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<mods:title id="84A557D4D7D09AD687BB14AB181DD89E">Generic revision of the staphylinid beetle tribe Hoplandriini (Insecta: Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<paragraph id="B7319717506FFFD8C195FE04FD27FE3C" blockId="34.[222,679,431,455]" box="[222,679,431,455]" pageId="34" pageNumber="117">
<heading id="EC79207B506FFFD8C195FE04FD27FE3C" box="[222,679,431,455]" centered="true" fontSize="9" level="2" pageId="34" pageNumber="117" reason="2">
SUBGENUS
<taxonomicName id="708EEC94506FFFD8C02AFE1BFD17FE3D" ID-CoL="8XKYT" authority="PACE, 1984" authorityName="PACE" authorityYear="1984" box="[353,663,431,455]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B05506FFFD8C02AFE1BFD8CFE3C" box="[353,524,432,455]" italics="true" pageId="34" pageNumber="117">DITROPANDRIA</emphasis>
PACE, 1984
</taxonomicName>
C
</heading>
</paragraph>
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<subSubSection id="FF94C49C506FFFD8C1C5FE73FE86FDF7" pageId="34" pageNumber="117" type="reference_group">
<paragraph id="B7319717506FFFD8C1C5FE73FE86FDF7" blockId="34.[142,758,472,525]" pageId="34" pageNumber="117">
<treatmentCitationGroup id="979EB039506FFFD8C1C5FE73FE86FDF7" pageId="34" pageNumber="117">
<treatmentCitation id="362FB106506FFFD8C1C5FE73FE59FE14" author="Pace R" box="[142,473,472,494]" page="75" pageId="34" pageNumber="117" year="1984">
<taxonomicName id="708EEC94506FFFD8C1C5FE73FE59FE14" ID-CoL="8XKYT" authority="Pace, 1984 c: 75" authorityName="PACE" authorityPageNumber="75" authorityYear="1984" box="[142,473,472,494]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B05506FFFD8C1C5FE73FEA5FE17" box="[142,293,472,493]" italics="true" pageId="34" pageNumber="117">Ditropandria</emphasis>
<bibRefCitation id="D31FEAE6506FFFD8C064FE73FE59FE14" author="Pace R" box="[303,473,472,494]" pageId="34" pageNumber="117" pagination="61 - 79" refId="ref38871" refString="Pace R. 1984 c. Aleocharinae del Madagascar riportate dal Prof. Franz (Coleoptera Staphylinidae) (LX contributo alla conoscenza delle Aleocharinae). Redia 67: 61 - 79." type="journal article" year="1984">Pace, 1984c:75</bibRefCitation>
</taxonomicName>
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.
<treatmentCitation id="362FB106506FFFD8C0ADFE73FD13FE17" author="Pace R" box="[486,659,472,493]" page="301" pageId="34" pageNumber="117" year="1991">
<bibRefCitation id="D31FEAE6506FFFD8C0ADFE73FD13FE17" author="Pace R" box="[486,659,472,493]" pageId="34" pageNumber="117" pagination="291 - 305" refId="ref38999" refString="Pace R. 1991. Nuove Aleocharinae del Madagascar (Coleoptera, Staphylinidae). Rendiconti, Classe di Scienze Fisiche, Matematiche e Naturali, Atti della Accademia Nazionale Dei Lincei 9 (2): 291 - 305." type="journal article" year="1991">Pace, 1991:301</bibRefCitation>
</treatmentCitation>
. Hanley, in press b.
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</paragraph>
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<subSubSection id="FF94C49C506FFFD8C1C5FD8EFB55FEED" pageId="34" pageNumber="117" type="materials_examined">
<paragraph id="B7319717506FFFD8C1C5FD8EFDB6FDA3" blockId="34.[142,757,549,601]" pageId="34" pageNumber="117">
<emphasis id="85FA4B05506FFFD8C1C5FD8EFE9FFDC0" box="[142,287,549,570]" italics="true" pageId="34" pageNumber="117">
<typeStatus id="683529B5506FFFD8C1C5FD8EFF43FDC0" box="[142,195,549,570]" pageId="34" pageNumber="117">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="708EEC94506FFFD8C067FD8EFE9EFDA3" authority="Pace (1984 c)" authorityName="Pace" authorityYear="1984" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="species" species="paupercula" subGenus="Ditropandria">
<emphasis id="85FA4B05506FFFD8C067FD8EFD75FDC0" box="[300,757,549,570]" italics="true" pageId="34" pageNumber="117">Hoplandria (Ditropandria) paupercula</emphasis>
<bibRefCitation id="D31FEAE6506FFFD8C1C5FDEFFE9EFDA3" author="Pace R" box="[142,286,579,601]" pageId="34" pageNumber="117" pagination="61 - 79" refId="ref38871" refString="Pace R. 1984 c. Aleocharinae del Madagascar riportate dal Prof. Franz (Coleoptera Staphylinidae) (LX contributo alla conoscenza delle Aleocharinae). Redia 67: 61 - 79." type="journal article" year="1984">Pace (1984c)</bibRefCitation>
</taxonomicName>
by original designation.
</paragraph>
<paragraph id="B7319717506FFFD8C1C5FDDAFE4EFA18" blockId="34.[142,758,625,1506]" pageId="34" pageNumber="117">
<emphasis id="85FA4B05506FFFD8C1C5FDDAFE23FD7C" box="[142,419,625,646]" italics="true" pageId="34" pageNumber="117">Abbreviated description:</emphasis>
Length of adults
<quantity id="70763AF2506FFFD8C3D2FDDAFD72FD7D" box="[665,754,625,647]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="34" pageNumber="117" unit="mm" value="2.2">2.2 mm</quantity>
. Body broadly fusiform, widest area at apex of elytra; pubescence variably dense on pronotum, elytra, and abdominal terga and sterna; sparse on head. Head slightly broader than long, eyes moderately small, about 0.4 times length of head. Labium with ligula very long, length subequal to length of labial palps, slightly broadened apically and bilobed apically with fork about 1/5 length of ligula; two short, medial setae of prementum present, about 0.3 times length of ligula, insertion of setae nearly adjacent; median pseudopore field narrow and linear without pseudopores, hypoglossal lobes moderately long, reaching to middle of ligula, with relatively short, subequal to slightly longer than width of lobe, comb-like, straight setae along entire length of adoral margin. Labial palpi elongate, overall length 67 times longer than greatest width, with 3 distinct articles. Pronotum slightly transverse, about 1.1 times wider than long. Elytra moderately broad, wider apically than basally; apico-lateral angles sinuate; elytra together about 1.5 times wider than long. Legs with tarsal formula 4-5-5. Abdomen elongate, more or less parallel-sided, tapering apically to broadly pointed apex. Aedeagus with bulb of median lobe elongate, generally subequal to length of tube. Spermatheca slightly bent with three primary divisions: basal bulb, simple and broadly rounded at base; neck, very long with 23 distinct bends of 90120∞; and tube.
</paragraph>
<paragraph id="B7319717506FFFD8C1C5FA51FEEDF9EA" blockId="34.[142,365,1530,1552]" box="[142,365,1530,1552]" pageId="34" pageNumber="117">
<emphasis id="85FA4B05506FFFD8C1C5FA51FF6CF9F5" box="[142,236,1530,1551]" italics="true" pageId="34" pageNumber="117">Habitat:</emphasis>
Unknown.
</paragraph>
<paragraph id="B7319717506FFFD8C1C5F983FB55FEED" blockId="34.[142,758,1576,1905]" lastBlockId="34.[806,1422,196,279]" pageId="34" pageNumber="117">
<emphasis id="85FA4B05506FFFD8C1C5F983FE8CF9C7" box="[142,268,1576,1597]" italics="true" pageId="34" pageNumber="117">Comments:</emphasis>
Specimens of representing species of
<emphasis id="85FA4B05506FFFD8C1C5F9ECFECDF9A6" box="[142,333,1607,1628]" italics="true" pageId="34" pageNumber="117">
<taxonomicName id="708EEC94506FFFD8C1C5F9ECFEA5F9A6" authorityName="PACE" authorityYear="1984" box="[142,293,1607,1628]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="genus" sensu="stricto">Ditropandria</taxonomicName>
<taxonomicNameLabel id="9EC9F67E506FFFD8C060F9ECFECDF9A6" box="[299,333,1607,1628]" pageId="34" pageNumber="117" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
were not examined during the course of this study since none are available for loan. Repeated efforts to borrow these specimens ended in repeated failure. As such, this subgenus cannot be distinguished from the
<taxonomicName id="708EEC94506FFFD8C0C1F96AFDA1F92C" authorityName="PACE" authorityYear="1984" box="[394,545,1729,1750]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="85FA4B05506FFFD8C0C1F96AFDA1F92C" box="[394,545,1729,1750]" italics="true" pageId="34" pageNumber="117">Ditropandria</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="708EEC94506FFFD8C3ECF96AFF3EF90F" authorityName="Pace" authorityYear="1991" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="subGenus" subGenus="Pedandria">
<emphasis id="85FA4B05506FFFD8C3ECF96AFF3EF90F" italics="true" pageId="34" pageNumber="117">Pedandria</emphasis>
</taxonomicName>
with confidence and thus no diagnosis is provided. The above abbreviated description was derived from morphological information extracted from
<bibRefCitation id="D31FEAE6506FFFD8C187F897FEDEF8A8" author="Pace R" box="[204,350,1852,1874]" pageId="34" pageNumber="117" pagination="61 - 79" refId="ref38871" refString="Pace R. 1984 c. Aleocharinae del Madagascar riportate dal Prof. Franz (Coleoptera Staphylinidae) (LX contributo alla conoscenza delle Aleocharinae). Redia 67: 61 - 79." type="journal article" year="1984">Pace (1984c)</bibRefCitation>
, who provided a very brief description and few illustrations of
<taxonomicName id="708EEC94506FFFD8C0AFF8F0FDFBF88A" authorityName="PACE" authorityYear="1984" box="[484,635,1883,1904]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="85FA4B05506FFFD8C0AFF8F0FDFBF88A" box="[484,635,1883,1904]" italics="true" pageId="34" pageNumber="117">Ditropandria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9EC9F67E506FFFD8C3CDF8F0FD29F88A" box="[646,681,1883,1904]" pageId="34" pageNumber="117" sensu="stricto">
<emphasis id="85FA4B05506FFFD8C3CDF8F0FD21F88A" box="[646,673,1883,1904]" italics="true" pageId="34" pageNumber="117">s.s</emphasis>
.
</taxonomicNameLabel>
<taxonomicName id="708EEC94506FFFD8C3F9F8F0FC02FF23" authorityName="PACE" authorityYear="1984" box="[690,898,196,1904]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="85FA4B05506FFFD8C3F9F8F0FC02FF23" box="[690,898,196,1904]" italics="true" pageId="34" pageNumber="117">Ditropandria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9EC9F67E506FFFD8C2D5FF6EFC40FF23" box="[926,960,196,218]" pageId="34" pageNumber="117" sensu="stricto">
<emphasis id="85FA4B05506FFFD8C2D5FF6EFC39FF20" box="[926,953,197,218]" italics="true" pageId="34" pageNumber="117">s.s</emphasis>
.
</taxonomicNameLabel>
contains three described species,
<taxonomicName id="708EEC94506FFFD8C26DFF48FB1AFF02" authority="Pace" authorityName="Pace" authorityYear="1991" box="[806,1178,227,248]" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="species" species="alternata" subGenus="Ditropandria">
<emphasis id="85FA4B05506FFFD8C26DFF48FCB9FF02" box="[806,825,227,248]" italics="true" pageId="34" pageNumber="117">D</emphasis>
. (
<emphasis id="85FA4B05506FFFD8C205FF48FC65FF02" box="[846,997,227,248]" italics="true" pageId="34" pageNumber="117">Ditropandria</emphasis>
)
<emphasis id="85FA4B05506FFFD8C2BFFF48FBDDFF02" box="[1012,1117,227,248]" italics="true" pageId="34" pageNumber="117">alternata</emphasis>
Pace
</taxonomicName>
,
<taxonomicName id="708EEC94506FFFD8C5EEFF48FCEDFEED" authority="(Pace)" baseAuthorityName="Pace" class="Insecta" family="Staphylinidae" genus="Ditropandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="species" species="dauphinensis" subGenus="Ditropandria">
<emphasis id="85FA4B05506FFFD8C5EEFF48FB38FF02" box="[1189,1208,227,248]" italics="true" pageId="34" pageNumber="117">D</emphasis>
. (
<emphasis id="85FA4B05506FFFD8C586FF48FA0DFF02" box="[1229,1421,227,248]" italics="true" pageId="34" pageNumber="117">D.) dauphinensis</emphasis>
(Pace)
</taxonomicName>
, and
<emphasis id="85FA4B05506FFFD8C2E6FEA9FC40FEED" box="[941,960,258,279]" italics="true" pageId="34" pageNumber="117">D</emphasis>
.
<taxonomicName id="708EEC94506FFFD8C286FEAAFB51FEED" authority="(Pace)" baseAuthorityName="Pace" box="[973,1233,257,279]" class="Insecta" family="Staphylinidae" genus="Hoplandria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="117" phylum="Arthropoda" rank="species" species="paupercula" subGenus="Ditropandria">
(
<emphasis id="85FA4B05506FFFD8C29EFEA9FC68FEED" box="[981,1000,258,279]" italics="true" pageId="34" pageNumber="117">D</emphasis>
.)
<emphasis id="85FA4B05506FFFD8C54BFEAAFB00FEEC" box="[1024,1152,257,278]" italics="true" pageId="34" pageNumber="117">paupercula</emphasis>
(Pace)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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