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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="13132970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E7836FFF4FF013B29AB0DEBE6" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Riotintobolus Wesener &amp; Enghoff &amp; Sierwald 2009" docType="key" docVersion="9" lastPageNumber="51" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="50" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2009</mods:date>
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<mods:number>2009-09-04</mods:number>
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<mods:number>19</mods:number>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791421" ID-GBIF-Taxon="163747635" ID-Zenodo-Dep="3791421" LSID="urn:lsid:plazi:treatment:1772122E7836FFF4FF013B29AB0DEBE6" httpUri="http://treatment.plazi.org/id/1772122E7836FFF4FF013B29AB0DEBE6" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
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Key to the
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[270,431,1116,1143]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[270,431,1116,1143]" italics="true" pageId="49" pageNumber="50">Riotintobolus</emphasis>
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species:
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1 Tarsal pads absent (
<figureCitation box="[474,578,1187,1214]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="49" pageNumber="50">Fig. 27F</figureCitation>
). Eyes with 20 or less ocelli (
<figureCitation box="[922,1029,1187,1213]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="49" pageNumber="50">Figs 26A</figureCitation>
,
<figureCitation box="[1044,1095,1188,1214]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="49" pageNumber="50">27E</figureCitation>
). Anal valves with a deep groove between anterior half and sharp-edged lips (
<figureCitation box="[253,357,1258,1284]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="49" pageNumber="50">Figs 26B</figureCitation>
,
<figureCitation box="[369,421,1258,1284]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="49" pageNumber="50">27G</figureCitation>
) .........................................................................................
<emphasis bold="true" box="[1092,1107,1257,1283]" pageId="49" pageNumber="50">2</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="49.[140,1108,1187,1672]" pageId="49" pageNumber="50">
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At least male legs 37 with tarsal pads (
<figureCitation box="[703,809,1293,1319]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., AC, male (MNHN), DG male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="49" pageNumber="50">Fig. 29C</figureCitation>
). Eyes with more than 25 ocelli. Anal valves with neither a groove, nor sharp-edged lips (
<figureCitation box="[994,1103,1328,1354]" captionStart="Figure 28" captionStartId="53.[140,212,1558,1580]" captionTargetBox="[146,1101,1197,1532]" captionTargetId="figure@53.[140,1108,1193,1526]" captionTargetPageId="53" captionText="Figure 28. Riotintobolus aridus sp. n., male holotype, A telson B left antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view.av = anal valves; Cx = coxite; ho = horn; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767308" httpUri="https://zenodo.org/record/3767308/files/figure.png" pageId="49" pageNumber="50">Figs 28A</figureCitation>
,
<figureCitation box="[246,295,1364,1390]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., AC, male (MNHN), DG male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="49" pageNumber="50">29A</figureCitation>
) ..........................................................................................................
<emphasis bold="true" box="[1092,1107,1363,1389]" pageId="49" pageNumber="50">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="49" pageNumber="50">
<paragraph blockId="49.[140,1108,1187,1672]" pageId="49" pageNumber="50">
<keyLead pageId="49" pageNumber="50">
<specimenCount box="[140,361,1399,1425]" pageId="49" pageNumber="50" type="generic">2 Specimens</specimenCount>
<quantity box="[369,493,1398,1425]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="4.3" metricValueMin="3.5" pageId="49" pageNumber="50" unit="mm" value="39.0" valueMax="43.0" valueMin="35.0">3543 mm</quantity>
long. Colour laterally black, dorsally with a thick, light brown stripe (
<figureCitation box="[395,493,1434,1460]" captionStart="Figure 24" captionStartId="44.[140,212,1590,1612]" captionTargetBox="[140,1108,447,1570]" captionTargetId="figure@44.[140,1108,446,1570]" captionTargetPageId="44" captionText="Figure 24. A Riotintobolus mandenensis sp. n., living specimens B pseudosteppe and burned shrubs dominating the area between the littoral forests of Mandena and Sainte Luce, SE Madagascar C Sainte Luce littoral forest, manioc plantation and destroyed trees inside conservation zone S8." figureDoi="http://doi.org/10.5281/zenodo.3767302" httpUri="https://zenodo.org/record/3767302/files/figure.png" pageId="49" pageNumber="50">Fig. 24A</figureCitation>
). Telopodite of posterior gonopod laterally with one large and second small finger-shaped process (
<figureCitation box="[684,788,1469,1496]" captionStart="Figure 26" captionStartId="46.[140,212,1398,1420]" captionTargetBox="[184,1064,165,1374]" captionTargetId="figure@46.[183,1065,165,1100]" captionTargetPageId="46" captionText="Figure 26. Riotintobolus mandenensis sp. n., A, B, IM male holotype, CH female paratype. A head, lateral view B telson C head, detail of incisura lateralis D head, frontal view E second leg with vulva F gnathochilarium, overview G gnathochilarium, detail of mentum H left mandible, mesal view I anterior gonopod, anterior view Į left posterior gonopod, anterior view K anterior gonopod, posterior view L telopodite of left posterior gonopod, posterior view M left posterior gonopod, overview, posterior view. 3iT = 3-combed internal tooth; av = anal valves; Co = collum; Cx = coxite; eT = external tooth; Gn = gnathochilarium; Gr = groove; IL = incisura lateralis; L = ledge; LL = lamella linguales; LT = lateral tooth; Md = mandible; Me = mentum; mp = molar plate; pL = pectinate lamella; Pre = preanal ring; sf = setiferous foveolae; St = sternite; ST = stipites; sub = subanal scale; T = telopodite; Vu = vulva. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767304" httpUri="https://zenodo.org/record/3767304/files/figure.png" pageId="49" pageNumber="50">Fig. 26M</figureCitation>
)........
<emphasis bold="true" box="[859,1107,1468,1495]" pageId="49" pageNumber="50">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[859,1038,1468,1495]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="mandenensis" status="sp. n.">
<emphasis bold="true" box="[859,1038,1468,1495]" italics="true" pageId="49" pageNumber="50">R. mandenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1045,1107,1469,1495]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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Specimens shorter than
<quantity box="[516,603,1504,1531]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="49" pageNumber="50" unit="mm" value="25.0">25 mm</quantity>
. Colour laterally black, dorsally with a thick, red stripe (
<figureCitation box="[364,466,1539,1566]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="49" pageNumber="50">Fig. 27A</figureCitation>
). Telopodite of posterior gonopod laterally with two large finger-shaped processes of equal size (
<figureCitation box="[667,770,1575,1601]" captionStart="Figure 27" captionStartId="47.[140,212,1494,1516]" captionTargetBox="[181,1066,165,1466]" captionTargetId="figure@47.[181,1067,165,1211]" captionTargetPageId="47" captionText="Figure 27. Riotintobolus minutus sp. n., AE female paratype(s), FK male holotype. A two living specimens B habitus C gnathochilarium, overview; D midbody ring with leg pair E head, detail of incisura lateralis F legs 15, ventral view G telson H anterior gonopod, anterior view I left posterior gonopod, anterior view Į anterior gonopod, posterior view K left posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; d-g = ozopores; Gr = groove; IL = incisura lateralis; L = ledge; Md = mandible; Pe = “penes”; Pre = preanal ring; St = sternite; T = telopodite; Te = telson. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767306" httpUri="https://zenodo.org/record/3767306/files/figure.png" pageId="49" pageNumber="50">Fig. 27K</figureCitation>
).................
<emphasis bold="true" box="[904,1108,1575,1601]" pageId="49" pageNumber="50">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[904,1036,1575,1601]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="minutus" status="sp. n.">
<emphasis bold="true" box="[904,1036,1575,1601]" italics="true" pageId="49" pageNumber="50">R. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,1575,1601]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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3 Telson with an elongated, protruding preanal process, lips on anal valves well-developed (
<figureCitation box="[429,532,1645,1671]" captionStart="Figure 28" captionStartId="53.[140,212,1558,1580]" captionTargetBox="[146,1101,1197,1532]" captionTargetId="figure@53.[140,1108,1193,1526]" captionTargetPageId="53" captionText="Figure 28. Riotintobolus aridus sp. n., male holotype, A telson B left antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view.av = anal valves; Cx = coxite; ho = horn; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767308" httpUri="https://zenodo.org/record/3767308/files/figure.png" pageId="49" pageNumber="50">Fig. 28A</figureCitation>
). Only male legs 37 with tarsal pads. Apical mem- brane on posterior gonopod weakly developed (
<figureCitation box="[782,888,167,193]" captionStart="Figure 28" captionStartId="53.[140,212,1558,1580]" captionTargetBox="[146,1101,1197,1532]" captionTargetId="figure@53.[140,1108,1193,1526]" captionTargetPageId="53" captionText="Figure 28. Riotintobolus aridus sp. n., male holotype, A telson B left antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view.av = anal valves; Cx = coxite; ho = horn; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767308" httpUri="https://zenodo.org/record/3767308/files/figure.png" pageId="50" pageNumber="51">Fig. 28G</figureCitation>
). Telopodite at mesal margin with a suspicious horn (
<figureCitation box="[643,745,202,228]" captionStart="Figure 28" captionStartId="53.[140,212,1558,1580]" captionTargetBox="[146,1101,1197,1532]" captionTargetId="figure@53.[140,1108,1193,1526]" captionTargetPageId="53" captionText="Figure 28. Riotintobolus aridus sp. n., male holotype, A telson B left antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view.av = anal valves; Cx = coxite; ho = horn; Md = mandible; Pre = preanal ring; St = sternite; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767308" httpUri="https://zenodo.org/record/3767308/files/figure.png" pageId="50" pageNumber="51">Fig. 28F</figureCitation>
). Colour brown-blackish with a conspicuously wide light brown stripe..................................
<emphasis bold="true" box="[926,1108,236,263]" pageId="50" pageNumber="51">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[926,1035,236,263]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="aridus" status="sp. n.">
<emphasis bold="true" box="[926,1035,236,263]" italics="true" pageId="50" pageNumber="51">R. aridus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,237,263]" pageId="50" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="50.[140,1108,167,440]" pageId="50" pageNumber="51">
<keyLead pageId="50" pageNumber="51">
Telson preanal process not protruding, lips on anal valves weakly developed (
<figureCitation box="[254,359,307,334]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., AC, male (MNHN), DG male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="50" pageNumber="51">Fig. 29A</figureCitation>
). Male legs three and beyond with tarsal pads (
<figureCitation box="[907,1014,307,334]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., AC, male (MNHN), DG male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="50" pageNumber="51">Fig. 29C</figureCitation>
). Apical membrane on posterior gonopod large, widely protruding (
<figureCitation box="[908,1008,343,369]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., AC, male (MNHN), DG male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="50" pageNumber="51">Fig. 29F</figureCitation>
). Colour light grey, posterior margin of rings dark grey, dorsally with a conspicuous, wide blood-red stripe .....................................................
<emphasis bold="true" box="[887,1108,412,439]" pageId="50" pageNumber="51">
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[887,1036,412,439]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="anomalus" status="sp. n.">
<emphasis bold="true" box="[887,1036,412,439]" italics="true" pageId="50" pageNumber="51">R. anomalus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1108,413,439]" pageId="50" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>