291 lines
39 KiB
XML
291 lines
39 KiB
XML
<document id="0671FE922918B2A26A31FFBEC3DC4BEF" ID-CLB-Dataset="46747" ID-DOI="10.11646/zootaxa.3753.3.1" ID-GBIF-Dataset="220b1885-c8f8-46f6-9452-f1dd51baf651" ID-ISSN="1175-5326" ID-Zenodo-Dep="253310" ID-ZooBank="DE234954-1829-4277-9E17-78C4E5C18142" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460364433523" checkinUser="plazi" docAuthor="Maintenon, Marta J." docDate="2014" docId="03C0B65EFFD2FF83FF0F5724FAEDFC80" docLanguage="en" docName="zt03753p225.pdf" docOrigin="Zootaxa 3753 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parvanachis diminuta C.B Adams 1852" docType="treatment" docVersion="9" lastPageNumber="211" masterDocId="FFF9CE26FFDAFF89FF985116FFF0FF94" masterDocTitle="Taxonomic revision of the species of Parvanachis Radwin, 1968 (Gastropoda: Columbellidae) from the Gulf of Panama" masterLastPageNumber="225" masterPageNumber="201" pageNumber="209" updateTime="1698295016751" updateUser="ExternalLinkService">
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<mods:title id="433EED806699023072AE3E433F540063">Taxonomic revision of the species of Parvanachis Radwin, 1968 (Gastropoda: Columbellidae) from the Gulf of Panama</mods:title>
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<treatment id="03C0B65EFFD2FF83FF0F5724FAEDFC80" ID-DOI="http://doi.org/10.5281/zenodo.6141028" ID-GBIF-Taxon="119410431" ID-Zenodo-Dep="6141028" LSID="urn:lsid:plazi:treatment:03C0B65EFFD2FF83FF0F5724FAEDFC80" httpUri="http://treatment.plazi.org/id/03C0B65EFFD2FF83FF0F5724FAEDFC80" lastPageId="10" lastPageNumber="211" pageId="8" pageNumber="209">
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<subSubSection id="C37354C3FFD2FF81FF0F5724FE83F9FA" pageId="8" pageNumber="209" type="nomenclature">
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<paragraph id="8BD60748FFD2FF81FF0F5724FD65F9D9" blockId="8.[151,661,1586,1646]" box="[151,661,1586,1613]" pageId="8" pageNumber="209">
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<heading id="D09EB024FFD2FF81FF0F5724FD65F9D9" bold="true" box="[151,661,1586,1613]" fontSize="11" level="1" pageId="8" pageNumber="209" reason="1">
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<taxonomicName id="4C697CCBFFD2FF81FF0F5724FD65F9D9" ID-CoL="7656V" authority="C.B Adams, 1852" authorityName="C.B Adams" authorityYear="1852" box="[151,661,1586,1613]" class="Gastropoda" family="Columbellidae" genus="Parvanachis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="209" phylum="Mollusca" rank="species" species="diminuta">
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<emphasis id="B91DDB5AFFD2FF81FF0F5724FD65F9D9" bold="true" box="[151,661,1586,1613]" pageId="8" pageNumber="209">
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<emphasis id="B91DDB5AFFD2FF81FF0F5724FE52F9D8" bold="true" box="[151,418,1586,1612]" italics="true" pageId="8" pageNumber="209">Parvanachis diminuta</emphasis>
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(C.B
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<bibRefCitation id="EFF87AB9FFD2FF81FE735725FD7DF9D9" author="Adams" box="[491,653,1587,1613]" pageId="8" pageNumber="209" refString="Adams, C. B. (1852) Catalogue of shells collected at Panama, with notes on their synonymy, station, and geographical distribution. Annals of Lyceum of Natural History of New York, 5, 229 - 567." type="journal article" year="1852">Adams, 1852</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BD60748FFD2FF81FF0F5743FE83F9FA" blockId="8.[151,661,1586,1646]" box="[151,371,1621,1646]" pageId="8" pageNumber="209">
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<figureCitation id="13521BCDFFD2FF81FF0F5743FF0DF9FA" box="[151,253,1621,1646]" captionStart="FIGURE 1" captionStartId="4.[151,250,1675,1697]" captionTargetBox="[151,1436,301,1654]" captionTargetId="figure@4.[151,1436,301,1654]" captionTargetPageId="4" captionText="FIGURE 1. Parvanachis shells. B – D and F – G are preserved specimens, so dark areas on the body whorls of the shells are the coloring of the preserved animal visible through the shell. Scale bar = 1 mm. A. Parvanachis obesa, SBMNH 56943. B. Parvanachis diminuta, USNM 1231301, Isla Venado. C. Parvanachis dichroma, holotype, USNM 1231302, Chumical. D. Parvanachis dichroma, paratype, USNM 1231303, Chumical. E. Anachis albonodosa, paratype, USNM 716233, Mazatlan. F. Parvanachis minibrunnea, holotype, USNM 1231307, Isla Venado. G. Parvanachis minibrunnea, paratype, USNM 1231308, Isla Venado." httpUri="https://zenodo.org/record/253311/files/figure.png" pageId="8" pageNumber="209">Figures 1</figureCitation>
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B, 5A, 5D
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</paragraph>
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</subSubSection>
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<subSubSection id="C37354C3FFD2FF80FF0F578BFBEFFE1C" lastPageId="9" lastPageNumber="210" pageId="8" pageNumber="209" type="reference_group">
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<paragraph id="8BD60748FFD2FF81FF0F578BFF2AF946" blockId="8.[151,1436,1693,2022]" pageId="8" pageNumber="209">
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<taxonomicName id="4C697CCBFFD2FF81FF0F578BFDC3F927" ID-CoL="XBQT" authority="C.B. Adams, 1852" authorityName="C.B. Adams" authorityYear="1852" box="[151,563,1693,1715]" class="Gastropoda" family="Columbellidae" genus="Columbella" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="209" phylum="Mollusca" rank="species" species="diminuta">
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<emphasis id="B91DDB5AFFD2FF81FF0F578BFE9DF927" box="[151,365,1693,1715]" italics="true" pageId="8" pageNumber="209">Columbella diminuta</emphasis>
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<bibRefCitation id="EFF87AB9FFD2FF81FEEC578BFDC3F927" author="Adams" box="[372,563,1693,1715]" pageId="8" pageNumber="209" refString="Adams, C. B. (1852) Catalogue of shells collected at Panama, with notes on their synonymy, station, and geographical distribution. Annals of Lyceum of Natural History of New York, 5, 229 - 567." type="journal article" year="1852">C.B. Adams, 1852</bibRefCitation>
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</taxonomicName>
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: p. 85, not figured.
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<bibRefCitation id="EFF87AB9FFD2FF81FD625788FC8CF927" author="Reeve" box="[762,892,1693,1715]" pageId="8" pageNumber="209" refString="Reeve, L. A. (1858 - 1859) Monograph of the genus Columbella. Conchologica Iconica Vol. XI. London, 37 pls." type="journal volume" year="1858">Reeve, 1858</bibRefCitation>
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: sp. 115, pl. 20 fig. 115.
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<bibRefCitation id="EFF87AB9FFD2FF81FBEE5788FB0CF927" author="Turner" box="[1142,1276,1693,1715]" pageId="8" pageNumber="209" refString="Turner, R. D. (1956) The eastern Pacific marine mollusks described by C. B. Adams. Occasional Papers on Mollusks, Museum of Comparative Zoology, Harvard, 2 (20), 21 - 135, pls. 5 - 21." type="journal article" year="1956">Turner, 1956</bibRefCitation>
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||
: p. 45, pl. 7 fig. 2.
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</paragraph>
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<paragraph id="8BD60748FFD2FF81FF0F57CCFD3AF964" blockId="8.[151,1436,1693,2022]" box="[151,714,1754,1776]" pageId="8" pageNumber="209">
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<taxonomicName id="4C697CCBFFD2FF81FF0F57CCFD77F964" authority="Adams. Morch, 1860" authorityName="Adams. Morch" authorityYear="1860" box="[151,647,1754,1776]" class="Gastropoda" family="Columbellidae" genus="Mitrella" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="209" phylum="Mollusca" rank="species" species="diminuta" subGenus="Astyris">
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<emphasis id="B91DDB5AFFD2FF81FF0F57CCFE57F964" box="[151,423,1754,1776]" italics="true" pageId="8" pageNumber="209">Mitrella (Astyris) diminuta</emphasis>
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<bibRefCitation id="EFF87AB9FFD2FF81FE3557CCFD77F964" author="Adams" box="[429,647,1754,1776]" pageId="8" pageNumber="209" refString="Adams, A. (1860) On some new genera and species of Mollusca from Japan. Annals and Magazine of Natural History (3 rd. ser.), 35, 331 - 337." type="journal article" year="1860">Adams. Mörch, 1860</bibRefCitation>
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</taxonomicName>
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: p. 94.
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</paragraph>
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<paragraph id="8BD60748FFD2FF81FF0F57EFFCF9F89B" blockId="8.[151,1436,1693,2022]" box="[151,777,1785,1807]" pageId="8" pageNumber="209">
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<taxonomicName id="4C697CCBFFD2FF81FF0F57EFFE0EF89B" ID-CoL="66J4B" authority="Carpenter, 1857" authorityName="Carpenter" authorityYear="1857" box="[151,510,1785,1807]" class="Gastropoda" family="Columbellidae" genus="Anachis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="209" phylum="Mollusca" rank="species" species="rufotincta">
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<emphasis id="B91DDB5AFFD2FF81FF0F57EFFEA3F89B" box="[151,339,1785,1807]" italics="true" pageId="8" pageNumber="209">Anachis rufotincta</emphasis>
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<bibRefCitation id="EFF87AB9FFD2FF81FEC157EFFE0EF89B" author="Carpenter" box="[345,510,1785,1807]" pageId="8" pageNumber="209" refString="Carpenter, P. P. (1857) Catalogue of the collection of Mazatlan Shells in the British Museum: collected by Philip Reigen. Oberlin Press, London, 552 pp." type="book" year="1857">Carpenter, 1857</bibRefCitation>
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</taxonomicName>
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: pp. 511–512, not figured.
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||
</paragraph>
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<paragraph id="8BD60748FFD2FF81FF0F560EFDC9F8D8" blockId="8.[151,1436,1693,2022]" pageId="8" pageNumber="209">
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||
<emphasis id="B91DDB5AFFD2FF81FF0F560EFEA1F8BA" box="[151,337,1816,1838]" italics="true" pageId="8" pageNumber="209">
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||
<taxonomicName id="4C697CCBFFD2FF81FF0F560EFEBCF8BA" ID-CoL="66J2J" box="[151,332,1816,1838]" class="Gastropoda" family="Columbellidae" genus="Anachis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="209" phylum="Mollusca" rank="species" species="diminuta">Anachis diminuta</taxonomicName>
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.
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</emphasis>
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||
<bibRefCitation id="EFF87AB9FFD2FF81FECF560EFDF9F8BA" author="Carpenter" box="[343,521,1816,1838]" pageId="8" pageNumber="209" refString="Carpenter, P. P. (1863) Review of Prof. C. B. Adams's Catalogue of the Shells of Panama, from the type specimens. Proceedings of the Zoological Society of London for 1863, 339 - 369." type="proceedings paper" year="1863">Carpenter (1863)</bibRefCitation>
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||
: p. 344, not figured.
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<bibRefCitation id="EFF87AB9FFD2FF81FD46560EFC29F8BA" author="Strong" box="[734,985,1816,1838]" pageId="8" pageNumber="209" refString="Strong, A. M. & Hertlein, L. G. (1939) Marine mollusks from Panama collected by the Allan Hancock Expedition to the Galapagos Islands, 1931 - 1932. Allan Hancock Foundation Publications of the University of Southern California, Ser. 1. Allan Hancock Pacific Expeditions, 2 (12), 177 - 245." type="journal article" year="1939">Strong & Hertlein, 1939</bibRefCitation>
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: p. 184, not figured.
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<bibRefCitation id="EFF87AB9FFD2FF81FB36560FFAD7F8BA" author="Keen" box="[1198,1319,1816,1838]" pageId="8" pageNumber="209" refString="Keen, A. M. (1958) Sea Shells of Tropical West America. Stanford University Press, 624 pp." type="book" year="1958">Keen, 1958</bibRefCitation>
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: p. 381, sp. 429 (illustration from
|
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<bibRefCitation id="EFF87AB9FFD2FF81FE3F5621FDDDF8D8" author="Turner" box="[423,557,1846,1868]" pageId="8" pageNumber="209" refString="Turner, R. D. (1956) The eastern Pacific marine mollusks described by C. B. Adams. Occasional Papers on Mollusks, Museum of Comparative Zoology, Harvard, 2 (20), 21 - 135, pls. 5 - 21." type="journal article" year="1956">Turner, 1956</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BD60748FFD2FF81FF0F5643FC01F8FF" blockId="8.[151,1436,1693,2022]" box="[151,1009,1877,1899]" pageId="8" pageNumber="209">
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<taxonomicName id="4C697CCBFFD2FF81FF0F5643FD8FF8FF" ID-CoL="XBQT" authority="C.B. Adams" authorityName="C.B. Adams" box="[151,639,1877,1899]" class="Gastropoda" family="Columbellidae" genus="Columbella" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="209" phylum="Mollusca" rank="species" species="diminuta" subGenus="Seminella">
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<emphasis id="B91DDB5AFFD2FF81FF0F5643FE18F8FF" box="[151,488,1877,1899]" italics="true" pageId="8" pageNumber="209">Columbella (Seminella) diminuta</emphasis>
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(C.B. Adams)
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</taxonomicName>
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.
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<bibRefCitation id="EFF87AB9FFD2FF81FD115640FCE3F8FF" author="Tryon" box="[649,787,1877,1899]" pageId="8" pageNumber="209" refString="Tryon, G. W. Jr. (1883) Manual of Conchology; Structural and Systematic. Vol. 5. Marginellidae, Olividae, Columbellidae. Academy of Natural Sciences, Philadelphia, pp. 100 - 276." type="book" year="1883">Tryon (1883)</bibRefCitation>
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: p. 177, pl. 58 fig. 44.
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</paragraph>
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||
<paragraph id="8BD60748FFD2FF81FF0F5662FBF9F81E" blockId="8.[151,1436,1693,2022]" box="[151,1033,1908,1930]" pageId="8" pageNumber="209">
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||
<taxonomicName id="4C697CCBFFD2FF81FF0F5662FD1DF81E" ID-CoL="XBQT" authority="C.B. Adams. Kobelt, 1897" authorityName="C.B. Adams. Kobelt" authorityYear="1897" box="[151,749,1908,1930]" class="Gastropoda" family="Columbellidae" genus="Columbella" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="209" phylum="Mollusca" rank="species" species="diminuta" subGenus="Anachis">
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<emphasis id="B91DDB5AFFD2FF81FF0F5662FE26F81E" box="[151,470,1908,1930]" italics="true" pageId="8" pageNumber="209">Columbella (Anachis) diminuta</emphasis>
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||
C.B. Adams.
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||
<bibRefCitation id="EFF87AB9FFD2FF81FDFD5662FD1DF81E" author="Kobelt" box="[613,749,1908,1930]" pageId="8" pageNumber="209" refString="Kobelt, W. (1897) Die Familie der Columbelliden. In: Systematisches Conchylien-Cabinet von Martini & Chemnitz. Nurnberg, Verlag von Bauer & Raspe. Bd. 3, Abt. 1 d, 344 pp., 44 pls." type="book" year="1897">Kobelt, 1897</bibRefCitation>
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</taxonomicName>
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: p. 163, taf. 22, figs. 13, 14.
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||
</paragraph>
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||
<paragraph id="8BD60748FFD2FF81FF0F5684FDC6F853" blockId="8.[151,1436,1693,2022]" pageId="8" pageNumber="209">
|
||
<taxonomicName id="4C697CCBFFD2FF81FF0F5684FD4DF83C" ID-CoL="66J2J" authority="C.B. Adams, 1852" authorityName="C.B. Adams" authorityYear="1852" box="[151,701,1938,1960]" class="Gastropoda" family="Columbellidae" genus="Anachis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="209" phylum="Mollusca" rank="species" species="diminuta" subGenus="Parvanachis">
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||
<emphasis id="B91DDB5AFFD2FF81FF0F5684FE13F83C" box="[151,483,1938,1960]" italics="true" pageId="8" pageNumber="209">Anachis (Parvanachis) diminuta</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF87AB9FFD2FF81FE6B5684FD47F83C" author="Adams" box="[499,695,1938,1960]" pageId="8" pageNumber="209" refString="Adams, C. B. (1852) Catalogue of shells collected at Panama, with notes on their synonymy, station, and geographical distribution. Annals of Lyceum of Natural History of New York, 5, 229 - 567." type="journal article" year="1852">C.B. Adams, 1852</bibRefCitation>
|
||
)
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||
</taxonomicName>
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||
.
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||
<bibRefCitation id="EFF87AB9FFD2FF81FD515685FCB3F83C" author="Keen" box="[713,835,1938,1960]" pageId="8" pageNumber="209" refString="Keen, A. M. (1971) Sea Shells of Tropical West America, 2 nd ed. Stanford University Press, 1064 pp." type="book" year="1971">Keen, 1971</bibRefCitation>
|
||
: p. 584, sp. 1204 (illustration from
|
||
<bibRefCitation id="EFF87AB9FFD2FF81FB2C5685FACCF83C" author="Turner" box="[1204,1340,1938,1960]" pageId="8" pageNumber="209" refString="Turner, R. D. (1956) The eastern Pacific marine mollusks described by C. B. Adams. Occasional Papers on Mollusks, Museum of Comparative Zoology, Harvard, 2 (20), 21 - 135, pls. 5 - 21." type="journal article" year="1956">Turner, 1956</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFF87AB9FFD2FF81FAD75684FF0DF853" author="Abbott" pageId="8" pageNumber="209" refString="Abbott, R. T. (1974) American Seashells, 2 nd ed. Van Nostrand Reinhold Co., 663 pp." type="book" year="1974">Abbott, 1974</bibRefCitation>
|
||
: p. 197, sp. 2064 (not figured).
|
||
</paragraph>
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||
<paragraph id="8BD60748FFD2FF81FF0F56C6FCCEF872" blockId="8.[151,1436,1693,2022]" box="[151,830,2000,2022]" pageId="8" pageNumber="209">
|
||
<taxonomicName id="4C697CCBFFD2FF81FF0F56C6FDBDF872" ID-CoL="7656V" authority="C.B. Adams, 1852" authorityName="C.B. Adams" authorityYear="1852" box="[151,589,2000,2022]" class="Gastropoda" family="Columbellidae" genus="Parvanachis" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="209" phylum="Mollusca" rank="species" species="diminuta">
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||
<emphasis id="B91DDB5AFFD2FF81FF0F56C6FE87F872" box="[151,375,2000,2022]" italics="true" pageId="8" pageNumber="209">Parvanachis diminuta</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF87AB9FFD2FF81FE1E56C6FDB6F872" author="Adams" box="[390,582,2000,2022]" pageId="8" pageNumber="209" refString="Adams, C. B. (1852) Catalogue of shells collected at Panama, with notes on their synonymy, station, and geographical distribution. Annals of Lyceum of Natural History of New York, 5, 229 - 567." type="journal article" year="1852">C.B. Adams, 1852</bibRefCitation>
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||
)
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFF87AB9FFD2FF81FDCF56C6FD0AF872" author="Skoglund" box="[599,762,2000,2022]" pageId="8" pageNumber="209" refString="Skoglund, C. (1992) Additions to the Panamic Province gastropod (Mollusca) literature 1971 to 1992. The Festivus, supplement 24, 1 - 169." type="journal article" year="1992">Skoglund, 1992</bibRefCitation>
|
||
: p. 89.
|
||
</paragraph>
|
||
<paragraph id="8BD60748FFD3FF80FF0F5181FA6CFF3B" blockId="9.[151,1436,151,933]" box="[151,1436,151,176]" pageId="9" pageNumber="210">
|
||
<emphasis id="B91DDB5AFFD3FF80FF0F5181FF13FF24" bold="true" box="[151,227,151,176]" pageId="9" pageNumber="210">
|
||
<typeStatus id="54D2B9EAFFD3FF80FF0F5181FF2FFF24" box="[151,223,151,176]" pageId="9" pageNumber="210">Types</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<typeStatus id="54D2B9EAFFD3FF80FF72518EFEADFF24" box="[234,349,152,176]" pageId="9" pageNumber="210" type="lectotype">Lectotype</typeStatus>
|
||
, MCZ 186401, from
|
||
<collectingCountry id="F37E47D8FFD3FF80FDD1518EFD55FF3B" box="[585,677,152,175]" name="Panama" pageId="9" pageNumber="210">Panama</collectingCountry>
|
||
. A photo of the
|
||
<typeStatus id="54D2B9EAFFD3FF80FCCE518EFC76FF24" box="[854,902,152,176]" pageId="9" pageNumber="210">type</typeStatus>
|
||
was available from W. Thorsson to confirm ID.
|
||
</paragraph>
|
||
<paragraph id="8BD60748FFD3FF80FF5F51AAFBEFFE1C" blockId="9.[151,1436,151,933]" pageId="9" pageNumber="210">
|
||
<emphasis id="B91DDB5AFFD3FF80FF5F51AAFE42FF41" bold="true" box="[199,434,188,213]" pageId="9" pageNumber="210">Taxonomic history.</emphasis>
|
||
Charles
|
||
<collectingCountry id="F37E47D8FFD3FF80FDBD51AAFD98FF41" box="[549,616,188,213]" name="United States Pacific Island Wildlife Refuges" pageId="9" pageNumber="210">Baker</collectingCountry>
|
||
Adams did not illustrate this species when he described it; Reeve illustrated it in 1858. In both cases its shell was described as pale, with a dark base, which is characteristic of this species, and its identity has not been in question.
|
||
<bibRefCitation id="EFF87AB9FFD3FF80FD4A5012FC64FE89" author="Carpenter" box="[722,916,260,285]" pageId="9" pageNumber="210" refString="Carpenter, P. P. (1863) Review of Prof. C. B. Adams's Catalogue of the Shells of Panama, from the type specimens. Proceedings of the Zoological Society of London for 1863, 339 - 369." type="proceedings paper" year="1863">Carpenter (1863)</bibRefCitation>
|
||
considered it a species of
|
||
<taxonomicName id="4C697CCBFFD3FF80FB5F5012FAD2FE88" ID-CoL="7NNYL" box="[1223,1314,260,284]" class="Gastropoda" family="Columbellidae" genus="Anachis" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="210" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91DDB5AFFD3FF80FB5F5012FAD2FE88" box="[1223,1314,260,284]" italics="true" pageId="9" pageNumber="210">Anachis</emphasis>
|
||
</taxonomicName>
|
||
(which he allied with
|
||
<taxonomicName id="4C697CCBFFD3FF80FE8D503EFE9AFEAB" box="[277,362,296,319]" class="Gastropoda" family="Buccinidae" genus="Pisania" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="210" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91DDB5AFFD3FF80FE8D503EFE9AFEAB" box="[277,362,296,319]" italics="true" pageId="9" pageNumber="210">Pisania</emphasis>
|
||
</taxonomicName>
|
||
based on the operculum) while
|
||
<bibRefCitation id="EFF87AB9FFD3FF80FD4B503EFC97FED4" author="Tryon" box="[723,871,296,320]" pageId="9" pageNumber="210" refString="Tryon, G. W. Jr. (1883) Manual of Conchology; Structural and Systematic. Vol. 5. Marginellidae, Olividae, Columbellidae. Academy of Natural Sciences, Philadelphia, pp. 100 - 276." type="book" year="1883">Tryon (1883)</bibRefCitation>
|
||
placed it in
|
||
<taxonomicName id="4C697CCBFFD3FF80FC6B503EFB0DFED4" ID-CoL="7NTCV" authority="(Seminella)" baseAuthorityName="Seminella" box="[1011,1277,296,320]" class="Gastropoda" family="Columbellidae" genus="Columbella" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="210" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91DDB5AFFD3FF80FC6B503EFB0DFED4" box="[1011,1277,296,320]" italics="true" pageId="9" pageNumber="210">Columbella (Seminella)</emphasis>
|
||
</taxonomicName>
|
||
and based his illustration on that of Reeve.
|
||
<taxonomicName id="4C697CCBFFD3FF80FE44505AFD5BFEF0" ID-CoL="66J4B" box="[476,683,332,356]" class="Gastropoda" family="Columbellidae" genus="Anachis" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="210" phylum="Mollusca" rank="species" species="rufotincta">
|
||
<emphasis id="B91DDB5AFFD3FF80FE44505AFD5BFEF0" box="[476,683,332,356]" italics="true" pageId="9" pageNumber="210">Anachis rufotincta</emphasis>
|
||
</taxonomicName>
|
||
(with a more reddish anterior end) was named by
|
||
<bibRefCitation id="EFF87AB9FFD3FF80FB44505AFA6CFEF1" author="Carpenter" box="[1244,1436,332,357]" pageId="9" pageNumber="210" refString="Carpenter, P. P. (1857) Catalogue of the collection of Mazatlan Shells in the British Museum: collected by Philip Reigen. Oberlin Press, London, 552 pp." type="book" year="1857">Carpenter (1857)</bibRefCitation>
|
||
from Mazatlan, and as reported by
|
||
<bibRefCitation id="EFF87AB9FFD3FF80FD875066FD44FE1C" author="Tryon" box="[543,692,368,392]" pageId="9" pageNumber="210" refString="Tryon, G. W. Jr. (1883) Manual of Conchology; Structural and Systematic. Vol. 5. Marginellidae, Olividae, Columbellidae. Academy of Natural Sciences, Philadelphia, pp. 100 - 276." type="book" year="1883">Tryon (1883)</bibRefCitation>
|
||
and later authors, is a synonym.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37354C3FFD3FF80FF5F5082FCD1FE44" pageId="9" pageNumber="210" type="diagnosis">
|
||
<paragraph id="8BD60748FFD3FF80FF5F5082FCD1FE44" blockId="9.[151,1436,151,933]" pageId="9" pageNumber="210">
|
||
<emphasis id="B91DDB5AFFD3FF80FF5F5082FEB1FE39" bold="true" box="[199,321,404,429]" pageId="9" pageNumber="210">Diagnosis.</emphasis>
|
||
Small species with a biconic axially ribbed shell, mottled pale tan with a narrow white spiral below the suture and a dark reddish or purple-brown anterior tip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37354C3FFD3FF83FF5F50CAFAEDFC80" lastPageId="10" lastPageNumber="211" pageId="9" pageNumber="210" type="materials_examined">
|
||
<paragraph id="8BD60748FFD3FF80FF5F50CAFB49FD83" blockId="9.[151,1436,151,933]" pageId="9" pageNumber="210">
|
||
<emphasis id="B91DDB5AFFD3FF80FF5F50CAFEC6FE61" bold="true" box="[199,310,476,501]" pageId="9" pageNumber="210">Material.</emphasis>
|
||
Over
|
||
<specimenCount id="9D6FCCC1FFD3FF80FE1950CAFDDAFE61" box="[385,554,476,501]" pageId="9" pageNumber="210" type="generic">100 specimens</specimenCount>
|
||
of this species were collected on the bottoms of rocks in muddy sand. Two were sectioned, two dissected.
|
||
<collectionCode id="ED789F8DFFD3FF80FE6A5316FDB3FD8C" LSID="urn:lsid:biocol.org:col:34871" box="[498,579,512,536]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="210">USNM</collectionCode>
|
||
has dry specimens from Baja California Sur to
|
||
<collectingCountry id="F37E47D8FFD3FF80FBC15316FB45FD83" box="[1113,1205,512,535]" name="Panama" pageId="9" pageNumber="210">Panama</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD60748FFD3FF80FF5F5332FC4DFCAC" blockId="9.[151,1436,151,933]" pageId="9" pageNumber="210">
|
||
<emphasis id="B91DDB5AFFD3FF80FF5F5332FF0FFDA8" box="[199,255,548,572]" italics="true" pageId="9" pageNumber="210">Shell</emphasis>
|
||
(
|
||
<figureCitation id="13521BCDFFD3FF80FE965332FEBFFDA8" box="[270,335,548,573]" captionStart="FIGURE 1" captionStartId="4.[151,250,1675,1697]" captionTargetBox="[151,1436,301,1654]" captionTargetId="figure@4.[151,1436,301,1654]" captionTargetPageId="4" captionText="FIGURE 1. Parvanachis shells. B – D and F – G are preserved specimens, so dark areas on the body whorls of the shells are the coloring of the preserved animal visible through the shell. Scale bar = 1 mm. A. Parvanachis obesa, SBMNH 56943. B. Parvanachis diminuta, USNM 1231301, Isla Venado. C. Parvanachis dichroma, holotype, USNM 1231302, Chumical. D. Parvanachis dichroma, paratype, USNM 1231303, Chumical. E. Anachis albonodosa, paratype, USNM 716233, Mazatlan. F. Parvanachis minibrunnea, holotype, USNM 1231307, Isla Venado. G. Parvanachis minibrunnea, paratype, USNM 1231308, Isla Venado." httpUri="https://zenodo.org/record/253311/files/figure.png" pageId="9" pageNumber="210">Fig. 1</figureCitation>
|
||
B): Shell small, biconic,
|
||
<quantity id="4C91AAADFFD3FF80FDF15332FCF7FDA8" box="[617,775,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.3" metricValueMin="3.5" pageId="9" pageNumber="210" unit="mm" value="3.9" valueMax="4.3" valueMin="3.5">3.5 to 4.3 mm</quantity>
|
||
long (avg.
|
||
<quantity id="4C91AAADFFD3FF80FC1F5332FC1DFDA8" box="[903,1005,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="3.84" pageId="9" pageNumber="210" unit="mm" value="3.84">3.84 mm</quantity>
|
||
) and
|
||
<quantity id="4C91AAADFFD3FF80FBB45332FB16FDA8" box="[1068,1254,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.95" metricValueMin="1.55" pageId="9" pageNumber="210" unit="mm" value="1.75" valueMax="1.95" valueMin="1.55">1.55 to 1.95 mm</quantity>
|
||
wide (avg.
|
||
<quantity id="4C91AAADFFD3FF80FAF35332FF34FDF4" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="210" unit="mm" value="1.7">1.70 mm</quantity>
|
||
) in
|
||
<specimenCount id="9D6FCCC1FFD3FF80FF68535EFE77FDF4" box="[240,391,583,608]" pageId="9" pageNumber="210" type="generic">12 specimens</specimenCount>
|
||
measured. Adult shells with 4 to 4.25 teleoconch whorls (avg. 4.00). Protoconch smooth, off white, 3.25 to 3.5 whorls (avg. 3.14). Dominant sculpture of axial ridges, except on first two teleoconch whorls where strong spiral ridges make the sculpture cancellate. One weak subsutural groove cuts across tops of axial ridges. Shell off-white with vague chestnut blotches that give an overall mottled tan affect, a dark purplish brown base and usually a narrow white band slightly below the suture. Aperture edge thickened, with a few denticles internally. Shallow posterior sinus present in aperture edge. Parietal wall denticulate, anterior edge of callus detached. Aperture white, with purple flush on anterior columellar wall.
|
||
</paragraph>
|
||
<paragraph id="8BD60748FFD3FF80FF5F5252FC52FCC9" blockId="9.[151,1436,151,933]" box="[199,930,836,861]" pageId="9" pageNumber="210">
|
||
<emphasis id="B91DDB5AFFD3FF80FF5F5252FE7DFCC9" bold="true" box="[199,397,836,861]" pageId="9" pageNumber="210">Body coloration.</emphasis>
|
||
Body cream colored with white specks overall.
|
||
</paragraph>
|
||
<paragraph id="8BD60748FFD3FF80FF5F527EFEA2FC31" blockId="9.[151,1436,151,933]" pageId="9" pageNumber="210">
|
||
<emphasis id="B91DDB5AFFD3FF80FF5F527EFEB4FC14" box="[199,324,872,896]" italics="true" pageId="9" pageNumber="210">Operculum</emphasis>
|
||
(
|
||
<figureCitation id="13521BCDFFD3FF80FECF5271FE6EFC14" box="[343,414,871,896]" captionStart="FIGURE 5" captionStartId="9.[151,250,1920,1942]" captionTargetBox="[151,1436,985,1898]" captionTargetId="figure@9.[151,1436,985,1898]" captionTargetPageId="9" captionText="FIGURE 5. Parvanachis operculae and radulae. Scale bars: A – C, 200 µm; D – F, 10 µm. A – C. Operculae. A. P. diminuta, 60 X. B. P. dichroma, 60 X. C. P. minibrunnea, 60 X. D – F. radulae. D. P. diminuta, 400 X. E. P. dichroma, 400 X. F. P. minibrunnea." httpUri="https://zenodo.org/record/253315/files/figure.png" pageId="9" pageNumber="210">Fig. 5</figureCitation>
|
||
A, tip broken): Operculum ovoid with a terminal nucleus, darker centrally, muscle scar bilobed, no keel.
|
||
</paragraph>
|
||
<caption id="DF1657C0FFD3FF80FF0F5696FA72F820" httpUri="https://zenodo.org/record/253315/files/figure.png" pageId="9" pageNumber="210" targetBox="[151,1436,985,1898]" targetPageId="9">
|
||
<paragraph id="8BD60748FFD3FF80FF0F5696FA72F820" blockId="9.[151,1435,1920,1973]" pageId="9" pageNumber="210">
|
||
<emphasis id="B91DDB5AFFD3FF80FF0F5696FEE2F802" bold="true" box="[151,274,1920,1942]" pageId="9" pageNumber="210">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C697CCBFFD3FF80FE815696FE69F802" box="[281,409,1920,1942]" class="Gastropoda" family="Columbellidae" genus="Parvanachis" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="210" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91DDB5AFFD3FF80FE815696FE69F802" box="[281,409,1920,1942]" italics="true" pageId="9" pageNumber="210">Parvanachis</emphasis>
|
||
</taxonomicName>
|
||
operculae and radulae. Scale bars: A–C, 200 µm; D–F, 10 µm. A–C. Operculae. A.
|
||
<taxonomicName id="4C697CCBFFD3FF80FB735697FAADF802" box="[1259,1373,1920,1942]" class="Gastropoda" family="Columbellidae" genus="Parvanachis" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="210" phylum="Mollusca" rank="species" species="diminuta">
|
||
<emphasis id="B91DDB5AFFD3FF80FB735697FAADF802" box="[1259,1373,1920,1942]" italics="true" pageId="9" pageNumber="210">P. diminuta</emphasis>
|
||
</taxonomicName>
|
||
, 60X. B.
|
||
<taxonomicName id="4C697CCBFFD3FF80FF2C56B6FEDDF820" box="[180,301,1950,1973]" class="Gastropoda" family="Columbellidae" genus="Parvanachis" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="210" phylum="Mollusca" rank="species" species="dichroma">
|
||
<emphasis id="B91DDB5AFFD3FF80FF2C56B6FEDDF820" box="[180,301,1950,1973]" italics="true" pageId="9" pageNumber="210">P. dichroma</emphasis>
|
||
</taxonomicName>
|
||
, 60X. C.
|
||
<taxonomicName id="4C697CCBFFD3FF80FE0856B6FDDAF820" box="[400,554,1950,1973]" class="Gastropoda" family="Columbellidae" genus="Parvanachis" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="210" phylum="Mollusca" rank="species" species="minibrunnea">
|
||
<emphasis id="B91DDB5AFFD3FF80FE0856B6FDDAF820" box="[400,554,1950,1973]" italics="true" pageId="9" pageNumber="210">P. minibrunnea</emphasis>
|
||
</taxonomicName>
|
||
, 60X. D–F. radulae. D.
|
||
<emphasis id="B91DDB5AFFD3FF80FC8456B6FC66F820" box="[796,918,1950,1973]" italics="true" pageId="9" pageNumber="210">
|
||
<taxonomicName id="4C697CCBFFD3FF80FC8456B6FC61F820" box="[796,913,1950,1973]" class="Gastropoda" family="Columbellidae" genus="Parvanachis" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="210" phylum="Mollusca" rank="species" species="diminuta">P. diminuta</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
400X. E.
|
||
<emphasis id="B91DDB5AFFD3FF80FC6556B6FB8CF820" box="[1021,1148,1950,1973]" italics="true" pageId="9" pageNumber="210">
|
||
<taxonomicName id="4C697CCBFFD3FF80FC6556B6FB89F820" box="[1021,1145,1950,1973]" class="Gastropoda" family="Columbellidae" genus="Parvanachis" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="210" phylum="Mollusca" rank="species" species="dichroma">P. dichroma</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
400X. F.
|
||
<taxonomicName id="4C697CCBFFD3FF80FB7856B6FA8EF820" box="[1248,1406,1950,1973]" class="Gastropoda" family="Columbellidae" genus="Parvanachis" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="210" phylum="Mollusca" rank="species" species="minibrunnea">P. minibrunnea</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD60748FFD0FF83FF5F518EFC48FF6C" blockId="10.[151,1437,151,789]" pageId="10" pageNumber="211">
|
||
<taxonomicName id="4C697CCBFFD0FF83FF5F518EFEE8FF24" box="[199,280,152,176]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="10" pageNumber="211" phylum="Bryophyta" rank="genus">
|
||
<emphasis id="B91DDB5AFFD0FF83FF5F518EFEE8FF24" box="[199,280,152,176]" italics="true" pageId="10" pageNumber="211">Radula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13521BCDFFD0FF83FEBE5181FE98FF24" box="[294,360,151,176]" captionStart="FIGURE 5" captionStartId="9.[151,250,1920,1942]" captionTargetBox="[151,1436,985,1898]" captionTargetId="figure@9.[151,1436,985,1898]" captionTargetPageId="9" captionText="FIGURE 5. Parvanachis operculae and radulae. Scale bars: A – C, 200 µm; D – F, 10 µm. A – C. Operculae. A. P. diminuta, 60 X. B. P. dichroma, 60 X. C. P. minibrunnea, 60 X. D – F. radulae. D. P. diminuta, 400 X. E. P. dichroma, 400 X. F. P. minibrunnea." httpUri="https://zenodo.org/record/253315/files/figure.png" pageId="10" pageNumber="211">Fig. 5</figureCitation>
|
||
D): One radula obtained whole had 123 tooth rows. Lateral teeth in adult specimens dissected 18 to 20 µm long, 7 µm wide. Lateral teeth with three pointed secondary cusps, the basal cusp pointed and curved toward the membrane, and separated from the distal two by a wide gap.
|
||
</paragraph>
|
||
<paragraph id="8BD60748FFD0FF83FF5F5012FB9BFE60" blockId="10.[151,1437,151,789]" pageId="10" pageNumber="211">
|
||
<emphasis id="B91DDB5AFFD0FF83FF5F5012FE32FE88" box="[199,450,260,284]" italics="true" pageId="10" pageNumber="211">Reproductive anatomy</emphasis>
|
||
: Male system typical for
|
||
<taxonomicName id="4C697CCBFFD0FF83FD7A5012FC9FFE88" box="[738,879,260,284]" class="Gastropoda" family="Columbellidae" genus="Parvanachis" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="211" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91DDB5AFFD0FF83FD7A5012FC9FFE88" box="[738,879,260,284]" italics="true" pageId="10" pageNumber="211">Parvanachis</emphasis>
|
||
</taxonomicName>
|
||
. Ciliated duct to mantle cavity present anterior to coiled seminal vesicle. Anterior spermiduct ciliated, moderately muscular, passes forward embedded in body wall from mantle cavity duct to penis base. Long spermiduct loop in hemocoel leaves body wall near penis base and runs back alongside proboscis, then doubles back and enters penis. Spermiduct of internal loop wide, with clustery mucoidal epithelium that continues along most of the length of the penis except for a non-secretory region at the base. Penis tip with subepithelial mucus glands in addition to secretory epithelium. Penis long, relatively thick, simple with a short filament tip. Penis curled back in the posterior mantle cavity at rest.
|
||
</paragraph>
|
||
<paragraph id="8BD60748FFD0FF83FF5F50E9FAEDFC80" blockId="10.[151,1437,151,789]" pageId="10" pageNumber="211">
|
||
Female system typical for
|
||
<taxonomicName id="4C697CCBFFD0FF83FE765316FD8BFD8C" box="[494,635,512,536]" class="Gastropoda" family="Columbellidae" genus="Parvanachis" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="211" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B91DDB5AFFD0FF83FE765316FD8BFD8C" box="[494,635,512,536]" italics="true" pageId="10" pageNumber="211">Parvanachis</emphasis>
|
||
</taxonomicName>
|
||
, including a single gland mass, with muscular gonopericardial duct and no bursa copulatrix. Gland mass stains red (eosinophilic) in the middle, and pale purple (with haematoxylin) on both ends. The female sectioned had an egg capsule in the vestibule, which was otherwise short and thin-walled. Gonopericardial duct wide and coiled, epithelium cuboidal with large basal nuclei; duct has a wide, very thickwalled medial region (
|
||
<figureCitation id="13521BCDFFD0FF83FE0B5399FE2FFD3C" box="[403,479,655,680]" captionStart="FIGURE 3" captionStartId="6.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@6.[151,1436,193,1830]" captionTargetPageId="6" captionText="FIGURE 3. Histological sections from the female reproductive tract of Parvanachis. Scale bars: A – E, 200 µm; F – G, 100 µm. A. Longitudinal section of female gland mass of P. di c hrom a. B. Pericardium of P. pygmaea. C, D, E. Serial cross sections of P. pygmaea showing oviducal glands and gonopericardial duct. F, G. Serial sections of gonopericardial duct of P. diminuta." httpUri="https://zenodo.org/record/253313/files/figure.png" pageId="10" pageNumber="211">Figs. 3</figureCitation>
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F, G) with a large mass of red-stained material on one side (
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<figureCitation id="13521BCDFFD0FF83FBE75399FB31FD3C" box="[1151,1217,655,680]" captionStart="FIGURE 3" captionStartId="6.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@6.[151,1436,193,1830]" captionTargetPageId="6" captionText="FIGURE 3. Histological sections from the female reproductive tract of Parvanachis. Scale bars: A – E, 200 µm; F – G, 100 µm. A. Longitudinal section of female gland mass of P. di c hrom a. B. Pericardium of P. pygmaea. C, D, E. Serial cross sections of P. pygmaea showing oviducal glands and gonopericardial duct. F, G. Serial sections of gonopericardial duct of P. diminuta." httpUri="https://zenodo.org/record/253313/files/figure.png" pageId="10" pageNumber="211">Fig. 3</figureCitation>
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G), which might be partially dissolved sperm. The epithelium in that region is different than the rest, and the epithelial characteristics are not readily discernible, but the tissue appears to be secretory. Gonopericardial-pallial duct (gpp) splits off in this same region (
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<figureCitation id="13521BCDFFD0FF83FEB653EAFE81FC80" box="[302,369,764,789]" captionStart="FIGURE 3" captionStartId="6.[151,250,1852,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@6.[151,1436,193,1830]" captionTargetPageId="6" captionText="FIGURE 3. Histological sections from the female reproductive tract of Parvanachis. Scale bars: A – E, 200 µm; F – G, 100 µm. A. Longitudinal section of female gland mass of P. di c hrom a. B. Pericardium of P. pygmaea. C, D, E. Serial cross sections of P. pygmaea showing oviducal glands and gonopericardial duct. F, G. Serial sections of gonopericardial duct of P. diminuta." httpUri="https://zenodo.org/record/253313/files/figure.png" pageId="10" pageNumber="211">Fig. 3</figureCitation>
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F). Pericardium in the specimen sectioned with a large amount of unoriented sperm.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |