treatments-xml/data/03/C0/5A/03C05A2AFFA9FFD51CCFF91259A6C58F.xml
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<document id="DACF7CA5A94CE32B0523FF6B75A093AB" ID-DOI="10.11646/zootaxa.4790.2.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3889595" ID-ZooBank="825280D9-C670-4A0C-890E-78ABF89B127C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591859699868" checkinUser="plazi" docAuthor="Behan-Pelletier, Valerie M. &amp; Ermilov, Sergey G." docDate="2020" docId="03C05A2AFFA9FFD51CCFF91259A6C58F" docLanguage="en" docName="zootaxa.4790.2.9.pdf" docOrigin="Zootaxa 4790 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sculpteremaeus Behan-Pelletier &amp; Ermilov 2020, gen. nov." docType="treatment" docVersion="3" lastPageNumber="343" masterDocId="FFF92252FFA8FFD71C58FF935B5EC068" masterDocTitle="Sculpteremaeus olszanowskii gen. nov., sp. nov. (Acari, Oribatida, Cymbaeremaeidae) from chaparral in California, USA, with a reassessment of Cymbaeremaeidae" masterLastPageNumber="357" masterPageNumber="341" pageNumber="342" updateTime="1700276622102" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2CEE807FFAB8A9F2D3E587379ED2E320">Sculpteremaeus olszanowskii gen. nov., sp. nov. (Acari, Oribatida, Cymbaeremaeidae) from chaparral in California, USA, with a reassessment of Cymbaeremaeidae</mods:title>
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<mods:namePart id="B3ED7D85144A81978A54A4844C185D5B">Behan-Pelletier, Valerie M.</mods:namePart>
<mods:affiliation id="BAD4776CA4BE99130AC44EB405DE8EBB">Invertebrate Biodiversity Program, Research Branch, Agriculture and Agri-Food Canada, K. W. Neatby Bldg., Ottawa, Ontario, Canada. behanpv @ gmail. com; https: // orcid. org / 0000 - 0002 - 9880 - 2380</mods:affiliation>
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<mods:namePart id="50CE265317031D9A9335A264E5270A9B">Ermilov, Sergey G.</mods:namePart>
<mods:affiliation id="0D226D5FCCB8BABB8FCB4D02A5BAEB13">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Russia.</mods:affiliation>
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<mods:date id="B46421DA092303945F616ADB6834625A">2020</mods:date>
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<subSubSection id="C373B8B7FFA9FFD61CCFF912597AC6F3" box="[151,548,1665,1692]" pageId="1" pageNumber="342" type="nomenclature">
<paragraph id="8BD6EB3CFFA9FFD61CCFF912597AC6F3" blockId="1.[151,548,1665,1692]" box="[151,548,1665,1692]" pageId="1" pageNumber="342">
<heading id="D09E5C50FFA9FFD61CCFF912597AC6F3" bold="true" box="[151,548,1665,1692]" fontSize="11" level="1" pageId="1" pageNumber="342" reason="1">
<emphasis id="B91D372EFFA9FFD61CCFF912597AC6F3" bold="true" box="[151,548,1665,1692]" pageId="1" pageNumber="342">
Genus
<taxonomicName id="4C6990BFFFA9FFD61CB6F9125AF0C6F4" ID-CoL="8T7TJ" authority="Behan-Pelletier &amp; Ermilov, 2020" authorityName="Behan-Pelletier &amp; Ermilov" authorityYear="2020" box="[238,430,1665,1692]" class="Arachnida" family="Cymbaeremaeidae" genus="Sculpteremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="342" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91D372EFFA9FFD61CB6F9125AF0C6F4" bold="true" box="[238,430,1665,1692]" italics="true" pageId="1" pageNumber="342">Sculpteremaeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E8A55FFA9FFD61DEEF912597AC6F3" box="[438,548,1665,1691]" pageId="1" pageNumber="342" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373B8B7FFA9FFD61CCFF95559FAC6B5" box="[151,676,1733,1758]" pageId="1" pageNumber="342" type="materials_examined">
<paragraph id="8BD6EB3CFFA9FFD61CCFF95559FAC6B5" blockId="1.[151,676,1733,1758]" box="[151,676,1733,1758]" pageId="1" pageNumber="342">
<typeStatus id="54D2559EFFA9FFD61CCFF9555B97C6B6" box="[151,201,1734,1758]" pageId="1" pageNumber="342">Type</typeStatus>
species:
<taxonomicName id="4C6990BFFFA9FFD61D7EF9565910C6B5" authority="Behan-Pelletier &amp; Ermilov, 2020" authorityName="Behan-Pelletier &amp; Ermilov" authorityYear="2020" box="[294,590,1733,1757]" class="Arachnida" family="Cymbaeremaeidae" genus="Sculpteremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="342" phylum="Arthropoda" rank="species" species="olszanowskii" status="sp. nov.">
<emphasis id="B91D372EFFA9FFD61D7EF9565910C6B5" box="[294,590,1733,1757]" italics="true" pageId="1" pageNumber="342">Sculpteremaeus olszanowskii</emphasis>
</taxonomicName>
<emphasis id="B91D372EFFA9FFD61E0CF95659FAC6B5" bold="true" box="[596,676,1733,1757]" pageId="1" pageNumber="342">
<taxonomicNameLabel id="A22E8A55FFA9FFD61E0CF95659FAC6B5" box="[596,676,1733,1757]" pageId="1" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C373B8B7FFA9FFD61CCFF8985946C7B2" pageId="1" pageNumber="342" type="diagnosis">
<paragraph id="8BD6EB3CFFA9FFD61CCFF8985946C7B2" blockId="1.[151,1437,1803,2010]" pageId="1" pageNumber="342">
<emphasis id="B91D372EFFA9FFD61CCFF8985A28C74E" bold="true" box="[151,374,1803,1830]" pageId="1" pageNumber="342">Generic diagnosis.</emphasis>
<emphasis id="B91D372EFFA9FFD61DD9F89F5AE0C74E" box="[385,446,1804,1830]" italics="true" pageId="1" pageNumber="342">Adult</emphasis>
. With character states of
<taxonomicName id="4C6990BFFFA9FFD61EB5F89F58E3C74E" authorityName="Sellnick" authorityYear="1928" box="[749,957,1804,1830]" class="Arachnida" family="Cymbaeremaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="342" phylum="Arthropoda" rank="family">Cymbaeremaeidae</taxonomicName>
(
<bibRefCitation id="EFF896CDFFA9FFD61F97F89F5FD5C74E" author="Grandjean, F." box="[975,1163,1803,1830]" pageId="1" pageNumber="342" pagination="421 - 446" refId="ref9305" refString="Grandjean, F. (1953) Essai de classification des Oribates (Acariens). Bulletin de la Societe zoologique de France, 78 (5 - 6), 421 - 446." type="journal article" year="1953">Grandjean 1953</bibRefCitation>
;
<bibRefCitation id="EFF896CDFFA9FFD618C3F89F5ECEC74E" author="Behan-Pelletier, V. M." box="[1179,1424,1804,1830]" pageId="1" pageNumber="342" pagination="301 - 307" refId="ref8433" refString="Behan-Pelletier, V. M. (1988) Systematic relationships of Ametroproctus, with a modified definition of Cymbaeremaeidae (Acari: Oribatida). In: Channabasavanna, G. P. &amp; Viraktamath, C. A. (Eds.), Progress in Acarology. Vol. 1. Oxford &amp; IBH Publishing Co. Pvt. Ltd., New Delhi, pp. 301 - 307." type="book chapter" year="1988">Behan-Pelletier 1988</bibRefCitation>
). Lamella located dorsolaterally on prodorsum, comparatively long and narrow, with small cusp lacking teeth. Tutorium simple, ridge-like. Notogaster with marginal tectum; anterior margin not covering opening of bothridium. Dorsosejugal scissure interrupted medially. Small area of notogaster, posterior of interrupted dorsosejugal scissure, without sculpturing, with same pigmentation as rest of notogaster. Ventral region tapered posteriorly. Genital and anal apertures narrowly separated.
</paragraph>
</subSubSection>
<subSubSection id="C373B8B7FFAAFFD51C9FFF0B5E2EC3EB" pageId="2" pageNumber="343" type="description">
<paragraph id="8BD6EB3CFFAAFFD51C9FFF0B5E2EC3EB" blockId="2.[151,1437,152,1511]" pageId="2" pageNumber="343">
<emphasis id="B91D372EFFAAFFD51C9FFF0B5A07C0DB" bold="true" box="[199,345,152,179]" pageId="2" pageNumber="343">Description.</emphasis>
<emphasis id="B91D372EFFAAFFD51D39FF095A81C0DB" box="[353,479,154,179]" italics="true" pageId="2" pageNumber="343">Integument</emphasis>
: Surface sculptured, other than small area of notogaster posterior of interrupted dorsosejugal scissure.
<emphasis id="B91D372EFFAAFFD51D33FF2E5AB6C0BF" box="[363,488,189,215]" italics="true" pageId="2" pageNumber="343">Prodorsum</emphasis>
: Rostrum rounded. Lamella located dorsolaterally on prodorsum, comparatively long and narrow, with small cusp lacking teeth. Lamellar setae inserted anteroventrally on lamellar cusps. Translamella, prolamella, sublamella and sublamellar porose area absent. Tutorium simple, ridge-like. Bothridial seta short, globose. Bothridium cup-shaped, without notch. Dorsosejugal porose area, dorsophragmata and pleurophragmata absent.
<emphasis id="B91D372EFFAAFFD51CA8FEDD5A32C10F" box="[240,364,334,359]" italics="true" pageId="2" pageNumber="343">Notogaster</emphasis>
: Without octotaxic system of porose organs. Dorsosejugal scissure interrupted medially. Notogaster with marginal tectum; anterior margin not covering opening of bothridium. Pteromorphs absent; small bulge present in humeral region anterior of lyrifissure
<emphasis id="B91D372EFFAAFFD51EA6FE05584AC1C7" box="[766,788,406,431]" italics="true" pageId="2" pageNumber="343">ia</emphasis>
. With strong topographic relief comprising relatively flat marginal zone and medial convexity extending from level of lyrifissure
<emphasis id="B91D372EFFAAFFD51FEBFE295891C1BB" box="[947,975,442,467]" italics="true" pageId="2" pageNumber="343">im</emphasis>
posteriorly, tapering to nearly reach level of seta
<emphasis id="B91D372EFFAAFFD51CB3FE4E5BA7C19F" box="[235,249,477,503]" italics="true" pageId="2" pageNumber="343">h</emphasis>
<subScript id="17EDE979FFAAFFD51CA1FE7D5A5FC194" attach="left" box="[249,257,494,508]" fontSize="6" pageId="2" pageNumber="343">2</subScript>
. Ten pairs of notogastral setae, all in marginal zone. Lyrifissure
<emphasis id="B91D372EFFAAFFD51FB8FE4D58A2C19F" box="[992,1020,478,503]" italics="true" pageId="2" pageNumber="343">im</emphasis>
most centrodorsal sensory structure, positioned at juncture of marginal zone and central convexity.
<taxonomicName id="4C6990BFFFAAFFD51F0EFD925FCEC273" authority=": Subcapitulum" authorityName="Subcapitulum" box="[854,1168,513,539]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="343" phylum="Nematoda" rank="genus">
<emphasis id="B91D372EFFAAFFD51F0EFD9258BBC273" box="[854,997,513,539]" italics="true" pageId="2" pageNumber="343">Gnathosoma</emphasis>
: Subcapitulum
</taxonomicName>
diarthric. Mentum with tectum. Palp with setation 0-2-1-3-9(+ω). Solenidion of palptarsus bacilliform, free; eupathidium positioned on tu- bercle. Axillary saccule absent. Chelicera chelate-dentate, with two acuminate setae and tapered Trägårdhs organ.
<emphasis id="B91D372EFFAAFFD51CCFFDFE5903C2EF" box="[151,605,621,647]" italics="true" pageId="2" pageNumber="343">Lateral podosomal and epimeral regions</emphasis>
: Pedotectum I represented by lamina of medium size, completely covering acetabulum I. Pedotectum II biconvex in dorso-ventral aspect, triangular in lateral aspect, with anterior lamina covering about half of acetabulum II. Custodium, circumpedal carina and humeral porose areas absent. Discidium present. Epimeral setal formula (I-IV) 3-1-2-2.
<emphasis id="B91D372EFFAAFFD51ED8FD4A5815C29B" box="[640,843,729,755]" italics="true" pageId="2" pageNumber="343">Anogenital region</emphasis>
: Ventral region tapered posteriorly. Genital and anal apertures narrowly separated. Four pairs of genital, one pair of aggenital, two pairs of anal and three pairs of adanal setae present;
<emphasis id="B91D372EFFAAFFD51D6CFCB25A0EC353" box="[308,336,801,827]" italics="true" pageId="2" pageNumber="343">ad</emphasis>
<subScript id="17EDE979FFAAFFD51D08FCA15A06C328" attach="left" box="[336,344,818,832]" fontSize="6" pageId="2" pageNumber="343">1</subScript>
and
<emphasis id="B91D372EFFAAFFD51DD4FCB25AF6C353" box="[396,424,801,827]" italics="true" pageId="2" pageNumber="343">ad</emphasis>
<subScript id="17EDE979FFAAFFD51DF0FCA15AEEC328" attach="left" box="[424,432,818,832]" fontSize="6" pageId="2" pageNumber="343">2</subScript>
postanal,
<emphasis id="B91D372EFFAAFFD51E78FCB25962C353" box="[544,572,801,827]" italics="true" pageId="2" pageNumber="343">ad</emphasis>
<subScript id="17EDE979FFAAFFD51E64FCA1591AC328" attach="left" box="[572,580,818,832]" fontSize="6" pageId="2" pageNumber="343">3</subScript>
closely lateral to anal aperture. Preanal organ hollow, cup-like.
<emphasis id="B91D372EFFAAFFD5195CFCB15E67C353" box="[1284,1337,802,827]" italics="true" pageId="2" pageNumber="343">Legs</emphasis>
: All pretarsi heterotridactylous. Tibia I with dorsodistal apophysis bearing solenidia; φ
<subScript id="17EDE979FFAAFFD51853FCC55F4DC30C" attach="left" box="[1035,1043,854,868]" fontSize="6" pageId="2" pageNumber="343">1</subScript>
distally, φ
<subScript id="17EDE979FFAAFFD518D1FCC55FCFC30C" attach="left" box="[1161,1169,854,868]" fontSize="6" pageId="2" pageNumber="343">2</subScript>
medially on apophysis, posteromedial to φ
<subScript id="17EDE979FFAAFFD51D32FCE95A2CC3E0" attach="left" box="[362,370,890,904]" fontSize="6" pageId="2" pageNumber="343">1</subScript>
. Tarsi, tibiae, femora and trochanters III, IV with porose areas.
<emphasis id="B91D372EFFAAFFD51862FCFA5FB5C3EB" box="[1082,1259,873,899]" italics="true" pageId="2" pageNumber="343">Juvenile instars</emphasis>
: Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C373B8B7FFAAFFD51C9FFC1E5982C3A3" pageId="2" pageNumber="343" type="etymology">
<paragraph id="8BD6EB3CFFAAFFD51C9FFC1E5982C3A3" blockId="2.[151,1437,152,1511]" pageId="2" pageNumber="343">
<emphasis id="B91D372EFFAAFFD51C9FFC1E5A10C3CF" bold="true" box="[199,334,909,935]" pageId="2" pageNumber="343">Etymology.</emphasis>
The name
<taxonomicName id="4C6990BFFFAAFFD51D94FC1E5922C3CF" authorityName="Behan-Pelletier &amp; Ermilov" authorityYear="2020" box="[460,636,909,935]" class="Arachnida" family="Cymbaeremaeidae" genus="Sculpteremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="B91D372EFFAAFFD51D94FC1E5922C3CF" box="[460,636,909,935]" italics="true" pageId="2" pageNumber="343">Sculpteremaeus</emphasis>
</taxonomicName>
refers to the presence of sculpturing on the body +
<emphasis id="B91D372EFFAAFFD51895FC1D5E69C3CF" box="[1229,1335,910,935]" italics="true" pageId="2" pageNumber="343">eremaeus</emphasis>
, a common suffix for generic names in
<taxonomicName id="4C6990BFFFAAFFD51E5CFC225986C3A3" authorityName="Sellnick" authorityYear="1928" box="[516,728,945,971]" class="Arachnida" family="Cymbaeremaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="family">Cymbaeremaeidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C373B8B7FFAAFFD51C9FFC4659A6C58F" pageId="2" pageNumber="343" type="discussion">
<paragraph id="8BD6EB3CFFAAFFD51C9FFC465F9FC4A0" blockId="2.[151,1437,152,1511]" pageId="2" pageNumber="343">
<emphasis id="B91D372EFFAAFFD51C9FFC465A65C387" bold="true" box="[199,315,981,1007]" pageId="2" pageNumber="343">Remarks.</emphasis>
Adults of
<taxonomicName id="4C6990BFFFAAFFD51DECFC46593AC387" authority="Behan-Pelletier &amp; Ermilov, 2020" authorityName="Behan-Pelletier &amp; Ermilov" authorityYear="2020" box="[436,612,981,1007]" class="Arachnida" family="Cymbaeremaeidae" genus="Sculpteremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91D372EFFAAFFD51DECFC46593AC387" box="[436,612,981,1007]" italics="true" pageId="2" pageNumber="343">Sculpteremaeus</emphasis>
</taxonomicName>
<emphasis id="B91D372EFFAAFFD51E34FC465988C387" bold="true" box="[620,726,981,1007]" pageId="2" pageNumber="343">
<taxonomicNameLabel id="A22E8A55FFAAFFD51E34FC465988C387" box="[620,726,981,1007]" pageId="2" pageNumber="343" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
are morphologically most similar to species of
<taxonomicName id="4C6990BFFFAAFFD518A9FC465ECBC387" baseAuthorityName="Behan-Pelletier" baseAuthorityYear="1987" box="[1265,1429,981,1007]" class="Arachnida" family="Ametroproctidae" genus="Ametroproctus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="B91D372EFFAAFFD518A9FC465ECBC387" box="[1265,1429,981,1007]" italics="true" pageId="2" pageNumber="343">Ametroproctus</emphasis>
</taxonomicName>
, and specifically those included in the subgenus
<taxonomicName id="4C6990BFFFAAFFD51EF5FC6A5F51C47B" authority="(Ametroproctus)" authorityName="Higgins &amp; Woolley" authorityYear="1968" baseAuthorityName="Ametroproctus" box="[685,1039,1017,1043]" class="Arachnida" family="Ametroproctidae" genus="Ametroproctus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="subGenus" subGenus="Ametroproctus">
<emphasis id="B91D372EFFAAFFD51EF5FC6A580FC47B" box="[685,849,1017,1043]" italics="true" pageId="2" pageNumber="343">Ametroproctus</emphasis>
(
<emphasis id="B91D372EFFAAFFD51F39FC6A5F5BC47B" box="[865,1029,1017,1043]" italics="true" pageId="2" pageNumber="343">Ametroproctus</emphasis>
)
</taxonomicName>
in the presence of short interlamellar setae; medially incomplete dorsosejugal scissure; small area of notogaster, posterior of interrupted dorsosejugal scissure, without sculpturing, with same or lighter pigmentation as rest of notogaster; 10 pairs of notogastral setae with centrodorsal region of notogaster glabrous; four pairs of genital setae; and genital and anal apertures closely adjacent. Adults differ from those of
<emphasis id="B91D372EFFAAFFD51E6AFB1A591DC4CB" box="[562,579,1161,1187]" italics="true" pageId="2" pageNumber="343">A</emphasis>
. (
<taxonomicName id="4C6990BFFFAAFFD51E02FB1A59A0C4CB" baseAuthorityName="Behan-Pelletier" baseAuthorityYear="1987" box="[602,766,1161,1187]" class="Arachnida" family="Ametroproctidae" genus="Ametroproctus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="B91D372EFFAAFFD51E02FB1A59A0C4CB" box="[602,766,1161,1187]" italics="true" pageId="2" pageNumber="343">Ametroproctus</emphasis>
</taxonomicName>
) species by having a thin lamella and small, separate lamellar cusps (versus well-developed lamella with large lamellar cusps, connected by translamella).
</paragraph>
<paragraph id="8BD6EB3CFFAAFFD51C9FFB425E50C53F" blockId="2.[151,1437,152,1511]" pageId="2" pageNumber="343">
It is unclear whether the small circular area of notogaster, posterior of the interrupted dorsosejugal scissure, which lacks sculpturing, but has the same pigmentation as rest of notogaster is a true lenticulus, such as is found in
<taxonomicName id="4C6990BFFFAAFFD51CCFFA8A5A6DC55B" authorityName="Colloff" authorityYear="2011" box="[151,307,1305,1331]" class="Arachnida" family="Cymbaeremaeidae" genus="Spineremaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="B91D372EFFAAFFD51CCFFA8A5A6DC55B" box="[151,307,1305,1331]" italics="true" pageId="2" pageNumber="343">Spineremaeus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6990BFFFAAFFD51D31FA8A599DC55B" authority="(Colloff 2011)" baseAuthorityName="Colloff" baseAuthorityYear="2011" box="[361,707,1305,1331]" class="Arachnida" family="Cymbaeremaeidae" genus="Scapheremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="B91D372EFFAAFFD51D31FA8A5946C55B" box="[361,536,1305,1331]" italics="true" pageId="2" pageNumber="343">Scapheremaeus</emphasis>
(
<bibRefCitation id="EFF896CDFFAAFFD51E7EFA8A59E5C55B" author="Colloff, M. J." box="[550,699,1305,1331]" pageId="2" pageNumber="343" pagination="19 - 37" refId="ref8925" refString="Colloff, M. J. (2011) A new genus of oribatid mite, Spineremaeus gen. nov. and three new species of Scapheremaeus (Acari: Oribatida: Cymbaeremaeidae) from Norfolk Island, South-west Pacific, and their boigeographical affinities. Zootaxa, 2828 (1), 19 - 37. https: // doi. org / 10.11646 / zootaxa. 2828.1.2" type="journal article" year="2011">Colloff 2011</bibRefCitation>
)
</taxonomicName>
. This area of
<taxonomicName id="4C6990BFFFAAFFD51F01FA8A5F57C55B" authorityName="Behan-Pelletier &amp; Ermilov" authorityYear="2020" box="[857,1033,1305,1331]" class="Arachnida" family="Cymbaeremaeidae" genus="Sculpteremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="B91D372EFFAAFFD51F01FA8A5F57C55B" box="[857,1033,1305,1331]" italics="true" pageId="2" pageNumber="343">Sculpteremaeus</emphasis>
</taxonomicName>
has the same morphology as that of
<taxonomicName id="4C6990BFFFAAFFD51CCFFAAE5A65C53F" baseAuthorityName="Behan-Pelletier" baseAuthorityYear="1987" box="[151,315,1341,1367]" class="Arachnida" family="Ametroproctidae" genus="Ametroproctus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="B91D372EFFAAFFD51CCFFAAE5A65C53F" box="[151,315,1341,1367]" italics="true" pageId="2" pageNumber="343">Ametroproctus</emphasis>
</taxonomicName>
(
<emphasis id="B91D372EFFAAFFD51D14FAAE5A03C53F" box="[332,349,1341,1367]" italics="true" pageId="2" pageNumber="343">A</emphasis>
.)
<emphasis id="B91D372EFFAAFFD51D2CFAAE5A8CC53F" box="[372,466,1341,1367]" italics="true" pageId="2" pageNumber="343">oresbios</emphasis>
<bibRefCitation id="EFF896CDFFAAFFD51D81FAAE59A9C53F" author="Higgins, H. G. &amp; Woolley, T. A." box="[473,759,1341,1367]" pageId="2" pageNumber="343" pagination="44 - 46" refId="ref9446" refString="Higgins, H. G. &amp; Woolley, T. A. (1968) Ametroproctus, a new genus of charassobatid mites from the United States (Acari: Cryptostigmata: Charassobatidae). Great Basin Naturalist, 28 (1), 44 - 46. https: // doi. org / 10.5962 / bhl. part. 28686" type="journal article" year="1968">Higgins &amp; Woolley, 1968</bibRefCitation>
and
<emphasis id="B91D372EFFAAFFD51F75FAAE5818C53F" box="[813,838,1341,1367]" italics="true" pageId="2" pageNumber="343">A.</emphasis>
(
<emphasis id="B91D372EFFAAFFD51F0EFAAE5839C53F" box="[854,871,1341,1367]" italics="true" pageId="2" pageNumber="343">A</emphasis>
.)
<emphasis id="B91D372EFFAAFFD51F26FAAE5F52C53F" box="[894,1036,1341,1367]" italics="true" pageId="2" pageNumber="343">tuberculosus</emphasis>
<bibRefCitation id="EFF896CDFFAAFFD5184BFAAE5E57C53F" author="Behan-Pelletier, V. M." box="[1043,1289,1341,1367]" pageId="2" pageNumber="343" pagination="505 - 535" refId="ref8387" refString="Behan-Pelletier, V. M. (1987) Redefinition of Ametroproctus (Acari: Oribatida) with descriptions of new species. The Canadian Entomologist, 119, 505 - 535. https: // doi. org / 10.4039 / Ent 119505 - 6" type="journal article" year="1987">Behan-Pelletier, 1987</bibRefCitation>
.
</paragraph>
<paragraph id="8BD6EB3CFFAAFFD51C9FFAF259A6C58F" blockId="2.[151,1437,152,1511]" pageId="2" pageNumber="343">
It is possible that with discovery of its immatures,
<taxonomicName id="4C6990BFFFAAFFD51F50FAF258E6C513" authorityName="Behan-Pelletier &amp; Ermilov" authorityYear="2020" box="[776,952,1377,1403]" class="Arachnida" family="Cymbaeremaeidae" genus="Sculpteremaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="B91D372EFFAAFFD51F50FAF258E6C513" box="[776,952,1377,1403]" italics="true" pageId="2" pageNumber="343">Sculpteremaeus</emphasis>
</taxonomicName>
may be considered a subgenus of
<emphasis id="B91D372EFFAAFFD5191AFAF25BACC5F7" italics="true" pageId="2" pageNumber="343">
<taxonomicName id="4C6990BFFFAAFFD5191AFAF25BB0C5F7" baseAuthorityName="Behan-Pelletier" baseAuthorityYear="1987" class="Arachnida" family="Ametroproctidae" genus="Ametroproctus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">Ametroproctus</taxonomicName>
.
</emphasis>
<bibRefCitation id="EFF896CDFFAAFFD51CAFFA165AA8C5F7" author="Behan-Pelletier, V. M." box="[247,502,1413,1439]" pageId="2" pageNumber="343" pagination="505 - 535" refId="ref8387" refString="Behan-Pelletier, V. M. (1987) Redefinition of Ametroproctus (Acari: Oribatida) with descriptions of new species. The Canadian Entomologist, 119, 505 - 535. https: // doi. org / 10.4039 / Ent 119505 - 6" type="journal article" year="1987">Behan-Pelletier (1987)</bibRefCitation>
placed
<emphasis id="B91D372EFFAAFFD51E14FA165987C5F7" box="[588,729,1413,1439]" italics="true" pageId="2" pageNumber="343">Coropoculia</emphasis>
in synonymy with
<taxonomicName id="4C6990BFFFAAFFD51FF6FA165F0CC5F7" baseAuthorityName="Behan-Pelletier" baseAuthorityYear="1987" box="[942,1106,1413,1439]" class="Arachnida" family="Ametroproctidae" genus="Ametroproctus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="B91D372EFFAAFFD51FF6FA165F0CC5F7" box="[942,1106,1413,1439]" italics="true" pageId="2" pageNumber="343">Ametroproctus</emphasis>
</taxonomicName>
, and recognized 2 subgenera, on the basis of immatures, specifically the tritonymph of
<taxonomicName id="4C6990BFFFAAFFD51F4CFA3A5FF6C5AB" authority="Behan-Pelletier, 1987" authorityName="Behan-Pelletier" authorityYear="1987" box="[788,1192,1449,1475]" class="Arachnida" family="Ametroproctidae" genus="Ametroproctus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="aridus" subGenus="Ametroproctus">
<emphasis id="B91D372EFFAAFFD51F4CFA3A5872C5AB" box="[788,812,1449,1475]" italics="true" pageId="2" pageNumber="343">A.</emphasis>
(
<emphasis id="B91D372EFFAAFFD51F64FA3A580AC5AB" box="[828,852,1449,1475]" italics="true" pageId="2" pageNumber="343">A.</emphasis>
)
<emphasis id="B91D372EFFAAFFD51F3CFA3A58F2C5AB" box="[868,940,1449,1475]" italics="true" pageId="2" pageNumber="343">aridus</emphasis>
<bibRefCitation id="EFF896CDFFAAFFD51FEAFA3A5FF6C5AB" author="Behan-Pelletier, V. M." box="[946,1192,1449,1475]" pageId="2" pageNumber="343" pagination="505 - 535" refId="ref8387" refString="Behan-Pelletier, V. M. (1987) Redefinition of Ametroproctus (Acari: Oribatida) with descriptions of new species. The Canadian Entomologist, 119, 505 - 535. https: // doi. org / 10.4039 / Ent 119505 - 6" type="journal article" year="1987">Behan-Pelletier, 1987</bibRefCitation>
</taxonomicName>
, which shows character states that are expressed in species of
<emphasis id="B91D372EFFAAFFD51E3BFA5E59A6C58F" box="[611,760,1485,1511]" italics="true" pageId="2" pageNumber="343">Coropoculia.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>