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<document id="3ADBDCB2E6DA7055AFB185CA64DA95A4" ID-DOI="10.1080/00222930601088040" ID-ISSN="1464-5262" ID-Zenodo-Dep="5230689" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629461471506" checkinUser="felipe" docAuthor="Guerrieri, Emilio" docDate="2006" docId="DC317F54FF845F4FFE4994D13915FC13" docLanguage="en" docName="JNatHist.40.41-43.2395-2401.pdf" docOrigin="Journal of Natural History 40 (41 - 43)" docSource="http://dx.doi.org/10.1080/00222930601088040" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Cheiloneurus javanus Perkins 1912" docType="treatment" docVersion="2" lastPageNumber="2400" masterDocId="2008072CFF825F48FF9F940B3A0CFF91" masterDocTitle="Review of the encyrtid (Hymenoptera, Chalcidoidea, Encyrtidae) parasitoids of Dryinidae (Hymenoptera, Chrysidoidea, Dryinidae): second contribution and description of a new species of Cheiloneurus" masterLastPageNumber="2401" masterPageNumber="2395" pageNumber="2399" updateTime="1699074071542" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="97CE968C2DAAA4F33322E57792800901">Review of the encyrtid (Hymenoptera, Chalcidoidea, Encyrtidae) parasitoids of Dryinidae (Hymenoptera, Chrysidoidea, Dryinidae): second contribution and description of a new species of Cheiloneurus</mods:title>
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<mods:namePart id="DB1BF2BFEC51AA49BA971416E108B9EF">Guerrieri, Emilio</mods:namePart>
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<mods:title id="6C4CA04789214ED0EAFA31AA6B7A2474">Journal of Natural History</mods:title>
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<mods:date id="CB61BC67BAE63CC849C71ACC710A3246">2006</mods:date>
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<mods:number id="B2E75AA611D50DDFB59C60F49103B7B5">2006-12-28</mods:number>
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<mods:number id="A82B1BDA18754D999CEA7648E6286ED5">40</mods:number>
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<mods:detail id="96116C6642FA1AE33954CA125241B16A" type="issue">
<mods:number id="66A0C6F2BD5667C12CDDCA8A67B51572">41 - 43</mods:number>
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<mods:classification id="86000375D2269C5659FAF7D765B68A0B">journal article</mods:classification>
<mods:identifier id="F583C7665F1FC2EF539E4E6BEF05250A" type="DOI">10.1080/00222930601088040</mods:identifier>
<mods:identifier id="60D550919E5E51CA6B8667E271175337" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="28A613AC890526FC6DA8C3E3626B2BB4" type="Zenodo-Dep">5230689</mods:identifier>
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<treatment id="DC317F54FF845F4FFE4994D13915FC13" LSID="urn:lsid:plazi:treatment:DC317F54FF845F4FFE4994D13915FC13" httpUri="http://treatment.plazi.org/id/DC317F54FF845F4FFE4994D13915FC13" lastPageId="7" lastPageNumber="2400" pageId="6" pageNumber="2399">
<subSubSection id="1C829DC9FF845F4EFE4994D13951FF63" box="[470,861,218,242]" pageId="6" pageNumber="2399" type="nomenclature">
<paragraph id="5427CE42FF845F4EFE4994D13951FF63" blockId="6.[138,1185,218,340]" box="[470,861,218,242]" pageId="6" pageNumber="2399">
<heading id="0F6F792EFF845F4EFE4994D13951FF63" box="[470,861,218,242]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="2399" reason="2">
<taxonomicName id="9398B5C1FF845F4EFE4994D13951FF63" ID-CoL="5XRJ6" authority="Perkins" authorityName="Perkins" authorityYear="1912" box="[470,861,218,242]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="2399" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="66EC1250FF845F4EFE4994D13951FF63" bold="true" box="[470,861,218,242]" pageId="6" pageNumber="2399">
<emphasis id="66EC1250FF845F4EFE4994D138FEFF63" bold="true" box="[470,754,218,242]" italics="true" pageId="6" pageNumber="2399">Cheiloneurus javanus</emphasis>
Perkins
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1C829DC9FF845F4EFDF794F038D0FE82" box="[616,732,251,275]" pageId="6" pageNumber="2399" type="description">
<paragraph id="5427CE42FF845F4EFDF794F038D0FE82" blockId="6.[138,1185,218,340]" box="[616,732,251,275]" pageId="6" pageNumber="2399">
(
<figureCitation id="CCA3D2C7FF845F4EFDEF94F038DFFE82" box="[624,723,251,275]" captionStart="Figure 2" captionStartId="6.[319,379,1665,1684]" captionTargetBox="[342,982,862,1628]" captionTargetId="figure-216@6.[342,982,862,1628]" captionTargetPageId="6" captionText="Figure 2. Cheiloneurus javanus ♀: (5) antenna and (6) fore wing venation." figureDoi="http://doi.org/10.5281/zenodo.5230693" httpUri="https://zenodo.org/record/5230693/files/figure.png" pageId="6" pageNumber="2399">Figure 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1C829DC9FF845F4EFF1595173B11FEC5" pageId="6" pageNumber="2399" type="reference_group">
<paragraph id="5427CE42FF845F4EFF1595173B11FEC5" blockId="6.[138,1185,218,340]" pageId="6" pageNumber="2399">
<treatmentCitationGroup id="7488E96CFF845F4EFF1595173B11FEC5" pageId="6" pageNumber="2399">
<treatmentCitation id="D539E853FF845F4EFF1595173855FEA5" author="Perkins RCL" box="[138,601,284,308]" page="17" pageId="6" pageNumber="2399" year="1912">
<taxonomicName id="9398B5C1FF845F4EFF1595173855FEA5" ID-CoL="5XRJ6" authority="Perkins, 1912: 17" authorityName="Perkins" authorityPageNumber="17" authorityYear="1912" box="[138,601,284,308]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="2399" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="66EC1250FF845F4EFF1595173B75FEA5" box="[138,377,284,308]" italics="true" pageId="6" pageNumber="2399">Cheiloneurus javanus</emphasis>
<materialsCitation id="E4F0C41FFF845F4EFE1695173855FEA5" box="[393,601,284,308]" collectionCode="BBMH" country="United States of America" location="USA" pageId="6" pageNumber="2399" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<bibRefCitation id="3009B3B3FF845F4EFE1695173855FEA5" author="Perkins RCL" box="[393,601,284,308]" pageId="6" pageNumber="2399" pagination="1 - 20" refId="ref2833" refString="Perkins RCL. 1912. Parasites of the family Dryinidae. Report of the work of the Experimental Station of the Hawaiian Planters' Association. Bulletin of the Hawaiian Sugar Planters' Association Experiment Station, Entomology Series 11: 1 - 20." type="book chapter" year="1912">Perkins, 1912:17</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="8B2370E0FF845F4EFDF3951738D3FEA5" box="[620,735,284,308]" pageId="6" pageNumber="2399" type="lectotype">Lectotype</typeStatus>
<specimenCount id="429E05CBFF845F4EFD71951038F0FEA5" box="[750,764,283,308]" pageId="6" pageNumber="2399" type="female">♀</specimenCount>
,
<collectingCountry id="2C8F8ED2FF845F4EFC8C95173940FEA5" box="[787,844,284,308]" name="United States of America" pageId="6" pageNumber="2399">USA</collectingCountry>
(BBMH), here designated, examined.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1C829DC9FF845F4FFF1595863915FC13" lastPageId="7" lastPageNumber="2400" pageId="6" pageNumber="2399" type="description">
<paragraph id="5427CE42FF845F4EFF1595863B14FE34" blockId="6.[138,280,397,421]" box="[138,280,397,421]" pageId="6" pageNumber="2399">
<emphasis id="66EC1250FF845F4EFF1595863B14FE34" box="[138,280,397,421]" italics="true" pageId="6" pageNumber="2399">Redescription</emphasis>
</paragraph>
<paragraph id="5427CE42FF845F4EFF1595B539D5FD69" blockId="6.[138,1185,446,826]" pageId="6" pageNumber="2399">
<emphasis id="66EC1250FF845F4EFF1595B53AEDFE47" box="[138,225,446,470]" italics="true" pageId="6" pageNumber="2399">Female.</emphasis>
<typeStatus id="8B2370E0FF845F4EFF7195B53B6AFE47" box="[238,358,446,470]" pageId="6" pageNumber="2399" type="lectotype">Lectotype</typeStatus>
: length
<quantity id="936063A7FF845F4EFE5295B5381EFE47" box="[461,530,446,470]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="2399" unit="mm" value="1.0">1 mm</quantity>
. Head yellow, frontovertex brown with faint metallic lustre; antenna (
<figureCitation id="CCA3D2C7FF845F4EFED595D53BA1FE67" box="[330,429,478,502]" captionStart="Figure 2" captionStartId="6.[319,379,1665,1684]" captionTargetBox="[342,982,862,1628]" captionTargetId="figure-216@6.[342,982,862,1628]" captionTargetPageId="6" captionText="Figure 2. Cheiloneurus javanus ♀: (5) antenna and (6) fore wing venation." figureDoi="http://doi.org/10.5281/zenodo.5230693" httpUri="https://zenodo.org/record/5230693/files/figure.png" pageId="6" pageNumber="2399">Figure 2</figureCitation>
(5)) with scape and pedicel light brown, funicle white and clava black; mesoscutum brown with metallic reflections, axillae, scutellum (except brown apex), tegulae, metanotum and propodeum yellow; fore wing strongly infuscate save speculum and a band running from postmarginal vein to posterior margin of wing; legs yellow, apex of tarsi brown; gaster brown, paler in the middle. Head 5× as broad as frontovertex; ocelli forming a strongly acute angle of about 30
<emphasis id="66EC1250FF845F4EFDF29674387BFD06" bold="true" box="[621,631,639,663]" italics="true" pageId="6" pageNumber="2399">°</emphasis>
; antenna (
<figureCitation id="CCA3D2C7FF845F4EFD6F9674395FFD06" box="[752,851,639,663]" captionStart="Figure 2" captionStartId="6.[319,379,1665,1684]" captionTargetBox="[342,982,862,1628]" captionTargetId="figure-216@6.[342,982,862,1628]" captionTargetPageId="6" captionText="Figure 2. Cheiloneurus javanus ♀: (5) antenna and (6) fore wing venation." figureDoi="http://doi.org/10.5281/zenodo.5230693" httpUri="https://zenodo.org/record/5230693/files/figure.png" pageId="6" pageNumber="2399">Figure 2</figureCitation>
(5)) with scape 6× as long as broad; F1 subquadrate and 0.5× as long as pedicel; clava obliquely truncate at apex (truncation 0.5× as long as clava length); linear sensilla on F4–F6 and on clava. Relative measurements: HW 35, FV 7, OOL 1, OCL 4.5, POL 3, SL 24, SW 4.
</paragraph>
<paragraph id="5427CE42FF845F4FFF3A970A3AD8FF51" blockId="6.[138,1185,446,826]" lastBlockId="7.[125,1172,137,192]" lastPageId="7" lastPageNumber="2400" pageId="6" pageNumber="2399">
Fore wing about 3× as long as broad, venation as in
<figureCitation id="CCA3D2C7FF845F4EFCA6970A3996FC88" box="[825,922,769,793]" captionStart="Figure 2" captionStartId="6.[319,379,1665,1684]" captionTargetBox="[342,982,862,1628]" captionTargetId="figure-216@6.[342,982,862,1628]" captionTargetPageId="6" captionText="Figure 2. Cheiloneurus javanus ♀: (5) antenna and (6) fore wing venation." figureDoi="http://doi.org/10.5281/zenodo.5230693" httpUri="https://zenodo.org/record/5230693/files/figure.png" pageId="6" pageNumber="2399">Figure 2</figureCitation>
(6). Relative measurements: FWL 91, FWW 30, SMV 35, MV 19, PMV 5, SV 7. Gaster with ovipositor slightly exserted, a little longer than mid tibia. Relative measurements: MT 102, OL 90, GL 26.
</paragraph>
<caption id="00E79ECAFF845F4EFEA0928A39E1F905" ID-DOI="http://doi.org/10.5281/zenodo.5230693" ID-Zenodo-Dep="5230693" box="[319,1005,1665,1684]" httpUri="https://zenodo.org/record/5230693/files/figure.png" pageId="6" pageNumber="2399" startId="6.[319,379,1665,1684]" targetBox="[342,982,862,1628]" targetPageId="6">
<paragraph id="5427CE42FF845F4EFEA0928A39E1F905" blockId="6.[319,1005,1665,1684]" box="[319,1005,1665,1684]" pageId="6" pageNumber="2399">
Figure 2.
<taxonomicName id="9398B5C1FF845F4EFE00928A3855F905" authorityName="Perkins" authorityYear="1912" box="[415,601,1665,1684]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="2399" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="66EC1250FF845F4EFE00928A3855F905" box="[415,601,1665,1684]" italics="true" pageId="6" pageNumber="2399">Cheiloneurus javanus</emphasis>
</taxonomicName>
♀: (5) antenna and (6) fore wing venation.
</paragraph>
</caption>
<paragraph id="5427CE42FF855F4FFFE294F23B40FE80" blockId="7.[125,332,249,273]" box="[125,332,249,273]" pageId="7" pageNumber="2400">
<emphasis id="66EC1250FF855F4FFFE294F23B40FE80" box="[125,332,249,273]" italics="true" pageId="7" pageNumber="2400">Material examined</emphasis>
</paragraph>
<paragraph id="5427CE42FF855F4FFFE295213B80FEF3" blockId="7.[125,1172,297,354]" pageId="7" pageNumber="2400">
<materialsCitation id="E4F0C41FFF855F4FFFE2952139BFFED3" box="[125,947,297,322]" collectionCode="BBMH" collectorName="Java &amp; Pekalogan F. M." pageId="7" pageNumber="2400" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<typeStatus id="8B2370E0FF855F4FFFE295213AF9FED3" box="[125,245,298,322]" pageId="7" pageNumber="2400" type="lectotype">Lectotype</typeStatus>
:
<specimenCount id="429E05CBFF855F4FFE9D95223B1CFED3" box="[258,272,297,322]" pageId="7" pageNumber="2400" type="female">♀</specimenCount>
,
<collectorName id="F96DAB94FF855F4FFEBE95213B58FED3" box="[289,340,298,322]" pageId="7" pageNumber="2400">Java</collectorName>
,
<collectorName id="F96DAB94FF855F4FFEFE95213825FED3" box="[353,553,298,322]" pageId="7" pageNumber="2400">Pekalogan F. M.</collectorName>
, 1907, here designated (
<collectionCode id="32895687FF855F4FFCD3952139A6FED3" box="[844,938,298,322]" pageId="7" pageNumber="2400">BBMH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E4F0C41FFF855F4FFC2095213B85FEF3" collectionCode="BBMH" collectorName="Java &amp; Pekalogan F. M." pageId="7" pageNumber="2400" specimenCount="4" specimenCount-female="4" typeStatus="paralectotype">
<typeStatus id="8B2370E0FF855F4FFC2095213E67FED3" box="[959,1131,298,322]" pageId="7" pageNumber="2400" type="paralectotype">Paralectotypes</typeStatus>
:
<specimenCount id="429E05CBFF855F4FFBE795213E98FED3" box="[1144,1172,297,322]" pageId="7" pageNumber="2400" type="female">4♀</specimenCount>
same data as lectotype
</materialsCitation>
.
</paragraph>
<paragraph id="5427CE42FF855F4FFFE295903AE2FE22" blockId="7.[125,238,411,435]" box="[125,238,411,435]" pageId="7" pageNumber="2400">
<emphasis id="66EC1250FF855F4FFFE295903AE2FE22" box="[125,238,411,435]" italics="true" pageId="7" pageNumber="2400">Comments</emphasis>
</paragraph>
<paragraph id="5427CE42FF855F4FFFE295C73915FC13" blockId="7.[125,1173,460,898]" pageId="7" pageNumber="2400">
I have examined a card-mounted specimen from BBMH that bears the number ‘‘352’’ handwritten on it, a label handwritten by Perkins with the name: ‘‘
<taxonomicName id="9398B5C1FF855F4FFC0C95E73E98FD95" authorityName="Perkins" authorityYear="1912" box="[915,1172,492,516]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="2400" phylum="Arthropoda" rank="species" species="javanus">Cheiloneurus javanus</taxonomicName>
type’’ and a typewritten label with the following indications: ‘‘Java Pekalogan F. M. 1907’’. On the card, five encyrtids (arranged in two rows) and the remnants of what seems to be an adult of
<taxonomicName id="9398B5C1FF855F4FFF7B96403B88FDF2" authorityName="Rafinesque" authorityYear="1815" box="[228,388,587,611]" class="Insecta" family="Membracidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="2400" phylum="Arthropoda" rank="family">Membracidae</taxonomicName>
are mounted. As the card data and those reported in the original description are coherent, in the interest of nomenclatural stability, I designate the first specimen top left as the
<typeStatus id="8B2370E0FF855F4FFE0E96803832FD32" box="[401,574,651,675]" pageId="7" pageNumber="2400" type="lectotype">LECTOTYPE</typeStatus>
of
<taxonomicName id="9398B5C1FF855F4FFDFD968038D5FD32" authorityName="Perkins" authorityYear="1912" box="[610,729,651,675]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="2400" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="66EC1250FF855F4FFDFD968038D5FD32" box="[610,729,651,675]" italics="true" pageId="7" pageNumber="2400">C. javanus</emphasis>
</taxonomicName>
. The female of
<taxonomicName id="9398B5C1FF855F4FFC1196803E09FD32" authorityName="Perkins" authorityYear="1912" box="[910,1029,651,675]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="2400" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="66EC1250FF855F4FFC1196803E09FD32" box="[910,1029,651,675]" italics="true" pageId="7" pageNumber="2400">C. javanus</emphasis>
</taxonomicName>
is extremely similar to that of
<taxonomicName id="9398B5C1FF855F4FFED196A03BFDFD52" authorityName="Perkins" authorityYear="1906" box="[334,497,683,707]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="2400" phylum="Arthropoda" rank="species" species="gonatopodis">
<emphasis id="66EC1250FF855F4FFED196A03BFDFD52" box="[334,497,683,707]" italics="true" pageId="7" pageNumber="2400">C. gonatopodis</emphasis>
</taxonomicName>
with which it shares the pattern of hyaline and infuscate areas on the fore wing (see Figure
<quantity id="936063A7FF855F4FFD8F96C03840FD72" box="[528,588,715,739]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="7" pageNumber="2400" unit="in" value="14.0">14 in</quantity>
<bibRefCitation id="3009B3B3FF855F4FFDCB96C039ADFD72" author="Guerrieri E &amp; Viggiani G" box="[596,929,715,739]" pageId="7" pageNumber="2400" pagination="305 - 317" refId="ref2712" refString="Guerrieri E, Viggiani G. 2005. A review of the encyrtid (Hymenoptera: Chalcidoidea, Encyrtidae) parasitoids of Dryinidae (Hymenoptera: Chrysidoidea) with description of Cheiloneurus olmii sp. nov. Systematics and Biodiversity 2: 305 - 317." type="journal article" year="2005">Guerrieri and Viggiani 2005</bibRefCitation>
). Females of the two species can be separated by the distribution of linear sensilla on the funicle and by the structure of the clava. In
<taxonomicName id="9398B5C1FF855F4FFE379700382DFCB2" authorityName="Perkins" authorityYear="1912" box="[424,545,779,803]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="2400" phylum="Arthropoda" rank="species" species="javanus">
<emphasis id="66EC1250FF855F4FFE379700382DFCB2" box="[424,545,779,803]" italics="true" pageId="7" pageNumber="2400">C. javanus</emphasis>
</taxonomicName>
, linear sensilla are present on F4–F6 and the clava is obliquely truncate at the apex (for nearly half of its length) and nearly as long as the funicle. In
<taxonomicName id="9398B5C1FF855F4FFF0197413B32FCF3" authorityName="Perkins" authorityYear="1906" box="[158,318,842,866]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="2400" phylum="Arthropoda" rank="species" species="gonatopodis">
<emphasis id="66EC1250FF855F4FFF0197413B32FCF3" box="[158,318,842,866]" italics="true" pageId="7" pageNumber="2400">C. gonatopodis</emphasis>
</taxonomicName>
, linear sensilla are present only on F5 and F6 and the clava is transversely truncate at apex and it is clearly shorter than the funicle.
</paragraph>
</subSubSection>
</treatment>
</document>
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