; Foell, 1986. Specimen CCZ _ 018: A, B in situ images C tentacle ossicles. Scale bars: 100 µm (C). Image attribution: Durden and Smith (A, B); Bribiesca-Contreras (C)." figureDoi="10.3897/zookeys.1113.82172.figure46" httpUri="https://binary.pensoft.net/fig/716372" pageId="0" pageNumber="1">Fig. 46</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation accessionNumber="ON400681, ON406621" collectingDate="2018-05-25" collectorName="Smith, Durden, GenBank, Voucher" elevation="4860" latitude="5.1042" longLatPrecision="7" longitude="-141.8861" specimenCount="1">
Clarion-Clipperton Zone •
<specimenCount type="generic">1 specimen</specimenCount>
; APEI 7;
<geoCoordinate degrees="5.1042" direction="north" orientation="latitude" precision="5" value="5.1042">5.1042°N</geoCoordinate>
,
<geoCoordinate degrees="141.8861" direction="west" orientation="longitude" precision="5" value="-141.8861">141.8861°W</geoCoordinate>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="4.86" unit="m" value="4860.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="4.86" unit="m" value="4860.0">4860 m</elevation>
</quantity>
deep;
<collectingDate value="2018-05-25">25 May. 2018</collectingDate>
;
<collectorName>Smith</collectorName>
&amp;
<collectorName>Durden</collectorName>
leg.;
<collectorName>GenBank</collectorName>
:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/ON400681">ON400681</accessionNumber>
(COI),
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/ON406621">ON406621</accessionNumber>
(16S); NHMUK 2022.61;
<collectorName>Voucher</collectorName>
code: CCZ_018
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Single specimen observed swimming (Fig.
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Peniagone leander Pawson &amp; Foell, 1986. Specimen CCZ _ 018: A, B in situ images C tentacle ossicles. Scale bars: 100 µm (C). Image attribution: Durden and Smith (A, B); Bribiesca-Contreras (C)." figureDoi="10.3897/zookeys.1113.82172.figure46" httpUri="https://binary.pensoft.net/fig/716372" pageId="0" pageNumber="1">46A</figureCitation>
). Specimen was severely damaged during collection, with only a few tentacles recovered, and hence description of morphological characters is based on in situ images. Body ovoid, slightly&gt; 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as it is wide. Velum composed of two pairs of fully fused papillae. Tube feet four pairs; three posteriormost pairs fused together forming a posterior swimming lobe; tube feet from the anteriormost pair very short.
</paragraph>
<caption doi="10.3897/zookeys.1113.82172.figure46" httpUri="https://binary.pensoft.net/fig/716372" pageId="0" pageNumber="1" start="Figure 46" startId="F46">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 46.</emphasis>
<taxonomicName authorityName="Pawson &amp; Foell" authorityYear="1986" class="Holothuroidea" family="Elpidiidae" genus="Peniagone" kingdom="Animalia" lsidName="Peniagone leander" order="Scalpelliformes" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leander">
<emphasis italics="true" pageId="0" pageNumber="1">Peniagone leander</emphasis>
</taxonomicName>
Pawson &amp; Foell, 1986. Specimen CCZ_018:
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
in situ images
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
tentacle ossicles. Scale bars: 100
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
). Image attribution: Durden and Smith (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
); Bribiesca-Contreras (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specimen collected during the DeepCCZ expedition was recovered in bits, so no morphological features can be distinguished. Only four reddish orange tentacles were recovered, which are embedded in a transparent skin where ossicles are evident. However,
<taxonomicName genus="P." lsidName="P. leander" pageId="0" pageNumber="1" rank="species" species="leander">
<emphasis italics="true" pageId="0" pageNumber="1">P. leander</emphasis>
</taxonomicName>
is one of the few species that can be identified from images. The external morphological characters evident in in situ images from the CCZ specimen are in accordance with the species description. The species was originally described from in situ images and video footage collected across the eastern CCZ (
<bibRefCitation author="Pawson, DL" journalOrPublisher="Bulletin of Marine Science" pageId="0" pageNumber="1" pagination="293 - 299" refId="B78" refString="Pawson, DL, Foell, EJ, 1986. Peniagone leander new species, an abyssal benthopelagic sea cucumber (Echinodermata: Holothuroidea) from the eastern central Pacific Ocean. Bulletin of Marine Science 38: 293 - 299" title="Peniagone leander new species, an abyssal benthopelagic sea cucumber (Echinodermata: Holothuroidea) from the eastern central Pacific Ocean." volume="38" year="1986">Pawson and Foell 1986</bibRefCitation>
) and subsequently observed in the area (e.g.,
<bibRefCitation DOI="https://doi.org/10.3897/BDJ.5.e11794" author="Amon, DJ" journalOrPublisher="Scientific Reports" pageId="0" pageNumber="1" refId="B6" refString="Amon, DJ, Ziegler, AF, Kremenetskaia, A, Mah, CL, Mooi, R, O'Hara, T, Pawson, DL, Roux, M, Smith, CR, 2017b. Megafauna of the UKSRL exploration contract area and eastern Clarion-Clipperton Zone in the Pacific Ocean: Echinodermata. Biodiversity Data Journal 11794: e11794. https://doi.org/10.3897/BDJ.5.e11794" title="Megafauna of the UKSRL exploration contract area and eastern Clarion-Clipperton Zone in the Pacific Ocean: Echinodermata. Biodiversity Data Journal 11794: e 11794." url="https://doi.org/10.3897/BDJ.5.e11794" year="2017 b">Amon et al. 2017b</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
In the phylogenetic tree, the CCZ specimen was recovered in a well-supported clade with other species of
<taxonomicName authorityName="Theel" authorityYear="1882" class="Holothuroidea" family="Elpidiidae" genus="Peniagone" kingdom="Animalia" lsidName="Peniagone" order="Scalpelliformes" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Peniagone</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Phylogenetic tree of the class Holothuroidea. Concatenated (12 S, 16 S, 18 S, 28 S, COI, and H 3) median consensus BEAST tree with posterior probability (PP) and bootstrap (BS) values indicated. Only values of PP&gt; 0.70 and BS&gt; 50 are shown, with values of PP&gt; 0.95 and BS&gt; 90 indicated with a circle. Nodes not recovered on the RAxML tree are indicated with a hyphen. Sequences generated in this study are highlighted in violet." figureDoi="10.3897/zookeys.1113.82172.figure34" httpUri="https://binary.pensoft.net/fig/716360" pageId="0" pageNumber="1">34</figureCitation>
). It was recovered together with a sequence of
<taxonomicName genus="P." lsidName="P. leander" pageId="0" pageNumber="1" rank="species" species="leander">
<emphasis italics="true" pageId="0" pageNumber="1">P. leander</emphasis>
</taxonomicName>
, which was recently rediscovered and collected for the first time in the Mariana Trench (
<bibRefCitation DOI="https://doi.org/10.1007/s00343-020-0067-9" author="Gong, L" journalOrPublisher="Journal of Oceanology and Limnology" pageId="0" pageNumber="1" pagination="1319 - 1327" refId="B35" refString="Gong, L, Li, X, Xiao, N, He, L, Zhang, H, Wang, Y, 2020. Rediscovery of the abyssal species Peniagone leander Pawson &amp; Foell, 1986 (Holothuroidea: Elasipodida: Elpidiidae): the first record from the Mariana Trench area. Journal of Oceanology and Limnology 38 (4): 1319 - 1327, DOI: https://doi.org/10.1007/s00343-020-0067-9" title="Rediscovery of the abyssal species Peniagone leander Pawson &amp; Foell, 1986 (Holothuroidea: Elasipodida: Elpidiidae): the first record from the Mariana Trench area." url="https://doi.org/10.1007/s00343-020-0067-9" volume="38" year="2020">Gong et al. 2020</bibRefCitation>
), both close to
<taxonomicName genus="P." lsidName="P. diaphana" pageId="0" pageNumber="1" rank="species" species="diaphana">
<emphasis italics="true" pageId="0" pageNumber="1">P. diaphana</emphasis>
</taxonomicName>
as reported by
<bibRefCitation DOI="https://doi.org/10.1007/s00343-020-0067-9" author="Gong, L" journalOrPublisher="Journal of Oceanology and Limnology" pageId="0" pageNumber="1" pagination="1319 - 1327" refId="B35" refString="Gong, L, Li, X, Xiao, N, He, L, Zhang, H, Wang, Y, 2020. Rediscovery of the abyssal species Peniagone leander Pawson &amp; Foell, 1986 (Holothuroidea: Elasipodida: Elpidiidae): the first record from the Mariana Trench area. Journal of Oceanology and Limnology 38 (4): 1319 - 1327, DOI: https://doi.org/10.1007/s00343-020-0067-9" title="Rediscovery of the abyssal species Peniagone leander Pawson &amp; Foell, 1986 (Holothuroidea: Elasipodida: Elpidiidae): the first record from the Mariana Trench area." url="https://doi.org/10.1007/s00343-020-0067-9" volume="38" year="2020">Gong et al. (2020)</bibRefCitation>
. The 16S sequence of the CCZ specimen is similar (K2P distance = 2%) to the only available sequence from
<taxonomicName genus="P." lsidName="P. leander" pageId="0" pageNumber="1" rank="species" species="leander">
<emphasis italics="true" pageId="0" pageNumber="1">P. leander</emphasis>
</taxonomicName>
, but no COI sequence was made available. Our COI sequence is&gt; 12% divergent (K2P distance) from other species within the genus. The COI gene seems to be highly divergent between species in this genus. Using the data provided in
<bibRefCitation DOI="https://doi.org/10.3390/d13110541" author="Kremenetskaia, A" journalOrPublisher="Marine Biodiversity" pageId="0" pageNumber="1" refId="B56" refString="Kremenetskaia, A, Gebruk, A, Alt, CHS, Budaeva, N, 2021. New and Poorly Known Species of Peniagone (Holothuroidea, Elpidiidae) from the Northwest Pacific Ocean with Discussion on Phylogeny of the Genus. Diversity (Basel) 13(11): e541. https://doi.org/10.3390/d13110541" title="New and Poorly Known Species of Peniagone (Holothuroidea, Elpidiidae) from the Northwest Pacific Ocean with Discussion on Phylogeny of the Genus. Diversity (Basel) 13 (11): e 541." url="https://doi.org/10.3390/d13110541" year="2021">Kremenetskaia et al. (2021)</bibRefCitation>
and including the CCZ sequence of
<taxonomicName genus="P." lsidName="P. leander" pageId="0" pageNumber="1" rank="species" species="leander">
<emphasis italics="true" pageId="0" pageNumber="1">P. leander</emphasis>
</taxonomicName>
, COI mean interspecific divergence in the genus is 15.9% (min = 2.5% and max = 22.7%), with our sequence of
<taxonomicName genus="P." lsidName="P. leander" pageId="0" pageNumber="1" rank="species" species="leander">
<emphasis italics="true" pageId="0" pageNumber="1">P. leander</emphasis>
</taxonomicName>
being 14.5%-21.2% divergent from other species within the genus. Intraspecific divergence for species in the genus was estimated between 0.9%-3.0%.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">The specimen was found swimming near the sediment surface on an abyssal plain in APEI 7 at 4860 m depth.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="comparisons">
<paragraph pageId="0" pageNumber="1">Comparison with image-based catalogue.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pawson &amp; Foell" authorityYear="1986" class="Holothuroidea" family="Elpidiidae" genus="Peniagone" kingdom="Animalia" lsidName="Peniagone leander" order="Scalpelliformes" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leander">
<emphasis italics="true" pageId="0" pageNumber="1">Peniagone leander</emphasis>
</taxonomicName>
(HOL_028) has been commonly encountered in seabed image surveys conducted across the eastern CCZ (e.g.,
<bibRefCitation DOI="https://doi.org/10.3897/BDJ.5.e11794" author="Amon, DJ" journalOrPublisher="Scientific Reports" pageId="0" pageNumber="1" refId="B6" refString="Amon, DJ, Ziegler, AF, Kremenetskaia, A, Mah, CL, Mooi, R, O'Hara, T, Pawson, DL, Roux, M, Smith, CR, 2017b. Megafauna of the UKSRL exploration contract area and eastern Clarion-Clipperton Zone in the Pacific Ocean: Echinodermata. Biodiversity Data Journal 11794: e11794. https://doi.org/10.3897/BDJ.5.e11794" title="Megafauna of the UKSRL exploration contract area and eastern Clarion-Clipperton Zone in the Pacific Ocean: Echinodermata. Biodiversity Data Journal 11794: e 11794." url="https://doi.org/10.3897/BDJ.5.e11794" year="2017 b">Amon et al. 2017b</bibRefCitation>
) and in abyssal areas of the Kiribati EEZ, usually swimming above the seabed but sometimes creeping on it. Body colour appears to be variable; bright red, semi-transparent, purplish, and whitish HOL_028 specimens have been encountered in seabed image surveys across the CCZ.
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</subSubSection>
</treatment>
</document>
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