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<document id="98F10B522876144821B8900594CB6D98" ID-DOI="10.37520/fi.2023.004" ID-ISSN="2533-4069" ID-Zenodo-Dep="10481035" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704924026152" checkinUser="felipe" docAuthor="Smith, MacKenzie A., Greenwalt, Dale E. &amp; Manchester, Steven R." docDate="2023" docId="4D2487A3EF43827FFC17FD7B6A7BF7F0" docLanguage="en" docName="FossilImprint.79.1.37-88.pdf" docOrigin="Fossil Imprint 79 (1)" docSource="http://dx.doi.org/10.37520/fi.2023.004" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Quadrasubulaflora kishenehnensis Smith &amp; Greenwalt &amp; Manchester 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="65" masterDocId="B11DFFDBEF5F8263FF8CFFB16F30FFE9" masterDocTitle="Diverse Fruits And Seeds Of The Mid-Eocene Kishenehn Formation, Northwestern Montana, Usa, And Their Implications For Biogeography" masterLastPageNumber="88" masterPageNumber="37" pageNumber="65" updateTime="1704925034431" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="973904EF4D0F1249F1C4A113EC17D94B">Diverse Fruits And Seeds Of The Mid-Eocene Kishenehn Formation, Northwestern Montana, Usa, And Their Implications For Biogeography</mods:title>
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<mods:namePart id="CC3AF46B92885AC7071DD2ED6D3CA942">Smith, MacKenzie A.</mods:namePart>
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<mods:namePart id="FCCD5BDE2258E03D057EC346C85C9BAD">Greenwalt, Dale E.</mods:namePart>
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<mods:namePart id="AA0CFD70A1399CE3EADC1FF71A1C57D1">Manchester, Steven R.</mods:namePart>
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<mods:date id="6AA1B23EC575F88F265681063D3029D6">2023</mods:date>
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<mods:identifier id="8274BE8D0A7FB05582F994FD05A70F4D" type="ISSN">2533-4069</mods:identifier>
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<treatment id="4D2487A3EF43827FFC17FD7B6A7BF7F0" LSID="urn:lsid:plazi:treatment:4D2487A3EF43827FFC17FD7B6A7BF7F0" httpUri="http://treatment.plazi.org/id/4D2487A3EF43827FFC17FD7B6A7BF7F0" lastPageNumber="65" pageId="28" pageNumber="65">
<subSubSection id="8D97653EEF43827FFC17FD7B6BECFCEA" pageId="28" pageNumber="65" type="nomenclature">
<paragraph id="C53236B5EF43827FFC17FD7B6A4AFD0A" blockId="28.[923,1402,714,739]" box="[923,1402,714,739]" pageId="28" pageNumber="65">
<heading id="9E7A81D9EF43827FFC17FD7B6A4AFD0A" box="[923,1402,714,739]" centered="true" fontSize="8" level="2" pageId="28" pageNumber="65" reason="2">
<emphasis id="F7F9EAA7EF43827FFC17FD7B6A4AFD0A" bold="true" box="[923,1402,714,739]" pageId="28" pageNumber="65">
<taxonomicName id="028D4D36EF43827FFC17FD7B6A2DFD0B" authority="Smith &amp; Greenwalt &amp; Manchester, 2023" authorityName="Smith &amp; Greenwalt &amp; Manchester" authorityYear="2023" box="[923,1309,714,738]" class="Magnoliopsida" family="Compositae" genus="Quadrasubulaflora" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asterales" pageId="28" pageNumber="65" phylum="Tracheophyta" rank="species" species="kishenehnensis" status="sp. nov.">
<emphasis id="F7F9EAA7EF43827FFC17FD7B6A2DFD0B" bold="true" box="[923,1309,714,738]" italics="true" pageId="28" pageNumber="65">Quadrasubulaflora kishenehnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECCA57DCEF43827FFAA8FD7A6A4AFD0A" box="[1316,1402,715,739]" pageId="28" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C53236B5EF43827FFBB5FD5A6BECFCEA" blockId="28.[1081,1244,747,771]" box="[1081,1244,747,771]" pageId="28" pageNumber="65">
<figureCitation id="5DB62A30EF43827FFBB5FD5A6BECFCEA" box="[1081,1244,747,771]" captionStart="Text-fig" captionStartId="27.[125,206,1964,1985]" captionTargetBox="[125,1440,151,1949]" captionTargetId="figure-10@27.[501,830,610,1091]" captionTargetPageId="27" captionText="Text-fig. 12. a: USNM PAL 722530 Pteroheterochrosperma horseflyensis gen. et sp. nov. SMITH et MANCHESTER. Scale bar 2mm. b: USNM PAL 622695 Quadrasubulaflora kishenehnensis gen. et sp. nov. SMITH et MANCHESTER showing three outer parts and two stigma. Scale bar 1mm. c: USNM PAL 625447 Q. kishenehnensis showing three outer parts. Scale bar 1mm. d: USNM PAL 624649 Q. kishenehnensis showing three outer parts. Scale bar 1mm. e: USNM PAL 622402 Q. kishenehnensis. Scale bar 2mm. f: USNM" figureDoi="http://doi.org/10.5281/zenodo.10481067" httpUri="https://zenodo.org/record/10481067/files/figure.png" pageId="28" pageNumber="65">Text-fig. 12b–e</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8D97653EEF43827FFCFDFCAA6A7BF7F0" pageId="28" pageNumber="65" type="description">
<paragraph id="C53236B5EF43827FFCFDFCAA6AF5FCDA" blockId="28.[843,1477,795,851]" box="[881,1477,795,819]" pageId="28" pageNumber="65">P l a n t F o s s i l N a m e s R e g i s t r y. PFN003033</paragraph>
<paragraph id="C53236B5EF43827FFCC7FC8A6B3AFCBA" blockId="28.[843,1477,795,851]" box="[843,1034,827,851]" pageId="28" pageNumber="65">(for new species).</paragraph>
<paragraph id="C53236B5EF43827FFCFDFCDD6AFAFC6A" blockId="28.[843,1482,875,932]" box="[881,1482,875,900]" pageId="28" pageNumber="65">E t y m o l o g y. The specific epithet is for the for the</paragraph>
<paragraph id="C53236B5EF43827FFCC7FC3D6B0AFC4D" blockId="28.[843,1482,875,932]" box="[843,1082,908,932]" pageId="28" pageNumber="65">Kishenehn Formation.</paragraph>
<paragraph id="C53236B5EF43827FFCFDFC0D6AFAFC3A" blockId="28.[843,1482,955,1012]" box="[881,1482,955,979]" pageId="28" pageNumber="65">M a t e r i a l. USNM PAL 624649, 625447, 622695,</paragraph>
<paragraph id="C53236B5EF43827FFCC7FC6D6B6DFC1D" blockId="28.[843,1482,955,1012]" box="[843,1117,988,1012]" pageId="28" pageNumber="65">622402, 623987, 624154.</paragraph>
<paragraph id="C53236B5EF43827FFCFDFBBD6AF9FBCA" blockId="28.[843,1481,1035,1092]" box="[881,1481,1035,1060]" pageId="28" pageNumber="65">H o l o t y p e. USNM PAL 625447, National Museum of</paragraph>
<paragraph id="C53236B5EF43827FFCC7FB9D6A88FBAD" blockId="28.[843,1481,1035,1092]" box="[843,1464,1068,1092]" pageId="28" pageNumber="65">
Natural History,
<collectingRegion id="0749F857EF43827FFC71FB9D6BEDFBAD" box="[1021,1245,1068,1092]" country="United States of America" name="District of Columbia" pageId="28" pageNumber="65">District of Columbia</collectingRegion>
,
<collectingCountry id="BD9A7625EF43827FFB6BFB9D6A2FFBAD" box="[1255,1311,1068,1092]" name="United States of America" pageId="28" pageNumber="65">USA</collectingCountry>
,
<figureCitation id="5DB62A30EF43827FFAA5FB9D6A84FBAD" box="[1321,1460,1068,1092]" captionStart="Text-fig" captionStartId="27.[125,206,1964,1985]" captionTargetBox="[125,1440,151,1949]" captionTargetId="figure-10@27.[501,830,610,1091]" captionTargetPageId="27" captionText="Text-fig. 12. a: USNM PAL 722530 Pteroheterochrosperma horseflyensis gen. et sp. nov. SMITH et MANCHESTER. Scale bar 2mm. b: USNM PAL 622695 Quadrasubulaflora kishenehnensis gen. et sp. nov. SMITH et MANCHESTER showing three outer parts and two stigma. Scale bar 1mm. c: USNM PAL 625447 Q. kishenehnensis showing three outer parts. Scale bar 1mm. d: USNM PAL 624649 Q. kishenehnensis showing three outer parts. Scale bar 1mm. e: USNM PAL 622402 Q. kishenehnensis. Scale bar 2mm. f: USNM" figureDoi="http://doi.org/10.5281/zenodo.10481067" httpUri="https://zenodo.org/record/10481067/files/figure.png" pageId="28" pageNumber="65">Text-fig. 12c</figureCitation>
.
</paragraph>
<paragraph id="C53236B5EF43827FFCFDFBED6AF5FB9D" blockId="28.[843,1477,1116,1172]" box="[881,1477,1116,1140]" pageId="28" pageNumber="65">Ty p e h o r i z o n a n d t y p e l o c a l i t y. Spring</paragraph>
<paragraph id="C53236B5EF43827FFCC7FBCD6A14FB7D" blockId="28.[843,1477,1116,1172]" box="[843,1316,1148,1172]" pageId="28" pageNumber="65">Site in the Kishenehn Formation, ca. 44 Ma.</paragraph>
<paragraph id="C53236B5EF43827FFCFDFB1D6AFAFB2D" blockId="28.[843,1482,1196,1252]" box="[881,1482,1196,1220]" pageId="28" pageNumber="65">O t h e r l o c a l i t i e s. Dakin, Dishbow Creek and</paragraph>
<paragraph id="C53236B5EF43827FFCC7FB7D6CE4FB0D" blockId="28.[843,1482,1196,1252]" box="[843,980,1228,1252]" pageId="28" pageNumber="65">Spring Sites.</paragraph>
<paragraph id="C53236B5EF43827FFCFDFB4D6A38FA7F" blockId="28.[843,1482,1276,1430]" pageId="28" pageNumber="65">S p e c i f i c d i a g n o s i s. Flower square in cross section, elliptical in longitudinal section; pedicel slightly curved; awl-shaped and apices point outward with concave calyx lamina curving up to connect corners; hair-like projections protruding from near the axis.</paragraph>
<paragraph id="C53236B5EF43827FFCFDFA1F6BD4F9CE" blockId="28.[843,1482,1454,1575]" pageId="28" pageNumber="65">
S i z e. Flower
<quantity id="02759B50EF43827FFB99FA1F6BA6FA2C" box="[1045,1174,1454,1478]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="28" pageNumber="65" unit="mm" value="1.85" valueMax="2.2" valueMin="1.5">1.5–2.2 mm</quantity>
diagonal cross section, each side 1.0–
<quantity id="02759B50EF43827FFC20FA7F6CCCFA0F" box="[940,1020,1486,1510]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="65" unit="mm" value="1.5">1.5 mm</quantity>
long,
<quantity id="02759B50EF43827FFBCCFA7F6B8FFA0F" box="[1088,1215,1486,1510]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.1" metricValueMin="2.9" pageId="28" pageNumber="65" unit="mm" value="4.0" valueMax="5.1" valueMin="2.9">2.9–5.1 mm</quantity>
tall; pedicel 0.2–0.3 tall, basal corners of flower protrude outward
<quantity id="02759B50EF43827FFA8BFA5E6AB7F9EF" box="[1287,1415,1519,1543]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="28" pageNumber="65" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
wide; hair like projections
<quantity id="02759B50EF43827FFBABF9BE6B97F9CE" box="[1063,1191,1551,1575]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="28" pageNumber="65" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.5–0.8 mm</quantity>
long.
</paragraph>
<paragraph id="C53236B5EF43827FFCFDF9F16B83F991" blockId="28.[843,1482,1599,1656]" pageId="28" pageNumber="65">D e s c r i p t i o n. Hair-like projections are in the same plane as the corners of the flower.</paragraph>
<paragraph id="C53236B5EF43827FFCFDF9216B16F8E0" blockId="28.[843,1482,1679,1801]" pageId="28" pageNumber="65">
R e m a r k s. Possible affinities for this structure are flowers or fruits in tribe
<taxonomicName id="028D4D36EF43827FFBE9F9016BE9F921" box="[1125,1241,1712,1736]" class="Magnoliopsida" family="Apiaceae" kingdom="Plantae" order="Apiales" pageId="28" pageNumber="65" phylum="Tracheophyta" rank="tribe" tribe="Saniculeae">Saniculeae</taxonomicName>
and tribe
<taxonomicName id="028D4D36EF43827FFADEF9016AFAF921" authorityName="Sprengel" authorityYear="1820" box="[1362,1482,1712,1736]" class="Magnoliopsida" family="Apiaceae" kingdom="Plantae" order="Apiales" pageId="28" pageNumber="65" phylum="Tracheophyta" rank="tribe" tribe="Scandiceae">Scandiceae</taxonomicName>
subtribe
<taxonomicName id="028D4D36EF43827FFC21F9616B1EF901" box="[941,1070,1744,1768]" class="Magnoliopsida" family="Apiaceae" kingdom="Plantae" order="Apiales" pageId="28" pageNumber="65" phylum="Tracheophyta" rank="subTribe" subTribe="Scandicinae">Scandicinae</taxonomicName>
in
<taxonomicName id="028D4D36EF43827FFBD4F9616B8BF901" box="[1112,1211,1744,1768]" class="Magnoliopsida" family="Apiaceae" kingdom="Plantae" order="Apiales" pageId="28" pageNumber="65" phylum="Tracheophyta" rank="family">Apiaceae</taxonomicName>
because of their shape, size and projections.
</paragraph>
<paragraph id="C53236B5EF43827FFF0DF83E6A7BF7F0" blockId="28.[128,1482,1935,2073]" pageId="28" pageNumber="65">
<emphasis id="F7F9EAA7EF43827FFF0DF83E6A7BF7F0" bold="true" pageId="28" pageNumber="65">
PAL 722210 “KF1”. Scale bar
<quantity id="02759B50EF43827FFE3CF83E6ED8F84D" box="[432,488,1935,1956]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="65" unit="mm" value="2.0">2 mm</quantity>
. g: USNM PAL 624877 KF2. Scale bar
<quantity id="02759B50EF43827FFCE3F83E6C96F84D" box="[879,934,1935,1956]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="65" unit="mm" value="1.0">1mm</quantity>
. h: USNM PAL 624832 KF3. Scale bar
<quantity id="02759B50EF43827FFAA2F83E6A56F84D" box="[1326,1382,1935,1956]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="65" unit="mm" value="2.0">2 mm</quantity>
. i: USNM PAL 768303 KF4. Scale bar
<quantity id="02759B50EF43827FFE18F81D6EFCF828" box="[404,460,1964,1985]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="65" unit="mm" value="1.0">1 mm</quantity>
. j: USNM PAL 625629 KF4. Scale bar
<quantity id="02759B50EF43827FFCCBF81D6C4FF828" box="[839,895,1964,1985]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="65" unit="mm" value="3.0">3 mm</quantity>
. k: USNM PAL 624455 KF5. Scale bar
<quantity id="02759B50EF43827FFB73F81D6A07F828" box="[1279,1335,1964,1985]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="65" unit="mm" value="3.0">3 mm</quantity>
. l: USNM PAL 609598 KF6 seed. Scale bar
<quantity id="02759B50EF43827FFE16F8786EE2F837" box="[410,466,1993,2014]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="65" unit="mm" value="1.0">1 mm</quantity>
. m: USNM PAL 722298 KF7. Scale bar
<quantity id="02759B50EF43827FFCE8F8786CACF837" box="[868,924,1993,2014]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="65" unit="mm" value="1.0">1 mm</quantity>
. n: USNM PAL 625631 KF7. Scale bar
<quantity id="02759B50EF43827FFAA4F8786A50F837" box="[1320,1376,1993,2014]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="65" unit="mm" value="1.0">1 mm</quantity>
. o: USNM PAL 625736 KF8. Scale bar
<quantity id="02759B50EF43827FFE12F8566EE6F815" box="[414,470,2023,2044]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="65" unit="mm" value="2.0">2 mm</quantity>
. p: USNM PAL 625551 KF9 Scale bar
<quantity id="02759B50EF43827FFCECF8566CA8F815" box="[864,920,2023,2044]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="65" unit="mm" value="1.0">1 mm</quantity>
. q: USNM PAL 621685 KF9. Scale bar
<quantity id="02759B50EF43827FFAA4F8566A50F815" box="[1320,1376,2023,2044]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="65" unit="mm" value="1.0">1 mm</quantity>
. r: USNM PAL 620311 KF9. Scale bar
<quantity id="02759B50EF43827FFE14F7B56EE0F7F0" box="[408,464,2052,2073]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="65" unit="mm" value="1.0">1 mm</quantity>
. s: USNM PAL 626216 KF9. Scale bar
<quantity id="02759B50EF43827FFCD8F7B56CBCF7F0" box="[852,908,2052,2073]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="65" unit="mm" value="1.0">1 mm</quantity>
. t: USNM PAL 625608 KF9. Scale bar
<quantity id="02759B50EF43827FFA9CF7B56A77F7F0" box="[1296,1351,2052,2073]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="65" unit="mm" value="1.0">1mm</quantity>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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