213 lines
22 KiB
XML
213 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.43.80969" ID-Pensoft-Pub="1314-2615-43-61" ID-Pensoft-UUID="94B591976E9051C7AA220F5AB7F30AF7" ID-ZooBank="9B9E6A5AA0784AC0A1DF6B1D32FF4DB3" ModsDocID="1314-2615-43-61" checkinTime="1654082233325" checkinUser="pensoft" docAuthor="Sasakawa, Koji & Ito, Hirotaro" docDate="2022" docId="8CD5F85F6B595FA08552C1864653A65E" docLanguage="en" docName="SubterBiol 43: 61-71" docOrigin="Subterranean Biology 43" docPubDate="2022-05-31" docSource="http://dx.doi.org/10.3897/subtbiol.43.80969" docTitle="Pterostichus (Nialoe) asahinus Habu & Baba 1960" docType="treatment" docVersion="1" id="94B591976E9051C7AA220F5AB7F30AF7" lastPageNumber="61" masterDocId="94B591976E9051C7AA220F5AB7F30AF7" masterDocTitle="The identities of two species in the Pterostichus macrogenys species group of subterranean carabid beetles (Coleoptera, Carabidae) revealed by external morphometric analysis and comparative genital morphology" masterLastPageNumber="71" masterPageNumber="61" pageNumber="61" updateTime="1654082233325" updateUser="pensoft">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>The identities of two species in the Pterostichus macrogenys species group of subterranean carabid beetles (Coleoptera, Carabidae) revealed by external morphometric analysis and comparative genital morphology</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Sasakawa, Koji</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9246-5777</mods:nameIdentifier>
|
|
<mods:affiliation>Laboratory of Zoology, Department of Science Education, Faculty of Education, Chiba University, 1 - 33 Yayoi-cho, Inage-ku, Chiba-shi, Chiba 263 - 8522, Japan</mods:affiliation>
|
|
<mods:nameIdentifier type="email">ksasa@chiba-u.jp</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Ito, Hirotaro</mods:namePart>
|
|
<mods:affiliation>1 - 14 - 16 Awayama, Niigata-shi, Niigata 950 - 0843, Japan</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>Subterranean Biology</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2022</mods:date>
|
|
<mods:detail type="pubDate">
|
|
<mods:number>2022-05-31</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="volume">
|
|
<mods:number>43</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>61</mods:start>
|
|
<mods:end>71</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/subtbiol.43.80969</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/subtbiol.43.80969</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1314-2615-43-61</mods:identifier>
|
|
<mods:identifier type="ZooBank">9B9E6A5AA0784AC0A1DF6B1D32FF4DB3</mods:identifier>
|
|
<mods:identifier type="Pensoft-UUID">94B591976E9051C7AA220F5AB7F30AF7</mods:identifier>
|
|
</mods:mods>
|
|
<treatment LSID="urn:lsid:plazi:treatment:8CD5F85F6B595FA08552C1864653A65E" httpUri="http://treatment.plazi.org/id/8CD5F85F6B595FA08552C1864653A65E" lastPageNumber="61" pageId="0" pageNumber="61">
|
|
<subSubSection pageId="0" pageNumber="61" type="nomenclature">
|
|
<paragraph pageId="0" pageNumber="61">
|
|
<taxonomicName LSID="8CD5F85F-6B59-5FA0-8552-C1864653A65E" authority="Habu & Baba, 1960" authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus" subGenus="Nialoe">Pterostichus (Nialoe) asahinus Habu & Baba, 1960</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="61" type="description">
|
|
<paragraph pageId="0" pageNumber="61">
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pterostichus asahinus male from Miomote, on the right bank of the Miomotegawa River A habitus dorsal view B-E Endophallus left lateral (B) ventral (C) right lateral (D) and dorsal (E) views F-H right paramere left lateral (F) apical (G) and dorsal (H) views. go, gonopore; lal, left apical lobe; lpl, left preapical lobe; rpl, right preapical lobe." figureDoi="10.3897/subtbiol.43.80969.figure2" httpUri="https://binary.pensoft.net/fig/690341" pageId="0" pageNumber="61">Fig. 2</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="61" type="reference_group">
|
|
<paragraph pageId="0" pageNumber="61">
|
|
<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Paralianoe) macrogenys subsp. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="macrogenys" subGenus="Paralianoe" subSpecies="asahinus">Pterostichus (Paralianoe) macrogenys asahinus</taxonomicName>
|
|
:
|
|
<bibRefCitation author="Habu, A" journalOrPublisher="Entomological Review Japan" pageId="0" pageNumber="61" refId="B5" refString="Habu, A, Baba, K, 1960. A new subspecies of Pterostichus macrogenys Bates from Niigata Prefecture, Japan. Akitu 9: 62." title="A new subspecies of Pterostichus macrogenys Bates from Niigata Prefecture, Japan. Akitu 9: 62." year="1960">Habu and Baba (1960)</bibRefCitation>
|
|
: 62 (original description), holotype ♀: "Mt. Dorokujin, Mts. Asahi, Niigata Pref." [Miomote, Mt.
|
|
<normalizedToken originalValue="Dôrokujinpô">Dorokujinpo</normalizedToken>
|
|
, Murakami-shi, Niigata Prefecture, Japan];
|
|
<bibRefCitation author="Habu, A" journalOrPublisher="Transactions of Essa Entomological Society" pageId="0" pageNumber="61" pagination="1 - 58" refId="B6" refString="Habu, A, Baba, K, 1972. List of Carabidae (excluding Carabini) from Niigata Prefecture, Japan. Transactions of Essa Entomological Society 41: 1 - 58" title="List of Carabidae (excluding Carabini) from Niigata Prefecture, Japan." volume="41" year="1972">Habu and Baba (1972)</bibRefCitation>
|
|
: 19.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="61">
|
|
<taxonomicName authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Paralianoe) asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus" subGenus="Paralianoe">Pterostichus (Paralianoe) asahinus</taxonomicName>
|
|
:
|
|
<bibRefCitation author="Habu, A" journalOrPublisher="Entomological Review Japan" pageId="0" pageNumber="61" pagination="13 - 17" refId="B4" refString="Habu, A, 1977. On two Pterostichus species found in caves in Niigata Prefecture, Japan (Coleoptera, Carabidae). Entomological Review Japan 30: 13 - 17" title="On two Pterostichus species found in caves in Niigata Prefecture, Japan (Coleoptera, Carabidae)." volume="30" year="1977">Habu (1977)</bibRefCitation>
|
|
: 14 (part).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="61">
|
|
<taxonomicName authorityName="Sasakawa & Itô" authorityYear="2022" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus macrogenys" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="macrogenys">Pterostichus macrogenys</taxonomicName>
|
|
:
|
|
<bibRefCitation author="Tanaka, K" editor="Ueno, S" journalOrPublisher="Vol. II. Hoikusha, Osaka" pageId="0" pageNumber="61" pagination="105 - 138" refId="B21" refString="Tanaka, K, 1985. Carabidae (Pterostichinae, Zabrinae). In: Ueno, S, Kurosawa, Y, Sato, M, Eds., The Coleoptera of Japan in Color. Vol. II. Hoikusha, Osaka: 105 - 138" title="Carabidae (Pterostichinae, Zabrinae)." volumeTitle="The Coleoptera of Japan in Color." year="1985">Tanaka (1985)</bibRefCitation>
|
|
: 114 (part?).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="61">
|
|
<taxonomicName authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus" subGenus="Nialoe">Pterostichus (Nialoe) asahinus</taxonomicName>
|
|
:
|
|
<bibRefCitation author="Bousquet, Y" editor="Loebl, I" journalOrPublisher="1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden" pageId="0" pageNumber="61" pagination="675 - 755" refId="B2" refString="Bousquet, Y, 2017. Tribe Pterostichini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Vol. 1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden: 675 - 755" title="Tribe Pterostichini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol." year="2017">Bousquet (2017)</bibRefCitation>
|
|
: 724.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="61">
|
|
<taxonomicName authorityName="Sasakawa" authorityYear="2005" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Nialoe) falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus" subGenus="Nialoe">Pterostichus (Nialoe) falcispinus</taxonomicName>
|
|
:
|
|
<bibRefCitation author="Sasakawa, K" journalOrPublisher="Biogeography" pageId="0" pageNumber="61" pagination="69 - 78" refId="B12" refString="Sasakawa, K, 2005. Pterostichus macrogenys Bates, 1883 (Coleoptera, Carabidae) and its allied species of northern Japan. Biogeography 7: 69 - 78" title="Pterostichus macrogenys Bates, 1883 (Coleoptera, Carabidae) and its allied species of northern Japan." volume="7" year="2005">Sasakawa (2005)</bibRefCitation>
|
|
: 75 (original description), holotype ♂: "Cave Ishikiri, Nakajo-Machi, N-Echigo" [Ishikiri Cave, Mt. Ishikiriyama, Haguro, Tainai-shi, Niigata Prefecture, Japan];
|
|
<bibRefCitation author="Bousquet, Y" editor="Loebl, I" journalOrPublisher="1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden" pageId="0" pageNumber="61" pagination="675 - 755" refId="B2" refString="Bousquet, Y, 2017. Tribe Pterostichini Bonelli, 1810. In: Loebl, I, Loebl, D, Eds., Catalogue of Palaearctic Coleoptera, Vol. 1, Archostemata-Myxophaga-Adephaga, Revised and Updated Edition. Brill, Leiden: 675 - 755" title="Tribe Pterostichini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Vol." year="2017">Bousquet (2017)</bibRefCitation>
|
|
: 724;
|
|
<bibRefCitation DOI="https://doi.org/10.3390/insects11110803" author="Sasakawa, K" journalOrPublisher="Applied Entomology and Zoology" pageId="0" pageNumber="61" refId="B17" refString="Sasakawa, K, Mitsuduka, Y, Ito, H, 2020. Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803. https://doi.org/10.3390/insects11110803" title="Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803." url="https://doi.org/10.3390/insects11110803" year="2020">Sasakawa et al. (2020)</bibRefCitation>
|
|
: 7. Syn. nov.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="61" type="specimen examined">
|
|
<paragraph pageId="0" pageNumber="61">Specimen examined.</paragraph>
|
|
<paragraph pageId="0" pageNumber="61">
|
|
1♂, Miomote, alt. 276 m, on the right bank of Miomotegawa River, Murakami-shi, Niigata Prefecture, Japan (
|
|
<geoCoordinate degrees="38.273211" direction="north" orientation="latitude" precision="1" value="38.273212">38.273211°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="139.779922" direction="east" orientation="longitude" precision="1" value="139.77992">139.779922°E</geoCoordinate>
|
|
), 12.vi.-17.vii.2021,
|
|
<normalizedToken originalValue="Hirotarô">Hirotaro</normalizedToken>
|
|
<normalizedToken originalValue="Itô">Ito</normalizedToken>
|
|
leg., in the collection of HI.
|
|
</paragraph>
|
|
<caption doi="10.3897/subtbiol.43.80969.figure2" httpUri="https://binary.pensoft.net/fig/690341" pageId="0" pageNumber="61" start="Figure 2" startId="F2">
|
|
<paragraph pageId="0" pageNumber="61">
|
|
<emphasis bold="true" pageId="0" pageNumber="61">Figure 2.</emphasis>
|
|
<taxonomicName authorityName="Habu & Baba" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="61">Pterostichus asahinus</emphasis>
|
|
</taxonomicName>
|
|
male from Miomote, on the right bank of the Miomotegawa River
|
|
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
|
|
habitus dorsal view
|
|
<emphasis bold="true" pageId="0" pageNumber="61">B-E</emphasis>
|
|
Endophallus left lateral (
|
|
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
|
|
) ventral (
|
|
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
|
|
) right lateral (
|
|
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
|
|
) and dorsal (
|
|
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
|
|
) views
|
|
<emphasis bold="true" pageId="0" pageNumber="61">F-H</emphasis>
|
|
right paramere left lateral (
|
|
<emphasis bold="true" pageId="0" pageNumber="61">F</emphasis>
|
|
) apical (
|
|
<emphasis bold="true" pageId="0" pageNumber="61">G</emphasis>
|
|
) and dorsal (
|
|
<emphasis bold="true" pageId="0" pageNumber="61">H</emphasis>
|
|
) views.
|
|
<emphasis bold="true" pageId="0" pageNumber="61">go</emphasis>
|
|
, gonopore;
|
|
<emphasis bold="true" pageId="0" pageNumber="61">lal</emphasis>
|
|
, left apical lobe;
|
|
<emphasis bold="true" pageId="0" pageNumber="61">lpl</emphasis>
|
|
, left preapical lobe;
|
|
<emphasis bold="true" pageId="0" pageNumber="61">rpl</emphasis>
|
|
, right preapical lobe.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="61" type="notes">
|
|
<paragraph pageId="0" pageNumber="61">Notes.</paragraph>
|
|
<paragraph pageId="0" pageNumber="61">
|
|
<bibRefCitation DOI="https://doi.org/10.3390/insects11110803" author="Sasakawa, K" journalOrPublisher="Applied Entomology and Zoology" pageId="0" pageNumber="61" refId="B17" refString="Sasakawa, K, Mitsuduka, Y, Ito, H, 2020. Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803. https://doi.org/10.3390/insects11110803" title="Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803." url="https://doi.org/10.3390/insects11110803" year="2020">Sasakawa et al. (2020)</bibRefCitation>
|
|
suggested that
|
|
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
|
</taxonomicName>
|
|
might be conspecific. However, this hypothesis remained untested because males from the type locality of
|
|
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
|
</taxonomicName>
|
|
have not been examined. The male specimen examined here was obtained from a valley southwest of Mt.
|
|
<normalizedToken originalValue="Dôrokujinpô">Dorokujinpo</normalizedToken>
|
|
, which can virtually be regarded as the type locality (Fig.
|
|
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the Pterostichus macrogenys species group in northern Chubu, northern Kanto, and southern Tohoku Districts, Honshu, based on specimens with unambiguous identities, modified from Sasakawa et al. (2020) and Sato (2021) 1 P. chokaisanus Sasakawa, 2009 2 P. kurikomasanus Sasakawa, 2005 3 P. asahinus Habu and Baba, 1960 4 P. takadateyamanus Sasakawa, 2009 5 P. adatarasanus Sasakawa, 2005 6 P. gassanus Sasakawa, 2009 7 P. shinbodakensis Sasakawa & Ito, 2017 8 P. awashimaensis sp. nov. 9 P. iwakiensis Sasakawa, 2009 10 P. eboshiyamanus Sasakawa, 2009 11 P. tateishiyamanus Sasakawa & Ito, 2017 12 P. monolineatus Sasakawa, Mitsuduka, & Ito, 2020 13 P. yamizosanus Sasakawa, 2005 14 P. yahikosanus Sasakawa, 2009 15 P. ohsawacavus Sasakawa, 2005 16 P. sumondakensis Sasakawa, 2005 17 P. isolatus Sasakawa, 2005 18 P. nakamiyorinus Morita, Ohkawa & Kurihara, 2013 19 P. momuranus Morita, Ohkawa & Kurihara, 2013 20 P. macrogenys Bates, 1883 21 P. nagasawai Ito & Ogai, 2015. Red letters denote the type localities of each species. The blue letter indicates where the male specimen of P. asahinus newly examined here was collected." figureDoi="10.3897/subtbiol.43.80969.figure1" httpUri="https://binary.pensoft.net/fig/690340" pageId="0" pageNumber="61">1</figureCitation>
|
|
). The structures of the endophallus and right paramere of this specimen are identical to those of the eastern type of
|
|
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
|
</taxonomicName>
|
|
in
|
|
<bibRefCitation DOI="https://doi.org/10.3390/insects11110803" author="Sasakawa, K" journalOrPublisher="Applied Entomology and Zoology" pageId="0" pageNumber="61" refId="B17" refString="Sasakawa, K, Mitsuduka, Y, Ito, H, 2020. Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803. https://doi.org/10.3390/insects11110803" title="Unexpected species identities and interspecific relationships in a subterranean beetle lineage, the Pterostichus macrogenys species group (Coleoptera, Carabidae), revealed by fine-scale field sampling and detailed morphological comparisons. Insects 11: 803." url="https://doi.org/10.3390/insects11110803" year="2020">Sasakawa et al. (2020)</bibRefCitation>
|
|
. Its body length is intermediate between that of the eastern and western types of
|
|
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
|
</taxonomicName>
|
|
as follows: body length from mandible apices to elytral end (BLm) 16.32 mm, that from anterior margin of labrum (BLl) 14.72 mm, and that from clypeal apex (BLc) 14.26 mm. Based on these results, we regard
|
|
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. falcispinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="falcispinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="61">P. falcispinus</emphasis>
|
|
</taxonomicName>
|
|
as conspecific. The results of the discriminant analysis support this conclusion. The
|
|
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
|
</taxonomicName>
|
|
holotype was classified in this newly defined
|
|
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
|
</taxonomicName>
|
|
in the discriminant function analysis, and was within the area of
|
|
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. asahinus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="asahinus">
|
|
<emphasis italics="true" pageId="0" pageNumber="61">P. asahinus</emphasis>
|
|
</taxonomicName>
|
|
on the scatterplot of the first two canonical variates (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scatterplot of the first two canonical variates obtained from the discriminant analysis of four measurements of the female pronotum." figureDoi="10.3897/subtbiol.43.80969.figure3" httpUri="https://binary.pensoft.net/fig/690342" pageId="0" pageNumber="61">3</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/subtbiol.43.80969.figure3" httpUri="https://binary.pensoft.net/fig/690342" pageId="0" pageNumber="61" start="Figure 3" startId="F3">
|
|
<paragraph pageId="0" pageNumber="61">
|
|
<emphasis bold="true" pageId="0" pageNumber="61">Figure 3.</emphasis>
|
|
Scatterplot of the first two canonical variates obtained from the discriminant analysis of four measurements of the female pronotum.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |