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<mods:title id="C7FFF7787A2E0B0B5AA703E3045D04C3">Generic revision of the staphylinid beetle tribe Hoplandriini (Insecta: Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<treatment id="3F272601506FFFDEC21EFE2AFE38FA7F" LSID="urn:lsid:plazi:treatment:3F272601506FFFDEC21EFE2AFE38FA7F" httpUri="http://treatment.plazi.org/id/3F272601506FFFDEC21EFE2AFE38FA7F" lastPageId="36" lastPageNumber="119" pageId="34" pageNumber="117">
<subSubSection id="FF94C49C506FFFD8C21EFE2AFADFFE63" box="[853,1375,385,409]" pageId="34" pageNumber="117" type="nomenclature">
<paragraph id="B7319717506FFFD8C21EFE2AFADFFE63" blockId="34.[853,1375,385,409]" box="[853,1375,385,409]" pageId="34" pageNumber="117">
<heading id="EC79207B506FFFD8C21EFE2AFADFFE63" box="[853,1375,385,409]" centered="true" fontSize="9" level="2" pageId="34" pageNumber="117" reason="2">
SUBGENUS
<taxonomicName id="708EEC94506FFFD8C292FE29FBDDFE62" ID-CoL="8XRD2" authorityName="Pace" authorityYear="1991" box="[985,1117,386,409]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="119" phylum="Arthropoda" rank="subGenus" subGenus="Pedandria">
<emphasis id="85FA4B05506FFFD8C292FE29FBDDFE62" box="[985,1117,386,409]" italics="true" pageId="34" pageNumber="117">PEDANDRIA</emphasis>
</taxonomicName>
<bibRefCitation id="D31FEAE6506FFFD8C52FFE2AFB72FE63" author="Pace R" box="[1124,1266,385,409]" pageId="34" pageNumber="117" pagination="291 - 305" refId="ref38999" refString="Pace R. 1991. Nuove Aleocharinae del Madagascar (Coleoptera, Staphylinidae). Rendiconti, Classe di Scienze Fisiche, Matematiche e Naturali, Atti della Accademia Nazionale Dei Lincei 9 (2): 291 - 305." type="journal article" year="1991">PACE (1991)</bibRefCitation>
(
<figureCitation id="2FB58B92506FFFD8C448FE2AFAD6FE63" box="[1283,1366,385,409]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="34" pageNumber="117">FIG. 18</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FF94C49C506FFFD8C26DFE01FB00FE25" pageId="34" pageNumber="117" type="reference_group">
<paragraph id="B7319717506FFFD8C26DFE01FB00FE25" blockId="34.[806,1422,426,479]" pageId="34" pageNumber="117">
<treatmentCitationGroup id="979EB039506FFFD8C26DFE01FB00FE25" pageId="34" pageNumber="117">
<treatmentCitation id="362FB106506FFFD8C26DFE01FAA1FE45" author="Pace R" box="[806,1313,426,447]" page="301" pageId="34" pageNumber="117" year="1991">
<taxonomicName id="708EEC94506FFFD8C26DFE01FAA1FE45" ID-CoL="8XRD2" authority="Pace, 1991: 301" authorityName="Pace" authorityPageNumber="301" authorityYear="1991" box="[806,1313,426,447]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="subGenus" subGenus="Pedandria">
<emphasis id="85FA4B05506FFFD8C26DFE01FBD7FE45" box="[806,1111,426,447]" italics="true" pageId="34" pageNumber="117">Ditropandria (Pedandria)</emphasis>
<bibRefCitation id="D31FEAE6506FFFD8C521FE01FAA1FE45" author="Pace R" box="[1130,1313,426,447]" pageId="34" pageNumber="117" pagination="291 - 305" refId="ref38999" refString="Pace R. 1991. Nuove Aleocharinae del Madagascar (Coleoptera, Staphylinidae). Rendiconti, Classe di Scienze Fisiche, Matematiche e Naturali, Atti della Accademia Nazionale Dei Lincei 9 (2): 291 - 305." type="journal article" year="1991">Pace, 1991:301</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<treatmentCitation id="362FB106506FFFD8C47CFE01FC1EFE24" author="Hanley RS" page="314" pageId="34" pageNumber="117" year="2002">
<bibRefCitation id="D31FEAE6506FFFD8C47CFE01FC1EFE24" author="Hanley RS" pageId="34" pageNumber="117" pagination="301 - 321" refId="ref37747" refString="Hanley RS. 2002 a. Phylogeny and higher classification of Hoplandriini (Coleoptera: Staphylinidae: Aleocharinae). Systematic Entomology 27: 301 - 321." type="journal article" year="2002">Hanley, 2002a:314</bibRefCitation>
</treatmentCitation>
. Hanley, in press b.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="FF94C49C506FFFDEC26DFE5CFE38FA7F" lastPageId="36" lastPageNumber="119" pageId="34" pageNumber="117" type="materials_examined">
<paragraph id="B7319717506FFFD8C26DFE5CFB06FDD1" blockId="34.[806,1422,503,555]" pageId="34" pageNumber="117">
<emphasis id="85FA4B05506FFFD8C26DFE5CFC37FDF6" box="[806,951,503,524]" italics="true" pageId="34" pageNumber="117">
<typeStatus id="683529B5506FFFD8C26DFE5CFCDBFDF6" box="[806,859,503,524]" pageId="34" pageNumber="117">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="708EEC94506FFFD8C28FFE5CFCEDFDD1" authority="Pace (1991)" authorityName="Pace" authorityYear="1991" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="species" species="absurda" subGenus="Pedandria">
<emphasis id="85FA4B05506FFFD8C28FFE5CFAD1FDF6" box="[964,1361,503,524]" italics="true" pageId="34" pageNumber="117">Ditropandria (Pedandria) absurda</emphasis>
<bibRefCitation id="D31FEAE6506FFFD8C412FE5CFCEDFDD1" author="Pace R" pageId="34" pageNumber="117" pagination="291 - 305" refId="ref38999" refString="Pace R. 1991. Nuove Aleocharinae del Madagascar (Coleoptera, Staphylinidae). Rendiconti, Classe di Scienze Fisiche, Matematiche e Naturali, Atti della Accademia Nazionale Dei Lincei 9 (2): 291 - 305." type="journal article" year="1991">Pace (1991)</bibRefCitation>
</taxonomicName>
by original designation.
</paragraph>
<paragraph id="B7319717506FFFD8C26DFDEFFADDFB99" blockId="34.[806,1422,579,1123]" pageId="34" pageNumber="117">
<emphasis id="85FA4B05506FFFD8C26DFDEFFC1DFDA3" box="[806,925,580,601]" italics="true" pageId="34" pageNumber="117">Diagnosis:</emphasis>
This subgenus is distinguishable from the other genera of
<taxonomicName id="708EEC94506FFFD8C2AAFDC9FBF3FD82" authorityName="HANLEY" authorityYear="2002" box="[993,1139,610,632]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="subTribe" subTribe="Platandriina">Platandriina</taxonomicName>
by the following combination of characters: body more or less small,
<quantity id="70763AF2506FFFD8C26DFD34FC2DFD4F" box="[806,941,671,693]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="34" pageNumber="117" unit="mm" value="2.55" valueMax="2.9" valueMin="2.2">2.2–2.9 mm</quantity>
; eyes average in size, 0.4–0.6 times length of head; apex of galea densely covered with rows of short, fine hairs, giving more or less stubble appearance; labial palps with two articles instead of three, articles 1 + 2 fused; mesosternum with a distinct, heavy ridge with setae; mesocoxal cavities continuous, only partially separated by meso- and metasternal processes; empodial bristle of hind legs slightly shorter than tarsal claws; apices of abdominal terga with typical macrosetae, not large, heavy, black setae; abdominal tergum IX with a distinct crescent-shaped setal pattern; males with secondary sexual features of abdominal tergum VII often with four, small, narrow denticles along midline and tergum
<collectionCode id="D19F0FD2506FFFD8C59FFB85FA84FBB9" box="[1236,1284,1070,1091]" pageId="34" pageNumber="117">VIII</collectionCode>
with many distinct, setiferous denticles along apical margin.
</paragraph>
<paragraph id="B7319717506FFFD8C26DFBD0FC23FB17" blockId="34.[806,1422,1147,1261]" pageId="34" pageNumber="117">
<emphasis id="85FA4B05506FFFD8C26DFBD0FC2EFB6A" box="[806,942,1147,1168]" italics="true" pageId="34" pageNumber="117">Description:</emphasis>
In agreement with
<taxonomicName id="708EEC94506FFFD8C5E2FBD0FAA6FB6A" authorityName="Casey" authorityYear="1893" box="[1193,1318,1147,1168]" class="Insecta" family="Staphylinidae" genus="Platandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B05506FFFD8C5E2FBD0FAA6FB6A" box="[1193,1318,1147,1168]" italics="true" pageId="34" pageNumber="117">Platandria</emphasis>
</taxonomicName>
description, except for the following characters. [1] Lengths of adults
<quantity id="70763AF2506FFFD8C23CFB13FB80FB37" box="[887,1024,1208,1229]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="34" pageNumber="117" unit="mm" value="2.55" valueMax="2.9" valueMin="2.2">2.2–2.9 mm</quantity>
. [2] Body broadly fusiform to parallel-sided.
</paragraph>
<paragraph id="B7319717506FFFD8C26DFAAEFA0CFA28" blockId="34.[806,1422,1284,1491]" pageId="34" pageNumber="117">
<emphasis id="85FA4B05506FFFD8C26DFAAEFCEAFAE0" box="[806,874,1285,1306]" italics="true" pageId="34" pageNumber="117">Head:</emphasis>
[11] Eyes average in size, 0.4–0.6 times length of head. [13] Infraorbital carina more or less strongly developed and incomplete. Antenna (
<figureCitation id="2FB58B92506FFFD8C5AEFAE9FAC5FAA2" box="[1253,1349,1346,1368]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="34" pageNumber="117">Fig. 18B</figureCitation>
) with [15] articles 1–3 elongate, 1.5–2.3 times longer than wide; [16] segment 4 about as long as wide; [17] articles 5–10 quadrate to short, 1.0–1.2 times wider than long, [18] coeloconical sensilla present in segment 11.
</paragraph>
<paragraph id="B7319717506FFFD9C26DFA40FAF3FA1B" blockId="34.[806,1422,1515,1905]" lastBlockId="35.[819,1435,1299,1506]" lastPageId="35" lastPageNumber="118" pageId="34" pageNumber="117">
<emphasis id="85FA4B05506FFFD8C26DFA40FC32F9FA" box="[806,946,1515,1536]" italics="true" pageId="34" pageNumber="117">Mouthparts:</emphasis>
Labrum with epipharyngeal area with [20] poorly delimited, longitudinal sensory field; [21] pores generally absent between medial pore region and lateral sclerotized. Mandible (
<figureCitation id="2FB58B92506FFFD8C5AEF9ECFAC5F9A7" box="[1253,1349,1607,1629]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="34" pageNumber="117">Fig. 18A</figureCitation>
) [22] appearing asymmetrical with right with median tooth; [23] apex more or less acute and curved adorally; [24] condylar molar patch small, width less than 1/5 of basal mandibular width, [25] composed of very small denticles, [26] densely arranged in irregular transverse rows; dorsobasal ‘velvety patch’ [28] large, wider than half of mandibular base, generally composed of [29] numerous, very fine hairs or spinules. Maxilla with [32] lacinia more or less acute apically, [33] teeth on adoral margin relatively long, about 5–10 times longer than wide, and [34] more or less closely placed; [37] galea relatively narrow, subequal to narrower as lacinia at widest area, mostly parallel-sided and broadly rounded apically, [38] membranous in apical 1/2–1/3, [39] densely covered with rows of short, fine hairs, giving stubble appearance. Labium with ligula [45] long, subequal in length to combined lengths of labial palpi 1 and 2, [46] slightly broadened apically and [47] bilobed apically with fork about 1/5 length of ligula, [48] with 2 short spines apically and numerous sensory pores on each lobe, and 2 relatively short spines on apical half; [49] two short, medial setae of prementum present, about 0.8 times as long as ligula, [50] insertion of setae nearly adjacent; [54] lateral pore field typically with a single short setose pore, two asetose pores, and numerous, small pseudopores; [55] hypoglossal lobes moderately long, reaching to middle of ligula, [56] with more or less short, about 1.4 times longer than width of lobe, comb-like, straight setae along [57] entire length of adoral margin. Labial palpi [58] elongate, overall length 8–10 times longer than greatest width, with [59] 2 distinct articles, articles 1 and 2 appearing fused, [60] with distal pore field composed of two small pores; [62] articles 1 and 2 appearing fused, apical segment homologous with segment 3.
<collectingRegion id="754A59F5506EFFD9C278FA67FC14FA1B" box="[819,916,1484,1505]" country="Myanmar" name="Chin" pageId="35" pageNumber="118">Mentum</collectingRegion>
with [65] apical margin slightly concave.
</paragraph>
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<paragraph id="B7319717506EFFD9C1D0FBD3FC34FB3C" blockId="35.[155,1435,1144,1222]" pageId="35" pageNumber="118">
<emphasis id="85FA4B05506EFFD9C1D0FBD3FE90FB71" bold="true" box="[155,272,1144,1163]" pageId="35" pageNumber="118">Figure 18.</emphasis>
<taxonomicName id="708EEC94506EFFD9C050FBD3FD60FB71" authority="Pace." authorityName="Pace." box="[283,736,1144,1163]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="118" phylum="Arthropoda" rank="species" species="ambrensis" subGenus="Pedandria">
<emphasis id="85FA4B05506EFFD9C050FBD3FE26FB71" box="[283,422,1144,1163]" italics="true" pageId="35" pageNumber="118">Ditropandria</emphasis>
(
<emphasis id="85FA4B05506EFFD9C0FCFBD3FDA4FB71" box="[439,548,1144,1163]" italics="true" pageId="35" pageNumber="118">Pedandria</emphasis>
)
<emphasis id="85FA4B05506EFFD9C37EFBD3FD20FB71" box="[565,672,1144,1163]" italics="true" pageId="35" pageNumber="118">ambrensis</emphasis>
Pace.
</taxonomicName>
(A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca.
</paragraph>
</caption>
<paragraph id="B7319717506EFFDEC278F9A2FD1EFEAF" blockId="35.[819,1435,1545,1905]" lastBlockId="36.[142,759,195,341]" lastPageId="36" lastPageNumber="119" pageId="35" pageNumber="118">
<emphasis id="85FA4B05506EFFD9C278F9A2FC0BF9E4" box="[819,907,1545,1566]" italics="true" pageId="35" pageNumber="118">Thorax:</emphasis>
Pronotum (
<figureCitation id="2FB58B92506EFFD9C569F9A2FB01F9E5" box="[1058,1153,1545,1567]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="35" pageNumber="118">Fig. 18C</figureCitation>
) [69] about 1.2 times wider than long, [69a] distinctly wider posteriorly than apically or in middle. Setae [72] directed primarily anterolaterally from midline. Elytra (
<figureCitation id="2FB58B92506EFFD9C47AF9CEFA13F981" box="[1329,1427,1637,1659]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="35" pageNumber="118">Fig. 18D</figureCitation>
) with [75] apico-lateral angles generally deeply sinuate; [76] elytra together about 1.4 times wider than long. Mesosternum (
<figureCitation id="2FB58B92506EFFD9C572F96AFB19F92D" box="[1081,1177,1729,1751]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="35" pageNumber="118">Fig. 18E</figureCitation>
) [80] with distinct medial ridge with numerous small setae. [83] Meso- and metasternal processes contiguous, with distinct flange present touching apex of metasternal process, isthmus absent; [84] mesosternal process more or less truncate to acute at apex, generally forming to contour of apex of metasternal process. Metasternum [85] length about 0.5 times greatest width of mesocoxae. Macrosetae [89] present on metasternum. Legs with [93] empodial bristle slightly shorter than tarsal claws; [94,95] articles 1–4 subequal in length.
</paragraph>
<paragraph id="B73197175069FFDEC1C5FEC6FE7CFE5B" blockId="36.[142,757,364,417]" pageId="36" pageNumber="119">
<emphasis id="85FA4B055069FFDEC1C5FEC6FF7FFE78" box="[142,255,365,386]" italics="true" pageId="36" pageNumber="119">Abdomen:</emphasis>
[101] Tergum IX with distinct crescentshaped setal pattern (
<figureCitation id="2FB58B925069FFDEC0C7FE20FE70FE5B" box="[396,496,395,417]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="36" pageNumber="119">Fig. 18G</figureCitation>
).
</paragraph>
<paragraph id="B73197175069FFDEC1C5FE12FEF1FDB0" blockId="36.[142,758,441,586]" pageId="36" pageNumber="119">
<emphasis id="85FA4B055069FFDEC1C5FE12FD88FE34" box="[142,520,441,462]" italics="true" pageId="36" pageNumber="119">Secondary sexual characteristics:</emphasis>
Males somewhat variable, with [108] tergum VII often with four small, narrow denticles along midline (
<figureCitation id="2FB58B925069FFDEC35BFE5CFDF0FDF7" box="[528,624,503,525]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="36" pageNumber="119">Fig. 18F</figureCitation>
); [109] tergum
<collectionCode id="D19F0FD25069FFDEC182FDBDFF79FDD1" box="[201,249,534,555]" pageId="36" pageNumber="119">VIII</collectionCode>
with distinct longitudinal row of setiferous denticles (
<figureCitation id="2FB58B925069FFDEC04FFD9FFEE4FDB0" box="[260,356,564,586]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="36" pageNumber="119">Fig. 18F</figureCitation>
).
</paragraph>
<paragraph id="B73197175069FFDEC1C5FDC9FD1FFC97" blockId="36.[142,758,609,877]" pageId="36" pageNumber="119">
<emphasis id="85FA4B055069FFDEC1C5FDC9FE81FD8D" box="[142,257,610,631]" italics="true" pageId="36" pageNumber="119">Aedeagus:</emphasis>
(
<figureCitation id="2FB58B925069FFDEC05EFDC9FEF8FD82" box="[277,376,610,632]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="36" pageNumber="119">Fig. 18H</figureCitation>
) Bulb of median lobe [110] short, generally shorter than tube, [111] with distinct, straight, ventral projection, [111a] ventral projection not extended away from apex; internal sac [112] with numerous spinules, few distinct scleritites and distinct flagellum; parameres (
<figureCitation id="2FB58B925069FFDEC08CFD50FD9FFCEB" box="[455,543,763,785]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="36" pageNumber="119">Fig. 18I</figureCitation>
) [114] with 2 short setae at apex; [115] paramerite anterior margin straight with distinct bend; [117a] apex of paramerite with distinct, straight area directed ventrally.
</paragraph>
<paragraph id="B73197175069FFDEC1C5FC2EFF71FBEC" blockId="36.[142,758,901,1046]" pageId="36" pageNumber="119">
<emphasis id="85FA4B055069FFDEC1C5FC2EFEA9FC60" box="[142,297,901,922]" italics="true" pageId="36" pageNumber="119">Spermatheca:</emphasis>
(
<figureCitation id="2FB58B925069FFDEC075FC2EFE18FC61" box="[318,408,901,923]" captionStart="Figure 18" captionStartId="35.[155,233,1144,1163]" captionTargetBox="[155,1431,196,1111]" captionTargetId="figure-402@35.[154,1431,196,1111]" captionTargetPageId="35" captionText="Figure 18. Ditropandria (Pedandria) ambrensis Pace. (A) Mandible, ventral view. (B) Antenna. (C) Pronotum. (D) Left elytron. (E) Meso- and metathoracic region. (F) Abdominal tergum VII (male) (arrows to denticles). (G) Abdominal tergum IX. (H) Aedeagus, lateral view. (I) Paramere, lateral view. (J) Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5437830" httpUri="https://zenodo.org/record/5437830/files/figure.png" pageId="36" pageNumber="119">Fig. 18J</figureCitation>
) [118] Generally long and straight, neck [120] typically very long, straight generally with two distinct bends of about 90–120∞; and tube [121] typically membranous, [122] more or less straight.
</paragraph>
<paragraph id="B73197175069FFDEC1C5FB85FEEDFBBE" blockId="36.[142,365,1070,1092]" box="[142,365,1070,1092]" pageId="36" pageNumber="119">
<emphasis id="85FA4B055069FFDEC1C5FB85FF6CFBB9" box="[142,236,1070,1091]" italics="true" pageId="36" pageNumber="119">Habitat:</emphasis>
Unknown.
</paragraph>
<paragraph id="B73197175069FFDEC1C5FBF7FE38FA7F" blockId="36.[142,758,1116,1414]" pageId="36" pageNumber="119">
<emphasis id="85FA4B055069FFDEC1C5FBF7FE8CFB8B" box="[142,268,1116,1137]" italics="true" pageId="36" pageNumber="119">Comments:</emphasis>
This subgenus is composed of two species,
<emphasis id="85FA4B055069FFDEC1C5FBD0FF21FB6A" box="[142,161,1147,1168]" italics="true" pageId="36" pageNumber="119">D</emphasis>
.
<taxonomicName id="708EEC945069FFDEC1E5FBD0FEF6FB6A" authority="Pace" authorityName="Pace" authorityYear="1991" box="[174,374,1147,1168]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="119" phylum="Arthropoda" rank="species" species="absurda" subGenus="Pedandria">
(
<emphasis id="85FA4B055069FFDEC1FDFBD0FF46FB6A" box="[182,198,1147,1168]" italics="true" pageId="36" pageNumber="119">P</emphasis>
.)
<emphasis id="85FA4B055069FFDEC197FBD0FEBAFB6A" box="[220,314,1147,1168]" italics="true" pageId="36" pageNumber="119">absurda</emphasis>
Pace
</taxonomicName>
and
<emphasis id="85FA4B055069FFDEC0E5FBD0FE41FB6A" box="[430,449,1147,1168]" italics="true" pageId="36" pageNumber="119">D</emphasis>
.
<taxonomicName id="708EEC945069FFDEC085FBD0FD33FB6A" authority="Pace." authorityName="Pace." box="[462,691,1147,1168]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="119" phylum="Arthropoda" rank="species" species="ambrensis" subGenus="Pedandria">
(
<emphasis id="85FA4B055069FFDEC09DFBD0FE66FB6A" box="[470,486,1147,1168]" italics="true" pageId="36" pageNumber="119">P</emphasis>
.)
<emphasis id="85FA4B055069FFDEC0B7FBD0FDF1FB6A" box="[508,625,1147,1168]" italics="true" pageId="36" pageNumber="119">ambrensis</emphasis>
Pace.
</taxonomicName>
Specimens of
<taxonomicName id="708EEC945069FFDEC1BEFB31FE35FB54" box="[245,437,1177,1199]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="119" phylum="Arthropoda" rank="species" species="ambrensis" subGenus="Pedandria">
<emphasis id="85FA4B055069FFDEC1BEFB31FE8EFB55" box="[245,270,1178,1199]" italics="true" pageId="36" pageNumber="119">D.</emphasis>
(
<emphasis id="85FA4B055069FFDEC056FB31FEB2FB55" box="[285,306,1178,1199]" italics="true" pageId="36" pageNumber="119">P.</emphasis>
)
<emphasis id="85FA4B055069FFDEC00BFB32FE35FB54" box="[320,437,1177,1198]" italics="true" pageId="36" pageNumber="119">ambrensis</emphasis>
</taxonomicName>
, the
<typeStatus id="683529B55069FFDEC0A5FB31FD9FFB55" box="[494,543,1178,1199]" pageId="36" pageNumber="119">type</typeStatus>
species of the subgenus
<taxonomicName id="708EEC945069FFDEC191FB13FED1FB37" authorityName="Pace" authorityYear="1991" box="[218,337,1208,1229]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="119" phylum="Arthropoda" rank="subGenus" subGenus="Pedandria">
<emphasis id="85FA4B055069FFDEC191FB13FED1FB37" box="[218,337,1208,1229]" italics="true" pageId="36" pageNumber="119">Pedandria</emphasis>
</taxonomicName>
, were determined from
<bibRefCitation id="D31FEAE65069FFDEC321FB13FD72FB34" author="Pace R" box="[618,754,1208,1230]" pageId="36" pageNumber="119" pagination="291 - 305" refId="ref38999" refString="Pace R. 1991. Nuove Aleocharinae del Madagascar (Coleoptera, Staphylinidae). Rendiconti, Classe di Scienze Fisiche, Matematiche e Naturali, Atti della Accademia Nazionale Dei Lincei 9 (2): 291 - 305." type="journal article" year="1991">Pace (1991)</bibRefCitation>
.
<typeStatus id="683529B55069FFDEC1C5FB7CFF46FB16" box="[142,198,1239,1260]" pageId="36" pageNumber="119">Type</typeStatus>
specimens of this species were not examined because they were unavailable for loan, even after repeated efforts. The above description is comparable to that of
<taxonomicName id="708EEC945069FFDEC04DFA98FE03FAB2" authorityName="Casey" authorityYear="1893" box="[262,387,1331,1352]" class="Insecta" family="Staphylinidae" genus="Platandria" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055069FFDEC04DFA98FE03FAB2" box="[262,387,1331,1352]" italics="true" pageId="36" pageNumber="119">Platandria</emphasis>
</taxonomicName>
, while the abbreviated descriptions of the genus
<taxonomicName id="708EEC945069FFDEC031FAFAFD91FA9C" authorityName="PACE" authorityYear="1984" box="[378,529,1361,1382]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B055069FFDEC031FAFAFD91FA9C" box="[378,529,1361,1382]" italics="true" pageId="36" pageNumber="119">Ditropandria</emphasis>
</taxonomicName>
and the subgenus
<taxonomicName id="708EEC945069FFDEC1C5FADBFEA5FA7F" authorityName="PACE" authorityYear="1984" box="[142,293,1392,1413]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="119" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="85FA4B055069FFDEC1C5FADBFEA5FA7F" box="[142,293,1392,1413]" italics="true" pageId="36" pageNumber="119">Ditropandria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9EC9F67E5069FFDEC065FADAFED8FA7F" box="[302,344,1392,1414]" pageId="36" pageNumber="119" sensu="stricto">
<emphasis id="85FA4B055069FFDEC065FADAFED1FA7C" box="[302,337,1393,1414]" italics="true" pageId="36" pageNumber="119">s. s</emphasis>
.
</taxonomicNameLabel>
are not.
</paragraph>
</subSubSection>
</treatment>
</document>
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