treatments-xml/data/95/37/87/953787B9FF84FFE45CF0F899870FFD89.xml
2024-06-21 12:44:36 +02:00

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<mods:title id="FB1D4B56754EC6F594C8F5801D5A9DC5">Deep-sea (&gt; 1000 m) Goniasteridae (Valvatida; Asteroidea) from the North Pacific, including an overview of Sibogaster, Bathyceramaster n. gen. and three new species</mods:title>
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<mods:date id="9AEB9C888C5FE404A83A734AE1440A95">2016</mods:date>
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<treatment id="953787B9FF84FFE45CF0F899870FFD89" ID-DOI="http://doi.org/10.5281/zenodo.6055203" ID-GBIF-Taxon="124996954" ID-Zenodo-Dep="6055203" LSID="urn:lsid:plazi:treatment:953787B9FF84FFE45CF0F899870FFD89" httpUri="http://treatment.plazi.org/id/953787B9FF84FFE45CF0F899870FFD89" lastPageId="12" lastPageNumber="113" pageId="11" pageNumber="112">
<subSubSection id="55846524FF84FFE35CF0F8998740F8D4" pageId="11" pageNumber="112" type="nomenclature">
<paragraph id="1D2136AFFF84FFE35CF0F89984BDF892" blockId="11.[151,454,1807,1834]" box="[151,454,1807,1834]" pageId="11" pageNumber="112">
<heading id="466981C3FF84FFE35CF0F89984BDF892" bold="true" box="[151,454,1807,1834]" fontSize="11" level="1" pageId="11" pageNumber="112" reason="1">
<taxonomicName id="DA9E4D2CFF84FFE35CF0F89984BDF892" ID-CoL="7NS9D" authority="Verrill, 1899" authorityName="Verrill" authorityYear="1899" box="[151,454,1807,1834]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="112" phylum="Echinodermata" rank="genus">
<emphasis id="2FEAEABDFF84FFE35CF0F89984BDF892" bold="true" box="[151,454,1807,1834]" pageId="11" pageNumber="112">
<emphasis id="2FEAEABDFF84FFE35CF0F8998458F891" bold="true" box="[151,291,1807,1833]" italics="true" pageId="11" pageNumber="112">Ceramaster</emphasis>
<bibRefCitation id="790F4B5EFF84FFE35D4DF88684BDF892" author="Verrill" box="[298,454,1808,1834]" pageId="11" pageNumber="112" refString="Verrill, A. E. (1899) Revision of certain genera and species of starfishes, with descriptions of new forms. Transactions of the Connecticut Academy of Arts and Sciences, 10 (1), 145 - 234." type="journal article" year="1899">Verrill, 1899</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="1D2136AFFF84FFE35CF0F8C08740F8D4" blockId="11.[151,1436,1878,2023]" box="[151,571,1878,1901]" pageId="11" pageNumber="112">
<bibRefCitation id="790F4B5EFF84FFE35CF0F8C08461F8D4" author="Verrill" box="[151,282,1878,1900]" pageId="11" pageNumber="112" refString="Verrill, A. E. (1899) Revision of certain genera and species of starfishes, with descriptions of new forms. Transactions of the Connecticut Academy of Arts and Sciences, 10 (1), 145 - 234." type="journal article" year="1899">Verrill, 1899</bibRefCitation>
: 161 [as
<taxonomicName id="DA9E4D2CFF84FFE35D13F8CE8748F8D4" authority="(Ceramaster)" baseAuthorityName="Ceramaster" box="[372,563,1878,1901]" class="Asteroidea" family="Goniasteridae" genus="Tosia" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="112" phylum="Echinodermata" rank="genus">
<emphasis id="2FEAEABDFF84FFE35D13F8CE84D3F8D5" box="[372,424,1880,1901]" italics="true" pageId="11" pageNumber="112">Tosia</emphasis>
(
<emphasis id="2FEAEABDFF84FFE35DD4F8C18751F8D4" box="[435,554,1879,1900]" italics="true" pageId="11" pageNumber="112">Ceramaster</emphasis>
)
</taxonomicName>
]
</paragraph>
</subSubSection>
<subSubSection id="55846524FF84FFE45CF0F8E384A4FF74" lastPageId="12" lastPageNumber="113" pageId="11" pageNumber="112" type="description">
<paragraph id="1D2136AFFF84FFE35CF0F8E386E1F85F" blockId="11.[151,1436,1878,2023]" pageId="11" pageNumber="112">
<bibRefCitation id="790F4B5EFF84FFE35CF0F8E38466F833" author="Fisher" box="[151,285,1909,1931]" pageId="11" pageNumber="112" refString="Fisher, W. K. (1906) The starfishes of the Hawaiian islands. Bulletin of the United States Fish Commission, 23, 987 - 1130." type="journal article" year="1906">Fisher, 1906</bibRefCitation>
: 1054; 1911: 162, 204;
<bibRefCitation id="790F4B5EFF84FFE35E42F8E387D6F833" author="Verrill" box="[549,685,1909,1931]" pageId="11" pageNumber="112" refString="Verrill, A. E. (1914) Monograph of the shallow-water starfishes of the North Pacific coast from the Arctic Ocean to California. Harriman Alaska series: US National Museum, 14, 1 - 408 pp." type="journal article" year="1914">Verrill, 1914</bibRefCitation>
: 289;
<bibRefCitation id="790F4B5EFF84FFE35E93F8E386F6F833" author="Koehler" box="[756,909,1909,1931]" pageId="11" pageNumber="112" refString="Koehler, R. (1924) Les Echiondermes des Mers d'Europe. 1. Generalites, Asterides, Ophiurides. xii. + 360 pp., 9 pls. Paris, Librarie Octave Doin." type="book" year="1924">Koehler, 1924</bibRefCitation>
: 173;
<bibRefCitation id="790F4B5EFF84FFE35FB3F8E081F3F833" author="Mortensen" box="[980,1160,1909,1931]" pageId="11" pageNumber="112" refString="Mortensen, T. (1927) Handbook of the Echinoderms of the British Isles. ix + 471 pp. 269 figs. Oxford University Press, London." type="book" year="1927">Mortensen, 1927</bibRefCitation>
: 80;
<bibRefCitation id="790F4B5EFF84FFE358A5F8E38016F833" author="Djakonov" box="[1218,1389,1909,1931]" pageId="11" pageNumber="112" refString="Djakonov, A. M. (1950) Morskie Zvezdy Morei SSSR T U 34, 1 - 203. [Translated as Dyakonov. A. M. 1968. Sea stars (Asteroids) of the USSR Seas. Keys to the Fauna of the USSR 34. Zoological Institute of the Academy of Sciences of the USSR, ed. By Strelkov, A. A., Israel Program for scientific translations Ltd. Jerusalem, 183 pp.]" type="book chapter" year="1950">Djakonov, 1950</bibRefCitation>
: 38;
<bibRefCitation id="790F4B5EFF84FFE35CA0F803877AF812" author="Tortonese" box="[199,513,1940,1962]" pageId="11" pageNumber="112" refString="Tortonese, E. &amp; Clark, A. M. (1956) On the generic position of the asteroid Goniodiscus placenta Muller &amp; Troschel. Annals and Magazine of Natural History, 9 (12), 347 - 352." type="journal article" year="1956">Tortonese &amp; A.M. Clark, 1956</bibRefCitation>
: 347;
<bibRefCitation id="790F4B5EFF84FFE35E59F80287A5F812" author="Halpern" box="[574,734,1940,1962]" pageId="11" pageNumber="112" refString="Halpern, J. A. (1970 b) A monographic revision of the goniasterid sea stars of the North Atlantic. 253 pp. Unpublished PhD thesis: University of Miami. [Abstract in Dissertation bstracts International 31 B, 5713 (1971)]" type="book" year="1970" yearSuffix="b">Halpern, 1970b</bibRefCitation>
: 62; 1970c: 212; 1970: 62;
<bibRefCitation id="790F4B5EFF84FFE35F94F80381F1F812" author="Downey" box="[1011,1162,1940,1962]" pageId="11" pageNumber="112" refString="Downey, M. E. (1973) Starfishes from the Caribbean and the Gulf of Mexico. Smithsonian Contributions to Zoology 126, 1 - 158." type="journal article" year="1973">Downey, 1973</bibRefCitation>
: 49;
<bibRefCitation id="790F4B5EFF84FFE358DDF802801EF812" author="McKnight" box="[1210,1381,1940,1962]" pageId="11" pageNumber="112" refString="McKnight, D. G. (1973) Additions to the asteroid fauna of New Zealand: family Goniasteridae. New Zealand Oceanographic Institute Records, 1 (13), 171 - 195." type="journal article" year="1973">McKnight, 1973</bibRefCitation>
: 178;
<bibRefCitation id="790F4B5EFF84FFE35CA0F8258425F870" author="Downey" box="[199,350,1970,1992]" pageId="11" pageNumber="112" refString="Downey, M. E. (1973) Starfishes from the Caribbean and the Gulf of Mexico. Smithsonian Contributions to Zoology 126, 1 - 158." type="journal article" year="1973">Downey, 1973</bibRefCitation>
: 49;
<bibRefCitation id="790F4B5EFF84FFE35DF7F8258674F870" author="Clark" box="[400,783,1970,1992]" pageId="11" pageNumber="112" refString="Clark AM &amp; Courtman-Stock J. (1976) The Echinoderms of Southern Africa. British Museum of Natural History, London. 277 pp." type="book" year="1976">A.M. Clark &amp; Courtman-Stock, 1976</bibRefCitation>
: 61; Downey in
<bibRefCitation id="790F4B5EFF84FFE35FDFF82481D2F870" author="Clark" box="[952,1193,1970,1992]" pageId="11" pageNumber="112" refString="Clark, A. M. &amp; Downey, M. E. (1992) Starfishes of the Atlantic. Chapman and Hall, London." type="book" year="1992">Clark &amp; Downey, 1992</bibRefCitation>
: 231;
<bibRefCitation id="790F4B5EFF84FFE35880F82580E3F870" author="Clark" box="[1255,1432,1970,1992]" pageId="11" pageNumber="112" refString="Clark, A. M. (1993) An index of names of recent Asteroidea - Part 2: Valvatida. E chinoderm Studies, 4, 187 - 366." type="journal article" year="1993">A.M. Clark 1993</bibRefCitation>
: 247;
<bibRefCitation id="790F4B5EFF84FFE35C9DF847873BF85F" author="Clark" box="[250,576,2001,2023]" pageId="11" pageNumber="112" refString="Clark, H. E. S. &amp; McKnight, D. G. (2001) The Marine Fauna of New Zealand: Echinodermata: Asteroidea (sea-stars), Order Valvatida. NIWA Biodiversity Memoir, 117, 1 - 270 pp." type="journal article" year="2001">H.E.S. Clark &amp; McKnight 2001</bibRefCitation>
: 32, Mah 2011: 5 (as
<taxonomicName id="DA9E4D2CFF84FFE35F7CF84486E9F85F" box="[795,914,2002,2023]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="11" pageNumber="112" phylum="Echinodermata" rank="genus">
<emphasis id="2FEAEABDFF84FFE35F7CF84486E9F85F" box="[795,914,2002,2023]" italics="true" pageId="11" pageNumber="112">Ceramaster</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="1D2136AFFF83FFE45CF0FF0E84A4FF74" blockId="12.[151,1212,152,205]" pageId="12" pageNumber="113">
<bibRefCitation id="790F4B5EFF83FFE45CF0FF0E8450FF16" author="Koehler" box="[151,299,152,174]" pageId="12" pageNumber="113" refString="Koehler, R. (1909) An account of the deep-sea Asteroidea collected by the Royal Indian Marine Survey Ship Investigator. Echinoderma of the Indian Museum, Part 5, 115 - 131." type="book chapter" year="1909">Koehler, 1909</bibRefCitation>
: 78 [type species
<taxonomicName id="DA9E4D2CFF83FFE45DBAFF0F8175FF16" authority="Koehler, 1909" authorityName="Koehler" authorityYear="1909" box="[477,1038,152,174]" class="Asteroidea" family="Goniasteridae" genus="Pentagonaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="species" species="mortenseni" subGenus="Philonaster">
<emphasis id="2FEAEABDFF83FFE45DBAFF0F870AFF16" box="[477,625,153,174]" italics="true" pageId="12" pageNumber="113">Pentagonaster</emphasis>
(
<emphasis id="2FEAEABDFF83FFE45EE7FF0E878DFF16" box="[640,758,152,174]" italics="true" pageId="12" pageNumber="113">Philonaster</emphasis>
)
<emphasis id="2FEAEABDFF83FFE45F63FF0F860EFF16" box="[772,885,153,174]" italics="true" pageId="12" pageNumber="113">mortenseni</emphasis>
<bibRefCitation id="790F4B5EFF83FFE45F1BFF0E8175FF16" author="Koehler" box="[892,1038,152,174]" pageId="12" pageNumber="113" refString="Koehler, R. (1909) An account of the deep-sea Asteroidea collected by the Royal Indian Marine Survey Ship Investigator. Echinoderma of the Indian Museum, Part 5, 115 - 131." type="book chapter" year="1909">Koehler, 1909</bibRefCitation>
</taxonomicName>
] (as
<taxonomicName id="DA9E4D2CFF83FFE45859FF0E81CFFF16" box="[1086,1204,152,174]" class="Asteroidea" family="Goniasteridae" genus="Philonaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="genus">
<emphasis id="2FEAEABDFF83FFE45859FF0E81CFFF16" box="[1086,1204,152,174]" italics="true" pageId="12" pageNumber="113">Philonaster</emphasis>
</taxonomicName>
)
<bibRefCitation id="790F4B5EFF83FFE45CF0FF208461FF74" author="Verrill" box="[151,282,182,204]" pageId="12" pageNumber="113" refString="Verrill, A. E. (1914) Monograph of the shallow-water starfishes of the North Pacific coast from the Arctic Ocean to California. Harriman Alaska series: US National Museum, 14, 1 - 408 pp." type="journal article" year="1914">Verrill, 1914</bibRefCitation>
: 292 (As
<taxonomicName id="DA9E4D2CFF83FFE45D1CFF2E84ACFF75" box="[379,471,184,205]" class="Asteroidea" family="Goniasteridae" genus="Tosiaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="genus">
<emphasis id="2FEAEABDFF83FFE45D1CFF2E84ACFF75" box="[379,471,184,205]" italics="true" pageId="12" pageNumber="113">Tosiaster</emphasis>
</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection id="55846524FF83FFE45CF0FF6F870FFD89" pageId="12" pageNumber="113" type="discussion">
<paragraph id="1D2136AFFF83FFE45CF0FF6F87B7FE8E" blockId="12.[151,1437,249,562]" pageId="12" pageNumber="113">
<emphasis id="2FEAEABDFF83FFE45CF0FF6F8464FEAA" bold="true" box="[151,287,249,274]" pageId="12" pageNumber="113">Comments.</emphasis>
Several issues surround the genus
<taxonomicName id="DA9E4D2CFF83FFE45EDBFF6C8644FEA9" box="[700,831,250,273]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="genus">
<emphasis id="2FEAEABDFF83FFE45EDBFF6C8644FEA9" box="[700,831,250,273]" italics="true" pageId="12" pageNumber="113">Ceramaster</emphasis>
</taxonomicName>
, including uncertainty regarding its monophyly and overlap with such genera as
<taxonomicName id="DA9E4D2CFF83FFE45DB2FE8B87B3FE8E" authority="Mah 2011" authorityName="Mah" authorityYear="2011" box="[469,712,285,310]" class="Asteroidea" family="Goniasteridae" genus="Peltaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="genus">
<emphasis id="2FEAEABDFF83FFE45DB2FE8B8741FE8E" box="[469,570,285,310]" italics="true" pageId="12" pageNumber="113">Peltaster</emphasis>
(Mah 2011)
</taxonomicName>
.
</paragraph>
<paragraph id="1D2136AFFF83FFE45CA0FED480EEFE7D" blockId="12.[151,1437,249,562]" pageId="12" pageNumber="113">
I argue that species described herein most closely resembles the type species,
<taxonomicName id="DA9E4D2CFF83FFE4584AFED481AFFEE2" box="[1069,1236,321,346]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="species" species="grenadensis">
<emphasis id="2FEAEABDFF83FFE4584AFED481AFFEE2" box="[1069,1236,321,346]" italics="true" pageId="12" pageNumber="113">C. grenadensis</emphasis>
</taxonomicName>
and share several fundamental synapomorphies, including the presence of wide (W&gt;L) inferomarginal plates, strongly abactinal facing superomarginal plates, and a flattened, pentagonal body shape (R/r&lt;2.0). These characters serve to separate
<emphasis id="2FEAEABDFF83FFE45CF0FE388670FE7E" box="[151,779,429,454]" italics="true" pageId="12" pageNumber="113">
<taxonomicName id="DA9E4D2CFF83FFE45CF0FE388580FE7E" box="[151,251,429,454]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="species" species="clarki">C. clarki</taxonomicName>
,
<taxonomicName id="DA9E4D2CFF83FFE45D62FE3884D5FE7E" box="[261,430,429,454]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="species" species="grenadensis">C. grenadensis</taxonomicName>
,
<taxonomicName id="DA9E4D2CFF83FFE45DDEFE388732FE7D" box="[441,585,430,453]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="species" species="japonicus">C. japonicus</taxonomicName>
,
<taxonomicName id="DA9E4D2CFF83FFE45E34FE388670FE7E" box="[595,779,429,454]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="species" species="leptoceramus">C. leptoceramus</taxonomicName>
</emphasis>
,
<taxonomicName id="DA9E4D2CFF83FFE45F70FE388607FE7E" box="[791,892,429,454]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="species" species="smithi">
<emphasis id="2FEAEABDFF83FFE45F70FE388607FE7E" box="[791,892,429,454]" italics="true" pageId="12" pageNumber="113">C. smithi</emphasis>
</taxonomicName>
and
<taxonomicName id="DA9E4D2CFF83FFE45FD6FE388132FE7D" box="[945,1097,430,453]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="species" species="pointsurae" status="sp. nov.">
<emphasis id="2FEAEABDFF83FFE45FD6FE388132FE7D" box="[945,1097,430,453]" italics="true" pageId="12" pageNumber="113">C. pointsurae</emphasis>
</taxonomicName>
<emphasis id="2FEAEABDFF83FFE45828FE3B81F6FE7E" bold="true" box="[1103,1165,429,454]" pageId="12" pageNumber="113">
<taxonomicNameLabel id="34D957C6FF83FFE45828FE3B81F6FE7E" box="[1103,1165,429,454]" pageId="12" pageNumber="113" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="DA9E4D2CFF83FFE458B6FE3B80F4FE7E" box="[1233,1423,429,454]" class="Asteroidea" family="Goniasteridae" genus="Bathyceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="118" phylum="Echinodermata" rank="genus" status="gen. nov.">
<emphasis id="2FEAEABDFF83FFE458B6FE3B80F4FE7E" box="[1233,1423,429,454]" italics="true" pageId="12" pageNumber="113">Bathyceramaster</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="1D2136AFFF83FFE45CA0FE47870FFD89" blockId="12.[151,1437,249,562]" pageId="12" pageNumber="113">
Descriptions of some
<taxonomicName id="DA9E4D2CFF83FFE45DA4FE44873CFE51" box="[451,583,466,489]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="genus">
<emphasis id="2FEAEABDFF83FFE45DA4FE44873CFE51" box="[451,583,466,489]" italics="true" pageId="12" pageNumber="113">Ceramaster</emphasis>
</taxonomicName>
sp., including
<taxonomicName id="DA9E4D2CFF83FFE45E93FE4481EBFE51" authority="McKnight 1993" authorityName="McKnight" authorityYear="1993" box="[756,1168,465,490]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="species" species="glasbyi">
<emphasis id="2FEAEABDFF83FFE45E93FE4486A8FE52" box="[756,979,465,490]" italics="true" pageId="12" pageNumber="113">Ceramaster glasbyi</emphasis>
<bibRefCitation id="790F4B5EFF83FFE45FBBFE4781EBFE51" author="McKnight" box="[988,1168,465,490]" pageId="12" pageNumber="113" refString="McKnight, D. G. (1993) Records of echinoderms (excluding holothurians) from the Norfolk Ridge and Three Kings Rise north of New Zealand. New Zealand Journal of Zoology, 20, 165 - 190." type="journal article" year="1993">McKnight 1993</bibRefCitation>
</taxonomicName>
display a more stellate shape and lack fasciolar grooves. These will need to be considered more thoroughly pending an overview of the 16 species nominally assigned to
<emphasis id="2FEAEABDFF83FFE45D8CFD8C870FFD89" box="[491,628,538,561]" italics="true" pageId="12" pageNumber="113">
<taxonomicName id="DA9E4D2CFF83FFE45D8CFD8C870BFD89" box="[491,624,538,561]" class="Asteroidea" family="Goniasteridae" genus="Ceramaster" kingdom="Animalia" order="Valvatida" pageId="12" pageNumber="113" phylum="Echinodermata" rank="genus">Ceramaster</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>