treatments-xml/data/8F/91/82/8F9182709A1D596CA19DF5D6BC236333.xml

<document id="9A25D8A831E882CF04B4578BC62A9889" ID-DOI="10.3897/phytokeys.250.136139" ID-publisher-id="136139" URI-arpha="163F7496-5C12-5455-AB48-30AD78073DF0" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1734707767354" checkinUser="pensoft" docAuthor="Sukhorukov, Alexander P., Kushunina, Maria, Nilova, Maya V., Baider, Cláudia &amp; Sennikov, Alexander N." docDate="2024" docId="8F9182709A1D596CA19DF5D6BC236333" docLanguage="en" docName="PhytoKeys 250: 21-94" docOrigin="PhytoKeys 250" docSource="https://phytokeys.pensoft.net/article/136139/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Achyranthes seychellensis Sukhor. 2024, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="94" masterDocId="163F74965C125455AB4830AD78073DF0" masterDocTitle="Africa and Arabia encompass a much greater species diversity in the Achyranthes aspera aggregate (Amaranthaceae, achyranthoid clade): Evidence from morphological and chorological data" masterLastPageNumber="94" masterPageNumber="21" pageNumber="21" updateTime="1734739432256" updateUser="admin">
<mods:mods id="93BF8E2E7E3A92A6E3032AAB7D7E1E90" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0536C355E026B322EE1D96F545951C83">
<mods:title id="3925D648C63392F945A4B0C4325FD7C7">Africa and Arabia encompass a much greater species diversity in the Achyranthes aspera aggregate (Amaranthaceae, achyranthoid clade): Evidence from morphological and chorological data</mods:title>
</mods:titleInfo>
<mods:name id="5001ADBF08992BFA6739BC5D19944D99" type="personal">
<mods:role id="859F476C2FC5207051CAA1CCC798369F">
<mods:roleTerm id="0E3A635E5BEB1AC5731C191C8A0C67EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CA87BBFE476E530CE4252D2123F1149E">Sukhorukov, Alexander P.</mods:namePart>
<mods:nameIdentifier id="CE6DE15694207743B9088E28196FE0AF" type="ORCID">0000-0003-2220-826X</mods:nameIdentifier>
<mods:affiliation id="BA641DC5D195346B31C1315823BF8A8C">Department of Higher Plants, Biological Faculty, Lomonosov Moscow State University, 119234, Moscow, Russia &amp; Laboratory Herbarium (TK), Tomsk State University, Lenin Ave. 36, 634050, Tomsk, Russia</mods:affiliation>
</mods:name>
<mods:name id="33A7C155711A87D168DAE48687B42E92" type="personal">
<mods:role id="5BB0D0472A00D11FB5B6D1CCFBEFCC8C">
<mods:roleTerm id="03A258A4274B570E58426931C5B7C4C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7CA8A1D0B5FA91DB2C22CA1DB5F77745">Kushunina, Maria</mods:namePart>
<mods:nameIdentifier id="526B0070117F17D011B19178B9F60D7D" type="ORCID">0000-0003-2541-6494</mods:nameIdentifier>
<mods:affiliation id="F11658365A0D79797358E4A6C5A7BB23">Department of Higher Plants, Biological Faculty, Lomonosov Moscow State University, 119234, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name id="349B3C43B6D6B471CE1BACC9F7BC713A" type="personal">
<mods:role id="232A2E4C246D24C86C6638E8F9DF12A4">
<mods:roleTerm id="A10AB3B075D6066D9273E8F9C9A5B765">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C04DD70E9D00A93733D116E5492E70C1">Nilova, Maya V.</mods:namePart>
<mods:nameIdentifier id="6B07343865AC7354E9EC2DF9308B4DAE" type="ORCID">0000-0001-5340-5915</mods:nameIdentifier>
<mods:affiliation id="8B01DB746C0CAF554B98C4713B4C1464">Department of Higher Plants, Biological Faculty, Lomonosov Moscow State University, 119234, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name id="7E1830623E8AEC12F60195FFB7DB42EC" type="personal">
<mods:role id="9ABC680F488B661D8F81717BF0836DFE">
<mods:roleTerm id="C43A71005A298B87CD41872A40480299">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="92CEDF405F1F2C1F0CA2084207F0C41E">Baider, Cláudia</mods:namePart>
<mods:nameIdentifier id="5FA6EDB21D76517C61DAB39321E21DD6" type="ORCID">0000-0002-2203-2076</mods:nameIdentifier>
<mods:affiliation id="994AC53DA2AD0B751534BE86E5B4842F">Department of Plant Physiology, Biological Faculty, Lomonosov Moscow State University, 119234, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name id="211E01975C5BB6DD68EE84D34697AF16" type="personal">
<mods:role id="16EEDAB4B799659429A48CA519FC6C3F">
<mods:roleTerm id="63B5713088AC1859FBABB8011E102545">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="89157B2E6AAE6BE88FF1EFB4E9FBFADD">Sennikov, Alexander N.</mods:namePart>
<mods:nameIdentifier id="B727F681BD443446EB232F1E8BFA839C" type="ORCID">0000-0001-6664-7657</mods:nameIdentifier>
<mods:affiliation id="EEAA9426E49C424DB2429B883BBED209">The Mauritius Herbarium, RE Vaughan Building, Agricultural Services, Ministry of Agro-Industry and Food Security, Réduit, 80835, Mauritius</mods:affiliation>
</mods:name>
<mods:typeOfResource id="01A94859F63338E2FFE83CFC31AC62F2">text</mods:typeOfResource>
<mods:relatedItem id="9920C9CE0B28E8569DCF01B265E6FA3E" type="host">
<mods:titleInfo id="0245D1D907201FE89E77F093B43641B6">
<mods:title id="6A6D9E8EC0F7466A3AFBD26FA4483286">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="5CB98F2620C85473895624CEDFCFEF68">
<mods:date id="C909B230F503FBB417C6DC923272BAAF">2024</mods:date>
<mods:detail id="97D5C456F6A1E03DBA0359C7C9AF20BC" type="pubDate">
<mods:number id="D2F36DC9B2ADC9E8DD74F1574D0E4486">2024-12-20</mods:number>
</mods:detail>
<mods:detail id="B7648390D5A49D2889A29C7E5145793C" type="volume">
<mods:number id="98B6B151A37ABBE0CA82B1379B32F931">250</mods:number>
</mods:detail>
<mods:extent id="5A9337593D362827A26D01FFA1D29790" unit="page">
<mods:start id="697A57EAB3716401C9D366A3C08BBF97">21</mods:start>
<mods:end id="87DFA61FCB3A927172F07AFB9571F14A">94</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="790DA7E7C91CCF82003253BF197130DE">journal article</mods:classification>
<mods:identifier id="8F8B23F18C480E0F1BFE04DE1A738B55" type="DOI">10.3897/phytokeys.250.136139</mods:identifier>
</mods:mods>
<treatment id="8F9182709A1D596CA19DF5D6BC236333" ID-DOI="http://doi.org/10.5281/zenodo.14536763" ID-Zenodo-Dep="14536763" ID-arpha="8F918270-9A1D-596C-A19D-F5D6BC236333" ID-ipni="urn:lsid:ipni.org:names:77353796-1" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333" LSID-IPNI="urn:lsid:ipni.org:names:77353796-1" httpUri="http://treatment.plazi.org/id/8F9182709A1D596CA19DF5D6BC236333" scope_family="Amaranthaceae">
<subSubSection id="4913E2B6892B18D34C68DD555E6E734B" type="nomenclature">
<paragraph id="3A74DE2A6FD5184E5EC4DE0A8D34DEBC">
<taxonomicName id="68153C49A0801D914A1A9FF5B11388C2" authority="Sukhor." authorityName="Sukhor." authorityYear="2024" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="seychellensis" status="sp. nov.">
<emphasis id="E4E946BBBAAD5F99CE495836C0B7719F" italics="true">Achyranthes seychellensis</emphasis>
Sukhor.
</taxonomicName>
<taxonomicNameLabel id="14107B57AACC037A1236CB241CD27299" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3PBK" type="Holotype">
<paragraph id="25CCB6B6E9889C2DBE84AF9EB99516E7">
<heading id="8F4736D3E48848AC5A4B015A8CDB0F28" reason="title">
<typeStatus id="07DF6412861FE2E953C6DF5EB52E2F6A" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="099DBAF0BE95A8CF122156BC6C805F0E">
<materialsCitation id="B2384AD1CC843A1DB12A895E95533D6C" collectedFrom="coconut plantation" collectingDate="1984-08-18" collectionCode="G" collectorName="G. Ogureeva" country="Seychelles" location="Farquhar Atoll" specimenCode="MW 0586306, LE 00019370" specimenCount="1" typeStatus="holotype">
<collectingCountry id="4704C609837A3976D67015C747283E13" name="Seychelles">Seychelles</collectingCountry>
, 
<location id="CB1C96CA7B287379A474EB362889741E" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:CB1C96CA7B287379A474EB362889741E" country="Seychelles" name="Farquhar Atoll">Farquhar Atoll</location>
, 
<collectedFrom id="BFD90D09A3CD2241E0E583FF08069DE4">coconut plantation</collectedFrom>
, 
<date id="502EA263305D2F2293A197EA8A49753C" value="1984-08-18">
<collectingDate id="6A1E58D055B503CF15408692CE633FD6" value="1984-08-18">18 August 1984</collectingDate>
</date>
, 
<emphasis id="5DA5E6C2A89C64976778FC9797DF9275" italics="true">
<collectorName id="1E40579D92AE8C7D723940BDDDDAF730">G. Ogureeva</collectorName>
608
</emphasis>
(
<specimenCode id="8837A63B11625A6E8ADC1229FFD37858" collectionCode="MW" country="Netherlands" name="Museum Wasmann">MW 0586306</specimenCode>
!, 
<typeStatus id="BC38DC80F44121A747AAD885CFDBC6DE" type="isotype">isotype</typeStatus>
<specimenCode id="ABAB45E2EE4529E4D3D416A58A476FD1" collectionCode="LE" country="Brazil" name="Servico de Microbiologia e Imunologia">LE 00019370</specimenCode>
!).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEQBK" type="description">
<paragraph id="69AA8F28C9C53A4EDB72F4C7C3349F93">
<heading id="45A6630D5C4DD1AFD54AF60EB1B931CB" reason="title">Description.</heading>
</paragraph>
<paragraph id="4D3C27BFB2626FDDBF47368E32430225">
(Fig. 
<figureCitation id="AE98D3F036232AC704B239E1A56E5002" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Holotype of Achyranthes seychellensis Sukhor., sp. nov. (Seychelles, Farquhar Atoll, coconut plantation, 18 Aug 1984, G. Ogureeva 608, MW 0586306)." figureDoi="10.3897/phytokeys.250.136139.figure16" httpUri="https://binary.pensoft.net/fig/1203860">16</figureCitation>
). Annual herb up to 
<quantity id="691D281E923A91D4B7DCB0F1AF6BFC2A" metricMagnitude="0" metricUnit="m" metricValue="1.5" unit="m" value="1.5">1.5 m</quantity>
tall; stem stout, basally ± roundish, four-angled in the inflorescence; leaf petioles 
<quantity id="C0F128FD474C03BCF9110C67BB95E1C5" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" unit="mm" value="17.5" valueMax="25.0" valueMin="10.0">10–25 mm</quantity>
long, blades 80–120 × 
<quantity id="89EE217DDC4528817BFED13FCCADDE74" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" unit="mm" value="45.0" valueMax="60.0" valueMin="30.0">30–60 mm</quantity>
, cuneate, tip shortly acuminate, green above and pale green below, slightly pubescent on veins; inflorescence up to 
<quantity id="094849CAF4B4B15D727CEA83D37F7FD2" metricMagnitude="-1" metricUnit="m" metricValue="2.5" unit="cm" value="25.0">25 cm</quantity>
long with hirsute axis; bract 3.5–4.0 mm long; bracteoles 3.2–4.0 mm long; perianth 
<quantity id="4216898A3B3DE3854CE956917389294D" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.5–5.5 mm</quantity>
long, ± equal, deep rose or purple inside; stamens 5, anthers 0.75–1.0 mm long; pseudostaminodes 1.25–2.0 mm long, brownish or white when dry, fimbriate; style with stigma 
<quantity id="7F283E16F6151C7104A6CFA9CFA146D6" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2–2.7 mm</quantity>
long; fruit (without style) ± 2.0 mm long.
</paragraph>
<caption id="D9EAC5C3ADD9CB23DD28AAB2A373CCDB" ID-DOI="10.3897/phytokeys.250.136139.figure16" ID-arpha="346ED6EA-EE5F-5B23-896C-A1DFCEFC177B" httpUri="https://binary.pensoft.net/fig/1203860" startId="F16">
<paragraph id="FE95B2F081221B02B0EACBD0D1C15CBA">
<label id="E5CAFAA9A977E0BBF0F23D22A21985F7">Figure 16.</label>
</paragraph>
<paragraph id="B5FD4EAE9B1BA8E3BFFE12F7366D71E1">
Holotype of 
<taxonomicName id="125E0999A689E771677C46DB9A43E91F" authority="Sukhor." authorityName="Sukhor." authorityYear="2024" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="seychellensis" status="sp. nov.">
<emphasis id="81F248477E6E18EC75C567AAE9848620" italics="true">Achyranthes seychellensis</emphasis>
Sukhor.
</taxonomicName>
, 
<taxonomicNameLabel id="950AFEEED0BCFF8ACAEA5F3AD05CC91B" rank="species">sp. nov.</taxonomicNameLabel>
(Seychelles, Farquhar Atoll, coconut plantation, 18 Aug 1984, 
<emphasis id="05F4FE55C3080F8D958E2BEFFF013B11" italics="true">G. Ogureeva 608</emphasis>
, MW 0586306).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EIRBK" type="habitat">
<paragraph id="168F9E889FA65C487EB4FB791A416786">
<heading id="FA1822655EB157A901506EEA36ABEDCD" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="F0B6BC85AE1AE1597754D6DB7E8A7BD7">
Calcareous and rocky areas at elevations of 
<quantity id="86DE0EE66522F4B24D8B87AA06F57D64" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" unit="m" value="100.0" valueMax="200.0" valueMin="0.0">0–200 m</quantity>
a. s. l. It can be a common weed in coconut plantations.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENRBK" type="Flowering at fruiting">
<paragraph id="F0BF92C7A87047481EC37B9BD4B92484">
<heading id="E69EC410E83E1C138008E72F4086DFC6" reason="title">Flowering at fruiting.</heading>
</paragraph>
<paragraph id="D49F8B6E4FB7FD3804FE5496DD3A14F5">Around the year.</paragraph>
</subSubSection>
<subSubSection id="SECID0ESRBK" type="Additional specimens seen">
<paragraph id="5F224384B3F5C2DCEA2C873A873F7353">
<heading id="3DF7EBC8BB2D603254A650AAFCC44C94" reason="title">Additional specimens seen.</heading>
</paragraph>
<paragraph id="1E5EB93B3E1FC8CCE842508FE9548E98">
<materialsCitation id="1EBB4C68F0869B5EE38645CB396A63AD" collectingDate="1938-01-23" collectionCode="K" collectorName="Ves" country="Seychelles" location="Seche Island" specimenCount="1">
<emphasis id="287B51931F9F5BD1B0A041F1F99FE53C" bold="true">
<collectingCountry id="CE35A23419E111818F57F73DD7CDAA7B" name="Seychelles">Seychelles</collectingCountry>
</emphasis>
: 
<location id="12DF81216EBC2304EFCC9A2C7F1A9033" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:12DF81216EBC2304EFCC9A2C7F1A9033" country="Seychelles" name="Seche Island">Sèche Island</location>
, 
<date id="26FCF5F7774A73FF089657ACDE5E24C4" value="1938-01-23">
<collectingDate id="6CC8212B3CD9CEEBE194810829B0374B" value="1938-01-23">23 Jan 1938</collectingDate>
</date>
, 
<emphasis id="E3A8F8ABC3C64E8EE7E847E0DAC6D50A" italics="true">
<collectorName id="2378FD4A90E9AE1413DD07905922F168">Ves</collectorName>
6149
</emphasis>
(
<collectionCode id="79DE9FB3C0A6E4056ECF534D5F5CDDF5" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="7DFADCF24D870C774B7657EFAFEA56FE" collectingDate="1960-10" collectionCode="K" collectorName="C. J. Pigeott" country="Seychelles" location="St. Pierre Island" specimenCount="1">
<location id="476D0F0C235A3176F7855F6940A3D062" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:476D0F0C235A3176F7855F6940A3D062" name="St. Pierre Island">St. Pierre Island</location>
, 
<date id="D81F1C79AEE28543BF54EB69561161BC" value="1960-10">
<collectingDate id="A07F897FCE60144525C61D0E9AC0309B" value="1960-10">Oct 1960</collectingDate>
</date>
, 
<emphasis id="0FA6F75460C4C41AD91577B019C51F6B" italics="true">
<collectorName id="7FC1749EB2EF2BE3DA109EE481873F94">C. J. Pigeott</collectorName>
s. n.
</emphasis>
(
<collectionCode id="474DD91B0AD753479A9FA178C559FA3B" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="011B6562E126DA9857760A9AC52B3B78" collectingDate="1983-02" collectionCode="P" collectorName="F. Friedmann" country="Seychelles" location="Aride Island" specimenCode="P 00804851" specimenCount="1">
<location id="82B28B9D9A4694AE86FA912CD874C1CB" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:82B28B9D9A4694AE86FA912CD874C1CB" name="Aride Island">Aride Island</location>
, 
<date id="7D55139286B607A37AA0AF8E47E77C97" value="1983-02">
<collectingDate id="D72502EC290F2D51396DE1772EEB09DF" value="1983-02">Feb 1983</collectingDate>
</date>
, 
<emphasis id="421805658A37E7CFDB2B777FD5CC29F7" italics="true">
<collectorName id="C65F21D70E4A2408095BD9C8703BC9F5">F. Friedmann</collectorName>
4469
</emphasis>
(
<specimenCode id="25EE5EFEA199793F0D93F3AFD56DCA47">P 00804851</specimenCode>
)
</materialsCitation>
&amp; 
<materialsCitation id="8AD750A2AE32AC60D4FBA5752AC2480C" collectingDate="1987-04" collectionCode="P" collectorName="F. Friedmann" country="Seychelles" location="Aride Island" specimenCode="P 00804847" specimenCount="1">
<date id="290FF7CC5FCB45E498913D8524B56966" value="1987-04">
<collectingDate id="2D51BD469DB782899DF1A512599CDDE9" value="1987-04">Apr 1987</collectingDate>
</date>
, 
<emphasis id="4C8A71726A9699A8AE2246D2502FAB30" italics="true">
<collectorName id="A64731015CDF9A77697E1B4B49D08087">F. Friedmann</collectorName>
5455
</emphasis>
(
<specimenCode id="3A4ED66EBF3C90AE00F03D7DD7FB0669">P 00804847</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="7737EF11811A1B14FB362E1C8A70F27E" collectingDate="1952-09-28" collectionCode="BM" collectorName="E. S. Brown" country="Seychelles" location="Bird Island" specimenCount="1">
<location id="F4FBD189A61FBEFF3F2D4613FD120D66" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:F4FBD189A61FBEFF3F2D4613FD120D66" name="Bird Island">Bird Island</location>
, 
<date id="C3C8325810109959F5D47E8423137005" value="1952-09-28">
<collectingDate id="1C245B2E82248F72D75C490D94B8C187" value="1952-09-28">28 Sep 1952</collectingDate>
</date>
, 
<emphasis id="E516CD4633B0A7B16FCCAC8124FBECD9" italics="true">
<collectorName id="0D9EF0DCF6005DB69CF6D8CD576BDF79">E. S. Brown</collectorName>
s. n
</emphasis>
. (
<collectionCode id="3CF306DA2293451715BA214109A83174" country="United Kingdom" name="Bristol Museum">BM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="A205F22DCC7BFAC491ED03434EBD5E7C" collectingDate="1970-01-25" collectionCode="K" collectorName="F. R. Fosberg" country="Seychelles" location="Cousin Island" specimenCount="1">
<location id="482701E360D1561C2D159138351032E1" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:482701E360D1561C2D159138351032E1" name="Cousin Island">Cousin Island</location>
, 
<date id="A306E1BC97ED4A003DA1DEE04FFE03E6" value="1970-01-25">
<collectingDate id="46EBC80D36B31184F46FF5D501315A79" value="1970-01-25">25 Jan 1970</collectingDate>
</date>
, 
<emphasis id="6E5265681FF8061C22045C290338B00F" italics="true">
<collectorName id="E8544FAA8D0123AFB194C573873B1B19">F. R. Fosberg</collectorName>
52173
</emphasis>
(
<collectionCode id="0873A87AD4B5D0DE8A4D1D2BE4A65529" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="646A3F37AD4B018B74F1B2BD3F054AF1" collectingDate="1976-04-05" collectionCode="K" collectorName="D. R. Stoddart" country="Seychelles" location="Fouquet Island" specimenCount="1">
<location id="E8781FD6F62A52E007CE3BAED32765C8" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:E8781FD6F62A52E007CE3BAED32765C8" name="Fouquet Island">Fouquet Island</location>
, 
<date id="0AFA9C79B6202DCEF5393F99E9F1C526" value="1976-04-05">
<collectingDate id="35A8E3C14707DE2A799D65F8CF672AF3" value="1976-04-05">5 Apr 1976</collectingDate>
</date>
, 
<emphasis id="995ABBE1E93A04986F7765B954B41967" italics="true">
<collectorName id="863C6B241050545553ED55500685A127">D. R. Stoddart</collectorName>
7148
</emphasis>
(
<collectionCode id="2D01376CD597C3092DD9A61BA8D48ED0" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="9D947A6AC87619149A06680BFB63E8D9" collectingDate="1967-10-02" collectionCode="K" collectorName="Gwynne &amp; Wood" country="Seychelles" location="Farquhar Atoll" specimenCount="1">
<location id="E35C9ED9BEC3D6589E75F1F54735C8E3" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:E35C9ED9BEC3D6589E75F1F54735C8E3" name="Farquhar Atoll">Farquhar Atoll</location>
(South), 
<date id="2D96CA6BDFD388485E672F9326EFDE1E" value="1967-10-02">
<collectingDate id="B49888F256A066D83A09629F21A9DAF7" value="1967-10-02">2 Oct 1967</collectingDate>
</date>
, 
<emphasis id="598BFE9F0E0DFF06B8CF6219F2E92237" italics="true">
<collectorName id="7F82A6C561DF9C8126B6C0D4A02DFBA7">Gwynne</collectorName>
&amp; 
<collectorName id="2BDE2C64AF87A692682F85E2667CFF4D">Wood</collectorName>
1189
</emphasis>
(
<collectionCode id="CD07863A7B7E88497AF400153E1064E5" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="31C87A65613CBCB359F18B5C1E9D65B7" collectingDate="1968-09-19" collectionCode="K" collectorName="D. R. Stoddart" country="Seychelles" location="Farquhar Atoll" specimenCount="1">
<location id="0D7D43B671ED2D75B5FB81DB76618F56" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:0D7D43B671ED2D75B5FB81DB76618F56" name="Farquhar Atoll">Farquhar Atoll</location>
(North), 
<date id="DDCA129869A458439C4C8A6D385628E4" value="1968-09-19">
<collectingDate id="1A6411A55586C81B0C83C61D45916857" value="1968-09-19">19 Sep 1968</collectingDate>
</date>
, 
<emphasis id="50D61F8CE9AEF12AEFE4EBC7C6E70CE6" italics="true">
<collectorName id="9AFFCF06A43B9A996B1FAF3CACB02771">D. R. Stoddart</collectorName>
1376
</emphasis>
(
<collectionCode id="9A410399A6A9D69EA0B57B8532D0BF38" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3228CC3CF41420E0B8619D1F39AC8682" collectingDate="1975-07" collectionCode="K" collectorName="Warman" country="Seychelles" location="Aride Island" specimenCount="1">
<location id="B080206258888F7800950EEF7CB05686" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:B080206258888F7800950EEF7CB05686" name="Aride Island">Aride Island</location>
, 
<date id="E0CD5E9F70B3D01B1610D8B364A4E6F0" value="1975-07">
<collectingDate id="CF9E594FF25C043BB5F5FAF994D900E3" value="1975-07">Jul 1975</collectingDate>
</date>
, 
<emphasis id="69D40FF191B4CE217C75744178219E11" italics="true">
<collectorName id="C6504B83BA865929B88C2797E5F92974">Warman</collectorName>
23
</emphasis>
(
<collectionCode id="ECF486189FCC6E4F18E33D87C96D7650" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="F64ED9476650961B746582F10C77F4C0" collectingDate="1976-10-25" collectionCode="K" collectorName="S. A. Robertson" country="Seychelles" location="Remire [Eagle] Island" specimenCount="1">
<location id="7E8BB095B76056AA6E9E3421340626E1" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:7E8BB095B76056AA6E9E3421340626E1" name="Remire [Eagle] Island">Rémire [Eagle] Island</location>
, 
<date id="66C99F18A25F950D19E05675BBEF3240" value="1976-10-25">
<collectingDate id="C5ADB1A33686AC7D655A265E0B14A47B" value="1976-10-25">25 Oct 1976</collectingDate>
</date>
, 
<emphasis id="9007D1737BFD1B61922230C167E40CEB" italics="true">
<collectorName id="29D5ECAADA22A2507B4EEB68CD8E1882">S. A. Robertson</collectorName>
2319
</emphasis>
(
<collectionCode id="F069D5A2A52D8700D90694D119908B32" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="7EFB594D01A80D58534AA766044054F7" collectingDate="1976-03" collectionCode="K" collectorName="D. R. Stoddart" country="Seychelles" location="Bird Island" specimenCount="1">
<location id="788380DAF1BD94FCBEAE029AD37D23A7" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:788380DAF1BD94FCBEAE029AD37D23A7" name="Bird Island">Bird Island</location>
, 
<date id="3A1C635C7B19716093E9ACA69CE0D4D2" value="1976-03">
<collectingDate id="C739D2F1352E18D5B44F85CB44A7ED97" value="1976-03">Mar 1976</collectingDate>
</date>
, 
<emphasis id="61D787F10DB9C18E68D408301CA56F91" italics="true">
<collectorName id="6112F2E743E279E452F8490B2F0DF296">D. R. Stoddart</collectorName>
7079
</emphasis>
(
<collectionCode id="4CAB3FB4CEB4AA88D89EBC22F6C141BB" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="372B4E29C084AF184486019D56E8EBCC" collectingDate="1977-12-10" collectionCode="K" collectorName="D. R. Stoddart" country="Seychelles" location="Denis Island" specimenCount="1">
<location id="39ECC75BE614AFAD33F86F6544CA7510" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:39ECC75BE614AFAD33F86F6544CA7510" name="Denis Island">Denis Island</location>
, 
<date id="9CD87486904059DCBB12F5097739545A" value="1977-12-10">
<collectingDate id="1920EA8828BE7F78BB7AA3166F555B90" value="1977-12-10">10 Dec 1977</collectingDate>
</date>
, 
<emphasis id="961818278C9B36A0DFF52E83E8856595" italics="true">
<collectorName id="2DE81E6C5FF00C3B36124864CDF34CE5">D. R. Stoddart</collectorName>
8086
</emphasis>
(
<collectionCode id="6581D37D29D2C2B2C672945ED1B5EDDA" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="479724EAECBA1BEB936761858FAB18F6" collectingDate="1980-02-26" collectionCode="K" collectorName="S. A. Robertson" country="Seychelles" location="Platte Island" specimenCount="1">
<location id="294C497D52374277D402C6A710A73F05" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:294C497D52374277D402C6A710A73F05" name="Platte Island">Platte Island</location>
, 
<date id="ECE74F6A2588114A691F5F0F84CB111D" value="1980-02-26">
<collectingDate id="E343A4E4B32E8AD5CABEBC00C3E56EC5" value="1980-02-26">26 Feb 1980</collectingDate>
</date>
, 
<emphasis id="CEE76B43C8E35DC5D807999AA8E93137" italics="true">
<collectorName id="28F4F6F98F89C8F76FCF5505553CBBA7">S. A. Robertson</collectorName>
3021
</emphasis>
(
<collectionCode id="D18143D135AE62D2ABA18B90F140A169" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="D8C29888B205CDD593E3D1E7A9015A3D" collectingDate="1984-08-06" collectionCode="LE, MHA" collectorName="L. Averyanov" country="Seychelles" location="Poivre Island" specimenCount="1">
<location id="CC96DA6EA5EC9ED35418B7F52C5F5501" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:CC96DA6EA5EC9ED35418B7F52C5F5501" name="Poivre Island">Poivre Island</location>
, 
<date id="C16E7424CA1D9A3D0C2C4FF356448C08" value="1984-08-06">
<collectingDate id="B660D59E9FBC4BE62DB076967347B1D0" value="1984-08-06">6 Aug 1984</collectingDate>
</date>
, 
<emphasis id="E08CCA84490CF327CCEE84707720E5E9" italics="true">
<collectorName id="E8846826EC108CFF439D05370DA5375C">L. Averyanov</collectorName>
&amp; al. s. n
</emphasis>
. (
<collectionCode id="CC1A3BE298250AF456B04794D6236D4F" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
, 
<collectionCode id="7FB4490264006532635189F8E12DC509" country="Russia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15585" lsid="urn:lsid:biocol.org:col:15585" name="Main Botanical Garden of the Russian Academy of Sciences" type="Herbarium">MHA</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="7F9985CE8C9F565E803B236C4400B0A6" collectingDate="1984-08-16" collectionCode="LE" collectorName="L. Averyanov" country="Seychelles" location="Farquhar Atoll" specimenCount="1">
<location id="07C2284EF51C72C2CC69A1FC454C0444" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:07C2284EF51C72C2CC69A1FC454C0444" name="Farquhar Atoll">Farquhar Atoll</location>
, 
<date id="23F11D58C0CF1895A3226055D5804BBD" value="1984-08-16">
<collectingDate id="5AEF4FCC7662AAC2D9AE681F981BAAA0" value="1984-08-16">16 Aug 1984</collectingDate>
</date>
, 
<emphasis id="C0315158D1C0AF6C798B3DC23E33B968" italics="true">
<collectorName id="D3CA811F8B878DCC4E0452578EDB92D7">L. Averyanov</collectorName>
&amp; al. 580
</emphasis>
(
<collectionCode id="56F9F594D35C1D484D5B386C52450C03" country="Brazil" name="Servico de Microbiologia e Imunologia">LE</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A8F965BBF7D745185F835584BA5286B7">
<materialsCitation id="1AB8946F7D3495CC9165186B3F46BA81" collectingDate="1967-07-26" collectionCode="K" collectorName="D. R. Stoddart" country="Seychelles" county="Chagos Archipelago" location="East Island" municipality="Diego Garcia Atoll" specimenCount="1">
<emphasis id="9B48A90C34646A77335BFCFC6EBBA5F9" bold="true">
<collectingCountry id="7B02A71DBAB3304F2AD2115BBEEE08E0" name="Mauritius">Mauritius</collectingCountry>
</emphasis>
: 
<collectingCounty id="C90912FF6944660965A0C08B079B992B">Chagos Archipelago</collectingCounty>
, 
<collectingMunicipality id="1EF38DB87CE5CB00BB0762C68387D4C5">Diego Garcia Atoll</collectingMunicipality>
, 
<location id="C2327B93B2CEDD576EE6DA9C91899288" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:C2327B93B2CEDD576EE6DA9C91899288" country="Mauritius" county="Chagos Archipelago" municipality="Diego Garcia Atoll" name="East Island">East Island</location>
, 
<date id="669005FA70BFA44E55077CB155F160F1" value="1967-07-26">
<collectingDate id="AA303971204D5E134834DA5E353A8790" value="1967-07-26">26 Jul 1967</collectingDate>
</date>
, 
<emphasis id="4F91350D81343539B68F2E1CDB4FF794" italics="true">
<collectorName id="3B0CF20EFB2EA42E45E92EBFBFCD9E60">D. R. Stoddart</collectorName>
875
</emphasis>
(
<collectionCode id="E05C634DF2A69D0C2C14D2A63E8C507E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B5AA92E15B460503CBB0D6B1FF6E97F7" collectingDate="1971-03-30" collectionCode="BM" collectorName="A. M. Hutson" country="Seychelles" location="Diego Garcia" specimenCount="1">
<location id="5F514C06A9DE833993BC8B5AF26CBC9C" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:5F514C06A9DE833993BC8B5AF26CBC9C" name="Diego Garcia">Diego Garcia</location>
, near north coast, 
<date id="D16C2A0845A5B4C5D11B164C83F74541" value="1971-03-30">
<collectingDate id="9F9BB2EF31706ED0FEF696B3B7336C86" value="1971-03-30">30 Mar 1971</collectingDate>
</date>
, 
<emphasis id="665956768160769D06F9FEDFDA693100" italics="true">
<collectorName id="A81EF2087326952842D9E701BE7BE6F4">A. M. Hutson</collectorName>
15
</emphasis>
(
<collectionCode id="135F036EC1A4DBEFD7DBE5B094D9A214" country="United Kingdom" name="Bristol Museum">BM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="1285FBC3ED6C5790B5FC0AA838A48646" collectingDate="1975-02-21" collectionCode="K" collectorName="M. J. D. Hirons" country="Seychelles" location="Sea Cow Island" specimenCount="1">
<location id="9C88607EDFC8F7E6D6E77F924DD159EC" LSID="urn:lsid:plazi:treatment:8F9182709A1D596CA19DF5D6BC236333:9C88607EDFC8F7E6D6E77F924DD159EC" name="Sea Cow Island">Sea Cow Island</location>
, 
<date id="ECA317503F0B0D4B9D6860AA4138C187" value="1975-02-21">
<collectingDate id="31C19C9289474BDC48ECC178B714469B" value="1975-02-21">21 Feb 1975</collectingDate>
</date>
, 
<emphasis id="9DB7917D1718440FAA1EA6B27B235D44" italics="true">
<collectorName id="C126E2A3EECB0A51092EE295F1C59569">M. J. D. Hirons</collectorName>
50
</emphasis>
(
<collectionCode id="D710A91FECFE519190F12EE6FFA87817" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYTBK" type="Relationship">
<paragraph id="9B9D0FB3E350984C1756EE17F50D57B5">
<heading id="7AFE676D11EC7B622A5EA3F2422DA258" reason="title">Relationship.</heading>
</paragraph>
<paragraph id="A460EF5AFBDE6D2B018483384724920C">
<bibRefCitation id="D562B5A20832826B192CF4D6FC1EF058" author="Friedmann F" firstAuthor="Friedmann" journalOrPublisher="Orstom, Paris" refId="B39" refString="Friedmann F (1994) Flore de Seychelles. Dicotylédones. Orstom, Paris, 663 pp." title="Flore de Seychelles. Dicotylédones." year="1994">Friedmann (1994)</bibRefCitation>
reported three varieties of 
<taxonomicName id="ED30B0EE91823DD14498F380786703F3" baseAuthorityName="Ghazanfar" baseAuthorityYear="1992" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="aspera">
<emphasis id="0D8E41F7AF3053822CD6DD617E94F8E9" italics="true">A. aspera</emphasis>
</taxonomicName>
: 
<emphasis id="BDB8DCFC83F1B2A090E1E5F1B208D164" italics="true">
var. 
<taxonomicName id="DF1A97B4663075CFCD214F997EE9FEF8" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="velutina">
aspera, 
<emphasis id="686961EE422DE759583A0DF1EDDAA7FA" italics="true">var. velutina</emphasis>
</taxonomicName>
</emphasis>
, and 
<taxonomicName id="0F12BEAAF74D5E635141BBC25381AB2D" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="fruticosa">
<emphasis id="BD9774F45F8878FC018B7B1CF44F1E02" italics="true">var. fruticosa</emphasis>
</taxonomicName>
from the 
<collectingCountry id="3C5B136748A9C94DAD60DD97753DD8A2" name="Seychelles">Seychelles</collectingCountry>
. The specimens seen were identified as 
<taxonomicName id="97212000222A8EE6AFF5AD9C9F49BAD7" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="aspera">
<emphasis id="CABAF1A262E08263275B65BF6C76EC00" italics="true">var. aspera</emphasis>
</taxonomicName>
or 
<taxonomicName id="29D67E98B1DFA05354B1E88B9DC91843" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="velutina">
<emphasis id="492563F63563B503AD38EBAF204DD594" italics="true">var. velutina</emphasis>
</taxonomicName>
. From 
<taxonomicName id="E72BE7FAEFCF7C0DE326DC96F336B409" baseAuthorityName="Ghazanfar" baseAuthorityYear="1992" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="aspera">
<emphasis id="5551F6940ECA04A021C0457DCE3D56D1" italics="true">A. aspera</emphasis>
</taxonomicName>
, the new species differs by longer leaf blades (80–120 × 
<quantity id="3777862ABD6606978D1B29402A9F8FF9" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" unit="mm" value="45.0" valueMax="60.0" valueMin="30.0">30–60 mm</quantity>
vs. 15–110 × 
<quantity id="1661D946024A86DC3505FDA7763431A1" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" unit="mm" value="50.0" valueMax="80.0" valueMin="20.0">20–80 mm</quantity>
), and longer anthers (0.75–1.0 mm vs. 
<quantity id="734F7DA3514CDD7BB113F377658A8199" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.4–0.6 mm</quantity>
) and styles (
<quantity id="CC780B98ACF4D45D8EAB6C9E171EC546" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2–2.7 mm</quantity>
vs. 1.0– 
<quantity id="06550D7FEA3B6485F8DE22B4BB525269" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
). The true 
<taxonomicName id="E6B4C72EE0C61B10141127E960745181" authority="Hook. &amp; Arn." authorityName="Hook. &amp; Arn." class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="velutina">
<emphasis id="FD5B273737557B3C3310A2E267F55AE4" italics="true">A. velutina</emphasis>
Hook. &amp; Arn.
</taxonomicName>
from the Pacific (
<collectingCountry id="B18EEE9703029CE6B00934CE68E90AED" name="Samoa">Samoa</collectingCountry>
, Tahiti, Tuamotu etc.) is a hirsute shrub with longer (ca. 
<quantity id="25E72F9A051DD5639E81CDE726E3ABA0" metricMagnitude="-3" metricUnit="m" metricValue="7.0" unit="mm" value="7.0">7 mm</quantity>
long) perianths, but some other characters, i. e. length of anthers and styles are similar. Several specimens from the Aldabra Islands identified as 
<taxonomicName id="70D16B229BB2805705EADF2360FEE0D9" class="Magnoliopsida" family="Amaranthaceae" genus="Achyranthes" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="aspera" variety="fruticosa">
‘ 
<emphasis id="78C07917C575B8F496E37B6916CD62F9" italics="true">var. fruticosa</emphasis>
</taxonomicName>
’ represent plant fragments with a woody base and need further evalution.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4XBK" type="distribution">
<paragraph id="6FDCA95CD19B108D0AFFC19D8ED281AE">
<heading id="DDD55EE73ECE391FD48189DD0275708F" reason="title">General distribution.</heading>
</paragraph>
<paragraph id="8F0E06AC05864D250A501234B05A11D0">
Endemic to the 
<collectingCountry id="83B08BDFD4D5CE8DECC1448D01751D9A" name="Seychelles">Seychelles</collectingCountry>
and Chagos Archipelago.
</paragraph>
</subSubSection>
</treatment>
</document>