189 lines
20 KiB
XML
189 lines
20 KiB
XML
<document id="33211F123502C7415B0260AEAE539689" ID="10.11646/zootaxa.4175.2.1" ID-DOI="10.11646/zootaxa.4175.2.1" ID-GBIF-Dataset="d7cbc23a-7a4c-4616-83fd-059fb458f7b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="257286" ID-ZooBank="B264C215-000D-42C5-8AC9-B801872CD182" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476339245459" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2016" docId="953787B9FF9FFFF95CF0FA7A805FFF40" docLanguage="en" docName="zootaxa.4175.2.1.pdf" docOrigin="Zootaxa 4175 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hippasteria phrygiana Parelius 1768" docType="treatment" docVersion="7" lastPageNumber="117" masterDocId="690EFFC1FF8FFFE85C67FF96857BFFB8" masterDocTitle="Deep-sea (> 1000 m) Goniasteridae (Valvatida; Asteroidea) from the North Pacific, including an overview of Sibogaster, Bathyceramaster n. gen. and three new species" masterLastPageNumber="141" masterPageNumber="101" pageNumber="117" updateTime="1698389676701" updateUser="plazi">
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<mods:mods id="EE5ABD6817A8E04502523C7FA51981DF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B7757E11AF7599737042A71D72A0BC54">
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<mods:title id="0D1D48EE759B3A4D18D2B7A6CB191E5D">Deep-sea (> 1000 m) Goniasteridae (Valvatida; Asteroidea) from the North Pacific, including an overview of Sibogaster, Bathyceramaster n. gen. and three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="B6A077BB71CE70DBA7A3A0BE10AE5A27" type="personal">
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<mods:roleTerm id="1DCE48DAC8F259293C8794CB8C131183">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8D56B6E32217D434A3545C916D308EE4">Mah, Christopher L.</mods:namePart>
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<mods:typeOfResource id="D4915A760CE8661DD5394151BA205AD9">text</mods:typeOfResource>
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<mods:title id="5D7D8E3242F92FB5B412EE2ED13A0339">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0F879A767DC52B3E48F14B44E6C880D8">
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<mods:date id="E92DC7711F0745763F7287402703CD04">2016</mods:date>
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<mods:detail id="75D14AB0975AD1D40F9661636340916B" type="volume">
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<mods:number id="62B743BAA422CE63F67D22D08DDDF2B1">4175</mods:number>
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</mods:detail>
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<mods:detail id="EC3783D78CF34D2DF3BA1B35689CE775" type="issue">
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<mods:number id="E26B78CE4936BC57F97BC19A2327551F">2</mods:number>
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<mods:start id="55A03BB110B54544C6C9A7CDD88DF1E0">101</mods:start>
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<mods:end id="80B54432E6CA2A459D13A4B943149C3F">141</mods:end>
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</mods:part>
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<mods:classification id="8B7B4378A9879124B86E151BDB014625">journal article</mods:classification>
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<mods:identifier id="331A033F32C6F93CA49DFDA21020B729" type="DOI">10.11646/zootaxa.4175.2.1</mods:identifier>
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<mods:identifier id="6F32C7CD35B79FCE8F66B0A6E00E62A6" type="ISSN">1175-5326</mods:identifier>
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<treatment id="953787B9FF9FFFF95CF0FA7A805FFF40" ID-DOI="http://doi.org/10.5281/zenodo.6055235" ID-GBIF-Taxon="124996928" ID-Zenodo-Dep="6055235" LSID="urn:lsid:plazi:treatment:953787B9FF9FFFF95CF0FA7A805FFF40" httpUri="http://treatment.plazi.org/id/953787B9FF9FFFF95CF0FA7A805FFF40" lastPageId="17" lastPageNumber="117" pageId="16" pageNumber="117">
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<subSubSection id="55846524FF9FFFF85CF0FA7A863EF9F0" pageId="16" pageNumber="117" type="nomenclature">
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<paragraph id="1D2136AFFF9FFFF85CF0FA7A8711F9BE" blockId="16.[151,618,1516,1542]" box="[151,618,1516,1542]" pageId="16" pageNumber="117">
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<heading id="466981C3FF9FFFF85CF0FA7A8711F9BE" bold="true" box="[151,618,1516,1542]" fontSize="11" level="1" pageId="16" pageNumber="117" reason="1">
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<taxonomicName id="DA9E4D2CFF9FFFF85CF0FA7A8711F9BE" ID-CoL="3LZ4X" authority="Parelius 1768" authorityName="Parelius" authorityYear="1768" box="[151,618,1516,1542]" class="Asteroidea" family="Goniasteridae" genus="Hippasteria" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="117" phylum="Echinodermata" rank="species" species="phrygiana">
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<emphasis id="2FEAEABDFF9FFFF85CF0FA7A8711F9BE" bold="true" box="[151,618,1516,1542]" pageId="16" pageNumber="117">
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<emphasis id="2FEAEABDFF9FFFF85CF0FA7A84D3F9BE" bold="true" box="[151,424,1516,1542]" italics="true" pageId="16" pageNumber="117">Hippasteria phrygiana</emphasis>
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(Parelius 1768)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="1D2136AFFF9FFFF85CF0F9A4863EF9F0" blockId="16.[151,837,1586,1608]" box="[151,837,1586,1608]" pageId="16" pageNumber="117">
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Mah
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<emphasis id="2FEAEABDFF9FFFF85CABF9A58584F9F0" box="[204,255,1586,1608]" italics="true" pageId="16" pageNumber="117">et al.</emphasis>
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2014: 441 (as
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<taxonomicName id="DA9E4D2CFF9FFFF85DF3F9A58760F9F0" box="[404,539,1586,1608]" class="Asteroidea" family="Goniasteridae" genus="Hippasteria" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="117" phylum="Echinodermata" rank="species" species="phrygiana">
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<emphasis id="2FEAEABDFF9FFFF85DF3F9A58760F9F0" box="[404,539,1586,1608]" italics="true" pageId="16" pageNumber="117">H. phrygiana</emphasis>
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</taxonomicName>
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) covers complete synonymy.
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</paragraph>
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</subSubSection>
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<subSubSection id="55846524FF9FFFF95CF0F9E3805FFF40" lastPageId="17" lastPageNumber="118" pageId="16" pageNumber="117" type="discussion">
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<paragraph id="1D2136AFFF9FFFF85CF0F9E381B5F832" blockId="16.[151,1436,1653,2038]" pageId="16" pageNumber="117">
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<emphasis id="2FEAEABDFF9FFFF85CF0F9E38464F936" bold="true" box="[151,287,1653,1678]" pageId="16" pageNumber="117">Comments.</emphasis>
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<bibRefCitation id="790F4B5EFF9FFFF85D42F9E38494F936" box="[293,495,1653,1678]" pageId="16" pageNumber="117" refString="Foltz, D., Fatland, S., Eleaume, M., Markello, K., Howell, K., Neil, K. & Mah, C. L. (2013) Global population divergence of the sea star Hippasteria phrygiana corresponds to onset of the last glacial period of the Pleistocene. Marine Biology, 160 (5), 1285 - 1296." type="journal article">
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Foltz
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<emphasis id="2FEAEABDFF9FFFF85D01F9E084E5F936" box="[358,414,1653,1678]" italics="true" pageId="16" pageNumber="117">et al.</emphasis>
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(2013)
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</bibRefCitation>
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and Mah
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<emphasis id="2FEAEABDFF9FFFF85E39F9E087EDF936" box="[606,662,1653,1678]" italics="true" pageId="16" pageNumber="117">et al.</emphasis>
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(2014) have extensively examined the global phylogeography of this species and concluded that several Pacific
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<taxonomicName id="DA9E4D2CFF9FFFF85EE5F90C8673F909" box="[642,776,1690,1713]" class="Asteroidea" family="Goniasteridae" genus="Hippasteria" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="117" phylum="Echinodermata" rank="genus">
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<emphasis id="2FEAEABDFF9FFFF85EE5F90C8673F909" box="[642,776,1690,1713]" italics="true" pageId="16" pageNumber="117">Hippasteria</emphasis>
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</taxonomicName>
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species, including
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<taxonomicName id="DA9E4D2CFF9FFFF85F81F90C81B1F909" box="[998,1226,1690,1713]" class="Asteroidea" family="Goniasteridae" genus="Hippasteria" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="117" phylum="Echinodermata" rank="species" species="spinosa">
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<emphasis id="2FEAEABDFF9FFFF85F81F90C81B1F909" box="[998,1226,1690,1713]" italics="true" pageId="16" pageNumber="117">Hippasteria spinosa</emphasis>
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</taxonomicName>
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, which have been studied at SCUBA depths in the Northwest Pacific (e.g.,
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<bibRefCitation id="790F4B5EFF9FFFF85F77F92B868EF96E" author="Birkeland" box="[784,1013,1725,1750]" pageId="16" pageNumber="117" refString="Birkeland, C. (1974) Interactions between a sea pen and seven of its predators. Ecological Monographs, 44, 211 - 232." type="journal article" year="1974">Birkeland et a. 1974</bibRefCitation>
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;
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<bibRefCitation id="790F4B5EFF9FFFF85867F92881A3F96E" author="Mauzey" box="[1024,1240,1725,1750]" pageId="16" pageNumber="117" refString="Mauzey, K. P., Birkeland, C. & Dayton, P. K. (1968) Feeding behavior of asteroids and escape responses of their prey in the Puget Sound region. Ecology, 49, 503 - 619." type="journal article" year="1968">
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Mauzey
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<emphasis id="2FEAEABDFF9FFFF85806F92881E3F96E" box="[1121,1176,1725,1750]" italics="true" pageId="16" pageNumber="117">et al.</emphasis>
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1968
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</bibRefCitation>
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) were synonyms. Although
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<taxonomicName id="DA9E4D2CFF9FFFF85D6AF974876AF942" box="[269,529,1761,1786]" class="Asteroidea" family="Goniasteridae" genus="Hippasteria" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="117" phylum="Echinodermata" rank="species" species="phyrgiana">
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<emphasis id="2FEAEABDFF9FFFF85D6AF974876AF942" box="[269,529,1761,1786]" italics="true" pageId="16" pageNumber="117">Hippasteria phyrgiana</emphasis>
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</taxonomicName>
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has not been recorded from below
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<quantity id="DA669B4AFF9FFFF85FA5F9748167F941" box="[962,1052,1762,1786]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="117" unit="m" value="1000.0">1000 m</quantity>
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in the Pacific Northwest, it has apparently been collected from deeper depths elsewhere (e.g, Maluf 1987). Thus, it is possible that
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<taxonomicName id="DA9E4D2CFF9FFFF85962F89080E7F8A6" box="[1285,1436,1797,1822]" class="Asteroidea" family="Goniasteridae" genus="Hippasteria" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="117" phylum="Echinodermata" rank="species" species="phrygiana">
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<emphasis id="2FEAEABDFF9FFFF85962F89080E7F8A6" box="[1285,1436,1797,1822]" italics="true" pageId="16" pageNumber="117">H. phrygiana</emphasis>
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</taxonomicName>
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could occur at deeper depths from the region covered herein. Alternatively it is also possible that there is morphological convergence as there was some morphological similarity between
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<taxonomicName id="DA9E4D2CFF9FFFF85850F8D881B5F8DE" box="[1079,1230,1869,1894]" class="Asteroidea" family="Goniasteridae" genus="Hippasteria" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="117" phylum="Echinodermata" rank="species" species="phrygiana">
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<emphasis id="2FEAEABDFF9FFFF85850F8D881B5F8DE" box="[1079,1230,1869,1894]" italics="true" pageId="16" pageNumber="117">H. phrygiana</emphasis>
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</taxonomicName>
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and specimens of
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<taxonomicName id="DA9E4D2CFF9FFFF85CF0F8E4844FF832" box="[151,308,1905,1930]" class="Asteroidea" family="Goniasteridae" genus="Hippasteria" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="117" phylum="Echinodermata" rank="species" species="californica">
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<emphasis id="2FEAEABDFF9FFFF85CF0F8E4844FF832" box="[151,308,1905,1930]" italics="true" pageId="16" pageNumber="117">H. californica</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="DA9E4D2CFF9FFFF85D0DF8E4849EF832" box="[362,485,1905,1930]" class="Asteroidea" family="Goniasteridae" genus="Hippasteria" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="117" phylum="Echinodermata" rank="species" species="tiburoni">
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<emphasis id="2FEAEABDFF9FFFF85D0DF8E4849EF832" box="[362,485,1905,1930]" italics="true" pageId="16" pageNumber="117">H. tiburoni</emphasis>
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</taxonomicName>
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, all different species but showing very similar diagnostic features.
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</paragraph>
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<paragraph id="1D2136AFFF9FFFF95CA0F803805FFF40" blockId="16.[151,1436,1653,2038]" lastBlockId="17.[151,1437,151,248]" lastPageId="17" lastPageNumber="118" pageId="16" pageNumber="117">
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<materialsCitation id="ADF63CF2FF9FFFF85CA0F80381E4F816" ID-GBIF-Occurrence="1316556628" box="[199,1183,1941,1966]" country="Japan" location="Aleutian Islands" municipality="North Pacific" pageId="16" pageNumber="117" specimenCount="1">
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<emphasis id="2FEAEABDFF9FFFF85CA0F8038422F816" bold="true" box="[199,345,1941,1966]" pageId="16" pageNumber="117">Occurrence.</emphasis>
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(data taken from Mah
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<emphasis id="2FEAEABDFF9FFFF85E38F80087E3F816" box="[607,664,1941,1966]" italics="true" pageId="16" pageNumber="117">et al.</emphasis>
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, 2014)
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<collectingMunicipality id="FD45ACD5FF9FFFF85E97F80386F0F816" box="[752,907,1941,1966]" pageId="16" pageNumber="117">North Pacific</collectingMunicipality>
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:
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<collectingCountry id="6589763FFF9FFFF85FF1F80086A2F815" box="[918,985,1942,1965]" name="Japan" pageId="16" pageNumber="117">Japan</collectingCountry>
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,
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<location id="18416074FF9FFFF85F82F80381E4F816" LSID="urn:lsid:plazi:treatment:953787B9FF9FFFF95CF0FA7A805FFF40:18416074FF9FFFF85F82F80381E4F816" box="[997,1183,1941,1966]" country="Japan" municipality="North Pacific" name="Aleutian Islands" pageId="16" pageNumber="117">Aleutian Islands</location>
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</materialsCitation>
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,
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<materialsCitation id="ADF63CF2FF9FFFF858CBF8038767F86A" ID-GBIF-Occurrence="1316556611" country="Canada" location="North America" pageId="16" pageNumber="117" specimenCount="1" stateProvince="Alaska">
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<collectingRegion id="DF5AF84DFF9FFFF858CBF8038187F816" box="[1196,1276,1941,1966]" country="United States of America" name="Alaska" pageId="16" pageNumber="117">Alaska</collectingRegion>
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, west coast of
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<location id="18416074FF9FFFF85CF0F82F843FF86A" LSID="urn:lsid:plazi:treatment:953787B9FF9FFFF95CF0FA7A805FFF40:18416074FF9FFFF85CF0F82F843FF86A" box="[151,324,1977,2002]" country="Canada" name="North America" pageId="16" pageNumber="117" stateProvince="Alaska">North America</location>
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, including
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<collectingCountry id="6589763FFF9FFFF85DA2F82F8767F86A" box="[453,540,1977,2002]" name="Canada" pageId="16" pageNumber="117">Canada</collectingCountry>
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</materialsCitation>
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||
,
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<materialsCitation id="ADF63CF2FF9FFFF85E4EF82F87CAF86A" ID-GBIF-Occurrence="1316556625" box="[553,689,1977,2002]" country="Canada" location="Washington" pageId="16" pageNumber="117" specimenCount="1" stateProvince="Washington">
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<collectingRegion id="DF5AF84DFF9FFFF85E4EF82F87CAF86A" box="[553,689,1977,2002]" country="United States of America" name="Washington" pageId="16" pageNumber="117">Washington</collectingRegion>
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</materialsCitation>
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,
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<materialsCitation id="ADF63CF2FF9FFFF85ED8F82C8477F84E" ID-GBIF-Occurrence="1316556612" country="Canada" location="Oregon" pageId="16" pageNumber="117" specimenCount="1" stateProvince="Oregon">
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<collectingRegion id="DF5AF84DFF9FFFF85ED8F82C866DF86A" box="[703,790,1978,2002]" country="United States of America" name="Oregon" pageId="16" pageNumber="117">Oregon</collectingRegion>
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, west to President Jackson Seamount, south to
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<collectingRegion id="DF5AF84DFF9FFFF85950F82F8477F84E" isEnumeration="true" pageId="16" pageNumber="117">Southern California</collectingRegion>
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||
</materialsCitation>
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||
.
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<materialsCitation id="ADF63CF2FF9FFFF85D7CF84B8109F84E" ID-GBIF-Occurrence="1316556627" box="[283,1138,2013,2038]" country="New Zealand" location="Solomon Islands" municipality="South Pacific" pageId="16" pageNumber="117" specimenCount="1">
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||
<collectingMunicipality id="FD45ACD5FF9FFFF85D7CF84B84C1F84E" box="[283,442,2013,2038]" pageId="16" pageNumber="117">South Pacific</collectingMunicipality>
|
||
:
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<collectingCountry id="6589763FFF9FFFF85DAEF848871DF84E" box="[457,614,2013,2038]" name="New Zealand" pageId="16" pageNumber="117">New Zealand</collectingCountry>
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||
,
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<location id="18416074FF9FFFF85E11F84B8643F84E" LSID="urn:lsid:plazi:treatment:953787B9FF9FFFF95CF0FA7A805FFF40:18416074FF9FFFF85E11F84B8643F84E" box="[630,824,2013,2038]" country="New Zealand" municipality="South Pacific" name="Solomon Islands" pageId="16" pageNumber="117">Solomon Islands</location>
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||
, Straits of Magellan-Chile
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||
</materialsCitation>
|
||
.
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||
<materialsCitation id="ADF63CF2FF9FFFF958E7F84B80E3FF08" ID-GBIF-Occurrence="1316556633" country="South Africa" county="South Indian" lastPageId="17" lastPageNumber="118" location="North America" municipality="Kerguelen Islands" pageId="16" pageNumber="117" specimenCount="1" stateProvince="Cape">
|
||
<collectingCounty id="F4404E23FF9FFFF858E7F84B8062F84E" box="[1152,1305,2013,2038]" pageId="16" pageNumber="117">South Indian</collectingCounty>
|
||
:
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<collectingMunicipality id="FD45ACD5FF9FFFF9594FF84B8592FF08" lastPageId="17" lastPageNumber="118" pageId="16" pageNumber="117">Kerguelen Islands</collectingMunicipality>
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||
, Crozet, Marion and Bouvet Isles,
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||
<collectingCountry id="6589763FFF9EFFF95E17FF018678FF08" box="[624,771,151,176]" name="South Africa" pageId="17" pageNumber="118">South Africa</collectingCountry>
|
||
. Atlantic: East coast of
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||
<location id="18416074FF9EFFF95869FF0181C2FF08" LSID="urn:lsid:plazi:treatment:953787B9FF9FFFF95CF0FA7A805FFF40:18416074FF9EFFF95869FF0181C2FF08" box="[1038,1209,151,176]" country="South Africa" county="South Indian" municipality="Kerguelen Islands" name="North America" pageId="17" pageNumber="118" stateProvince="Cape">North America</location>
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||
, south to
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||
<collectingRegion id="DF5AF84DFF9EFFF95941FF0E801BFF08" box="[1318,1376,152,176]" country="South Africa" name="Western Cape" pageId="17" pageNumber="118">Cape</collectingRegion>
|
||
Cod
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="ADF63CF2FF9EFFF95CF0FF2A81BDFF6C" ID-GBIF-Occurrence="1316556635" box="[151,1222,188,213]" country="Ireland" location="northern Scotland" municipality="Northeastern Atlantic-North Sea" pageId="17" pageNumber="118" specimenCount="1">
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<collectingMunicipality id="FD45ACD5FF9EFFF95CF0FF2A877FFF6C" box="[151,516,188,213]" pageId="17" pageNumber="118">Northeastern Atlantic-North Sea</collectingMunicipality>
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, Kattegat,
|
||
<location id="18416074FF9EFFF95E1BFF2A8638FF6D" LSID="urn:lsid:plazi:treatment:953787B9FF9FFFF95CF0FA7A805FFF40:18416074FF9EFFF95E1BFF2A8638FF6D" box="[636,835,188,213]" country="Ireland" municipality="Northeastern Atlantic-North Sea" name="northern Scotland" pageId="17" pageNumber="118">northern Scotland</location>
|
||
and northernmost
|
||
<collectingCountry id="6589763FFF9EFFF95872FF2A811CFF6D" box="[1045,1127,188,213]" name="Ireland" pageId="17" pageNumber="118">Ireland</collectingCountry>
|
||
, the UK
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="ADF63CF2FF9EFFF958B7FF2A805FFF40" ID-GBIF-Occurrence="1316556617" country="Ireland" county="South Atlantic-off Argentina" elevation="708" location="Lambert's Bay" municipality="South Africa" pageId="17" pageNumber="118" specimenCount="1" stateProvince="Cape">
|
||
<collectingCounty id="F4404E23FF9EFFF958B7FF2A8472FF40" pageId="17" pageNumber="118">South Atlantic-off Argentina</collectingCounty>
|
||
, west side of
|
||
<collectingMunicipality id="FD45ACD5FF9EFFF95DC6FF498748FF40" box="[417,563,223,248]" pageId="17" pageNumber="118">South Africa</collectingMunicipality>
|
||
,
|
||
<location id="18416074FF9EFFF95E58FF49879FFF40" LSID="urn:lsid:plazi:treatment:953787B9FF9FFFF95CF0FA7A805FFF40:18416074FF9EFFF95E58FF49879FFF40" box="[575,740,223,248]" country="Ireland" county="South Atlantic-off Argentina" municipality="South Africa" name="Lambert's Bay" pageId="17" pageNumber="118" stateProvince="Cape">Lambert’s Bay</location>
|
||
to
|
||
<collectingRegion id="DF5AF84DFF9EFFF95F6FFF76863AFF40" box="[776,833,224,248]" country="South Africa" name="Western Cape" pageId="17" pageNumber="118">Cape</collectingRegion>
|
||
Point.
|
||
<elevation id="96B3D19CFF9EFFF95FF5FF49816AFF4F" box="[914,1041,223,248]" metricMagnitude="2" metricUnit="m" metricValue="7.075" metricValueMax="14.05" metricValueMin="0.1" pageId="17" pageNumber="118" unit="m" value="707.5" valueMax="1405.0" valueMin="10.0">
|
||
<quantity id="DA669B4AFF9EFFF95FF5FF49816AFF4F" box="[914,1041,223,248]" metricMagnitude="2" metricUnit="m" metricValue="7.075" metricValueMax="14.05" metricValueMin="0.1" pageId="17" pageNumber="118" unit="m" value="707.5" valueMax="1405.0" valueMin="10.0">10–1405 m</quantity>
|
||
</elevation>
|
||
(but varies by location).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |