treatments-xml/data/23/05/FC/2305FC1F9369BB5095EC6F72E7C592A6.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.220.2178" ID-GBIF-Dataset="07cc435f-a623-4a17-b510-a38bf0569844" ID-PMC="PMC3459032" ID-Pensoft-Pub="1313-2970-220-1" ID-PubMed="23077429" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-220-1" ModsDocOrigin="ZooKeys 220" ModsDocTitle="Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae)" checkinTime="1451248758882" checkinUser="pensoft" docAuthor="Henry, Thomas J." docDate="2012" docId="2305FC1F9369BB5095EC6F72E7C592A6" docLanguage="en" docName="ZooKeys 220: 1-114" docOrigin="ZooKeys 220" docSource="http://dx.doi.org/10.3897/zookeys.220.2178" docTitle="Tytthus panamensis Carvalho &amp; Southwood" docType="treatment" docVersion="3" lastPageNumber="48" masterDocId="FFA2D657D204FFEEFF8DF82F5143FF91" masterDocTitle="Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae)" masterLastPageNumber="114" masterPageNumber="1" pageNumber="47" updateTime="1668154436991" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Henry, Thomas J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>220</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>114</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.220.2178</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.220.2178</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-220-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152036996" LSID="urn:lsid:plazi:treatment:2305FC1F9369BB5095EC6F72E7C592A6" httpUri="http://treatment.plazi.org/id/2305FC1F9369BB5095EC6F72E7C592A6" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<subSubSection pageId="46" pageNumber="47" type="nomenclature">
<paragraph pageId="46" pageNumber="47">
<taxonomicName LSID="http://species-id.net/wiki/Tytthus_panamensis" authority="Carvalho &amp; Southwood" class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus panamensis" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="panamensis">Tytthus panamensis Carvalho &amp; Southwood</taxonomicName>
Figs 30, 31
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="reference_group">
<paragraph pageId="46" pageNumber="47">
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus panamensis" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="panamensis">Tytthus panamensis</taxonomicName>
<bibRefCitation pageId="46" pageNumber="47">Carvalho and Southwood 1955</bibRefCitation>
: 32 (orig. descrip.); Carvalho 1958: 158 (cat.);
<bibRefCitation author="Carvalho, JCM" journalOrPublisher="Revista Brasileira de Biologia" pageId="81" pageNumber="82" pagination="7 - 16" title="Mirideos neotropicais, CCVIII: Sobre uma colecao enviada para Estudo Pela Academia de Ciencias da California (Hemiptera)." volume="37" year="1977">Carvalho and Afonso 1977</bibRefCitation>
: 9 (list);
<bibRefCitation author="Carvalho, JCM" journalOrPublisher="Journal of the New York Entomological Society" pageId="81" pageNumber="82" pagination="121 - 224" title="Taxonomic names proposed in the insect order Heteroptera by Jose Candido de Melo Carvalho from 1943 to January 1985, with type depositories." volume="95" year="1987">Carvalho and Froeschner 1987</bibRefCitation>
: 218 (list, type info.);
<bibRefCitation pageId="46" pageNumber="47">Schuh 1995</bibRefCitation>
: 249 (cat.);
<bibRefCitation author="Froeschner, RC" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="81" pageNumber="82" pagination="1 - 393" title="True bugs (Heteroptera) of Panama: A synoptic catalog as a contribution to the study of Panamanian biodiversity." volume="61" year="1999">Froeschner 1999</bibRefCitation>
: 133 (list, note).
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="diagnosis">
<paragraph pageId="46" pageNumber="47">Diagnosis.</paragraph>
<paragraph pageId="46" pageNumber="47">
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus panamensis" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="panamensis">Tytthus panamensis</taxonomicName>
is a small species distinguished by the combination of the fuscous head, pronotum, and scutellum; the fuscous antennal segment I, with only the apex pale; the pale yellowish brown antennal segment II; and the uniformly pale legs. All known specimens are macropterous.
</paragraph>
<paragraph pageId="46" pageNumber="47">
It is most similar to
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus pallidus" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pallidus">Tytthus pallidus</taxonomicName>
, sp. n. in overall size and color, including the reduced or apparent lack of an interocular spot on the head and the uniformly pale legs. It differs in having a dark brown antennal segment I, with only the apex pale, and a pale brown antennal segment II, whereas
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus pallidus" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pallidus">Tytthus pallidus</taxonomicName>
has a pale antennal segment I, with only a narrow dark ring at the base, and segment II is dark brown.
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus panamensis" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="panamensis">Tytthus panamensis</taxonomicName>
keys near
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus vagus" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="vagus">Tytthus vagus</taxonomicName>
based on the dark antennal segment I with only the apex pale, but can be distinguished by the uniformly pale yellowish brown antennal segment II and hind femur.
</paragraph>
</subSubSection>
<subSubSection lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="description">
<paragraph pageId="46" pageNumber="47">Description.</paragraph>
<paragraph pageId="46" pageNumber="47">Male (n =1, plus holotype in parentheses) (Fig. 30): Length to apex of hemelytron 2.21 mm (2.37 mm), length to base of cuneus 1.57 mm (1.63 mm), width across hemelytra 2.21 mm (2.37 mm). Head: Length 0.26 mm (0.29 mm), width across eyes 0.48 mm (0.51 mm), interocular width 0.24 mm (0.26 mm). Labium: Length 0.85 mm (0.93 mm). Antenna: Segment I length 0.30 mm (0.35 mm), II 0.88 mm (0.99 mm), III missing (0.56 mm), IV missing (missing). Pronotum: Length 0.22 mm (0.22 mm), basal width 0.58 mm (0.66 mm).</paragraph>
<paragraph lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
Coloration: Head: Uniformly dark brown, pale interocular spot found in all other species (except Tytthus pallidus) apparently absent; eyes dark brown to fuscous. Labium: Uniformly pale yellow. Antenna: Segment I dark brown, narrowly pale at apex; segment II pale yellowish brown; segments III and IV brown. Pronotum: Uniformly dark brown. Mesoscutum: Uniformly dark brown. Scutellum: Uniformly dark brown. Hemelytron: Uniformly pale translucent brown. Ostiolar evaporative area: Reddish brown. Ventral surface: Thorax and abdomen uniformly dark reddish brown. Legs: Procoxae
<pageBreakToken pageId="47" pageNumber="48" start="start">pale</pageBreakToken>
yellow; meso- and metacoxae pale yellow, with bases reddish brown; femora, tibiae, tarsi, and claws uniformly pale yellow.
</paragraph>
<paragraph pageId="47" pageNumber="48">Structure, texture, and vestiture: Head: Shiny, impunctate; buccula slender, tapering posteriorly, ending near level with middle of eye. Labium: Extending to apices of meso- or bases of metacoxae; segment I extending beyond base of head to anterior edge of xyphus before procoxae. Antenna: Segment I with relatively sparse, short, recumbent setae and two, long, subpical, bristlelike setae; segment II thickly set with short, recumbent setae, intermixed with a few more scattered, semierect setae. Pronotum: Shiny, impunctate; anterior angles weakly rounded; lateral margins straight, gradually widening to posterior angles; basal margin weakly sinuate; set with scattered recumbent and semierect setae. Mesoscutum: Shiny, impunctate, broadly exposed; set with a few short, semierect setae. Scutellum: Shiny, impunctate, equilateral; set with a few scattered, short, recumbent and semierect setae. Hemelytron: Macropterous, cuneus and membrane fully developed, extending posteriorly well beyond apex of abdomen; evenly set with relatively long, recumbent setae.</paragraph>
<paragraph pageId="47" pageNumber="48">Male genitalia: Not examined. See note below.</paragraph>
<paragraph pageId="47" pageNumber="48">Female (n = 1) (Fig. 31): Length to apex of hemelytron 2.91mm, length to base of cuneus 1.98 mm, width across hemelytra 0.80 mm. Head: Length 0.34 mm, width across eyes 0.59 mm, interocular width 0.29 mm. Labium: Length 1.20 mm. Antenna: Missing. Pronotum: Length 0.27 mm, basal width 0.74 mm.</paragraph>
<paragraph pageId="47" pageNumber="48">Similar to male in overall appearance, differing primarily in the broader body form.</paragraph>
</subSubSection>
<subSubSection pageId="47" pageNumber="48" type="host">
<paragraph pageId="47" pageNumber="48">Host.</paragraph>
<paragraph pageId="47" pageNumber="48">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="47" pageNumber="48" type="distribution">
<paragraph pageId="47" pageNumber="48">Distribution.</paragraph>
<paragraph pageId="47" pageNumber="48">Described and known only from Panama.</paragraph>
</subSubSection>
<subSubSection pageId="47" pageNumber="48" type="discussion">
<paragraph pageId="47" pageNumber="48">Discussion.</paragraph>
<paragraph pageId="47" pageNumber="48">The one male paratype previously had been dissected and the genitalia apparently were placed in a glass genitalia vial sealed with a cork stopper. Since that dissection, the cork has dried and crumbled and the genitalia are missing from the vial. I now have placed the glass vial inside a polyethylene vial, sealed it with a neoprene stopper, and reattached it to the specimen pin.</paragraph>
</subSubSection>
<subSubSection pageId="47" pageNumber="48" type="type material examined">
<paragraph pageId="47" pageNumber="48">Type material examined.</paragraph>
<paragraph pageId="47" pageNumber="48">
Holotype ♂ (00162200) (USNM): PANAMA:Canal Zone: Corozal,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.98844">8.98844°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-79.5719">79.5719°W</geoCoordinate>
,14 Apr 1912, A. Busck, at light. Paratypes: Same data as for holotype, 1 ♂ (00161395) (USNM), 1 ♀ (00161394) (USNM).
</paragraph>
</subSubSection>
<subSubSection pageId="47" pageNumber="48" type="other specimens examined">
<paragraph pageId="47" pageNumber="48">Other specimens examined.</paragraph>
<paragraph pageId="47" pageNumber="48">
PANAMA:Canal Zone:. Fort Gulick,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="9.31667">9.31667°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-79.86667">79.86667°W</geoCoordinate>
, 21 Aug 1952, F. S. Blanton, 1 ♀ (00162201) (USNM). Darien: Garachine,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.06472">8.06472°N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-78.36277">78.36277°W</geoCoordinate>
, 17 Feb 1953, F. S. Blanton, 1 ♀ (00161396) (USNM).
</paragraph>
</subSubSection>
</treatment>
</document>