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<document id="D2E14B8BFE78E03FE8F03F40192BDD54" ID-CLB-Dataset="23381" ID-DOI="10.5252/zoosystema2019v41a26" ID-GBIF-Dataset="cef5076b-424b-4cdc-8d13-6e472d861e09" ID-ISSN="1638-9387" ID-Zenodo-Dep="3726028" ID-ZooBank="urn:lsid:zoobank.org:pub:CF16A992-0401-44C8-BEEE-842CE7F1D27E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585071141162" checkinUser="felipe" docAuthor="Ortega, José Rafael &amp; Gofas, Serge" docDate="2019" docId="03E687A0B24CFF919B60FB04FEC3F995" docLanguage="en" docName="zoosystema.41.26.513-551.pdf.imf" docOrigin="Zoosystema 41 (26)" docStyle="DocumentStyle:AFAF64E709DD7C8340B70A24A5804402.4:Zoosystema.2018-2019.journal_article" docStyleId="AFAF64E709DD7C8340B70A24A5804402" docStyleName="Zoosystema.2018-2019.journal_article" docStyleVersion="4" docTitle="Marginella carmenae Ortega &amp; Gofas 2019, n. sp." docType="treatment" docUuid="6BE37985-54C1-4964-895B-B3435F50348A" docUuidSource="ZooBank" docVersion="8" lastPageNumber="533" masterDocId="FFDFFFD8B258FF879A52FFD1FFCDFFBB" masterDocTitle="The unknown bathyal of the Canaries: new species and new records of deep-sea Mollusca" masterLastPageNumber="551" masterPageNumber="513" pageNumber="531" updateTime="1698828230493" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="343F1DB44C43C028F8F7B896ADA278DA">The unknown bathyal of the Canaries: new species and new records of deep-sea Mollusca</mods:title>
</mods:titleInfo>
<mods:name id="97960838C6FAE0408845A0C3FF53E2C7" type="personal">
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<mods:namePart id="A8B8A99A43E167428531A1027A04BD5B">Ortega, José Rafael</mods:namePart>
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<mods:roleTerm id="C411DF956EF125351EE2F1C446CA7319">Author</mods:roleTerm>
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<mods:namePart id="4CED3757CC77E8A1E4CCF8B0A5C273F3">Gofas, Serge</mods:namePart>
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<mods:title id="068C5A0C5271F150868A9DA9F413B2BE">Zoosystema</mods:title>
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<mods:part id="251D7A8591843A26241EDED6B6608EB7">
<mods:date id="233B5C66182CF7CA0CB7081B1014BF11">2019</mods:date>
<mods:detail id="6B05F6C2330171805C4064CC36D51FFC" type="pubDate">
<mods:number id="DB88488787B897AB93EB6075AF8898BA">2019-11-28</mods:number>
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<mods:number id="16E3609B3AD8043BA3EA22C415ED8CF0">41</mods:number>
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<mods:classification id="02391DD9D41CE8C5ACA6506281972EB1">journal article</mods:classification>
<mods:identifier id="674053F2C855E3CC6668C9141CE38C04" type="CLB-Dataset">23381</mods:identifier>
<mods:identifier id="F22E5541777FDC8871EDDF067FC650BE" type="DOI">10.5252/zoosystema2019v41a26</mods:identifier>
<mods:identifier id="FE3340F0C97EE3A546A53C0622766C97" type="GBIF-Dataset">cef5076b-424b-4cdc-8d13-6e472d861e09</mods:identifier>
<mods:identifier id="75003B525960EB503941F9DF36A67AA1" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="F65F7B263E38AD3E905B6FF0D34CAB0E" type="Zenodo-Dep">3726028</mods:identifier>
<mods:identifier id="2CB7073C480089802B19FFE082867CEA" type="ZooBank">urn:lsid:zoobank.org:pub:CF16A992-0401-44C8-BEEE-842CE7F1D27E</mods:identifier>
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<treatment id="03E687A0B24CFF919B60FB04FEC3F995" ID-DOI="http://doi.org/10.5281/zenodo.3729418" ID-GBIF-Taxon="163283915" ID-Zenodo-Dep="3729418" LSID="urn:lsid:zoobank.org:act:6BE37985-54C1-4964-895B-B3435F50348A" httpUri="http://treatment.plazi.org/id/03E687A0B24CFF919B60FB04FEC3F995" lastPageId="22" lastPageNumber="533" pageId="20" pageNumber="531">
<subSubSection id="C355653DB24CFF939B60FB04FD97FB4B" box="[306,602,1237,1264]" pageId="20" pageNumber="531" type="nomenclature">
<paragraph id="8BF036B6B24CFF939B60FB04FD97FB4B" blockId="20.[306,602,1237,1296]" box="[306,602,1237,1264]" pageId="20" pageNumber="531">
<heading id="D0B881DAB24CFF939B60FB04FD97FB4B" box="[306,602,1237,1264]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="531" reason="2">
<taxonomicName id="4C4F4D35B24CFF939B60FB04FDD5FB4B" ID-CoL="7S2KQ" authority="Ortega &amp; Gofas, 2019" authorityName="Ortega &amp; Gofas" authorityYear="2019" box="[306,536,1237,1264]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="531" phylum="Mollusca" rank="species" species="carmenae" status="sp. nov.">
<emphasis id="B93BEAA4B24CFF939B60FB04FDD5FB4B" bold="true" box="[306,536,1237,1264]" italics="true" pageId="20" pageNumber="531">Marginella carmenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857DFB24CFF93984DFB07FD97FB4B" box="[543,602,1238,1264]" pageId="20" pageNumber="531" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C355653DB24CFF939B3EFB24FD25FAFF" pageId="20" pageNumber="531" type="description">
<paragraph id="8BF036B6B24CFF939B3EFB24FDD2FAAB" blockId="20.[306,602,1237,1296]" box="[364,543,1269,1296]" pageId="20" pageNumber="531">
(
<figureCitation id="13742A33B24CFF939B21FB24FE0DFAAB" box="[371,448,1269,1296]" captionStart="FIG" captionStartId="21.[132,143,1465,1483]" captionTargetBox="[132,1455,215,1406]" captionTargetId="figure@21.[132,1455,215,1406]" captionTargetPageId="21" captionText="FIG. 14. — A-F, Marginella carmenae n. sp.: A, B, holotype from DW128 (16.7 mm); C, D, paratype,same locality (14.7 mm);E, F, paratype,same locality (14.0 mm); G, H, Marginella belcheri Hinds, 1844, specimen from off Mauritania, Atlor sta. 49, 20°00’N, 17°44’W, 200 m, R/V Cornide de Saavedra (Museo Nacional de Ciencias naturales, Madrid) (17.8 mm)." figureDoi="http://doi.org/10.5281/zenodo.3726056" httpUri="https://zenodo.org/record/3726056/files/figure.png" pageId="20" pageNumber="531">Figs 14</figureCitation>
A-F; 15)
</paragraph>
<paragraph id="8BF036B6B24CFF939AF1FAE3FD25FAFF" blockId="20.[163,744,1330,1348]" box="[163,744,1330,1348]" pageId="20" pageNumber="531">
<uri id="FFDE3AB4B24CFF939AF1FAE3FD25FAFF" box="[163,744,1330,1348]" pageId="20" pageNumber="531">
urn:lsid:zoobank.org:act:
<uuid id="FFE90C63B24CFF939B3BFAE3FD25FAFF" box="[361,744,1330,1348]" pageId="20" pageNumber="531">6BE37985-54C1-4964-895B-B3435F50348A</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C355653DB24CFF939AD6FAB5FCC5FA77" pageId="20" pageNumber="531" type="materials_examined">
<paragraph id="8BF036B6B24CFF939AD6FAB5FE49FA2C" blockId="20.[132,776,1380,1484]" pageId="20" pageNumber="531">
<typeStatus id="54F48814B24CFF939AD6FAB5FF78FAC1" box="[132,181,1380,1404]" pageId="20" pageNumber="531">TYPE</typeStatus>
MATERIAL. —
<materialsCitation id="3B273CEBB24CFF939B69FAB5FE4DFA2C" ID-GBIF-Occurrence="2580042106" country="Djibouti" pageId="20" pageNumber="531" specimenCount="1" typeStatus="holotype">
<emphasis id="B93BEAA4B24CFF939B69FAB5FE53FAC7" bold="true" box="[315,414,1380,1404]" pageId="20" pageNumber="531">
<typeStatus id="54F48814B24CFF939B69FAB5FE57FAC7" box="[315,410,1380,1404]" pageId="20" pageNumber="531" type="holotype">Holotype</typeStatus>
.
</emphasis>
spm., MNHN-IM-2000-34261 from SEAMOUNT 2 DW128
</materialsCitation>
.
</paragraph>
<paragraph id="8BF036B6B24CFF939AD6FA48FCC5FA77" blockId="20.[132,776,1380,1484]" pageId="20" pageNumber="531">
<materialsCitation id="3B273CEBB24CFF939AD6FA48FD26FA09" ID-GBIF-Occurrence="2580042104" box="[132,747,1433,1458]" country="Djibouti" pageId="20" pageNumber="531" specimenCount="2" typeStatus="paratype">
<emphasis id="B93BEAA4B24CFF939AD6FA48FF13FA0A" bold="true" box="[132,222,1433,1457]" pageId="20" pageNumber="531">
<typeStatus id="54F48814B24CFF939AD6FA48FF13FA0A" box="[132,222,1433,1457]" pageId="20" pageNumber="531" type="paratype">Paratypes</typeStatus>
</emphasis>
. 2 spm., MNHN-IM-2000-34262 from the
<typeStatus id="54F48814B24CFF93982AFA4BFD52FA09" box="[632,671,1434,1458]" pageId="20" pageNumber="531">type</typeStatus>
locality.
</materialsCitation>
—
<materialsCitation id="3B273CEBB24CFF939AD6FA65FCCEFA77" ID-GBIF-Occurrence="2580005090" box="[132,771,1460,1484]" country="Djibouti" pageId="20" pageNumber="531" specimenCount="1">1 spm., MNHN-IM-2000-34497 from SEAMOUNT 2, DW126</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C355653DB24CFF939AD6FA32FDB0F9AD" pageId="20" pageNumber="531" type="distribution">
<paragraph id="8BF036B6B24CFF939AD6FA32FDB0F9AD" blockId="20.[132,774,1507,1558]" pageId="20" pageNumber="531">
<typeStatus id="54F48814B24CFF939AD6FA32FF77FA42" box="[132,186,1507,1531]" pageId="20" pageNumber="531">TYPE</typeStatus>
LOCALITY. — Off NW Gran Canaria,
<geoCoordinate id="EE7B5071B24CFF93982DFA35FCCEFA47" box="[639,771,1508,1532]" degrees="28" direction="north" minutes="08.26" orientation="latitude" pageId="20" pageNumber="531" precision="9" value="28.137667">28°08.26’N</geoCoordinate>
,
<geoCoordinate id="EE7B5071B24CFF939AD6FA2FFF33F9AD" box="[132,254,1534,1558]" degrees="15" direction="west" minutes="51.99" orientation="longitude" pageId="20" pageNumber="531" precision="9" value="-15.8665">15°51.99’W</geoCoordinate>
/
<geoCoordinate id="EE7B5071B24CFF939B54FA2FFEB6F9AD" box="[262,379,1534,1558]" degrees="28" direction="north" minutes="08.47" orientation="latitude" pageId="20" pageNumber="531" precision="9" value="28.141167">28°08.47’N</geoCoordinate>
,
<geoCoordinate id="EE7B5071B24CFF939BD7FA2FFE30F9AD" box="[389,509,1534,1558]" degrees="15" direction="west" minutes="51.66" orientation="longitude" pageId="20" pageNumber="531" precision="9" value="-15.861">15°51.66’W</geoCoordinate>
,
<quantity id="4CB79B53B24CFF939855FA2FFDB4F9AD" box="[519,633,1534,1558]" metricMagnitude="2" metricUnit="m" metricValue="4.775" metricValueMax="4.85" metricValueMin="4.7" pageId="20" pageNumber="531" unit="m" value="477.5" valueMax="485.0" valueMin="470.0">
470-
<specimenCount id="9D49FD3FB24CFF939867FA2FFDB4F9AD" box="[565,633,1534,1558]" pageId="20" pageNumber="531" type="generic">485 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C355653DB24CFF939AD6F9FCFD3BF9FD" box="[132,758,1581,1606]" pageId="20" pageNumber="531" type="etymology">
<paragraph id="8BF036B6B24CFF939AD6F9FCFD3BF9FD" blockId="20.[132,758,1581,1606]" box="[132,758,1581,1606]" pageId="20" pageNumber="531">ETYMOLOGY. — Dedicated to the second author’s wife Carmen.</paragraph>
</subSubSection>
<subSubSection id="C355653DB24CFF939AD6F98CFED4F911" pageId="20" pageNumber="531" type="materials_examined">
<paragraph id="8BF036B6B24CFF939AD6F98CFED4F911" blockId="20.[132,775,1629,1707]" pageId="20" pageNumber="531">
OTHER MATERIAL EXAMINED. —
<specimenCount id="9D49FD3FB24CFF939BE8F98FFE31F9CE" box="[442,508,1629,1654]" pageId="20" pageNumber="531" type="adult">2 adult</specimenCount>
sh. and
<specimenCount id="9D49FD3FB24CFF93981BF98FFD65F9CE" box="[585,680,1629,1654]" pageId="20" pageNumber="531" type="juvenile">1 juvenile</specimenCount>
, DW126;
<specimenCount id="9D49FD3FB24CFF939AD6F9A9FF07F92B" box="[132,202,1656,1680]" pageId="20" pageNumber="531" type="adult">1 adult</specimenCount>
spm., DW129;
<specimenCount id="9D49FD3FB24CFF939B3DF9A9FE2EF92B" box="[367,483,1656,1680]" pageId="20" pageNumber="531" type="immature">3 immature</specimenCount>
spm, 9 sh. in early postlarval stage, DW130.
</paragraph>
</subSubSection>
<subSubSection id="C355653DB24CFF939AD6F902FB28F9F4" pageId="20" pageNumber="531" type="description">
<paragraph id="8BF036B6B24CFF939AD6F902FED8F957" blockId="20.[132,775,1747,2030]" box="[132,277,1747,1773]" pageId="20" pageNumber="531">DESCRIPTION</paragraph>
<paragraph id="8BF036B6B24CFF939AD6F925FBA8FB6B" blockId="20.[132,775,1747,2030]" lastBlockId="20.[813,1456,1014,1615]" pageId="20" pageNumber="531">
Shell medium-sized, consisting of 4 1/2 whorls, greyish white, very solid, smooth and glossy, with a moderately elevated spire. Apex blunt and rounded, protoconch about
<quantity id="4CB79B53B24CFF9398FAF8E5FD24F8F5" box="[680,745,1844,1870]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="531" unit="mm" value="2.0">2 mm</quantity>
in diameter, consisting of approximately 1 3/4 whorl, not delimited from the teleoconch. Spire whorls very slightly convex, with a thin suture. Last whorl representing more than 80% of total height. Aperture elongate, narrow, parallel-sided except at the adapical end where it tapers. Outer lip smooth inside, bordered externally by a broad, well delimited, evenly thickened labial varix, very slightly receding at its adapical end, forming there a moderately pronounced shoulder against the penultimate whorl. There are four columellar plaits, stout with a flattened crest, decreasing in size towards the abapical part of the columella, occupying two-thirds of the aperture length. No columellar callus.
</paragraph>
<paragraph id="8BF036B6B24CFF939911FB07FB28F9F4" blockId="20.[813,1456,1014,1615]" pageId="20" pageNumber="531">
Animal (
<figureCitation id="13742A33B24CFF9399F1FB07FC3FFB4B" box="[931,1010,1237,1264]" captionStart="FIG" captionStartId="22.[132,143,1292,1310]" captionTargetBox="[226,680,220,1228]" captionTargetId="figure@22.[132,804,172,1280]" captionTargetPageId="22" captionText="FIG. 15. — Marginella carmenae n. sp., shipboard drawing of a living paratype from DW126 (shell 14.3 mm)." figureDoi="http://doi.org/10.5281/zenodo.3726058" httpUri="https://zenodo.org/record/3726058/files/figure.png" pageId="20" pageNumber="531">Fig. 15</figureCitation>
) colourless, with a broad and short foot extending flat when crawling, reaching about the same length as the shell longitudinally and about 120% of the shell breadth transversally; the propodium with a broad transverse flap and, behind this, an opaque white glandular area visible by transparency. Head bifid as usual in marginellids, with long, slender and tapering tentacles, lacking eyes but presenting a distinct bulge on each side, at the base of the tentacles where eyes should normally be situated. Siphon large, extending anteriorly about one-third of shell length. Dimensions of
<typeStatus id="54F48814B24CFF93997FF9C5FC40F995" box="[813,909,1556,1582]" pageId="20" pageNumber="531" type="holotype">holotype</typeStatus>
: 16.7 height ×
<quantity id="4CB79B53B24CFF939E76F9C4FBBBF994" box="[1060,1142,1557,1583]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="20" pageNumber="531" unit="mm" value="8.2">8.2 mm</quantity>
diameter; of
<typeStatus id="54F48814B24CFF939EACF9C4FAA9F994" box="[1278,1380,1557,1583]" pageId="20" pageNumber="531" type="paratype">paratypes</typeStatus>
, 14.1 ×
<quantity id="4CB79B53B24CFF93997FF9E5FC4BF9F4" box="[813,902,1588,1615]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="20" pageNumber="531" unit="mm" value="7.6">7.6 mm</quantity>
, 14.2 ×
<quantity id="4CB79B53B24CFF93998CF9E5FBFDF9F4" box="[990,1072,1588,1615]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="20" pageNumber="531" unit="mm" value="7.5">7.5mm</quantity>
, 14.3 ×
<quantity id="4CB79B53B24CFF939EDAF9E5FB2FF9F4" box="[1160,1250,1588,1615]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="20" pageNumber="531" unit="mm" value="7.5">7.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C355653DB24CFF91997FF9A5FEC3F995" lastPageId="22" lastPageNumber="533" pageId="20" pageNumber="531" type="discussion">
<paragraph id="8BF036B6B24CFF93997FF9A5FC5CF937" blockId="20.[813,1456,1652,2030]" box="[813,913,1652,1678]" pageId="20" pageNumber="531">REMARKS</paragraph>
<paragraph id="8BF036B6B24CFF93997FF944FB97F836" blockId="20.[813,1456,1652,2030]" pageId="20" pageNumber="531">
Large species of the family
<taxonomicName id="4C4F4D35B24CFF939E15F945FB12F915" authorityName="J. Fleming" authorityYear="1828" box="[1095,1247,1684,1710]" class="Gastropoda" family="Marginellidae" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="531" phylum="Mollusca" rank="family">Marginellidae</taxonomicName>
, widely represented in tropical eastern Atlantic waters, are placed in the genera
<taxonomicName id="4C4F4D35B24CFF93997FF905FB9DF955" authority="Lamarck, 1799" authorityName="Lamarck" authorityYear="1799" box="[813,1104,1748,1774]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="531" phylum="Mollusca" rank="genus">
<emphasis id="B93BEAA4B24CFF93997FF905FC6DF955" box="[813,928,1748,1774]" italics="true" pageId="20" pageNumber="531">Marginella</emphasis>
Lamarck, 1799
</taxonomicName>
,
<emphasis id="B93BEAA4B24CFF939E0FF905FB79F955" box="[1117,1204,1748,1774]" italics="true" pageId="20" pageNumber="531">Glabella</emphasis>
Swainson, 1840,
<taxonomicName id="4C4F4D35B24CFF939F28F904FBF3F8B5" authority="Cossmann, 1899" authorityName="Cossmann" authorityYear="1899" class="Gastropoda" family="Marginellidae" genus="Dentimargo" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="531" phylum="Mollusca" rank="genus">
<emphasis id="B93BEAA4B24CFF939F28F904FCB0F8B5" italics="true" pageId="20" pageNumber="531">Dentimargo</emphasis>
Cossmann, 1899
</taxonomicName>
,
<taxonomicName id="4C4F4D35B24CFF939E1AF924FA4DF8B4" authority="Herrmannsen, 1852" authorityName="Herrmannsen" authorityYear="1852" box="[1096,1408,1780,1807]" class="Gastropoda" family="Marginellidae" genus="Prunum" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="531" phylum="Mollusca" rank="genus">
<emphasis id="B93BEAA4B24CFF939E1AF924FB52F8B5" box="[1096,1183,1781,1806]" italics="true" pageId="20" pageNumber="531">Prunum</emphasis>
Herrmannsen, 1852
</taxonomicName>
and
<taxonomicName id="4C4F4D35B24CFF93997FF8C5FB3CF895" authority="Hinds, 1844" authorityName="Hinds" authorityYear="1844" box="[813,1265,1812,1838]" class="Gastropoda" family="Marginellidae" genus="Volvarina" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="531" phylum="Mollusca" rank="genus">
<emphasis id="B93BEAA4B24CFF93997FF8C5FC5FF895" box="[813,914,1812,1838]" italics="true" pageId="20" pageNumber="531">Volvarina</emphasis>
Hinds, 1844 (
<bibRefCitation id="EFDE4B47B24CFF939E63F8C5FB2BF895" author="COSSIGNANI T." box="[1073,1254,1812,1838]" pageId="20" pageNumber="531" refId="ref22467" refString="COSSIGNANI T. 2006. - Marginellidae &amp; Cystiscidae of the World. L'Informatore Piceno Ancona, 408 p." type="book" year="2006">Cossignani 2006</bibRefCitation>
)
</taxonomicName>
. The first of these genera includes species with a prominent spire and a relatively wide opening without a pronounced labial denticle and without axial sculpture.
</paragraph>
<paragraph id="8BF036B6B24CFF929911F842FAFAF955" blockId="20.[813,1456,1652,2030]" lastBlockId="21.[813,1456,1588,2030]" lastPageId="21" lastPageNumber="532" pageId="20" pageNumber="531">
The majority of
<taxonomicName id="4C4F4D35B24CFF9399B5F842FB9BF816" authorityName="Lamarck" authorityYear="1799" box="[999,1110,1939,1965]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="531" phylum="Mollusca" rank="genus">
<emphasis id="B93BEAA4B24CFF9399B5F842FB9BF816" box="[999,1110,1939,1965]" italics="true" pageId="20" pageNumber="531">Marginella</emphasis>
</taxonomicName>
species (illustrated in
<bibRefCitation id="EFDE4B47B24CFF939F67F842FCAAF875" author="COSSIGNANI T." pageId="20" pageNumber="531" refId="ref22467" refString="COSSIGNANI T. 2006. - Marginellidae &amp; Cystiscidae of the World. L'Informatore Piceno Ancona, 408 p." type="book" year="2006">Cossignani, 2006</bibRefCitation>
) live in shallow water, with a generally characteristic colour pattern. Only
<taxonomicName id="4C4F4D35B24CFF939E4BF802FA67F855" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1049,1450,2003,2030]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="531" phylum="Mollusca" rank="species" species="glabella">
<emphasis id="B93BEAA4B24CFF939E4BF802FB2EF856" box="[1049,1251,2003,2029]" italics="true" pageId="20" pageNumber="531">Marginella glabella</emphasis>
(Linnaeus, 1758)
</taxonomicName>
,
<taxonomicName id="4C4F4D35B24DFF929AD6F9E5FDD8F9F4" authority="Clover, 1990" authorityName="Clover" authorityYear="1990" box="[132,533,1588,1615]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="senegalensis">
<emphasis id="B93BEAA4B24DFF929AD6F9E5FEB4F9F5" box="[132,377,1588,1614]" italics="true" pageId="21" pageNumber="532">Marginella senegalensis</emphasis>
Clover, 1990
</taxonomicName>
,
<taxonomicName id="4C4F4D35B24DFF929870F9E5FED2F9D5" authority="Locard, 1897" authorityName="Locard" authorityYear="1897" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="marocana">
<emphasis id="B93BEAA4B24DFF929870F9E5FCC5F9F5" box="[546,776,1588,1614]" italics="true" pageId="21" pageNumber="532">Marginella marocana</emphasis>
<bibRefCitation id="EFDE4B47B24DFF929AD6F985FED2F9D5" author="LOCARD A." box="[132,287,1620,1646]" pageId="21" pageNumber="532" refId="ref23938" refString="LOCARD A. 1897 - 1898. - Expeditions scientifiques du Travailleur et du Talisman pendant les annees 1880, 1881, 1882 et 1883. Mollusques testaces. Masson, Paris, vol. 1 [1897], p. 1 - 516, pl. 1 - 22; vol 2 [1898], p. 1 - 515, pl. 1 - 18. https: // www. biodiversitylibrary. org / item / 41367" type="journal volume" year="1897">Locard, 1897</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C4F4D35B24DFF929B7FF985FD18F9D5" authority="P. Fischer, 1883" authorityName="P. Fischer" authorityYear="1883" box="[301,725,1620,1647]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="subturrita">
<emphasis id="B93BEAA4B24DFF929B7FF985FDD4F9D5" box="[301,537,1620,1646]" italics="true" pageId="21" pageNumber="532">Marginella subturrita</emphasis>
P. Fischer, 1883
</taxonomicName>
and
<taxonomicName id="4C4F4D35B24DFF929AD6F9A5FD01F935" authority="Espinosa, Ortea &amp; Moro, 2013" authorityName="Espinosa, Ortea &amp; Moro" authorityYear="2013" box="[132,716,1652,1679]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="gustavoi">
<emphasis id="B93BEAA4B24DFF929AD6F9A5FE96F935" box="[132,347,1652,1678]" italics="true" pageId="21" pageNumber="532">Marginella gustavoi</emphasis>
Espinosa, Ortea &amp; Moro, 2013
</taxonomicName>
have been cited in Canarian waters or nearby. The first two are from shallow water and very different from the species found in our material (
<bibRefCitation id="EFDE4B47B24DFF929B13F905FDF0F954" author="COSSIGNANI T." box="[321,573,1748,1775]" pageId="21" pageNumber="532" refId="ref22467" refString="COSSIGNANI T. 2006. - Marginellidae &amp; Cystiscidae of the World. L'Informatore Piceno Ancona, 408 p." type="book" year="2006">Cossignani 2006: 115</bibRefCitation>
, 127).
<taxonomicName id="4C4F4D35B24DFF9298C2F905FF27F8B5" authorityName="Locard" authorityYear="1897" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="marocana">
<emphasis id="B93BEAA4B24DFF9298C2F905FF27F8B5" italics="true" pageId="21" pageNumber="532">Marginella marocana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4F4D35B24DFF929B4DF924FE61F8B5" authorityName="P. Fischer" authorityYear="1883" box="[287,428,1780,1806]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="subturrita">
<emphasis id="B93BEAA4B24DFF929B4DF924FE61F8B5" box="[287,428,1780,1806]" italics="true" pageId="21" pageNumber="532">M. subturrita</emphasis>
</taxonomicName>
belong to a group of deep water species illustrated in
<bibRefCitation id="EFDE4B47B24DFF929B31F8C5FD2BF895" author="BOUCHET P. &amp; WAREN A." box="[355,742,1811,1838]" pageId="21" pageNumber="532" pagination="121 - 296" refId="ref21969" refString="BOUCHET P. &amp; WAREN A. 1985. - Revision of the Northeast Atlantic bathyal and abyssal Neogastropoda excluding Turridae (Mollusca, Gastropoda). Bollettino Malacologico supplement 1: 121 - 296. https: // doi. org / 10.5962 / bhl. title. 140763" type="journal article" year="1985">Bouchet &amp; Warén (1985: 274-275)</bibRefCitation>
, of relatively large size (
<quantity id="4CB79B53B24DFF929B0EF8E5FE18F8F5" box="[348,469,1844,1870]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="21" pageNumber="532" unit="mm" value="25.0" valueMax="30.0" valueMin="20.0">20-30 mm</quantity>
).
<taxonomicName id="4C4F4D35B24DFF929BBAF8E2FD08F8F6" authority="Ortega &amp; Gofas, 2019" authorityName="Ortega &amp; Gofas" authorityYear="2019" box="[488,709,1843,1869]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="carmenae" status="sp. nov.">
<emphasis id="B93BEAA4B24DFF929BBAF8E2FD08F8F6" box="[488,709,1843,1869]" italics="true" pageId="21" pageNumber="532">Marginella carmenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857DFB24DFF92989EF8E5FCCAF8F5" box="[716,775,1844,1870]" pageId="21" pageNumber="532" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13742A33B24DFF929ADEF882FF1AF8D5" box="[140,215,1875,1902]" captionStart="FIG" captionStartId="21.[132,143,1465,1483]" captionTargetBox="[132,1455,215,1406]" captionTargetId="figure@21.[132,1455,215,1406]" captionTargetPageId="21" captionText="FIG. 14. — A-F, Marginella carmenae n. sp.: A, B, holotype from DW128 (16.7 mm); C, D, paratype,same locality (14.7 mm);E, F, paratype,same locality (14.0 mm); G, H, Marginella belcheri Hinds, 1844, specimen from off Mauritania, Atlor sta. 49, 20°00’N, 17°44’W, 200 m, R/V Cornide de Saavedra (Museo Nacional de Ciencias naturales, Madrid) (17.8 mm)." figureDoi="http://doi.org/10.5281/zenodo.3726056" httpUri="https://zenodo.org/record/3726056/files/figure.png" pageId="21" pageNumber="532">Fig. 14</figureCitation>
A-F) is clearly differentiated by the shape of the lip of uniform thickness (swollen in the central part and thinning at the ends in
<taxonomicName id="4C4F4D35B24DFF929AAEF845FE48F816" authorityName="Locard" authorityYear="1897" box="[252,389,1940,1965]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="marocana">
<emphasis id="B93BEAA4B24DFF929AAEF845FE48F816" box="[252,389,1940,1965]" italics="true" pageId="21" pageNumber="532">M. marocana</emphasis>
</taxonomicName>
) and by the elongated, parallel-sided aperture (very wide, almost oval in
<taxonomicName id="4C4F4D35B24DFF929BADF865FD41F876" authorityName="P. Fischer" authorityYear="1883" box="[511,652,1971,1997]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="subturrita">
<emphasis id="B93BEAA4B24DFF929BADF865FD41F876" box="[511,652,1971,1997]" italics="true" pageId="21" pageNumber="532">M. subturrita</emphasis>
</taxonomicName>
). A further difference is that in
<taxonomicName id="4C4F4D35B24DFF929B03F805FE17F856" authorityName="Locard" authorityYear="1897" box="[337,474,2004,2029]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="marocana">
<emphasis id="B93BEAA4B24DFF929B03F805FE17F856" box="[337,474,2004,2029]" italics="true" pageId="21" pageNumber="532">M. marocana</emphasis>
</taxonomicName>
the abapical columellar plait is continued as a ridge to the abapical end of the shell whereas it fades out at a short distance in
<taxonomicName id="4C4F4D35B24DFF929EC0F985FAA3F9D5" authority="Ortega &amp; Gofas, 2019" authorityName="Ortega &amp; Gofas" authorityYear="2019" box="[1170,1390,1620,1646]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="carmenae" status="sp. nov.">
<emphasis id="B93BEAA4B24DFF929EC0F985FAA3F9D5" box="[1170,1390,1620,1646]" italics="true" pageId="21" pageNumber="532">Marginella carmenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857DFB24DFF929F27F984FA7DF9D4" box="[1397,1456,1621,1647]" pageId="21" pageNumber="532" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="4C4F4D35B24DFF92997FF9A5FC3FF935" box="[813,1010,1652,1678]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="gustavo">
<emphasis id="B93BEAA4B24DFF92997FF9A5FC3FF935" box="[813,1010,1652,1678]" italics="true" pageId="21" pageNumber="532">Marginella gustavo</emphasis>
</taxonomicName>
i, the only species so far with a Canarian deep-water
<typeStatus id="54F48814B24DFF9299E3F944FC2CF914" box="[945,993,1685,1711]" pageId="21" pageNumber="532">type</typeStatus>
locality, has a still more globose apex, a shorter and stouter spire and considerably thinner outer lip and columellar plait, with a very broad aperture.
</paragraph>
<caption id="DF30663EB24DFF929AD6FA68FE0BFA43" ID-DOI="http://doi.org/10.5281/zenodo.3726056" ID-Zenodo-Dep="3726056" httpUri="https://zenodo.org/record/3726056/files/figure.png" pageId="21" pageNumber="532" startId="21.[132,143,1465,1483]" targetBox="[132,1455,215,1406]" targetPageId="21">
<paragraph id="8BF036B6B24DFF929AD6FA68FE0BFA43" pageId="21" pageNumber="532">
FIG. 14. —
<emphasis id="B93BEAA4B24DFF929A8EFA68FF36FA70" bold="true" box="[220,251,1465,1483]" pageId="21" pageNumber="532">A -F</emphasis>
,
<emphasis id="B93BEAA4B24DFF929B56FA68FE63FA70" box="[260,430,1465,1483]" italics="true" pageId="21" pageNumber="532">Marginella carmenae</emphasis>
n. sp.:
<emphasis id="B93BEAA4B24DFF929BBAFA68FE38FA70" bold="true" box="[488,501,1465,1483]" pageId="21" pageNumber="532">A</emphasis>
,
<emphasis id="B93BEAA4B24DFF929BAFFA68FDC7FA70" bold="true" box="[509,522,1465,1483]" pageId="21" pageNumber="532">B</emphasis>
, holotype from DW128 (16.7 mm);
<emphasis id="B93BEAA4B24DFF929977FA68FCFEFA70" bold="true" box="[805,819,1465,1483]" pageId="21" pageNumber="532">C</emphasis>
,
<emphasis id="B93BEAA4B24DFF92996EFA68FC87FA70" bold="true" box="[828,842,1465,1483]" pageId="21" pageNumber="532">D</emphasis>
, paratype,same locality (14.7 mm);
<emphasis id="B93BEAA4B24DFF929E3CFA68FBB7FA70" bold="true" box="[1134,1146,1465,1483]" pageId="21" pageNumber="532">E</emphasis>
,
<emphasis id="B93BEAA4B24DFF929ED1FA68FB43FA70" bold="true" box="[1155,1166,1465,1483]" pageId="21" pageNumber="532">F</emphasis>
, paratype,same locality (14.0 mm);
<emphasis id="B93BEAA4B24DFF929AD6FA01FF5FFA59" bold="true" box="[132,146,1488,1506]" pageId="21" pageNumber="532">G</emphasis>
,
<emphasis id="B93BEAA4B24DFF929ACCFA01FF61FA59" bold="true" box="[158,172,1488,1506]" pageId="21" pageNumber="532">H</emphasis>
,
<emphasis id="B93BEAA4B24DFF929AEAFA01FE98FA59" box="[184,341,1488,1506]" italics="true" pageId="21" pageNumber="532">Marginella belcheri</emphasis>
Hinds, 1844, specimen from off Mauritania, Atlor sta. 49, 20°00’N, 17°44’W, 200 m, R/V
<emphasis id="B93BEAA4B24DFF929E1FFA1EFACDFA5A" box="[1101,1280,1487,1506]" italics="true" pageId="21" pageNumber="532">Cornide de Saavedra</emphasis>
(Museo Nacional de Ciencias naturales, Madrid) (17.8 mm).
</paragraph>
</caption>
<paragraph id="8BF036B6B24DFF929911F925FA4AF855" blockId="21.[813,1456,1588,2030]" pageId="21" pageNumber="532">
From the platform of NW Africa, comparable species would be
<taxonomicName id="4C4F4D35B24DFF9299C1F8C5FB70F895" authority="Bozzetti, 1994" authorityName="Bozzetti" authorityYear="1994" box="[915,1213,1812,1838]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="adamkusi">
<emphasis id="B93BEAA4B24DFF9299C1F8C5FBD6F895" box="[915,1051,1812,1838]" italics="true" pageId="21" pageNumber="532">M. adamkusi</emphasis>
Bozzetti, 1994
</taxonomicName>
, which differs by being smaller (
<quantity id="4CB79B53B24DFF9299DAF8E5FC16F8F5" box="[904,987,1844,1870]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="532" unit="mm" value="10.0">10 mm</quantity>
), with a prominent colour pattern and with a swelling in the upper third of the lip that could justify its assignment to
<taxonomicName id="4C4F4D35B24DFF929998F8A5FB8BF836" authorityName="Cossmann" authorityYear="1899" box="[970,1094,1908,1933]" class="Gastropoda" family="Marginellidae" genus="Dentimargo" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="genus">
<emphasis id="B93BEAA4B24DFF929998F8A5FB8BF836" box="[970,1094,1908,1933]" italics="true" pageId="21" pageNumber="532">Dentimargo</emphasis>
</taxonomicName>
.
<taxonomicName id="4C4F4D35B24DFF929E07F8A2FA62F836" authority="Hinds, 1844" authorityName="Hinds" authorityYear="1844" box="[1109,1455,1907,1933]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="532" phylum="Mollusca" rank="species" species="belcheri">
<emphasis id="B93BEAA4B24DFF929E07F8A2FAD3F836" box="[1109,1310,1907,1933]" italics="true" pageId="21" pageNumber="532">Marginella belcheri</emphasis>
Hinds, 1844
</taxonomicName>
(
<figureCitation id="13742A33B24DFF929966F842FC73F815" box="[820,958,1939,1966]" captionStart="FIG" captionStartId="21.[132,143,1465,1483]" captionTargetBox="[132,1455,215,1406]" captionTargetId="figure@21.[132,1455,215,1406]" captionTargetPageId="21" captionText="FIG. 14. — A-F, Marginella carmenae n. sp.: A, B, holotype from DW128 (16.7 mm); C, D, paratype,same locality (14.7 mm);E, F, paratype,same locality (14.0 mm); G, H, Marginella belcheri Hinds, 1844, specimen from off Mauritania, Atlor sta. 49, 20°00’N, 17°44’W, 200 m, R/V Cornide de Saavedra (Museo Nacional de Ciencias naturales, Madrid) (17.8 mm)." figureDoi="http://doi.org/10.5281/zenodo.3726056" httpUri="https://zenodo.org/record/3726056/files/figure.png" pageId="21" pageNumber="532">Fig. 14G, H</figureCitation>
) is larger (
<quantity id="4CB79B53B24DFF929E61F842FB67F815" box="[1075,1194,1939,1966]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="21" pageNumber="532" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">18-20 mm</quantity>
) with a different colour pattern, has a much more pointed apex and the outer lip is thinning out at the adapical end (
<bibRefCitation id="EFDE4B47B24DFF929ECAF802FABAF856" author="GOUD J. &amp; NEEFS J." box="[1176,1399,2003,2029]" pageId="21" pageNumber="532" pagination="25 - 50" refId="ref23083" refString="GOUD J. &amp; NEEFS J. 1996. - The larger marginelliform gastropods Cystiscidae and Marginellidae collected during the CANCAP and Mauritania expeditions in the south-eastern part of the north Atlantic Ocean The genera Persicula, Prunum, Dentimargo, Marginella and Glabella. Vita Marina 433 - 434: 25 - 50." type="journal article" year="1996">Goud &amp; Neefs 1996</bibRefCitation>
).
</paragraph>
<caption id="DF30663EB24EFF919AD6FADDFEBAFA8E" ID-DOI="http://doi.org/10.5281/zenodo.3726058" ID-Zenodo-Dep="3726058" httpUri="https://zenodo.org/record/3726058/files/figure.png" pageId="22" pageNumber="533" startId="22.[132,143,1292,1310]" targetBox="[226,680,220,1228]" targetPageId="22">
<paragraph id="8BF036B6B24EFF919AD6FADDFEBAFA8E" blockId="22.[132,775,1292,1333]" pageId="22" pageNumber="533">
FIG. 15. —
<taxonomicName id="4C4F4D35B24EFF919A8CFADDFE47FAA5" authority="Ortega &amp; Gofas, 2019" authorityName="Ortega &amp; Gofas" authorityYear="2019" box="[222,394,1292,1310]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="533" phylum="Mollusca" rank="species" species="carmenae" status="sp. nov.">
<emphasis id="B93BEAA4B24EFF919A8CFADDFE47FAA5" box="[222,394,1292,1310]" italics="true" pageId="22" pageNumber="533">Marginella carmenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857DFB24EFF919BC2FADDFE0FFAA5" box="[400,450,1292,1310]" pageId="22" pageNumber="533" rank="species">n. sp.</taxonomicNameLabel>
, shipboard drawing of a living paratype from DW126 (shell 14.3 mm).
</paragraph>
</caption>
<paragraph id="8BF036B6B24EFF919AC9FA47FEC3F995" blockId="22.[132,775,1429,1582]" pageId="22" pageNumber="533">
In
<taxonomicName id="4C4F4D35B24EFF919AEFFA47FE80FA14" authority="Ortega &amp; Gofas, 2019" authorityName="Ortega &amp; Gofas" authorityYear="2019" box="[189,333,1430,1455]" class="Gastropoda" family="Marginellidae" genus="Marginella" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="533" phylum="Mollusca" rank="species" species="carmenae" status="sp. nov.">
<emphasis id="B93BEAA4B24EFF919AEFFA47FE80FA14" box="[189,333,1430,1455]" italics="true" pageId="22" pageNumber="533">M. carmenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857DFB24EFF919B04FA47FE55FA0B" box="[342,408,1430,1456]" pageId="22" pageNumber="533" rank="species">n. sp.</taxonomicNameLabel>
, no definite limit could be seen between protoconch and teleoconch, neither on the live taken specimens nor on more or less worn shells. The approximate limit is taken where growth lines start to be apparent on the shell surface.
</paragraph>
</subSubSection>
</treatment>
</document>
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