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<document id="999D79C125DDA0381A27BFA5BBBCAB2C" ID-CLB-Dataset="5297" ID-DOI="10.5852/ejt.2021.760.1437" ID-GBIF-Dataset="9ecf4292-270f-4343-aeaa-04bc7b75de2b" ID-ISSN="2118-9773" ID-Zenodo-Dep="5121748" ID-ZooBank="02A1C7B1-CE07-4BB8-BC6C-3B596598A8C3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626959611123" checkinUser="felipe" docAuthor="Reverter-Gil, Oscar &amp; Souto, Javier" docDate="2021" docId="DF223750FFE7CC3DFDCEFAD6156B0BA5" docLanguage="en" docName="ejt-760_16-31.pdf" docOrigin="European Journal of Taxonomy 760" docSource="http://dx.doi.org/10.5852/ejt.2021.760.1437" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cradoscrupocellaria severoi Reverter-Gil &amp; Souto 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="22" masterDocId="231B4F28FFE6CC3BFFF1FFD91444096B" masterDocTitle="Two new species of cheilostomate Bryozoa from Iberian waters" masterLastPageNumber="31" masterPageNumber="16" pageNumber="17" updateTime="1699037029033" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7A6F625D57EAF901F480D686B6DBA5EF">Two new species of cheilostomate Bryozoa from Iberian waters</mods:title>
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<mods:namePart id="99DB5FAD0BD7BEA413DC59C7963CFA64">Reverter-Gil, Oscar</mods:namePart>
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<mods:affiliation id="24C8D478A399CF4A23B65F371D29A6A0">Museo de Historia Natural da Universidade de Santiago de Compostela, Parque Vista Alegre s / n, 15705 Santiago de Compostela, Spain.</mods:affiliation>
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<mods:namePart id="46659E45C2DA18FBE97EB3EA0C1B9E17">Souto, Javier</mods:namePart>
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<mods:affiliation id="1F38E42577E362FD16544FC794CAD5E4">Institut für Paläontologie, Fakultät für Geowissenschaften, Geographie und Astronomie, Geozentrum, Universität Wien, Althanstrasse 14, 1090 Wien, Austria.</mods:affiliation>
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<heading id="0C7C312AFFE7CC3AFDCEFAD617B10C41" box="[575,1013,1295,1322]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="17" reason="2">
<taxonomicName id="908BFDC5FFE7CC3AFDCEFAD617D70C41" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[575,915,1295,1322]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="1" pageNumber="17" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE7CC3AFDCEFAD617D70C41" bold="true" box="[575,915,1295,1322]" italics="true" pageId="1" pageNumber="17">Cradoscrupocellaria severoi</emphasis>
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<taxonomicNameLabel id="7ECCE72FFFE7CC3AFC6BFAC917B10C41" box="[922,1013,1296,1322]" pageId="1" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act: 
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</paragraph>
<paragraph id="57348646FFE7CC3AFD45FA8017C40C1F" blockId="1.[354,1233,1295,1396]" box="[692,896,1369,1396]" pageId="1" pageNumber="17">
<figureCitation id="CFB09AC3FFE7CC3AFD45FA8017590C1F" box="[692,797,1369,1396]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[189,232,1894,1920]" captionStartId-1="3.[191,234,1857,1883]" captionTargetBox-0="[189,1399,617,1854]" captionTargetBox-1="[189,1399,773,1816]" captionTargetId-0="figure-132@2.[189,1399,617,1854]" captionTargetId-1="figure-181@3.[189,1399,773,1816]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Cradoscrupocellaria severoi sp. nov. A. View of the colony (MNCN 25.03/3059: holotype). B. View of the colony (MNCN 25.03/3200: paratype). C. Branching pattern (MNCN 25.03/4234: paratype). D. Detail of a branch with large frontal avicularia, ovicells and scuta (MNCN 25.03/3059: holotype)." captionText-1="Fig. 2. Cradoscrupocellaria severoi sp. nov. A. Detail of branches with large frontal avicularia, ovicells, scuta and a lateral avicularium (centre left) (MNCN 25.03/3059: holotype). B. Detail of a branch with large frontal avicularia, ovicells and a lateral avicularium (bottom left) (MNCN 25.03/3200: paratype). C. Basal view of the branching pattern (MNCN 25.03/4234: paratype). D. Single axial vibraculum (MNCN 25.03/3059: holotype)." figureDoi-0="http://doi.org/10.5281/zenodo.5121752" figureDoi-1="http://doi.org/10.5281/zenodo.5121754" httpUri-0="https://zenodo.org/record/5121752/files/figure.png" httpUri-1="https://zenodo.org/record/5121754/files/figure.png" pageId="1" pageNumber="17">Figs 1–2</figureCitation>
; 
<tableCitation id="1A09B3FDFFE7CC3AFCD8FA8017C40C1F" box="[809,896,1369,1396]" captionStart="Table 1" captionStartId="5.[189,258,265,291]" captionTargetPageId="5" captionText="Table 1. Measurements (in mm) of Cradoscrupocellaria severoi sp. nov. (holotype + paratypes). Abbreviations: SD = standard deviation, N = number of measurements." httpUri="http://table.plazi.org/id/03F4D6CEFFE3CC3EFF4CFED017FD082C" pageId="1" pageNumber="17" tableUuid="03F4D6CEFFE3CC3EFF4CFED017FD082C">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="1F91D5CDFFE7CC3AFF4CFA4510E20F6B" pageId="1" pageNumber="17" type="reference_group">
<paragraph id="57348646FFE7CC3AFF4CFA4517660CDC" blockId="1.[189,1190,1436,1537]" box="[189,802,1436,1463]" pageId="1" pageNumber="17">
<treatmentCitationGroup id="779BA168FFE7CC3AFF4CFA4517660CDC" box="[189,802,1436,1463]" pageId="1" pageNumber="17">
<taxonomicName id="908BFDC5FFE7CC3AFF4CFA45158E0CDD" ID-CoL="4VZP6" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[189,458,1436,1462]" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="1" pageNumber="17" phylum="Bryozoa" rank="species" species="reptans">
<emphasis id="65FF5A54FFE7CC3AFF4CFA45158E0CDD" box="[189,458,1436,1462]" italics="true" pageId="1" pageNumber="17">Scrupocellaria reptans</emphasis>
</taxonomicName>
– 
<treatmentCitation id="D62AA057FFE7CC3AFE17FA45175B0CDC" author="Templado J. &amp; Calvo M. &amp; Garcia Carrascosa A. M. &amp; Boisset F. &amp; Jimenez J." box="[486,799,1436,1463]" page="203" pageId="1" pageNumber="17" year="2002">
<bibRefCitation id="331AFBB7FFE7CC3AFE17FA45175B0CDC" author="Templado J. &amp; Calvo M. &amp; Garcia Carrascosa A. M. &amp; Boisset F. &amp; Jimenez J." box="[486,799,1436,1463]" pageId="1" pageNumber="17" refId="ref8136" refString="Templado J., Calvo M., Garcia Carrascosa A. M., Boisset F. &amp; Jimenez J. 2002. Flora y Fauna de la Reserva Marina de las Islas Columbretes. Ministerio de Agricultura, Pesca y Alimentacion, Madrid." type="book" year="2002">
Templado 
<emphasis id="65FF5A54FFE7CC3AFD93FA4416D90CDD" box="[610,669,1436,1462]" italics="true" pageId="1" pageNumber="17">et al.</emphasis>
2002: 203
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="57348646FFE7CC3AFF4CFA1810670CB0" blockId="1.[189,1190,1436,1537]" box="[189,1059,1473,1500]" pageId="1" pageNumber="17">
<treatmentCitationGroup id="779BA168FFE7CC3AFF4CFA1810670CB0" box="[189,1059,1473,1500]" pageId="1" pageNumber="17">
? 
<taxonomicName id="908BFDC5FFE7CC3AFF20FA1816740CB0" ID-CoL="4VZNP" authorityName="Gautier" authorityYear="1962" box="[209,560,1473,1499]" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="1" pageNumber="17" phylum="Bryozoa" rank="species" species="macrorhyncha">
<emphasis id="65FF5A54FFE7CC3AFF20FA1816740CB0" box="[209,560,1473,1499]" italics="true" pageId="1" pageNumber="17">Scrupocellaria macrorhyncha</emphasis>
</taxonomicName>
– 
<treatmentCitation id="D62AA057FFE7CC3AFDBFFA18175A0CB0" author="Zabala M." box="[590,798,1473,1499]" page="318" pageId="1" pageNumber="17" year="1986">
<bibRefCitation id="331AFBB7FFE7CC3AFDBFFA18175A0CB0" author="Zabala M." box="[590,798,1473,1499]" pageId="1" pageNumber="17" pagination="1 - 836" refId="ref8492" refString="Zabala M. 1986. Fauna dels briozous dels Paisos Catalans. Arxius de la Seccio de Ciencies 84: 1 - 836." type="journal article" year="1986">Zabala 1986: 318</bibRefCitation>
</treatmentCitation>
, fig. 88, pl. 3 figs a–d.
</treatmentCitationGroup>
</paragraph>
<paragraph id="57348646FFE7CC3AFF4CFA3F10E20F6B" blockId="1.[189,1190,1436,1537]" box="[189,1190,1510,1537]" pageId="1" pageNumber="17">
<treatmentCitationGroup id="779BA168FFE7CC3AFF4CFA3F10E20F6B" box="[189,1190,1510,1537]" pageId="1" pageNumber="17">
? 
<taxonomicName id="908BFDC5FFE7CC3AFF20FA3F159A0F6B" ID-CoL="4VZP6" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[209,478,1510,1536]" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="1" pageNumber="17" phylum="Bryozoa" rank="species" species="reptans">
<emphasis id="65FF5A54FFE7CC3AFF20FA3F159A0F6B" box="[209,478,1510,1536]" italics="true" pageId="1" pageNumber="17">Scrupocellaria reptans</emphasis>
</taxonomicName>
– 
<treatmentCitation id="D62AA057FFE7CC3AFE0AFA3F17100F6B" author="Zabala M. &amp; Maluquer P." box="[507,852,1510,1536]" page="97" pageId="1" pageNumber="17" year="1988">
<bibRefCitation id="331AFBB7FFE7CC3AFE0AFA3F17100F6B" author="Zabala M. &amp; Maluquer P." box="[507,852,1510,1536]" pageId="1" pageNumber="17" pagination="1 - 294" refId="ref8516" refString="Zabala M. &amp; Maluquer P. 1988. Illustrated keys for the classification of Mediterranean Bryozoa. Treballs del Museu de Zoologia, Barcelona 4: 1 - 294." type="journal article" year="1988">Zabala &amp; Maluquer 1988: 97</bibRefCitation>
</treatmentCitation>
, figs 159–160, pl. 5 figs a–b.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1F91D5CDFFE7CC3AFF4CF9F016B10E40" pageId="1" pageNumber="17" type="diagnosis">
<paragraph id="57348646FFE7CC3AFF4CF9F0158D0F28" blockId="1.[189,457,1577,1603]" box="[189,457,1577,1603]" pageId="1" pageNumber="17">
<heading id="0C7C312AFFE7CC3AFF4CF9F0158D0F28" bold="true" box="[189,457,1577,1603]" fontSize="11" level="3" pageId="1" pageNumber="17" reason="3">
<emphasis id="65FF5A54FFE7CC3AFF4CF9F0158D0F28" bold="true" box="[189,457,1577,1603]" pageId="1" pageNumber="17">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="57348646FFE7CC3AFF4CF98016B10E40" blockId="1.[188,1399,1625,1836]" pageId="1" pageNumber="17">
<taxonomicName id="908BFDC5FFE7CC3AFF4CF98015F70F18" authorityName="Vieira, Spencer Jones &amp; Winston" authorityYear="2013" box="[189,435,1625,1651]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="1" pageNumber="17" phylum="Bryozoa" rank="genus">
<emphasis id="65FF5A54FFE7CC3AFF4CF98015F70F18" box="[189,435,1625,1651]" italics="true" pageId="1" pageNumber="17">Cradoscrupocellaria</emphasis>
</taxonomicName>
with erect, branched colonies, with internodes comprising 5–10 alternating autozooids. Chitinous joints passing across proximal gymnocyst of outer and inner zooids. Large scutum, stout and flat, highly branched, completely covering the opesia. Two inner spines and 3 outer spines. Small distolateral avicularium frequent. Large monomorphic frontal avicularium in outer zooids oriented upwards on the branch. Ovicells globular, wider than long, perforated by 10–18 rounded pores; ovicelled zooids with 2 inner and 2 outer spines.
</paragraph>
</subSubSection>
<subSubSection id="1F91D5CDFFE7CC3AFF4CF88C16290E83" pageId="1" pageNumber="17" type="etymology">
<paragraph id="57348646FFE7CC3AFF4CF88C15000E04" blockId="1.[189,324,1877,1903]" box="[189,324,1877,1903]" pageId="1" pageNumber="17">
<heading id="0C7C312AFFE7CC3AFF4CF88C15000E04" bold="true" box="[189,324,1877,1903]" fontSize="11" level="3" pageId="1" pageNumber="17" reason="3">
<emphasis id="65FF5A54FFE7CC3AFF4CF88C15000E04" bold="true" box="[189,324,1877,1903]" pageId="1" pageNumber="17">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="57348646FFE7CC3AFF4CF85C16290E83" blockId="1.[188,1399,1924,2024]" pageId="1" pageNumber="17">This species is dedicated to Severo Ochoa de Albornoz (1905–1993), Spanish physician and biochemist, winner of the 1959 Nobel Prize in Physiology or Medicine for the discovery of the mechanisms in the biological synthesis of nucleic acids.</paragraph>
</subSubSection>
<subSubSection id="1F91D5CDFFE4CC38FF4CFED1159C0B58" lastPageId="3" lastPageNumber="19" pageId="2" pageNumber="18" type="materials_examined">
<paragraph id="57348646FFE4CC39FF4CFED115EE0849" blockId="2.[189,426,264,290]" box="[189,426,264,290]" pageId="2" pageNumber="18">
<heading id="0C7C312AFFE4CC39FF4CFED115EE0849" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="2" pageNumber="18" reason="3">
<emphasis id="65FF5A54FFE4CC39FF4CFED115EE0849" bold="true" box="[189,426,264,290]" pageId="2" pageNumber="18">Material examined</emphasis>
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<paragraph id="57348646FFE4CC39FF1BFEEE108508F3" blockId="2.[189,1399,311,408]" pageId="2" pageNumber="18">
<materialsCitation id="E7E38C1BFFE4CC39FF1BFEEE108508F3" ID-GBIF-Occurrence="3326570307" collectionCode="MNCN" elevation="22" location="MEDITERRANEAN SPAIN" municipality="Columbretes Islands" pageId="2" pageNumber="18" specimenCode="MNCN 25.03/3059" specimenCount="1" typeStatus="holotype">
<emphasis id="65FF5A54FFE4CC39FF1BFEEE1518083A" bold="true" box="[234,348,311,337]" pageId="2" pageNumber="18">
<typeStatus id="883038E4FFE4CC39FF1BFEEE1518083A" box="[234,348,311,337]" pageId="2" pageNumber="18" type="holotype">Holotype</typeStatus>
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<location id="5254D09DFFE4CC39FF4CFE831651081F" LSID="urn:lsid:plazi:treatment:DF223750FFE7CC3DFDCEFAD6156B0BA5:5254D09DFFE4CC39FF4CFE831651081F" box="[189,533,346,372]" municipality="Columbretes Islands" name="MEDITERRANEAN SPAIN" pageId="2" pageNumber="18">MEDITERRANEAN SPAIN</location>
• colony; 
<collectingMunicipality id="B7501C3CFFE4CC39FD62FE8317CD081F" box="[659,905,346,372]" pageId="2" pageNumber="18">Columbretes Islands</collectingMunicipality>
; 
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, 
<geoCoordinate id="32BFE081FFE4CC39FBB6FE831093081E" box="[1095,1239,346,373]" pageId="2" pageNumber="18">0º66.817´E</geoCoordinate>
; stn 277B15; depth 
<quantity id="90732BA3FFE4CC39FEF4FEA7152508F3" box="[261,353,382,408]" metricMagnitude="1" metricUnit="m" metricValue="2.15" metricValueMax="4.0" metricValueMin="0.3" pageId="2" pageNumber="18" unit="m" value="21.5" valueMax="40.0" valueMin="3.0">
<elevation id="DCA66175FFE4CC39FEF4FEA7152508F3" box="[261,353,382,408]" metricMagnitude="1" metricUnit="m" metricValue="2.15" metricValueMax="4.0" metricValueMin="0.3" pageId="2" pageNumber="18" unit="m" value="21.5" valueMax="40.0" valueMin="3.0">3–40 m</elevation>
</quantity>
; 1996; Fauna Ibérica IV exped.; 
<specimenCode id="072D2E3DFFE4CC39FD13FEA7178208F3" box="[738,966,381,408]" pageId="2" pageNumber="18">MNCN 25.03/3059</specimenCode>
(
<figureCitation id="CFB09AC3FFE4CC39FC27FEA4101808F3" box="[982,1116,381,408]" captionStart="Fig" captionStartId="2.[189,232,1894,1920]" captionTargetBox="[189,1399,617,1854]" captionTargetId="figure-132@2.[189,1399,617,1854]" captionTargetPageId="2" captionText="Fig. 1. Cradoscrupocellaria severoi sp. nov. A. View of the colony (MNCN 25.03/3059: holotype). B. View of the colony (MNCN 25.03/3200: paratype). C. Branching pattern (MNCN 25.03/4234: paratype). D. Detail of a branch with large frontal avicularia, ovicells and scuta (MNCN 25.03/3059: holotype)." figureDoi="http://doi.org/10.5281/zenodo.5121752" httpUri="https://zenodo.org/record/5121752/files/figure.png" pageId="2" pageNumber="18">Figs 1A, D</figureCitation>
, 
<figureCitation id="CFB09AC3FFE4CC39FB99FEA710F708F3" box="[1128,1203,382,408]" captionStart="Fig" captionStartId="3.[191,234,1857,1883]" captionTargetBox="[189,1399,773,1816]" captionTargetId="figure-181@3.[189,1399,773,1816]" captionTargetPageId="3" captionText="Fig. 2. Cradoscrupocellaria severoi sp. nov. A. Detail of branches with large frontal avicularia, ovicells, scuta and a lateral avicularium (centre left) (MNCN 25.03/3059: holotype). B. Detail of a branch with large frontal avicularia, ovicells and a lateral avicularium (bottom left) (MNCN 25.03/3200: paratype). C. Basal view of the branching pattern (MNCN 25.03/4234: paratype). D. Single axial vibraculum (MNCN 25.03/3059: holotype)." figureDoi="http://doi.org/10.5281/zenodo.5121754" httpUri="https://zenodo.org/record/5121754/files/figure.png" pageId="2" pageNumber="18">2A, D</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="57348646FFE4CC38FF1BFE661132082D" blockId="2.[189,1400,447,579]" lastBlockId="3.[189,1399,265,327]" lastPageId="3" lastPageNumber="19" pageId="2" pageNumber="18">
<materialsCitation id="E7E38C1BFFE4CC39FF1BFE6617930B4B" ID-GBIF-Occurrence="3326570309" collectionCode="MNCN" elevation="42" location="MEDITERRANEAN SPAIN" municipality="Columbretes Islands" pageId="2" pageNumber="18" specimenCode="MNCN 25.03/2999" specimenCount="1" typeStatus="paratype">
<emphasis id="65FF5A54FFE4CC39FF1BFE66152308B2" bold="true" box="[234,359,447,473]" pageId="2" pageNumber="18">
<typeStatus id="883038E4FFE4CC39FF1BFE66152308B2" box="[234,359,447,473]" pageId="2" pageNumber="18" type="paratype">Paratypes</typeStatus>
</emphasis>
<location id="5254D09DFFE4CC39FF4CFE3B16500897" LSID="urn:lsid:plazi:treatment:DF223750FFE7CC3DFDCEFAD6156B0BA5:5254D09DFFE4CC39FF4CFE3B16500897" box="[189,532,482,508]" municipality="Columbretes Islands" name="MEDITERRANEAN SPAIN" pageId="2" pageNumber="18">MEDITERRANEAN SPAIN</location>
• 
<specimenCount id="418D4DCFFFE4CC39FDDEFE3B16DF0897" box="[559,667,482,508]" pageId="2" pageNumber="18">1 colony</specimenCount>
; 
<collectingMunicipality id="B7501C3CFFE4CC39FD56FE3B17D90897" box="[679,925,482,508]" pageId="2" pageNumber="18">Columbretes Islands</collectingMunicipality>
; 
<geoCoordinate id="32BFE081FFE4CC39FC58FE3B100F0896" box="[937,1099,482,509]" pageId="2" pageNumber="18">39º85.450´N</geoCoordinate>
, 
<geoCoordinate id="32BFE081FFE4CC39FBA9FE3B10AC0896" box="[1112,1256,482,509]" pageId="2" pageNumber="18">0º67.617´E</geoCoordinate>
; stn 285B1; depth 
<quantity id="90732BA3FFE4CC39FEFDFDDC150B0B4B" box="[268,335,517,544]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="18" unit="m" value="42.0">
<elevation id="DCA66175FFE4CC39FEFDFDDC150B0B4B" box="[268,335,517,544]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="18" unit="m" value="42.0">42 m</elevation>
</quantity>
; 1996; Fauna Ibérica IV exped.; 
<specimenCode id="072D2E3DFFE4CC39FD1CFDDC17930B4B" box="[749,983,517,544]" pageId="2" pageNumber="18">MNCN 25.03/2999</specimenCode>
</materialsCitation>
• 
<materialsCitation id="E7E38C1BFFE4CC38FC08FDDC16980848" ID-GBIF-Occurrence="3326570308" collectionCode="MNCN" elevation="8" lastPageId="3" lastPageNumber="19" location="Columbretes Islands" municipality="Columbretes Islands" pageId="2" pageNumber="18" specimenCode="MNCN 25.03/3057, 25.03/4234" specimenCount="1">
<specimenCount id="418D4DCFFFE4CC39FC08FDDC10950B74" box="[1017,1233,517,543]" pageId="2" pageNumber="18">several fragments</specimenCount>
; 
<collectingMunicipality id="B7501C3CFFE4CC39FB13FDDC15500B29" pageId="2" pageNumber="18">Columbretes Islands</collectingMunicipality>
; 
<geoCoordinate id="32BFE081FFE4CC39FED8FDF1158F0B28" box="[297,459,552,579]" pageId="2" pageNumber="18">39º87967´N</geoCoordinate>
, 
<geoCoordinate id="32BFE081FFE4CC39FE11FDF116340B28" box="[480,624,552,579]" pageId="2" pageNumber="18">0º66817´E</geoCoordinate>
; stn 277B14; depth 
<quantity id="90732BA3FFE4CC39FC71FDF1178B0B28" box="[896,975,552,579]" metricMagnitude="0" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="18" unit="m" value="7.5">
<elevation id="DCA66175FFE4CC39FC71FDF1178B0B28" box="[896,975,552,579]" metricMagnitude="0" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="18" unit="m" value="7.5">7.5 m</elevation>
</quantity>
; 1996; Fauna Ibérica IV exped.; 
<specimenCode id="072D2E3DFFE5CC38FF4CFED015E7084F" box="[189,419,265,292]" pageId="3" pageNumber="19">MNCN 25.03/3057</specimenCode>
, 
<specimenCode id="072D2E3DFFE5CC38FE5EFED01670084F" box="[431,564,265,292]" pageId="3" pageNumber="19">25.03/4234</specimenCode>
(
<figureCitation id="CFB09AC3FFE5CC38FDB2FED016E60848" box="[579,674,265,291]" captionStart="Fig" captionStartId="2.[189,232,1894,1920]" captionTargetBox="[189,1399,617,1854]" captionTargetId="figure-132@2.[189,1399,617,1854]" captionTargetPageId="2" captionText="Fig. 1. Cradoscrupocellaria severoi sp. nov. A. View of the colony (MNCN 25.03/3059: holotype). B. View of the colony (MNCN 25.03/3200: paratype). C. Branching pattern (MNCN 25.03/4234: paratype). D. Detail of a branch with large frontal avicularia, ovicells and scuta (MNCN 25.03/3059: holotype)." figureDoi="http://doi.org/10.5281/zenodo.5121752" httpUri="https://zenodo.org/record/5121752/files/figure.png" pageId="3" pageNumber="19">Figs 1C</figureCitation>
, 
<figureCitation id="CFB09AC3FFE5CC38FD5EFED016960848" box="[687,722,265,291]" captionStart="Fig" captionStartId="3.[191,234,1857,1883]" captionTargetBox="[189,1399,773,1816]" captionTargetId="figure-181@3.[189,1399,773,1816]" captionTargetPageId="3" captionText="Fig. 2. Cradoscrupocellaria severoi sp. nov. A. Detail of branches with large frontal avicularia, ovicells, scuta and a lateral avicularium (centre left) (MNCN 25.03/3059: holotype). B. Detail of a branch with large frontal avicularia, ovicells and a lateral avicularium (bottom left) (MNCN 25.03/3200: paratype). C. Basal view of the branching pattern (MNCN 25.03/4234: paratype). D. Single axial vibraculum (MNCN 25.03/3059: holotype)." figureDoi="http://doi.org/10.5281/zenodo.5121754" httpUri="https://zenodo.org/record/5121754/files/figure.png" pageId="3" pageNumber="19">2C</figureCitation>
)
</materialsCitation>
• 
<materialsCitation id="E7E38C1BFFE5CC38FD05FED01132082D" ID-GBIF-Occurrence="3326570301" collectionCode="MNCN" elevation="35" location="Columbretes Islands" municipality="Columbretes Islands" pageId="3" pageNumber="19" specimenCode="MNCN 25.03/3200" specimenCount="1">
<specimenCount id="418D4DCFFFE5CC38FD05FED017830848" box="[756,967,265,291]" pageId="3" pageNumber="19">several fragments</specimenCount>
; 
<collectingMunicipality id="B7501C3CFFE5CC38FC22FED010830848" box="[979,1223,265,291]" pageId="3" pageNumber="19">Columbretes Islands</collectingMunicipality>
; 
<geoCoordinate id="32BFE081FFE5CC38FB23FED011360848" box="[1234,1394,265,291]" pageId="3" pageNumber="19">39º89.800´N</geoCoordinate>
, 
<geoCoordinate id="32BFE081FFE5CC38FF4CFEF5150F082C" box="[189,331,300,327]" pageId="3" pageNumber="19">0º68.883´E</geoCoordinate>
; stn 275B5; depth 
<quantity id="90732BA3FFE5CC38FDD7FEF51621082C" box="[550,613,300,327]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="19" unit="m" value="35.0">
<elevation id="DCA66175FFE5CC38FDD7FEF51621082C" box="[550,613,300,327]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="19" unit="m" value="35.0">35 m</elevation>
</quantity>
; 1996; Fauna Ibérica IV exped.; 
<specimenCode id="072D2E3DFFE5CC38FC15FEF5108C082C" box="[996,1224,300,327]" pageId="3" pageNumber="19">MNCN 25.03/3200</specimenCode>
(
<figureCitation id="CFB09AC3FFE5CC38FB26FEF51172082D" box="[1239,1334,300,326]" captionStart="Fig" captionStartId="2.[189,232,1894,1920]" captionTargetBox="[189,1399,617,1854]" captionTargetId="figure-132@2.[189,1399,617,1854]" captionTargetPageId="2" captionText="Fig. 1. Cradoscrupocellaria severoi sp. nov. A. View of the colony (MNCN 25.03/3059: holotype). B. View of the colony (MNCN 25.03/3200: paratype). C. Branching pattern (MNCN 25.03/4234: paratype). D. Detail of a branch with large frontal avicularia, ovicells and scuta (MNCN 25.03/3059: holotype)." figureDoi="http://doi.org/10.5281/zenodo.5121752" httpUri="https://zenodo.org/record/5121752/files/figure.png" pageId="3" pageNumber="19">Figs 1B</figureCitation>
, 
<figureCitation id="CFB09AC3FFE5CC38FAB2FEF5112C082D" box="[1347,1384,300,326]" captionStart="Fig" captionStartId="3.[191,234,1857,1883]" captionTargetBox="[189,1399,773,1816]" captionTargetId="figure-181@3.[189,1399,773,1816]" captionTargetPageId="3" captionText="Fig. 2. Cradoscrupocellaria severoi sp. nov. A. Detail of branches with large frontal avicularia, ovicells, scuta and a lateral avicularium (centre left) (MNCN 25.03/3059: holotype). B. Detail of a branch with large frontal avicularia, ovicells and a lateral avicularium (bottom left) (MNCN 25.03/3200: paratype). C. Basal view of the branching pattern (MNCN 25.03/4234: paratype). D. Single axial vibraculum (MNCN 25.03/3059: holotype)." figureDoi="http://doi.org/10.5281/zenodo.5121754" httpUri="https://zenodo.org/record/5121754/files/figure.png" pageId="3" pageNumber="19">2B</figureCitation>
).
</materialsCitation>
</paragraph>
<caption id="03F4D6CEFFE4CC39FF4CF8BF15730E80" ID-DOI="http://doi.org/10.5281/zenodo.5121752" ID-Zenodo-Dep="5121752" httpUri="https://zenodo.org/record/5121752/files/figure.png" pageId="2" pageNumber="18" startId="2.[189,232,1894,1920]" targetBox="[189,1399,617,1854]" targetPageId="2">
<paragraph id="57348646FFE4CC39FF4CF8BF15730E80" blockId="2.[189,1399,1894,2027]" pageId="2" pageNumber="18">
<emphasis id="65FF5A54FFE4CC39FF4CF8BF154A0EEB" bold="true" box="[189,270,1894,1920]" pageId="2" pageNumber="18">Fig. 1.</emphasis>
<taxonomicName id="908BFDC5FFE4CC39FEE9F8BE16290EEB" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[280,621,1895,1921]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="18" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE4CC39FEE9F8BE16290EEB" box="[280,621,1895,1921]" italics="true" pageId="2" pageNumber="18">Cradoscrupocellaria severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE4CC39FD86F8BE16910EEA" box="[631,725,1895,1921]" pageId="2" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="65FF5A54FFE4CC39FD2EF8BF16B00EEB" bold="true" box="[735,756,1894,1920]" pageId="2" pageNumber="18">A</emphasis>
. View of the colony (MNCN 25.03/3059: holotype). 
<emphasis id="65FF5A54FFE4CC39FF4CF85314950ECF" bold="true" box="[189,209,1930,1956]" pageId="2" pageNumber="18">B</emphasis>
. View of the colony (MNCN 25.03/3200: paratype). 
<emphasis id="65FF5A54FFE4CC39FC9DF85317C50ECF" bold="true" box="[876,897,1930,1956]" pageId="2" pageNumber="18">C</emphasis>
. Branching pattern (MNCN 25.03/4234: paratype). 
<emphasis id="65FF5A54FFE4CC39FECDF87415150EAC" bold="true" box="[316,337,1965,1991]" pageId="2" pageNumber="18">D</emphasis>
. Detail of a branch with large frontal avicularia, ovicells and scuta (MNCN 25.03/3059: holotype).
</paragraph>
</caption>
<paragraph id="57348646FFE5CC38FF1BFEB415E208EC" blockId="3.[189,1399,365,428]" box="[234,422,365,391]" pageId="3" pageNumber="19">
<emphasis id="65FF5A54FFE5CC38FF1BFEB415E208EC" bold="true" box="[234,422,365,391]" pageId="3" pageNumber="19">Other material</emphasis>
</paragraph>
<paragraph id="57348646FFE5CC38FF4CFE48113308C7" blockId="3.[189,1399,365,428]" box="[189,1399,401,428]" pageId="3" pageNumber="19">
<typeStatus id="883038E4FFE5CC38FF4CFE48152C08C0" box="[189,360,401,427]" pageId="3" pageNumber="19" type="paralectotype">Paralectotypes</typeStatus>
of 
<taxonomicName id="908BFDC5FFE5CC38FE7AFE4817D408C0" authority="Gautier, 1962" authorityName="Gautier" authorityYear="1962" box="[395,912,401,427]" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="3" pageNumber="19" phylum="Bryozoa" rank="species" species="macrorhyncha">
<emphasis id="65FF5A54FFE5CC38FE7AFE4816AD08C0" box="[395,745,401,427]" italics="true" pageId="3" pageNumber="19">Scrupocellaria macrorhyncha</emphasis>
<bibRefCitation id="331AFBB7FFE5CC38FD01FE4817D408C0" author="Gautier Y. - V." box="[752,912,401,427]" pageId="3" pageNumber="19" pagination="1 - 435" refId="ref6642" refString="Gautier Y. - V. 1962. Recherches ecologiques sur les Bryozoaires Chilostomes en Mediterranee occidentale. Recueil des Travaux de la Station marine d'Endoume 38: 1 - 435." type="journal article" year="1962">Gautier, 1962</bibRefCitation>
</taxonomicName>
(here designated, see explanation below):
</paragraph>
<paragraph id="57348646FFE5CC38FF4CFE0B159C0B58" blockId="3.[188,1399,466,563]" pageId="3" pageNumber="19">
<materialsCitation id="E7E38C1BFFE5CC38FF4CFE0B162D0B7B" ID-GBIF-Occurrence="3326570306" collectingDate="1952-12-01" collectorName="Gautier" location="MEDITERRANEAN FRANCE" municipality="Marseille" pageId="3" pageNumber="19" specimenCode="MNHN-IB-2008-11319" specimenCount="1">
<location id="5254D09DFFE5CC38FF4CFE0B16700887" LSID="urn:lsid:plazi:treatment:DF223750FFE7CC3DFDCEFAD6156B0BA5:5254D09DFFE5CC38FF4CFE0B16700887" box="[189,564,466,492]" municipality="Marseille" name="MEDITERRANEAN FRANCE" pageId="3" pageNumber="19">MEDITERRANEAN FRANCE</location>
• 
<specimenCount id="418D4DCFFFE5CC38FDBBFE0B17580887" box="[586,796,466,492]" pageId="3" pageNumber="19">several fragments</specimenCount>
; 
<collectingMunicipality id="B7501C3CFFE5CC38FCD6FE0B17DD0887" box="[807,921,466,492]" pageId="3" pageNumber="19">Marseille</collectingMunicipality>
, 
<locationDeviation id="46869F64FFE5CC38FC52FE0B10300887" box="[931,1140,466,492]" pageId="3" pageNumber="19">SW Cap Caveaux</locationDeviation>
; stn 258; 
<date id="2335A086FFE5CC38FB13FE0B11360886" box="[1250,1394,466,493]" pageId="3" pageNumber="19" value="1952-12-01">
<collectingDate id="3371596EFFE5CC38FB13FE0B11360886" box="[1250,1394,466,493]" pageId="3" pageNumber="19" value="1952-12-01">1 Dec. 1952</collectingDate>
</date>
; 
<collectorName id="FA7EE390FFE5CC38FF4DFE2C15500B64" box="[188,276,501,527]" pageId="3" pageNumber="19">Gautier</collectorName>
leg.; 
<specimenCode id="072D2E3DFFE5CC38FEA3FE2F162D0B7B" box="[338,617,502,528]" pageId="3" pageNumber="19">MNHN-IB-2008-11319</specimenCode>
</materialsCitation>
• 
<materialsCitation id="E7E38C1BFFE5CC38FD8FFE2C159C0B58" ID-GBIF-Occurrence="3326570316" collectingDate="1952-12-01" collectionCode="MNHN" collectorName="Gautier" location="MEDITERRANEAN FRANCE" municipality="Marseille" pageId="3" pageNumber="19" specimenCode="MNHN- IB-2008-11321" specimenCount="1">
<specimenCount id="418D4DCFFFE5CC38FD8FFE2C17140B64" box="[638,848,501,527]" pageId="3" pageNumber="19">several fragments</specimenCount>
; same collection data as for preceding; 
<specimenCode id="072D2E3DFFE5CC38FAE3FE2F15340B58" pageId="3" pageNumber="19">MNHN- IB-2008-11321</specimenCode>
(
<figureCitation id="CFB09AC3FFE5CC38FE70FDC0158C0B58" box="[385,456,537,563]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[188,1398,911,1804]" captionTargetId="figure-190@5.[188,1398,911,1804]" captionTargetPageId="5" captionText="Fig. 3. Cradoscrupocellaria macrorhyncha (Gautier, 1962) (MNHN-IB-2008-11321: paralectotype), SW Cap Caveaux (Marseille, France), Gautier Coll. A. Frontal view of the branching pattern. B. Detail showing the scuta, ovicells, large frontal avicularia and a small frontal avicularium (bottom right). C. Lateral view showing the large frontal avicularia and basal vibracula. D. Basal view of the branching pattern." figureDoi="http://doi.org/10.5281/zenodo.5121756" httpUri="https://zenodo.org/record/5121756/files/figure.png" pageId="3" pageNumber="19">Fig. 3</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1F91D5CDFFE5CC3FFF4CFD8316EB0A76" lastPageId="4" lastPageNumber="20" pageId="3" pageNumber="19" type="description">
<paragraph id="57348646FFE5CC38FF4CFD83150A0B1F" blockId="3.[189,334,602,628]" box="[189,334,602,628]" pageId="3" pageNumber="19">
<heading id="0C7C312AFFE5CC38FF4CFD83150A0B1F" bold="true" box="[189,334,602,628]" fontSize="11" level="3" pageId="3" pageNumber="19" reason="3">
<emphasis id="65FF5A54FFE5CC38FF4CFD83150A0B1F" bold="true" box="[189,334,602,628]" pageId="3" pageNumber="19">Description</emphasis>
</heading>
</paragraph>
<paragraph id="57348646FFE5CC38FF4CFD5011370BAD" blockId="3.[186,1398,648,710]" pageId="3" pageNumber="19">
Colony erect, up to 
<quantity id="90732BA3FFE5CC38FE52FD50159F0BC8" box="[419,475,649,675]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="19" unit="cm" value="1.0">1 cm</quantity>
in height, branched, with internodes comprising 5–10 alternating autozooids. Internodes slender, almost straight, angled at axis, with acute bifurcating pattern; chitinous joints
</paragraph>
<caption id="03F4D6CEFFE5CC38FF4EF89816770E82" ID-DOI="http://doi.org/10.5281/zenodo.5121754" ID-Zenodo-Dep="5121754" httpUri="https://zenodo.org/record/5121754/files/figure.png" pageId="3" pageNumber="19" startId="3.[191,234,1857,1883]" targetBox="[189,1399,773,1816]" targetPageId="3">
<paragraph id="57348646FFE5CC38FF4EF89816770E82" blockId="3.[188,1400,1857,2025]" pageId="3" pageNumber="19">
<emphasis id="65FF5A54FFE5CC38FF4EF898154F0E30" bold="true" box="[191,267,1857,1883]" pageId="3" pageNumber="19">Fig. 2.</emphasis>
<taxonomicName id="908BFDC5FFE5CC38FEE1F89816240E30" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[272,608,1857,1883]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="3" pageNumber="19" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE5CC38FEE1F89816240E30" box="[272,608,1857,1883]" italics="true" pageId="3" pageNumber="19">Cradoscrupocellaria severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE5CC38FD97F89B16FB0E37" box="[614,703,1858,1884]" pageId="3" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="65FF5A54FFE5CC38FD35F898169D0E30" bold="true" box="[708,729,1857,1883]" pageId="3" pageNumber="19">A</emphasis>
. Detail of branches with large frontal avicularia, ovicells, scuta and a lateral avicularium (centre left) (MNCN 25.03/3059: holotype). 
<emphasis id="65FF5A54FFE5CC38FBB3F8BC10110E14" bold="true" box="[1090,1109,1893,1919]" pageId="3" pageNumber="19">B</emphasis>
. Detail of a branch with large frontal avicularia, ovicells and a lateral avicularium (bottom left) (MNCN 25.03/3200: paratype). 
<emphasis id="65FF5A54FFE5CC38FF4CF87214960EAE" bold="true" box="[189,210,1963,1989]" pageId="3" pageNumber="19">C</emphasis>
. Basal view of the branching pattern (MNCN 25.03/4234: paratype). 
<emphasis id="65FF5A54FFE5CC38FBDFF87210070EAE" bold="true" box="[1070,1091,1963,1989]" pageId="3" pageNumber="19">D</emphasis>
. Single axial vibraculum (MNCN 25.03/3059: holotype).
</paragraph>
</caption>
<paragraph id="57348646FFE2CC3FFF4CFED016EB0A76" blockId="4.[188,1400,265,797]" pageId="4" pageNumber="20">
passing across proximal gymnocyst of outer (C and D) and inner zooids (F and G) at bifurcation. Autozooids cylindrical with straight sides. Oval opesia occupying two thirds of zooidal length; cryptocyst reduced to narrow rim around opesia. Large scutum inserted at midline of the inner edge of opesia, stout but relatively flat, its tips highly branched, fully developed and completely covering entire opesia. Distal spines short, unbranched; 2 inner and 3 outer spines, with outer distal spines closer to scutum; axial zooid with 5 spines. Small distolateral avicularium frequent and easily visible in frontal view. Monomorphic frontal avicularium often present in outer zooids of each internode; large but variable in size, with elongate rostrum 
<quantity id="90732BA3FFE2CC3FFC7EFDDF107B0B4B" box="[911,1087,518,544]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="4" pageNumber="20" unit="mm" value="0.25" valueMax="0.28" valueMin="0.22">0.22–0.28 mm</quantity>
long, with fringed edges, proximally upright, with strongly hooked tip; mandible long, hooked distally. Vibracular chamber laterally placed on the basal surface of each zooid, visible in frontal view; chamber of vibraculum almost trapezoidal, with a proximal rhizoidal foramen; setal groove transverse to internode axis, straight, with smooth seta longer than one autozooid. Single axial vibraculum without rhizoidal foramen. Rhizoids tubular and smooth. Ovicells globular, wider than long, with an even, almost straight proximal rim; ectooecium perforated by 10–18 rounded pores; ovicelled zooids with 2 inner and 2 outer spines. Ancestrula unknown.
</paragraph>
</subSubSection>
<subSubSection id="1F91D5CDFFE2CC3DFF4CFC9C156B0BA5" lastPageId="6" lastPageNumber="22" pageId="4" pageNumber="20" type="discussion">
<paragraph id="57348646FFE2CC3FFF4CFC9C156B0A34" blockId="4.[189,303,837,863]" box="[189,303,837,863]" pageId="4" pageNumber="20">
<heading id="0C7C312AFFE2CC3FFF4CFC9C156B0A34" bold="true" box="[189,303,837,863]" fontSize="11" level="3" pageId="4" pageNumber="20" reason="3">
<emphasis id="65FF5A54FFE2CC3FFF4CFC9C156B0A34" bold="true" box="[189,303,837,863]" pageId="4" pageNumber="20">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="57348646FFE2CC3FFF4CFCAD10420F72" blockId="4.[186,1400,884,1561]" pageId="4" pageNumber="20">
The genus 
<taxonomicName id="908BFDC5FFE2CC3FFEB9FCAD160D0AE5" authorityName="Vieira, Spencer Jones &amp; Winston" authorityYear="2013" box="[328,585,884,910]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="genus">
<emphasis id="65FF5A54FFE2CC3FFEB9FCAD160D0AE5" box="[328,585,884,910]" italics="true" pageId="4" pageNumber="20">Cradoscrupocellaria</emphasis>
</taxonomicName>
has been recently described by 
<bibRefCitation id="331AFBB7FFE2CC3FFC29FCAD108E0AE4" author="Vieira L. M. &amp; Spencer Jones M. E. &amp; Winston J. E." box="[984,1226,884,911]" pageId="4" pageNumber="20" pagination="1 - 63" refId="ref8324" refString="Vieira L. M., Spencer Jones M. E. &amp; Winston J. E. 2013. Cradoscrupocellaria, a new bryozoan genus for Scrupocellaria bertholletii (Audouin) and related species (Cheilostomata, Candidae): taxonomy, biodiversity and distribution. Zootaxa 3707 (1): 1 - 63. https: // doi. org / 10.11646 / zootaxa. 3707.1.1" type="journal article" year="2013">
Vieira 
<emphasis id="65FF5A54FFE2CC3FFBDFFCAC10290AE5" box="[1070,1133,884,910]" italics="true" pageId="4" pageNumber="20">et al.</emphasis>
(2013)
</bibRefCitation>
. Nine already existing species were transferred to this genus, and the same authors described 18 new species.Among all these species, only 
<taxonomicName id="908BFDC5FFE2CC3FFE2DFC64101A0ABC" authority="Vieira, Spencer Jones &amp; Winston, 2013" authorityName="Vieira, Spencer Jones &amp; Winston" authorityYear="2013" box="[476,1118,957,983]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="gautieri">
<emphasis id="65FF5A54FFE2CC3FFE2DFC6416200ABD" box="[476,612,957,983]" italics="true" pageId="4" pageNumber="20">C. gautieri</emphasis>
<bibRefCitation id="331AFBB7FFE2CC3FFD9EFC64101A0ABC" author="Vieira L. M. &amp; Spencer Jones M. E. &amp; Winston J. E." box="[623,1118,957,983]" pageId="4" pageNumber="20" pagination="1 - 63" refId="ref8324" refString="Vieira L. M., Spencer Jones M. E. &amp; Winston J. E. 2013. Cradoscrupocellaria, a new bryozoan genus for Scrupocellaria bertholletii (Audouin) and related species (Cheilostomata, Candidae): taxonomy, biodiversity and distribution. Zootaxa 3707 (1): 1 - 63. https: // doi. org / 10.11646 / zootaxa. 3707.1.1" type="journal article" year="2013">Vieira, Spencer Jones &amp; Winston, 2013</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="908BFDC5FFE2CC3FFB51FC64153E0A97" authority="(Gautier, 1962)" baseAuthorityName="Gautier" baseAuthorityYear="1962" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="macrorhyncha">
<emphasis id="65FF5A54FFE2CC3FFB51FC6411330ABC" box="[1184,1399,957,983]" italics="true" pageId="4" pageNumber="20">C. macrorhyncha</emphasis>
(
<bibRefCitation id="331AFBB7FFE2CC3FFF34FC3815350A97" author="Gautier Y. - V." box="[197,369,993,1020]" pageId="4" pageNumber="20" pagination="1 - 435" refId="ref6642" refString="Gautier Y. - V. 1962. Recherches ecologiques sur les Bryozoaires Chilostomes en Mediterranee occidentale. Recueil des Travaux de la Station marine d'Endoume 38: 1 - 435." type="journal article" year="1962">Gautier, 1962</bibRefCitation>
)
</taxonomicName>
show similarities with 
<taxonomicName id="908BFDC5FFE2CC3FFD6FFC38175C0A90" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[670,792,993,1019]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE2CC3FFD6FFC38175C0A90" box="[670,792,993,1019]" italics="true" pageId="4" pageNumber="20">C. severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE2CC3FFCD0FC3817C70A90" box="[801,899,993,1019]" pageId="4" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
, mainly regarding the very stout scutum with truncate tips and the large frontal avicularia. The description of 
<taxonomicName id="908BFDC5FFE2CC3FFBF1FBDC10C00D74" authorityName="Vieira, Spencer Jones &amp; Winston" authorityYear="2013" box="[1024,1156,1029,1055]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="gautieri">
<emphasis id="65FF5A54FFE2CC3FFBF1FBDC10C00D74" box="[1024,1156,1029,1055]" italics="true" pageId="4" pageNumber="20">C. gautieri</emphasis>
</taxonomicName>
is based on a single sample collected by 
<bibRefCitation id="331AFBB7FFE2CC3FFE45FBF016230D2F" author="Gautier Y. - V." box="[436,615,1065,1092]" pageId="4" pageNumber="20" pagination="1 - 435" refId="ref6642" refString="Gautier Y. - V. 1962. Recherches ecologiques sur les Bryozoaires Chilostomes en Mediterranee occidentale. Recueil des Travaux de la Station marine d'Endoume 38: 1 - 435." type="journal article" year="1962">Gautier (1962)</bibRefCitation>
in 
<collectingCountry id="2F9CC6D6FFE2CC3FFD7AFBF016AF0D28" box="[651,747,1065,1091]" name="Algeria" pageId="4" pageNumber="20">Algeria</collectingCountry>
. 
<taxonomicName id="908BFDC5FFE2CC3FFD05FBF010E90D28" baseAuthorityName="Gautier" baseAuthorityYear="1962" box="[756,1197,1065,1091]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="macrorhyncha">
<emphasis id="65FF5A54FFE2CC3FFD05FBF010E90D28" box="[756,1197,1065,1091]" italics="true" pageId="4" pageNumber="20">Cradoscrupocellaria macrorhyncha</emphasis>
</taxonomicName>
was redescribed by 
<bibRefCitation id="331AFBB7FFE2CC3FFF2EFB9415870D0C" author="Vieira L. M. &amp; Spencer Jones M. E. &amp; Winston J. E." box="[223,451,1101,1127]" pageId="4" pageNumber="20" pagination="1 - 63" refId="ref8324" refString="Vieira L. M., Spencer Jones M. E. &amp; Winston J. E. 2013. Cradoscrupocellaria, a new bryozoan genus for Scrupocellaria bertholletii (Audouin) and related species (Cheilostomata, Candidae): taxonomy, biodiversity and distribution. Zootaxa 3707 (1): 1 - 63. https: // doi. org / 10.11646 / zootaxa. 3707.1.1" type="journal article" year="2013">
Vieira 
<emphasis id="65FF5A54FFE2CC3FFEDFFB97152D0D0C" box="[302,361,1101,1127]" italics="true" pageId="4" pageNumber="20">et al.</emphasis>
(2013)
</bibRefCitation>
based on the only original sample of Gautier deposited at NHMUK, designated as 
<typeStatus id="883038E4FFE2CC3FFF2FFBA815160DE0" box="[222,338,1137,1163]" pageId="4" pageNumber="20" type="lectotype">lectotype</typeStatus>
, and other Mediterranean samples by other authors. However, the MNHN collection includes two other original samples of Gautier from the same station as the 
<typeStatus id="883038E4FFE2CC3FFB99FB4C109C0DC4" box="[1128,1240,1173,1199]" pageId="4" pageNumber="20" type="lectotype">lectotype</typeStatus>
that seem to have been overlooked in the redescription of the species, perhaps because in the museum’s catalogue and in the external labels of the samples both are simply marked as “ 
<taxonomicName id="908BFDC5FFE2CC3FFBB2FB0711240D9C" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1091,1376,1246,1272]" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="reptans">
<emphasis id="65FF5A54FFE2CC3FFBB2FB0711240D9C" box="[1091,1376,1246,1272]" italics="true" pageId="4" pageNumber="20">Scrupocellaria reptans</emphasis>
</taxonomicName>
”: sample MNHN-IB-2008-11319 contains a handwritten label by Gautier stating “ 
<taxonomicName id="908BFDC5FFE2CC3FFB31FADB15530C2B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="reptans">
<emphasis id="65FF5A54FFE2CC3FFB31FADB15530C2B" italics="true" pageId="4" pageNumber="20">Scrupocellaria reptans</emphasis>
</taxonomicName>
group? Dry specimen”; being written in English, it is highly probable that this sample together with the 
<typeStatus id="883038E4FFE2CC3FFE64FA9316410C0F" box="[405,517,1354,1380]" pageId="4" pageNumber="20" type="lectotype">lectotype</typeStatus>
were sent by Gautier to A. Hastings at the NHMUK, and later returned. Sample MNHN-IB-2008-11321 (
<figureCitation id="CFB09AC3FFE2CC3FFD96FAB716FD0CE3" box="[615,697,1390,1416]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[188,1398,911,1804]" captionTargetId="figure-190@5.[188,1398,911,1804]" captionTargetPageId="5" captionText="Fig. 3. Cradoscrupocellaria macrorhyncha (Gautier, 1962) (MNHN-IB-2008-11321: paralectotype), SW Cap Caveaux (Marseille, France), Gautier Coll. A. Frontal view of the branching pattern. B. Detail showing the scuta, ovicells, large frontal avicularia and a small frontal avicularium (bottom right). C. Lateral view showing the large frontal avicularia and basal vibracula. D. Basal view of the branching pattern." figureDoi="http://doi.org/10.5281/zenodo.5121756" httpUri="https://zenodo.org/record/5121756/files/figure.png" pageId="4" pageNumber="20">Fig. 3</figureCitation>
) includes a label handwritten by Gautier indicating “ 
<taxonomicName id="908BFDC5FFE2CC3FFF3AFA4B15A60CC7" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[203,482,1426,1452]" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="reptans">
<emphasis id="65FF5A54FFE2CC3FFF3AFA4B15A60CC7" box="[203,482,1426,1452]" italics="true" pageId="4" pageNumber="20">Scrupocellaria reptans</emphasis>
</taxonomicName>
? grand avic.”. Since both samples actually correspond to 
<taxonomicName id="908BFDC5FFE2CC3FFB52FA4A11320CC7" baseAuthorityName="Gautier" baseAuthorityYear="1962" box="[1187,1398,1426,1453]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="macrorhyncha">
<emphasis id="65FF5A54FFE2CC3FFB52FA4A11320CC7" box="[1187,1398,1426,1453]" italics="true" pageId="4" pageNumber="20">C. macrorhyncha</emphasis>
</taxonomicName>
and were collected by Gautier at the type locality of the species and studied by him, they should be considered as part of the type series and consequently considered as 
<typeStatus id="883038E4FFE2CC3FFBE2FA0310800C9F" box="[1043,1220,1498,1524]" pageId="4" pageNumber="20" type="paralectotype">paralectotypes</typeStatus>
of the species according to Arts 73.2.2 and 74.1.3 of the ICZN Code (
<bibRefCitation id="331AFBB7FFE2CC3FFC97FA2617BD0F72" author="ICZN" box="[870,1017,1535,1561]" pageId="4" pageNumber="20" refId="ref7144" refString="ICZN. 1999. International Code of Zoological Nomenclature. International Trust for Zoological Nomenclature, London." type="book" year="1999">ICZN 1999</bibRefCitation>
).
</paragraph>
<paragraph id="57348646FFE2CC3FFF4CF99816B10E83" blockId="4.[187,1400,1601,2025]" pageId="4" pageNumber="20">
<taxonomicName id="908BFDC5FFE2CC3FFF4CF99816480F30" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[189,524,1601,1627]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE2CC3FFF4CF99816480F30" box="[189,524,1601,1627]" italics="true" pageId="4" pageNumber="20">Cradoscrupocellaria severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE2CC3FFDE0F99B162D0F37" box="[529,617,1602,1628]" pageId="4" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="908BFDC5FFE2CC3FFD0AF99817800F30" baseAuthorityName="Gautier" baseAuthorityYear="1962" box="[763,964,1601,1627]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="macrorhyncha">
<emphasis id="65FF5A54FFE2CC3FFD0AF99817800F30" box="[763,964,1601,1627]" italics="true" pageId="4" pageNumber="20">C. macrorhyncha</emphasis>
</taxonomicName>
and 
<taxonomicName id="908BFDC5FFE2CC3FFC08F99810320F30" authorityName="Vieira, Spencer Jones &amp; Winston" authorityYear="2013" box="[1017,1142,1601,1627]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="gautieri">
<emphasis id="65FF5A54FFE2CC3FFC08F99810320F30" box="[1017,1142,1601,1627]" italics="true" pageId="4" pageNumber="20">C. gautieri</emphasis>
</taxonomicName>
by several characters: the frontal avicularium is monomorphic in 
<taxonomicName id="908BFDC5FFE2CC3FFD4AF9BF17760F14" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[699,818,1638,1664]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE2CC3FFD4AF9BF17760F14" box="[699,818,1638,1664]" italics="true" pageId="4" pageNumber="20">C. severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE2CC3FFCC8F9BF17D00FEB" box="[825,916,1638,1664]" pageId="4" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
because the small columnar avicularium present in the other species is absent; this frontal avicularium is oriented upwards on the branch, not proximally (
<figureCitation id="CFB09AC3FFE2CC3FFEBEF97415970FAC" box="[335,467,1709,1735]" captionStart="Fig" captionStartId="3.[191,234,1857,1883]" captionTargetBox="[189,1399,773,1816]" captionTargetId="figure-181@3.[189,1399,773,1816]" captionTargetPageId="3" captionText="Fig. 2. Cradoscrupocellaria severoi sp. nov. A. Detail of branches with large frontal avicularia, ovicells, scuta and a lateral avicularium (centre left) (MNCN 25.03/3059: holotype). B. Detail of a branch with large frontal avicularia, ovicells and a lateral avicularium (bottom left) (MNCN 25.03/3200: paratype). C. Basal view of the branching pattern (MNCN 25.03/4234: paratype). D. Single axial vibraculum (MNCN 25.03/3059: holotype)." figureDoi="http://doi.org/10.5281/zenodo.5121754" httpUri="https://zenodo.org/record/5121754/files/figure.png" pageId="4" pageNumber="20">Figs 2A–B</figureCitation>
, 
<figureCitation id="CFB09AC3FFE2CC3FFE2EF97416410FAC" box="[479,517,1709,1735]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[188,1398,911,1804]" captionTargetId="figure-190@5.[188,1398,911,1804]" captionTargetPageId="5" captionText="Fig. 3. Cradoscrupocellaria macrorhyncha (Gautier, 1962) (MNHN-IB-2008-11321: paralectotype), SW Cap Caveaux (Marseille, France), Gautier Coll. A. Frontal view of the branching pattern. B. Detail showing the scuta, ovicells, large frontal avicularia and a small frontal avicularium (bottom right). C. Lateral view showing the large frontal avicularia and basal vibracula. D. Basal view of the branching pattern." figureDoi="http://doi.org/10.5281/zenodo.5121756" httpUri="https://zenodo.org/record/5121756/files/figure.png" pageId="4" pageNumber="20">3C</figureCitation>
). The lateral avicularium is more frequent and more visible (
<figureCitation id="CFB09AC3FFE2CC3FFB3AF97411080FAC" box="[1227,1356,1709,1735]" captionStart="Fig" captionStartId="3.[191,234,1857,1883]" captionTargetBox="[189,1399,773,1816]" captionTargetId="figure-181@3.[189,1399,773,1816]" captionTargetPageId="3" captionText="Fig. 2. Cradoscrupocellaria severoi sp. nov. A. Detail of branches with large frontal avicularia, ovicells, scuta and a lateral avicularium (centre left) (MNCN 25.03/3059: holotype). B. Detail of a branch with large frontal avicularia, ovicells and a lateral avicularium (bottom left) (MNCN 25.03/3200: paratype). C. Basal view of the branching pattern (MNCN 25.03/4234: paratype). D. Single axial vibraculum (MNCN 25.03/3059: holotype)." figureDoi="http://doi.org/10.5281/zenodo.5121754" httpUri="https://zenodo.org/record/5121754/files/figure.png" pageId="4" pageNumber="20">Fig. 2A–B</figureCitation>
). In the bifurcations of 
<taxonomicName id="908BFDC5FFE2CC3FFE50F90B165C0F80" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[417,536,1746,1772]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE2CC3FFE50F90B165C0F80" box="[417,536,1746,1772]" italics="true" pageId="4" pageNumber="20">C. severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE2CC3FFDEEF90B163A0F87" box="[543,638,1746,1772]" pageId="4" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
, the chitinous joints pass through the gymnocyst of the external zooids (C and D) instead of through their opesiae (
<figureCitation id="CFB09AC3FFE2CC3FFCC0F92F17D00E7B" box="[817,916,1782,1808]" captionStart="Fig" captionStartId="2.[189,232,1894,1920]" captionTargetBox="[189,1399,617,1854]" captionTargetId="figure-132@2.[189,1399,617,1854]" captionTargetPageId="2" captionText="Fig. 1. Cradoscrupocellaria severoi sp. nov. A. View of the colony (MNCN 25.03/3059: holotype). B. View of the colony (MNCN 25.03/3200: paratype). C. Branching pattern (MNCN 25.03/4234: paratype). D. Detail of a branch with large frontal avicularia, ovicells and scuta (MNCN 25.03/3059: holotype)." figureDoi="http://doi.org/10.5281/zenodo.5121752" httpUri="https://zenodo.org/record/5121752/files/figure.png" pageId="4" pageNumber="20">Figs 1C</figureCitation>
, 
<figureCitation id="CFB09AC3FFE2CC3FFC54F92F17880E7B" box="[933,972,1782,1808]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[188,1398,911,1804]" captionTargetId="figure-190@5.[188,1398,911,1804]" captionTargetPageId="5" captionText="Fig. 3. Cradoscrupocellaria macrorhyncha (Gautier, 1962) (MNHN-IB-2008-11321: paralectotype), SW Cap Caveaux (Marseille, France), Gautier Coll. A. Frontal view of the branching pattern. B. Detail showing the scuta, ovicells, large frontal avicularia and a small frontal avicularium (bottom right). C. Lateral view showing the large frontal avicularia and basal vibracula. D. Basal view of the branching pattern." figureDoi="http://doi.org/10.5281/zenodo.5121756" httpUri="https://zenodo.org/record/5121756/files/figure.png" pageId="4" pageNumber="20">3A</figureCitation>
). The ovicell is wider than long in 
<taxonomicName id="908BFDC5FFE2CC3FFF4CF8C315730E58" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[189,311,1818,1844]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE2CC3FFF4CF8C315730E58" box="[189,311,1818,1844]" italics="true" pageId="4" pageNumber="20">C. severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE2CC3FFEB2F8C315E00E5F" box="[323,420,1818,1844]" pageId="4" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
, usually uniformly perforated by small pseudopores (
<figureCitation id="CFB09AC3FFE2CC3FFBC2F8C310DC0E5F" box="[1075,1176,1818,1844]" captionStart="Fig" captionStartId="2.[189,232,1894,1920]" captionTargetBox="[189,1399,617,1854]" captionTargetId="figure-132@2.[189,1399,617,1854]" captionTargetPageId="2" captionText="Fig. 1. Cradoscrupocellaria severoi sp. nov. A. View of the colony (MNCN 25.03/3059: holotype). B. View of the colony (MNCN 25.03/3200: paratype). C. Branching pattern (MNCN 25.03/4234: paratype). D. Detail of a branch with large frontal avicularia, ovicells and scuta (MNCN 25.03/3059: holotype)." figureDoi="http://doi.org/10.5281/zenodo.5121752" httpUri="https://zenodo.org/record/5121752/files/figure.png" pageId="4" pageNumber="20">Figs 1D</figureCitation>
, 
<figureCitation id="CFB09AC3FFE2CC3FFB5BF8C310B70E5F" box="[1194,1267,1818,1844]" captionStart="Fig" captionStartId="3.[191,234,1857,1883]" captionTargetBox="[189,1399,773,1816]" captionTargetId="figure-181@3.[189,1399,773,1816]" captionTargetPageId="3" captionText="Fig. 2. Cradoscrupocellaria severoi sp. nov. A. Detail of branches with large frontal avicularia, ovicells, scuta and a lateral avicularium (centre left) (MNCN 25.03/3059: holotype). B. Detail of a branch with large frontal avicularia, ovicells and a lateral avicularium (bottom left) (MNCN 25.03/3200: paratype). C. Basal view of the branching pattern (MNCN 25.03/4234: paratype). D. Single axial vibraculum (MNCN 25.03/3059: holotype)." figureDoi="http://doi.org/10.5281/zenodo.5121754" httpUri="https://zenodo.org/record/5121754/files/figure.png" pageId="4" pageNumber="20">2A–B</figureCitation>
), whilst in 
<taxonomicName id="908BFDC5FFE2CC3FFF4CF8E715C80E33" baseAuthorityName="Gautier" baseAuthorityYear="1962" box="[189,396,1854,1880]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="macrorhyncha">
<emphasis id="65FF5A54FFE2CC3FFF4CF8E715C80E33" box="[189,396,1854,1880]" italics="true" pageId="4" pageNumber="20">C. macrorhyncha</emphasis>
</taxonomicName>
the ovicell is more spherical and usually exhibits an imperforate flat frontal area (
<figureCitation id="CFB09AC3FFE2CC3FFF37F8BB15670E17" box="[198,291,1890,1916]" captionStart="Fig" captionStartId="5.[189,232,1857,1883]" captionTargetBox="[188,1398,911,1804]" captionTargetId="figure-190@5.[188,1398,911,1804]" captionTargetPageId="5" captionText="Fig. 3. Cradoscrupocellaria macrorhyncha (Gautier, 1962) (MNHN-IB-2008-11321: paralectotype), SW Cap Caveaux (Marseille, France), Gautier Coll. A. Frontal view of the branching pattern. B. Detail showing the scuta, ovicells, large frontal avicularia and a small frontal avicularium (bottom right). C. Lateral view showing the large frontal avicularia and basal vibracula. D. Basal view of the branching pattern." figureDoi="http://doi.org/10.5281/zenodo.5121756" httpUri="https://zenodo.org/record/5121756/files/figure.png" pageId="4" pageNumber="20">Fig. 3B</figureCitation>
) (in 
<taxonomicName id="908BFDC5FFE2CC3FFE92F8BB15A20E17" authorityName="Vieira, Spencer Jones &amp; Winston" authorityYear="2013" box="[355,486,1890,1916]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="gautieri">
<emphasis id="65FF5A54FFE2CC3FFE92F8BB15A20E17" box="[355,486,1890,1916]" italics="true" pageId="4" pageNumber="20">C. gautieri</emphasis>
</taxonomicName>
the ovicell is unknown). The autozooids of 
<taxonomicName id="908BFDC5FFE2CC3FFBFBF8BB10C00E17" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[1034,1156,1890,1916]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE2CC3FFBFBF8BB10C00E17" box="[1034,1156,1890,1916]" italics="true" pageId="4" pageNumber="20">C. severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE2CC3FFB7EF8BB10A90E17" box="[1167,1261,1890,1916]" pageId="4" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
are clearly smaller than those of 
<taxonomicName id="908BFDC5FFE2CC3FFE47F85F16C40ECB" baseAuthorityName="Gautier" baseAuthorityYear="1962" box="[438,640,1926,1952]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="macrorhyncha">
<emphasis id="65FF5A54FFE2CC3FFE47F85F16C40ECB" box="[438,640,1926,1952]" italics="true" pageId="4" pageNumber="20">C. macrorhyncha</emphasis>
</taxonomicName>
, but similar in size to those of 
<taxonomicName id="908BFDC5FFE2CC3FFC17F85F10200ECB" authorityName="Vieira, Spencer Jones &amp; Winston" authorityYear="2013" box="[998,1124,1926,1952]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="gautieri">
<emphasis id="65FF5A54FFE2CC3FFC17F85F10200ECB" box="[998,1124,1926,1952]" italics="true" pageId="4" pageNumber="20">C. gautieri</emphasis>
</taxonomicName>
. Finally, the internodes are short in 
<taxonomicName id="908BFDC5FFE2CC3FFEB2F87215FC0EAF" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[323,440,1963,1989]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE2CC3FFEB2F87215FC0EAF" box="[323,440,1963,1989]" italics="true" pageId="4" pageNumber="20">C. severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE2CC3FFE4EF872165F0EAE" box="[447,539,1963,1989]" pageId="4" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
, formed by 5–10 autozooids (
<figureCitation id="CFB09AC3FFE2CC3FFC82F87317B60EAF" box="[883,1010,1962,1988]" captionStart="Fig" captionStartId="2.[189,232,1894,1920]" captionTargetBox="[189,1399,617,1854]" captionTargetId="figure-132@2.[189,1399,617,1854]" captionTargetPageId="2" captionText="Fig. 1. Cradoscrupocellaria severoi sp. nov. A. View of the colony (MNCN 25.03/3059: holotype). B. View of the colony (MNCN 25.03/3200: paratype). C. Branching pattern (MNCN 25.03/4234: paratype). D. Detail of a branch with large frontal avicularia, ovicells and scuta (MNCN 25.03/3059: holotype)." figureDoi="http://doi.org/10.5281/zenodo.5121752" httpUri="https://zenodo.org/record/5121752/files/figure.png" pageId="4" pageNumber="20">Fig. 1A–B</figureCitation>
), as in 
<taxonomicName id="908BFDC5FFE2CC3FFBB1F87210FA0EAF" authorityName="Vieira, Spencer Jones &amp; Winston" authorityYear="2013" box="[1088,1214,1963,1989]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="4" pageNumber="20" phylum="Bryozoa" rank="species" species="gautieri">
<emphasis id="65FF5A54FFE2CC3FFBB1F87210FA0EAF" box="[1088,1214,1963,1989]" italics="true" pageId="4" pageNumber="20">C. gautieri</emphasis>
</taxonomicName>
, but the scutum is flatter and not swollen as in that latter species.
</paragraph>
<caption id="03F4D6CEFFE3CC3EFF4CFED017FD082C" ID-Table-UUID="03F4D6CEFFE3CC3EFF4CFED017FD082C" httpUri="http://table.plazi.org/id/03F4D6CEFFE3CC3EFF4CFED017FD082C" pageId="5" pageNumber="21" startId="5.[189,258,265,291]" targetBox="[188,1362,380,774]" targetIsTable="true" targetPageId="5">
<paragraph id="57348646FFE3CC3EFF4CFED017FD082C" blockId="5.[189,1398,265,327]" pageId="5" pageNumber="21">
<emphasis id="65FF5A54FFE3CC3EFF4CFED0155B0848" bold="true" box="[189,287,265,291]" pageId="5" pageNumber="21">Table 1.</emphasis>
Measurements (in mm) of 
<taxonomicName id="908BFDC5FFE3CC3EFDA8FED017ED0848" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[601,937,265,291]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="21" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE3CC3EFDA8FED017ED0848" box="[601,937,265,291]" italics="true" pageId="5" pageNumber="21">Cradoscrupocellaria severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE3CC3EFC41FED3104E084F" box="[944,1034,266,292]" pageId="5" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
(holotype + paratypes). Abbreviations: SD = standard deviation, N = number of measurements.
</paragraph>
</caption>
<paragraph id="57348646FFE3CC3EFD98FEA411160A6D" pageId="5" pageNumber="21">
<table id="258B74E6FFE333C4FF4DFEA511160A6D" box="[188,1362,380,774]" colsContinueIn="8.[189,1364,378,891]" gridcols="6" gridrows="10" pageId="5" pageNumber="21">
<tr id="E9BB8404FFE333C4FF4DFEA5111608FC" box="[188,1362,380,407]" gridrow="0" pageId="5" pageNumber="21" rowspan-0="1">
<th id="AA6AED78FFE333C4FD98FEA516F508FC" box="[617,689,380,407]" gridcol="1" gridrow="0" pageId="5" pageNumber="21">
<emphasis id="65FF5A54FFE3CC3EFD98FEA416F508FC" bold="true" box="[617,689,381,407]" pageId="5" pageNumber="21">Mean</emphasis>
</th>
<th id="AA6AED78FFE333C4FCD0FEA5173608FC" box="[801,882,380,407]" gridcol="2" gridrow="0" pageId="5" pageNumber="21">
<emphasis id="65FF5A54FFE3CC3EFCC6FEA5171808FD" bold="true" box="[823,860,380,406]" pageId="5" pageNumber="21">SD</emphasis>
</th>
<th id="AA6AED78FFE333C4FC35FEA5100508FC" box="[964,1089,380,407]" gridcol="3" gridrow="0" pageId="5" pageNumber="21">
<emphasis id="65FF5A54FFE3CC3EFC35FEA4100508FC" bold="true" box="[964,1089,381,407]" pageId="5" pageNumber="21">Minimum</emphasis>
</th>
<th id="AA6AED78FFE333C4FB86FEA510BD08FC" box="[1143,1273,380,407]" gridcol="4" gridrow="0" pageId="5" pageNumber="21">
<emphasis id="65FF5A54FFE3CC3EFB86FEA410BD08FC" bold="true" box="[1143,1273,381,407]" pageId="5" pageNumber="21">Maximum</emphasis>
</th>
<th id="AA6AED78FFE333C4FAC4FEA5111608FC" box="[1333,1362,380,407]" gridcol="5" gridrow="0" pageId="5" pageNumber="21">
<emphasis id="65FF5A54FFE3CC3EFAC8FEA4110A08FC" bold="true" box="[1337,1358,381,407]" pageId="5" pageNumber="21">N</emphasis>
</th>
</tr>
<tr id="E9BB8404FFE333C4FF4DFE72111608AD" box="[188,1362,427,454]" gridrow="1" pageId="5" pageNumber="21">
<th id="AA6AED78FFE333C4FF4DFE7215E408AD" box="[188,416,427,454]" gridcol="0" gridrow="1" pageId="5" pageNumber="21">Autozooid length</th>
<td id="AA6AED78FFE333C4FD98FE7216F508AD" box="[617,689,427,454]" gridcol="1" gridrow="1" pageId="5" pageNumber="21">0.454</td>
<td id="AA6AED78FFE333C4FCD0FE72173608AD" box="[801,882,427,454]" gridcol="2" gridrow="1" pageId="5" pageNumber="21">0.0480</td>
<td id="AA6AED78FFE333C4FC35FE72100508AD" box="[964,1089,427,454]" gridcol="3" gridrow="1" pageId="5" pageNumber="21">0.370</td>
<td id="AA6AED78FFE333C4FB86FE7210BD08AD" box="[1143,1273,427,454]" gridcol="4" gridrow="1" pageId="5" pageNumber="21">0.565</td>
<td id="AA6AED78FFE333C4FAC4FE72111608AD" box="[1333,1362,427,454]" gridcol="5" gridrow="1" pageId="5" pageNumber="21">29</td>
</tr>
<tr id="E9BB8404FFE333C4FF4DFE0A11160885" box="[188,1362,467,494]" gridrow="2" pageId="5" pageNumber="21">
<th id="AA6AED78FFE333C4FF4DFE0A15E40885" box="[188,416,467,494]" gridcol="0" gridrow="2" pageId="5" pageNumber="21">Autozooid width</th>
<td id="AA6AED78FFE333C4FD98FE0A16F50885" box="[617,689,467,494]" gridcol="1" gridrow="2" pageId="5" pageNumber="21">0.191</td>
<td id="AA6AED78FFE333C4FCD0FE0A17360885" box="[801,882,467,494]" gridcol="2" gridrow="2" pageId="5" pageNumber="21">0.0154</td>
<td id="AA6AED78FFE333C4FC35FE0A10050885" box="[964,1089,467,494]" gridcol="3" gridrow="2" pageId="5" pageNumber="21">0.161</td>
<td id="AA6AED78FFE333C4FB86FE0A10BD0885" box="[1143,1273,467,494]" gridcol="4" gridrow="2" pageId="5" pageNumber="21">0.232</td>
<td id="AA6AED78FFE333C4FAC4FE0A11160885" box="[1333,1362,467,494]" gridcol="5" gridrow="2" pageId="5" pageNumber="21">29</td>
</tr>
<tr id="E9BB8404FFE333C4FF4DFE2211160B7D" box="[188,1362,507,534]" gridrow="3" pageId="5" pageNumber="21">
<th id="AA6AED78FFE333C4FF4DFE2215E40B7D" box="[188,416,507,534]" gridcol="0" gridrow="3" pageId="5" pageNumber="21">Sutum length</th>
<td id="AA6AED78FFE333C4FD98FE2216F50B7D" box="[617,689,507,534]" gridcol="1" gridrow="3" pageId="5" pageNumber="21">0.289</td>
<td id="AA6AED78FFE333C4FCD0FE2217360B7D" box="[801,882,507,534]" gridcol="2" gridrow="3" pageId="5" pageNumber="21">0.0176</td>
<td id="AA6AED78FFE333C4FC35FE2210050B7D" box="[964,1089,507,534]" gridcol="3" gridrow="3" pageId="5" pageNumber="21">0.258</td>
<td id="AA6AED78FFE333C4FB86FE2210BD0B7D" box="[1143,1273,507,534]" gridcol="4" gridrow="3" pageId="5" pageNumber="21">0.324</td>
<td id="AA6AED78FFE333C4FAC4FE2211160B7D" box="[1333,1362,507,534]" gridcol="5" gridrow="3" pageId="5" pageNumber="21">29</td>
</tr>
<tr id="E9BB8404FFE333C4FF4DFDFA11160B55" box="[188,1362,547,574]" gridrow="4" pageId="5" pageNumber="21">
<th id="AA6AED78FFE333C4FF4DFDFA15E40B55" box="[188,416,547,574]" gridcol="0" gridrow="4" pageId="5" pageNumber="21">Scutum width</th>
<td id="AA6AED78FFE333C4FD98FDFA16F50B55" box="[617,689,547,574]" gridcol="1" gridrow="4" pageId="5" pageNumber="21">0.164</td>
<td id="AA6AED78FFE333C4FCD0FDFA17360B55" box="[801,882,547,574]" gridcol="2" gridrow="4" pageId="5" pageNumber="21">0.0206</td>
<td id="AA6AED78FFE333C4FC35FDFA10050B55" box="[964,1089,547,574]" gridcol="3" gridrow="4" pageId="5" pageNumber="21">0.121</td>
<td id="AA6AED78FFE333C4FB86FDFA10BD0B55" box="[1143,1273,547,574]" gridcol="4" gridrow="4" pageId="5" pageNumber="21">0.207</td>
<td id="AA6AED78FFE333C4FAC4FDFA11160B55" box="[1333,1362,547,574]" gridcol="5" gridrow="4" pageId="5" pageNumber="21">29</td>
</tr>
<tr id="E9BB8404FFE333C4FF4DFD9211160B0D" box="[188,1362,587,614]" gridrow="5" pageId="5" pageNumber="21">
<th id="AA6AED78FFE333C4FF4DFD9215E40B0D" box="[188,416,587,614]" gridcol="0" gridrow="5" pageId="5" pageNumber="21">Frontal avic. length</th>
<td id="AA6AED78FFE333C4FD98FD9216F50B0D" box="[617,689,587,614]" gridcol="1" gridrow="5" pageId="5" pageNumber="21">0.259</td>
<td id="AA6AED78FFE333C4FCD0FD9217360B0D" box="[801,882,587,614]" gridcol="2" gridrow="5" pageId="5" pageNumber="21">0.0283</td>
<td id="AA6AED78FFE333C4FC35FD9210050B0D" box="[964,1089,587,614]" gridcol="3" gridrow="5" pageId="5" pageNumber="21">0.218</td>
<td id="AA6AED78FFE333C4FB86FD9210BD0B0D" box="[1143,1273,587,614]" gridcol="4" gridrow="5" pageId="5" pageNumber="21">0.281</td>
<td id="AA6AED78FFE333C4FAC4FD9211160B0D" box="[1333,1362,587,614]" gridcol="5" gridrow="5" pageId="5" pageNumber="21">4</td>
</tr>
<tr id="E9BB8404FFE333C4FF4DFDAA11160BE5" box="[188,1362,627,654]" gridrow="6" pageId="5" pageNumber="21">
<th id="AA6AED78FFE333C4FF4DFDAA15E40BE5" box="[188,416,627,654]" gridcol="0" gridrow="6" pageId="5" pageNumber="21">Frontal avic. width</th>
<td id="AA6AED78FFE333C4FD98FDAA16F50BE5" box="[617,689,627,654]" gridcol="1" gridrow="6" pageId="5" pageNumber="21">0.087</td>
<td id="AA6AED78FFE333C4FCD0FDAA17360BE5" box="[801,882,627,654]" gridcol="2" gridrow="6" pageId="5" pageNumber="21">0.0037</td>
<td id="AA6AED78FFE333C4FC35FDAA10050BE5" box="[964,1089,627,654]" gridcol="3" gridrow="6" pageId="5" pageNumber="21">0.082</td>
<td id="AA6AED78FFE333C4FB86FDAA10BD0BE5" box="[1143,1273,627,654]" gridcol="4" gridrow="6" pageId="5" pageNumber="21">0.093</td>
<td id="AA6AED78FFE333C4FAC4FDAA11160BE5" box="[1333,1362,627,654]" gridcol="5" gridrow="6" pageId="5" pageNumber="21">6</td>
</tr>
<tr id="E9BB8404FFE333C4FF4DFD4211160BDD" box="[188,1362,667,694]" gridrow="7" pageId="5" pageNumber="21">
<th id="AA6AED78FFE333C4FF4DFD4215E40BDD" box="[188,416,667,694]" gridcol="0" gridrow="7" pageId="5" pageNumber="21">Lateral avic. length</th>
<td id="AA6AED78FFE333C4FD98FD4216F50BDD" box="[617,689,667,694]" gridcol="1" gridrow="7" pageId="5" pageNumber="21">0.065</td>
<td id="AA6AED78FFE333C4FCD0FD4217360BDD" box="[801,882,667,694]" gridcol="2" gridrow="7" pageId="5" pageNumber="21">0.0080</td>
<td id="AA6AED78FFE333C4FC35FD4210050BDD" box="[964,1089,667,694]" gridcol="3" gridrow="7" pageId="5" pageNumber="21">0.055</td>
<td id="AA6AED78FFE333C4FB86FD4210BD0BDD" box="[1143,1273,667,694]" gridcol="4" gridrow="7" pageId="5" pageNumber="21">0.073</td>
<td id="AA6AED78FFE333C4FAC4FD4211160BDD" box="[1333,1362,667,694]" gridcol="5" gridrow="7" pageId="5" pageNumber="21">6</td>
</tr>
<tr id="E9BB8404FFE333C4FF4DFD1A11160BB5" box="[188,1362,707,734]" gridrow="8" pageId="5" pageNumber="21">
<th id="AA6AED78FFE333C4FF4DFD1A15E40BB5" box="[188,416,707,734]" gridcol="0" gridrow="8" pageId="5" pageNumber="21">Ovicell length</th>
<td id="AA6AED78FFE333C4FD98FD1A16F50BB5" box="[617,689,707,734]" gridcol="1" gridrow="8" pageId="5" pageNumber="21">0.163</td>
<td id="AA6AED78FFE333C4FCD0FD1A17360BB5" box="[801,882,707,734]" gridcol="2" gridrow="8" pageId="5" pageNumber="21">0.0160</td>
<td id="AA6AED78FFE333C4FC35FD1A10050BB5" box="[964,1089,707,734]" gridcol="3" gridrow="8" pageId="5" pageNumber="21">0.134</td>
<td id="AA6AED78FFE333C4FB86FD1A10BD0BB5" box="[1143,1273,707,734]" gridcol="4" gridrow="8" pageId="5" pageNumber="21">0.186</td>
<td id="AA6AED78FFE333C4FAC4FD1A11160BB5" box="[1333,1362,707,734]" gridcol="5" gridrow="8" pageId="5" pageNumber="21">18</td>
</tr>
<tr id="E9BB8404FFE333C4FF4DFD3211160A6D" box="[188,1362,747,774]" gridrow="9" pageId="5" pageNumber="21">
<th id="AA6AED78FFE333C4FF4DFD3215E40A6D" box="[188,416,747,774]" gridcol="0" gridrow="9" pageId="5" pageNumber="21">Ovicell width</th>
<td id="AA6AED78FFE333C4FD98FD3216F50A6D" box="[617,689,747,774]" gridcol="1" gridrow="9" pageId="5" pageNumber="21">0.213</td>
<td id="AA6AED78FFE333C4FCD0FD3217360A6D" box="[801,882,747,774]" gridcol="2" gridrow="9" pageId="5" pageNumber="21">0.0187</td>
<td id="AA6AED78FFE333C4FC35FD3210050A6D" box="[964,1089,747,774]" gridcol="3" gridrow="9" pageId="5" pageNumber="21">0.170</td>
<td id="AA6AED78FFE333C4FB86FD3210BD0A6D" box="[1143,1273,747,774]" gridcol="4" gridrow="9" pageId="5" pageNumber="21">0.245</td>
<td id="AA6AED78FFE333C4FAC4FD3211160A6D" box="[1333,1362,747,774]" gridcol="5" gridrow="9" pageId="5" pageNumber="21">18</td>
</tr>
</table>
</paragraph>
<caption id="03F4D6CEFFE3CC3EFF4CF89815520E82" ID-DOI="http://doi.org/10.5281/zenodo.5121756" ID-Zenodo-Dep="5121756" httpUri="https://zenodo.org/record/5121756/files/figure.png" pageId="5" pageNumber="21" startId="5.[189,232,1857,1883]" targetBox="[188,1398,911,1804]" targetPageId="5">
<paragraph id="57348646FFE3CC3EFF4CF89815520E82" blockId="5.[188,1399,1857,2025]" pageId="5" pageNumber="21">
<emphasis id="65FF5A54FFE3CC3EFF4CF898154A0E30" bold="true" box="[189,270,1857,1883]" pageId="5" pageNumber="21">Fig. 3.</emphasis>
<taxonomicName id="908BFDC5FFE3CC3EFEEBF89817C30E37" authority="(Gautier, 1962)" baseAuthorityName="Gautier" baseAuthorityYear="1962" box="[282,903,1857,1884]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="5" pageNumber="21" phylum="Bryozoa" rank="species" species="macrorhyncha">
<emphasis id="65FF5A54FFE3CC3EFEEBF89816800E30" box="[282,708,1857,1883]" italics="true" pageId="5" pageNumber="21">Cradoscrupocellaria macrorhyncha</emphasis>
(
<bibRefCitation id="331AFBB7FFE3CC3EFD29F898173B0E37" author="Gautier Y. - V." box="[728,895,1857,1884]" pageId="5" pageNumber="21" pagination="1 - 435" refId="ref6642" refString="Gautier Y. - V. 1962. Recherches ecologiques sur les Bryozoaires Chilostomes en Mediterranee occidentale. Recueil des Travaux de la Station marine d'Endoume 38: 1 - 435." type="journal article" year="1962">Gautier, 1962</bibRefCitation>
)
</taxonomicName>
(MNHN-IB-2008-11321: paralectotype), SW Cap Caveaux (Marseille, France), Gautier Coll. 
<emphasis id="65FF5A54FFE3CC3EFCD5F8BD177D0E15" bold="true" box="[804,825,1892,1918]" pageId="5" pageNumber="21">A</emphasis>
. Frontal view of the branching pattern. 
<emphasis id="65FF5A54FFE3CC3EFAFCF8BC11650E14" bold="true" box="[1293,1313,1893,1919]" pageId="5" pageNumber="21">B</emphasis>
. Detail showing the scuta, ovicells, large frontal avicularia and a small frontal avicularium (bottom right). 
<emphasis id="65FF5A54FFE3CC3EFF4CF87214960EAE" bold="true" box="[189,210,1963,1989]" pageId="5" pageNumber="21">C</emphasis>
. Lateral view showing the large frontal avicularia and basal vibracula. 
<emphasis id="65FF5A54FFE3CC3EFBFDF87210650EAE" bold="true" box="[1036,1057,1963,1989]" pageId="5" pageNumber="21">D</emphasis>
. Basal view of the branching pattern.
</paragraph>
</caption>
<paragraph id="57348646FFE0CC3DFF4CFED0156B0BA5" blockId="6.[189,1400,265,718]" pageId="6" pageNumber="22">
<taxonomicName id="908BFDC5FFE0CC3DFF4CFED016510848" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[189,533,265,291]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="22" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE0CC3DFF4CFED016510848" box="[189,533,265,291]" italics="true" pageId="6" pageNumber="22">Cradoscrupocellaria severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE0CC3DFDD3FED316C7084F" box="[546,643,266,292]" pageId="6" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
is known with certainty only from its 
<typeStatus id="883038E4FFE0CC3DFB89FED310EF084F" box="[1144,1195,266,292]" pageId="6" pageNumber="22">type</typeStatus>
localities at the Columbretes Islands (Mediterranean Spain) between 3 and 
<quantity id="90732BA3FFE0CC3DFC5CFEE917B50821" box="[941,1009,304,330]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="22" unit="m" value="42.0">42 m</quantity>
depth. 
<bibRefCitation id="331AFBB7FFE0CC3DFBA9FEE911410820" author="Zabala M." box="[1112,1285,304,331]" pageId="6" pageNumber="22" pagination="1 - 836" refId="ref8492" refString="Zabala M. 1986. Fauna dels briozous dels Paisos Catalans. Arxius de la Seccio de Ciencies 84: 1 - 836." type="journal article" year="1986">Zabala (1986)</bibRefCitation>
reported 
<taxonomicName id="908BFDC5FFE0CC3DFF4CFE8E1683081B" authority="Gautier, 1962" authorityName="Gautier" authorityYear="1962" box="[189,711,342,369]" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="22" phylum="Bryozoa" rank="species" species="macrorhyncha">
<emphasis id="65FF5A54FFE0CC3DFF4CFE8E1659081A" box="[189,541,343,369]" italics="true" pageId="6" pageNumber="22">Scrupocellaria macrorhyncha</emphasis>
<bibRefCitation id="331AFBB7FFE0CC3DFDD4FE8E1683081B" author="Gautier Y. - V." box="[549,711,342,369]" pageId="6" pageNumber="22" pagination="1 - 435" refId="ref6642" refString="Gautier Y. - V. 1962. Recherches ecologiques sur les Bryozoaires Chilostomes en Mediterranee occidentale. Recueil des Travaux de la Station marine d'Endoume 38: 1 - 435." type="journal article" year="1962">Gautier, 1962</bibRefCitation>
</taxonomicName>
from several Catalonian localities, but these records were later reassigned to 
<taxonomicName id="908BFDC5FFE0CC3DFE57FEA716B108F3" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[422,757,381,408]" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="22" phylum="Bryozoa" rank="species" species="reptans">
<emphasis id="65FF5A54FFE0CC3DFE57FEA7165A08FC" box="[422,542,382,408]" italics="true" pageId="6" pageNumber="22">S. reptans</emphasis>
(
<bibRefCitation id="331AFBB7FFE0CC3DFDC0FEA416A908F3" author="Linnaeus C." box="[561,749,381,408]" pageId="6" pageNumber="22" refId="ref7256" refString="Linnaeus C. 1758. Systemae naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differetiis, synonymis, locis, Ed. 10. Laurentii Salvii, Holmiae. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
by 
<bibRefCitation id="331AFBB7FFE0CC3DFCD8FEA4103208F3" author="Zabala M. &amp; Maluquer P." box="[809,1142,381,408]" pageId="6" pageNumber="22" pagination="1 - 294" refId="ref8516" refString="Zabala M. &amp; Maluquer P. 1988. Illustrated keys for the classification of Mediterranean Bryozoa. Treballs del Museu de Zoologia, Barcelona 4: 1 - 294." type="journal article" year="1988">Zabala &amp; Maluquer (1988)</bibRefCitation>
. 
<bibRefCitation id="331AFBB7FFE0CC3DFB75FEA4113608F3" author="Vieira L. M. &amp; Spencer Jones M. E. &amp; Winston J. E." box="[1156,1394,381,408]" pageId="6" pageNumber="22" pagination="1 - 63" refId="ref8324" refString="Vieira L. M., Spencer Jones M. E. &amp; Winston J. E. 2013. Cradoscrupocellaria, a new bryozoan genus for Scrupocellaria bertholletii (Audouin) and related species (Cheilostomata, Candidae): taxonomy, biodiversity and distribution. Zootaxa 3707 (1): 1 - 63. https: // doi. org / 10.11646 / zootaxa. 3707.1.1" type="journal article" year="2013">
Vieira 
<emphasis id="65FF5A54FFE0CC3DFB26FEA7115208FC" box="[1239,1302,381,407]" italics="true" pageId="6" pageNumber="22">et al.</emphasis>
(2013)
</bibRefCitation>
, however, stated that these specimens are distinct from 
<taxonomicName id="908BFDC5FFE0CC3DFCADFE7C106F08D5" authorityName="Gautier" authorityYear="1962" box="[860,1067,420,447]" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="22" phylum="Bryozoa" rank="species" species="macrorhyncha">
<emphasis id="65FF5A54FFE0CC3DFCADFE7C106F08D5" box="[860,1067,420,447]" italics="true" pageId="6" pageNumber="22">C. macrorhyncha</emphasis>
</taxonomicName>
and 
<taxonomicName id="908BFDC5FFE0CC3DFB9CFE7C10AE08D5" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1133,1258,421,447]" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="22" phylum="Bryozoa" rank="species" species="reptans">
<emphasis id="65FF5A54FFE0CC3DFB9CFE7C10AE08D5" box="[1133,1258,421,447]" italics="true" pageId="6" pageNumber="22">C. reptans</emphasis>
</taxonomicName>
, as well as from 
<taxonomicName id="908BFDC5FFE0CC3DFF0FFE15169C088E" authority="(Vieira &amp; Spencer Jones, 2012)" baseAuthorityName="Vieira &amp; Spencer Jones" baseAuthorityYear="2012" box="[254,728,459,486]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="22" phylum="Bryozoa" rank="species" species="ellisi">
<emphasis id="65FF5A54FFE0CC3DFF0FFE15151E088E" box="[254,346,459,486]" italics="true" pageId="6" pageNumber="22">C. ellisi</emphasis>
(
<bibRefCitation id="331AFBB7FFE0CC3DFE9BFE121694088E" author="Vieira L. M. &amp; Spencer Jones M. E." box="[362,720,459,486]" pageId="6" pageNumber="22" pagination="26 - 42" refId="ref8278" refString="Vieira L. M. &amp; Spencer Jones M. E. 2012. The identity of Sertularia reptans Linnaeus, 1758 (Bryozoa, Candidae). Zootaxa 3563: 26 - 42. https: // doi. org / 10.11646 / zootaxa 3563.1.2" type="journal article" year="2012">Vieira &amp; Spencer Jones, 2012</bibRefCitation>
)
</taxonomicName>
, and that they may belong to a new undescribed species. Figures by 
<bibRefCitation id="331AFBB7FFE0CC3DFEB0FE2B15A10B66" author="Zabala M." box="[321,485,498,525]" pageId="6" pageNumber="22" pagination="1 - 836" refId="ref8492" refString="Zabala M. 1986. Fauna dels briozous dels Paisos Catalans. Arxius de la Seccio de Ciencies 84: 1 - 836." type="journal article" year="1986">Zabala (1986)</bibRefCitation>
and 
<bibRefCitation id="331AFBB7FFE0CC3DFDEFFE2B171D0B67" author="Zabala M. &amp; Maluquer P." box="[542,857,498,524]" pageId="6" pageNumber="22" pagination="1 - 294" refId="ref8516" refString="Zabala M. &amp; Maluquer P. 1988. Illustrated keys for the classification of Mediterranean Bryozoa. Treballs del Museu de Zoologia, Barcelona 4: 1 - 294." type="journal article" year="1988">Zabala &amp; Maluquer (1988)</bibRefCitation>
are quite similar to 
<taxonomicName id="908BFDC5FFE0CC3DFBB0FE2B10F30B67" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[1089,1207,498,524]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="22" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE0CC3DFBB0FE2B10F30B67" box="[1089,1207,498,524]" italics="true" pageId="6" pageNumber="22">C. severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE0CC3DFB4FFE2B115F0B67" box="[1214,1307,498,524]" pageId="6" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
, but the description given (actually an almost literal translation of the original description of 
<taxonomicName id="908BFDC5FFE0CC3DFB41FDC0113C0B58" authorityName="Gautier" authorityYear="1962" box="[1200,1400,537,563]" class="Gymnolaemata" family="Candidae" genus="Scrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="22" phylum="Bryozoa" rank="species" species="macrorhyncha">
<emphasis id="65FF5A54FFE0CC3DFB41FDC0113C0B58" box="[1200,1400,537,563]" italics="true" pageId="6" pageNumber="22">S. macrorhyncha</emphasis>
</taxonomicName>
by Gautier) show several differences from the present description of the new species. A revision of the original material and collection of new material will be necessary to clarify which species was reported from 
<collectingRegion id="954F48A4FFE0CC3DFEF1FD5415370BCC" box="[256,371,653,679]" country="Spain" name="Catalunya" pageId="6" pageNumber="22">Catalonia</collectingRegion>
, as well as to determine the possible presence of 
<taxonomicName id="908BFDC5FFE0CC3DFC23FD5710080BCC" authority="Reverter-Gil &amp; Souto, 2021" authorityName="Reverter-Gil &amp; Souto" authorityYear="2021" box="[978,1100,654,680]" class="Gymnolaemata" family="Candidae" genus="Cradoscrupocellaria" kingdom="Animalia" order="Cheilostomatida" pageId="6" pageNumber="22" phylum="Bryozoa" rank="species" species="severoi" status="sp. nov.">
<emphasis id="65FF5A54FFE0CC3DFC23FD5710080BCC" box="[978,1100,654,680]" italics="true" pageId="6" pageNumber="22">C. severoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ECCE72FFFE0CC3DFBA7FD5710F70BC3" box="[1110,1203,654,680]" pageId="6" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
at other Iberian localities.
</paragraph>
</subSubSection>
</treatment>
</document>