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<document id="6B3498D033CF341A3250D41461246EA8" ID-CLB-Dataset="5367" ID-DOI="10.11646/zootaxa.3811.4.7" ID-GBIF-Dataset="f37862dc-d3d8-422d-8c39-debbbf81eb0c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4919029" ID-ZooBank="C550F505-5BE1-422D-A196-1081B4CB6C04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623268316781" checkinUser="felipe" docAuthor="Shepard, William D. &amp; Barr, Cheryl B." docDate="2014" docId="8C3587E0FF88AF00638FAD8FFDC5A1AF" docLanguage="en" docName="zt03811p568.pdf" docOrigin="Zootaxa 3811 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neoeubria inbionis Shepard &amp; Barr 2014, sp. n." docType="treatment" docVersion="7" lastPageNumber="567" masterDocId="700CFF98FF83AF0E6318A93EFF96A750" masterDocTitle="Neoeubria inbionis Shepard &amp; Barr, a new genus and new species of Neotropical water penny beetle (Coleoptera: Psephenidae: Eubriinae), with a key to the adult Eubriinae of the Neotropic Zone" masterLastPageNumber="568" masterPageNumber="553" pageNumber="564" updateTime="1699007713493" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="44FCB3558E8E414289B8CFEDEB7B3D06" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="900B465FB987CB1B67279FD17FE84C0C">
<mods:title id="670B941BAA7E2730E161226FCAF913A8">Neoeubria inbionis Shepard &amp; Barr, a new genus and new species of Neotropical water penny beetle (Coleoptera: Psephenidae: Eubriinae), with a key to the adult Eubriinae of the Neotropic Zone</mods:title>
</mods:titleInfo>
<mods:name id="3F8D899D01B9A87E27B7B62DA4894E3A" type="personal">
<mods:role id="6A984A084C5E4A3CFA4C91AA1E3A5725">
<mods:roleTerm id="36FF93EA8F2B263FE405158C8DE51054">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F6157799F60A009BB54DF88660365F4A">Shepard, William D.</mods:namePart>
<mods:nameIdentifier id="EA762B8DBE70124BBA0BA8B80CEDF6C4" type="email">william.shepard@csus.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="9800A9F79399E4F2488CBFC47B8A0E15" type="personal">
<mods:role id="033E29259CF8820F9230F141844FDBAC">
<mods:roleTerm id="2DC76E59707C702C12196D71FED72482">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8474713CF37FF8DB913A4C46FCC4F654">Barr, Cheryl B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0357C408F678FA9D7A75373F6432AEE3">text</mods:typeOfResource>
<mods:relatedItem id="8AD5BB0200CE19FA24B032981AECC6A9" type="host">
<mods:titleInfo id="40675C0853687D5ECC0E18C158AA71B8">
<mods:title id="DCC1B523A1C85F972EB2154A83B90DB3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7879576C62DEACF224B52E8630E38010">
<mods:date id="BF6E59CBE3D68F631643B7A67C771C19">2014</mods:date>
<mods:detail id="728075417892462FE5FC571FF7C26464" type="pubDate">
<mods:number id="C65582CD3E9DE601C10A277BD362FFA0">2014-06-05</mods:number>
</mods:detail>
<mods:detail id="7D4DE129E8ABA66B49965B04C4328503" type="volume">
<mods:number id="91D20EE4F2133233BB84B4D95DD258B5">3811</mods:number>
</mods:detail>
<mods:detail id="E5A613936ACF07EEFEA11C6242F6ACF9" type="issue">
<mods:number id="B1F1292545DC58CE14AC8A3298FEE0FA">4</mods:number>
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<mods:start id="2279838CA703FA6EB217C5D43A48EBD2">553</mods:start>
<mods:end id="904AB280EBD6F912BB187EBE4757F3DE">568</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="152595B1E10633F3218035D2BC2F69F8">journal article</mods:classification>
<mods:identifier id="C491FCE5C3181D9E14EA553109586404" type="CLB-Dataset">5367</mods:identifier>
<mods:identifier id="FF1130A0EF34B2C02CD1174F34C13915" type="DOI">10.11646/zootaxa.3811.4.7</mods:identifier>
<mods:identifier id="E27A95816F217406AE3F7ADBC6AD9F5E" type="GBIF-Dataset">f37862dc-d3d8-422d-8c39-debbbf81eb0c</mods:identifier>
<mods:identifier id="1E5FF8D5235F95BAB5957A2702B3C85A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F1936797C0A15D251034F97D999CA263" type="Zenodo-Dep">4919029</mods:identifier>
<mods:identifier id="ED0A975D9EE3D4129CE52A011E005B22" type="ZooBank">C550F505-5BE1-422D-A196-1081B4CB6C04</mods:identifier>
</mods:mods>
<treatment id="8C3587E0FF88AF00638FAD8FFDC5A1AF" ID-DOI="http://doi.org/10.5281/zenodo.5116331" ID-GBIF-Taxon="183709127" ID-Zenodo-Dep="5116331" LSID="urn:lsid:plazi:treatment:8C3587E0FF88AF00638FAD8FFDC5A1AF" httpUri="http://treatment.plazi.org/id/8C3587E0FF88AF00638FAD8FFDC5A1AF" lastPageId="14" lastPageNumber="567" pageId="11" pageNumber="564">
<subSubSection id="4C86657DFF88AF05638FAD8FFE5EA39B" box="[151,456,1201,1227]" pageId="11" pageNumber="564" type="nomenclature">
<paragraph id="042336F6FF88AF05638FAD8FFE5EA39B" blockId="11.[151,456,1201,1261]" box="[151,456,1201,1227]" pageId="11" pageNumber="564">
<heading id="5F6B819AFF88AF05638FAD8FFE5EA39B" bold="true" box="[151,456,1201,1227]" fontSize="11" level="1" pageId="11" pageNumber="564" reason="1">
<emphasis id="36E8EAE4FF88AF05638FAD8FFE5EA39B" bold="true" box="[151,456,1201,1227]" pageId="11" pageNumber="564">
<taxonomicName id="C39C4D75FF88AF05638FAD8FFEE9A39B" authority="Shepard &amp; Barr, 2014" authorityName="Shepard &amp; Barr" authorityYear="2014" box="[151,383,1201,1227]" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="564" phylum="Arthropoda" rank="species" species="inbionis" status="sp. nov.">
<emphasis id="36E8EAE4FF88AF05638FAD8FFEE9A39B" bold="true" box="[151,383,1201,1227]" italics="true" pageId="11" pageNumber="564">Neoeubria inbionis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2DDB579FFF88AF05629EAD8FFE5EA39B" box="[390,456,1201,1227]" pageId="11" pageNumber="564" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4C86657DFF88AF05638FADEAFEC2A3BD" box="[151,340,1236,1261]" pageId="11" pageNumber="564" type="description">
<paragraph id="042336F6FF88AF05638FADEAFEC2A3BD" blockId="11.[151,456,1201,1261]" box="[151,340,1236,1261]" pageId="11" pageNumber="564">
<figureCitation id="9CA72A73FF88AF05638FADEAFE97A3BD" box="[151,257,1236,1261]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1967,1989]" captionStartId-1="2.[151,250,1202,1224]" captionStartId-2="3.[151,250,1332,1354]" captionStartId-3="4.[151,250,948,970]" captionTargetId-0="figure-271@1.[409,1177,980,1943]" captionTargetId-1="figure-346@2.[409,1177,193,1181]" captionTargetId-2="figure-336@3.[307,1267,249,1324]" captionTargetId-3="figure-43@4.[335,1264,165,932]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Neoeubria inbionis, adult female, dorsal habitus." captionText-1="FIGURE 2. Neoeubria inbionis, adult female, ventral habitus." captionText-2="FIGURE 3. Neoeubria inbionis, adult male, frontal view." captionText-3="FIGURE 4. Neoeubria inbionis, adult female, frontal view." figureDoi-0="http://doi.org/10.5281/zenodo.5116305" figureDoi-1="http://doi.org/10.5281/zenodo.5116309" figureDoi-2="http://doi.org/10.5281/zenodo.5116311" figureDoi-3="http://doi.org/10.5281/zenodo.5116313" httpUri-0="https://zenodo.org/record/5116305/files/figure.png" httpUri-1="https://zenodo.org/record/5116309/files/figure.png" httpUri-2="https://zenodo.org/record/5116311/files/figure.png" httpUri-3="https://zenodo.org/record/5116313/files/figure.png" pageId="11" pageNumber="564">Figs. 1–4</figureCitation>
,
<figureCitation id="9CA72A73FF88AF056215ADEAFEC2A3BD" box="[269,340,1236,1261]" captionStart-0="FIGURE 25" captionStart-1="FIGURE 26" captionStart-2="FIGURE 27" captionStartId-0="12.[151,250,1652,1674]" captionStartId-1="13.[151,250,1721,1743]" captionStartId-2="14.[151,250,985,1007]" captionTargetBox-0="[315,1279,349,1618]" captionTargetBox-2="[281,1305,195,963]" captionTargetId-0="figure-233@12.[296,1291,337,1638]" captionTargetId-1="figure-201@13.[367,1220,289,1714]" captionTargetId-2="figure-438@14.[281,1305,195,963]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURE 25. Neoeubria inbionis, aedeagus, A— dorsal view, B—left lateral view." captionText-1="FIGURE 26. Neoeubria inbionis, ovipositor, A—dorsal view, B—left lateral view." captionText-2="FIGURE 27. Shallow, seep pool with waterlogged wood at Parque Nacional Rincón de la Vieja, type locality of Neoeubria inbionis." figureDoi-0="http://doi.org/10.5281/zenodo.4919047" figureDoi-1="http://doi.org/10.5281/zenodo.5116317" figureDoi-2="http://doi.org/10.5281/zenodo.4919049" httpUri-0="https://zenodo.org/record/4919047/files/figure.png" httpUri-1="https://zenodo.org/record/5116317/files/figure.png" httpUri-2="https://zenodo.org/record/4919049/files/figure.png" pageId="11" pageNumber="564">25–27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4C86657DFF88AF02638FAC22FE33A0BF" lastPageId="12" lastPageNumber="565" pageId="11" pageNumber="564" type="materials_examined">
<paragraph id="042336F6FF88AF05638FAC22FCF6A105" blockId="11.[151,1437,1308,1945]" pageId="11" pageNumber="564">
<emphasis id="36E8EAE4FF88AF05638FAC22FEDEA265" bold="true" box="[151,328,1308,1333]" inLineHeading="true" pageId="11" pageNumber="564" reason="1">Type material.</emphasis>
<materialsCitation id="B4F43CABFF88AF05624CAC22FA0AA209" ID-GBIF-Occurrence="3325438305" collectingDate="2001-06-14" collectorName="William D. Shepard" country="Costa Rica" county="Parque Nacional Rincon de la Vieja" location="Las Pailas Trail" municipality="Parque Nacional Rincon de la Vieja" pageId="11" pageNumber="564" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<emphasis id="36E8EAE4FF88AF05624CAC22FE2DA265" box="[340,443,1308,1333]" italics="true" pageId="11" pageNumber="564">
<typeStatus id="DB278854FF88AF05624CAC22FE2DA265" box="[340,443,1308,1333]" pageId="11" pageNumber="564" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<emphasis id="36E8EAE4FF88AF056139AC22FD44A265" bold="true" box="[545,722,1308,1333]" pageId="11" pageNumber="564">
<collectingCountry id="7C8B7666FF88AF056139AC22FD44A265" box="[545,722,1308,1333]" name="Costa Rica" pageId="11" pageNumber="564">COSTA RICA</collectingCountry>
</emphasis>
:
<collectingRegion id="C658F814FF88AF0561FDAC22FC38A264" box="[741,942,1308,1333]" country="Costa Rica" name="Guanacaste" pageId="11" pageNumber="564">Guanacaste Prov.</collectingRegion>
,
<collectingMunicipality id="E447AC8CFF88AF0560A5AC23FAF3A265" box="[957,1381,1308,1333]" pageId="11" pageNumber="564">
<collectingCounty id="ED424E7AFF88AF0560A5AC23FAF3A265" box="[957,1381,1308,1333]" pageId="11" pageNumber="564">Parque Nacional Rincón de la Vieja</collectingCounty>
</collectingMunicipality>
,
<location id="0143602DFF88AF05666CAC23FE89A209" LSID="urn:lsid:plazi:treatment:8C3587E0FF88AF00638FAD8FFDC5A1AF:0143602DFF88AF05666CAC23FE89A209" country="Costa Rica" county="Parque Nacional Rincon de la Vieja" municipality="Parque Nacional Rincon de la Vieja" name="Las Pailas Trail" pageId="11" pageNumber="564" stateProvince="Guanacaste">Las Pailas Trail</location>
,
<date id="70221036FF88AF056229AC7EFE2EA209" box="[305,440,1344,1369]" pageId="11" pageNumber="564" value="2001-06-14">
<collectingDate id="6066E9DEFF88AF056229AC7EFE2EA209" box="[305,440,1344,1369]" pageId="11" pageNumber="564" value="2001-06-14">14-VI-2001</collectingDate>
</date>
,
<collectorName id="A9695320FF88AF0562D2AC7EFD2FA209" box="[458,697,1344,1369]" pageId="11" pageNumber="564">William D. Shepard</collectorName>
, leg. // reared from pupa collected on wood in seep basin //
</materialsCitation>
<materialsCitation id="B4F43CABFF88AF05638FAC5AFBA7A22D" ID-GBIF-Occurrence="3325438306" box="[151,1073,1380,1405]" collectionCode="INBC" collectorName="Deposited" country="Costa Rica" pageId="11" pageNumber="564" specimenCount="1" typeStatus="holotype">
<typeStatus id="DB278854FF88AF05638FAC5AFEBDA22D" box="[151,299,1380,1405]" pageId="11" pageNumber="564" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="C39C4D75FF88AF05622BAC5AFD50A22C" authority="Shepard &amp; Barr" authorityName="Shepard &amp; Barr" box="[307,710,1380,1405]" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="564" phylum="Arthropoda" rank="species" species="inbionis">Neoeubria inbionis Shepard &amp; Barr</taxonomicName>
[red label].
<collectorName id="A9695320FF88AF05604BAC5AFC53A22D" box="[851,965,1380,1405]" pageId="11" pageNumber="564">Deposited</collectorName>
in
<collectionCode id="628DAE33FF88AF0560F2AC5AFBA7A22D" box="[1002,1073,1380,1405]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="11" pageNumber="564">INBC</collectionCode>
</materialsCitation>
.
<materialsCitation id="B4F43CABFF88AF056724AC5AFEF8A2F1" ID-GBIF-Occurrence="3325438302" collectionCode="INBC" collectorName="Deposited" country="Costa Rica" pageId="11" pageNumber="564" specimenCount="1" typeStatus="allotype">
<emphasis id="36E8EAE4FF88AF056724AC5AFB0FA22D" box="[1084,1177,1380,1405]" italics="true" pageId="11" pageNumber="564">
<typeStatus id="DB278854FF88AF056724AC5AFB0FA22D" box="[1084,1177,1380,1405]" pageId="11" pageNumber="564" type="allotype">Allotype</typeStatus>
</emphasis>
(female): locality data same as holotype //
</materialsCitation>
<materialsCitation id="B4F43CABFF88AF05626CACB6FA95A2F1" ID-GBIF-Occurrence="3325438303" box="[372,1283,1416,1441]" collectionCode="INBC" collectorName="Deposited" country="Costa Rica" pageId="11" pageNumber="564" specimenCount="1" typeStatus="allotype">
<typeStatus id="DB278854FF88AF05626CACB6FD93A2F1" box="[372,517,1416,1441]" pageId="11" pageNumber="564" type="allotype">ALLOTYPE</typeStatus>
<taxonomicName id="C39C4D75FF88AF056114ACB6FC0FA2F0" authority="Shepard &amp; Barr" authorityName="Shepard &amp; Barr" box="[524,921,1416,1441]" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="564" phylum="Arthropoda" rank="species" species="inbionis">Neoeubria inbionis Shepard &amp; Barr</taxonomicName>
[red label].
<collectorName id="A9695320FF88AF05673FACB6FB0FA2F1" box="[1063,1177,1416,1441]" pageId="11" pageNumber="564">Deposited</collectorName>
in
<collectionCode id="628DAE33FF88AF0567A4ACB6FA95A2F1" box="[1212,1283,1416,1441]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="11" pageNumber="564">INBC</collectionCode>
</materialsCitation>
.
<materialsCitation id="B4F43CABFF88AF056614ACB7FB84A2B9" ID-GBIF-Occurrence="3325438307" collectingDate="1989-12-15" collectionCode="CALIFORNIA, STATE, COLLN, AGRICULTURE" collectorName="JS Wasbauer, H &amp; Real" country="Ecuador" county="Huahua Sumaco" location="Hollin-Loreto Rd." municipality="Km" pageId="11" pageNumber="564" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
<emphasis id="36E8EAE4FF88AF056614ACB7FAEBA2F0" box="[1292,1405,1417,1440]" italics="true" pageId="11" pageNumber="564">
<typeStatus id="DB278854FF88AF056614ACB7FAEBA2F0" box="[1292,1405,1417,1440]" pageId="11" pageNumber="564" type="paratype">Paratypes</typeStatus>
</emphasis>
(2 M &amp; 5 F):
<emphasis id="36E8EAE4FF88AF05620EAC92FE35A295" bold="true" box="[278,419,1452,1477]" pageId="11" pageNumber="564">
<collectingCountry id="7C8B7666FF88AF05620EAC92FE35A295" box="[278,419,1452,1477]" name="Ecuador" pageId="11" pageNumber="564">ECUADOR</collectingCountry>
</emphasis>
:
<collectingRegion id="C658F814FF88AF0562ACAC93FDA0A294" box="[436,566,1453,1476]" country="Ecuador" name="Napo" pageId="11" pageNumber="564">Napo Prov.</collectingRegion>
,
<collectingCounty id="ED424E7AFF88AF05615BAC92FD69A295" box="[579,767,1452,1477]" pageId="11" pageNumber="564">Huahua Sumaco</collectingCounty>
,
<collectingMunicipality id="E447AC8CFF88AF056014AC93FCA0A294" box="[780,822,1453,1476]" pageId="11" pageNumber="564">Km</collectingMunicipality>
44 on
<location id="0143602DFF88AF056090AC92FBC0A295" LSID="urn:lsid:plazi:treatment:8C3587E0FF88AF00638FAD8FFDC5A1AF:0143602DFF88AF056090AC92FBC0A295" box="[904,1110,1452,1477]" country="Ecuador" county="Huahua Sumaco" municipality="Km" name="Hollin-Loreto Rd." pageId="11" pageNumber="564" stateProvince="Napo">Hollin-Loreto Rd.</location>
,
<date id="70221036FF88AF05677BAC92FB64A295" box="[1123,1266,1452,1477]" pageId="11" pageNumber="564" value="1989-12-15">
<collectingDate id="6066E9DEFF88AF05677BAC92FB64A295" box="[1123,1266,1452,1477]" pageId="11" pageNumber="564" value="1989-12-15">XII-15-1989</collectingDate>
</date>
,
<collectingMethod id="DDDD4EE1FF88AF056618AC92FA0EA294" box="[1280,1432,1452,1477]" pageId="11" pageNumber="564">Malaise Trap</collectingMethod>
, MS/
<collectorName id="A9695320FF88AF0563D0ACEEFEEBA2B8" box="[200,381,1488,1513]" pageId="11" pageNumber="564">JS Wasbauer, H</collectorName>
.
<collectorName id="A9695320FF88AF056291ACEEFE2AA2B9" box="[393,444,1488,1513]" pageId="11" pageNumber="564">Real</collectorName>
//
<collectionCode id="628DAE33FF88AF0562C5ACEEFB84A2B9" box="[477,1042,1488,1513]" pageId="11" pageNumber="564">CALIFORNIA STATE COLLN AGRICULTURE</collectionCode>
</materialsCitation>
//
<materialsCitation id="B4F43CABFF88AF056729ACEEFCCAA105" ID-GBIF-Occurrence="3325438304" collectingDate="1989-12-16" collectingDateMax="1989-12-22" collectingDateMin="1989-12-16" collectionCode="EMEC, CSCA, FF" country="Ecuador" pageId="11" pageNumber="564" specimenCount="1" typeStatus="paratype">
<typeStatus id="DB278854FF88AF056729ACEEFB2AA2B9" box="[1073,1212,1488,1513]" pageId="11" pageNumber="564" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="C39C4D75FF88AF0567DDACEEFEDAA15C" authority="Shepard &amp; Barr" authorityName="Shepard &amp; Barr" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="564" phylum="Arthropoda" rank="species" species="inbionis">Neoeubria inbionis Shepard &amp; Barr</taxonomicName>
[yellow label] (1 M) (
<collectionCode id="628DAE33FF88AF056148ACCAFD32A15D" box="[592,676,1524,1549]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="11" pageNumber="564" type="University or college">EMEC</collectionCode>
); data same, except
<date id="70221036FF88AF056095ACCAFB8CA15D" box="[909,1050,1524,1549]" pageId="11" pageNumber="564" value="1989-12-16">
<collectingDate id="6066E9DEFF88AF056095ACCAFB8CA15D" box="[909,1050,1524,1549]" pageId="11" pageNumber="564" value="1989-12-16">XII-16-1989</collectingDate>
</date>
(1 F) (
<collectionCode id="628DAE33FF88AF056775ACCAFB2AA15D" box="[1133,1212,1524,1549]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="11" pageNumber="564" type="Museum">CSCA</collectionCode>
); data same, except
<date id="70221036FF88AF05638FAF26FEB2A161" box="[151,292,1560,1585]" pageId="11" pageNumber="564" value="1989-12-18">
<collectingDate id="6066E9DEFF88AF05638FAF26FEB2A161" box="[151,292,1560,1585]" pageId="11" pageNumber="564" value="1989-12-18">XII-18-1989</collectingDate>
</date>
(1 M) (
<collectionCode id="628DAE33FF88AF056265AF26FE5AA161" box="[381,460,1560,1585]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="11" pageNumber="564" type="Museum">CSCA</collectionCode>
); data same, except
<date id="70221036FF88AF0561A8AF26FCABA161" box="[688,829,1560,1585]" pageId="11" pageNumber="564" value="1989-12-19">
<collectingDate id="6066E9DEFF88AF0561A8AF26FCABA161" box="[688,829,1560,1585]" pageId="11" pageNumber="564" value="1989-12-19">XII-19-1989</collectingDate>
</date>
(1 F) (
<collectionCode id="628DAE33FF88AF056094AF26FC4DA161" box="[908,987,1560,1585]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="11" pageNumber="564" type="Museum">CSCA</collectionCode>
); data same, except
<date id="70221036FF88AF0567A7AF26FADBA161" box="[1215,1357,1560,1585]" pageId="11" pageNumber="564" value="1989-12-21">
<collectingDate id="6066E9DEFF88AF0567A7AF26FADBA161" box="[1215,1357,1560,1585]" pageId="11" pageNumber="564" value="1989-12-21">XII-21-1989</collectingDate>
</date>
(2
<collectionCode id="628DAE33FF88AF05666CAF26FA03A161" box="[1396,1429,1560,1585]" pageId="11" pageNumber="564">FF</collectionCode>
) (
<collectionCode id="628DAE33FF88AF056387AF02FF7AA105" box="[159,236,1596,1621]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="11" pageNumber="564" type="Museum">CSCA</collectionCode>
,
<collectionCode id="628DAE33FF88AF0563EFAF02FEDFA105" box="[247,329,1596,1621]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="11" pageNumber="564" type="University or college">EMEC</collectionCode>
); data same, except
<date id="70221036FF88AF056132AF02FD2EA105" box="[554,696,1596,1621]" pageId="11" pageNumber="564" value="1989-12-22">
<collectingDate id="6066E9DEFF88AF056132AF02FD2EA105" box="[554,696,1596,1621]" pageId="11" pageNumber="564" value="1989-12-22">XII-22-1989</collectingDate>
</date>
(1 F) (
<collectionCode id="628DAE33FF88AF05601DAF02FCC2A105" box="[773,852,1596,1621]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="11" pageNumber="564" type="Museum">CSCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="042336F6FF88AF0563DFAF5EFB33A058" blockId="11.[151,1437,1308,1945]" pageId="11" pageNumber="564">
<emphasis id="36E8EAE4FF88AF0563DFAF5EFE32A129" bold="true" box="[199,420,1632,1657]" pageId="11" pageNumber="564">Adult Description.</emphasis>
Body oval; males (
<figureCitation id="9CA72A73FF88AF056166AF5EFD56A129" box="[638,704,1632,1657]" captionStart="FIGURE 3" captionStartId="3.[151,250,1332,1354]" captionTargetId="figure-336@3.[307,1267,249,1324]" captionTargetPageId="3" captionText="FIGURE 3. Neoeubria inbionis, adult male, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5116311" httpUri="https://zenodo.org/record/5116311/files/figure.png" pageId="11" pageNumber="564">Fig. 3</figureCitation>
) smaller than females (
<figureCitation id="9CA72A73FF88AF0560D0AF5EFBA5A129" box="[968,1075,1632,1657]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,250,1967,1989]" captionStartId-1="2.[151,250,1202,1224]" captionTargetId-0="figure-271@1.[409,1177,980,1943]" captionTargetId-1="figure-346@2.[409,1177,193,1181]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Neoeubria inbionis, adult female, dorsal habitus." captionText-1="FIGURE 2. Neoeubria inbionis, adult female, ventral habitus." figureDoi-0="http://doi.org/10.5281/zenodo.5116305" figureDoi-1="http://doi.org/10.5281/zenodo.5116309" httpUri-0="https://zenodo.org/record/5116305/files/figure.png" httpUri-1="https://zenodo.org/record/5116309/files/figure.png" pageId="11" pageNumber="564">Figs. 1–2</figureCitation>
,
<figureCitation id="9CA72A73FF88AF056727AF5EFBC6A129" box="[1087,1104,1632,1657]" captionStart="FIGURE 4" captionStartId="4.[151,250,948,970]" captionTargetId="figure-43@4.[335,1264,165,932]" captionTargetPageId="4" captionText="FIGURE 4. Neoeubria inbionis, adult female, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5116313" httpUri="https://zenodo.org/record/5116313/files/figure.png" pageId="11" pageNumber="564">4</figureCitation>
); males 4.6–5.0 mm long and
<quantity id="C3649B13FF88AF05638FAFBAFF6BA1CC" box="[151,253,1668,1693]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="11" pageNumber="564" unit="mm" value="2.75">2.75 mm</quantity>
wide, females 5.0–
<quantity id="C3649B13FF88AF0562F8AFBAFDAEA1CC" box="[480,568,1668,1693]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="11" pageNumber="564" unit="mm" value="5.6">5.6 mm</quantity>
long and
<quantity id="C3649B13FF88AF0561B6AFBAFCAEA1CC" box="[686,824,1668,1693]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="11" pageNumber="564" unit="mm" value="3.15" valueMax="3.5" valueMin="2.8">2.8–3.5 mm</quantity>
wide. Integument color medium brown, shiny where setae sparse; covered dorsally with a combination of different kinds of setae: widely-spaced, long, erect blond and dark brown setae; sparse, shorter, pale brown setae; and very dense, recumbent, pale blond setae forming a pattern of broad bands and large spots. Venter uniformly clothed in medium-length pale blond setae.
</paragraph>
<paragraph id="042336F6FF88AF0563DFAE2AFBA9A0C9" blockId="11.[151,1437,1308,1945]" pageId="11" pageNumber="564">
Aedeagus of trilobed
<typeStatus id="DB278854FF88AF0562A5AE2BFE7BA07D" box="[445,493,1813,1837]" pageId="11" pageNumber="564">type</typeStatus>
(
<figureCitation id="9CA72A73FF88AF0562E6AE2AFDC7A07D" box="[510,593,1812,1837]" captionStart="FIGURE 25" captionStartId="12.[151,250,1652,1674]" captionTargetBox="[315,1279,349,1618]" captionTargetId="figure-233@12.[296,1291,337,1638]" captionTargetPageId="12" captionText="FIGURE 25. Neoeubria inbionis, aedeagus, A— dorsal view, B—left lateral view." figureDoi="http://doi.org/10.5281/zenodo.4919047" httpUri="https://zenodo.org/record/4919047/files/figure.png" pageId="11" pageNumber="564">Fig. 25</figureCitation>
) and lightly sclerotized. Basal piece long, reduced to ventral plate basally with lateral flanges that clasp the base of parameres. Parameres long, widest at apical three-fourths of aedeagus; tips narrow, curved laterally; dorsally conjoined just anterior to midlength. Penis lanceolate; shorter than parameres; tip slightly curved ventrally and laterally compressed; base deeply cleft.
</paragraph>
<footnote id="67872AF8FF88AF05638FAEE5FA9EA0A1" box="[151,1288,2011,2033]" pageId="11" pageNumber="564">
<paragraph id="042336F6FF88AF05638FAEE5FA9EA0A1" blockId="11.[151,1288,2011,2033]" box="[151,1288,2011,2033]" pageId="11" pageNumber="564">
1.
<taxonomicName id="C39C4D75FF88AF0563D5AEE2FEB0A0A1" authorityName="LeConte" authorityYear="1853" box="[205,294,2012,2033]" class="Insecta" family="Psephenidae" genus="Ectopria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="564" phylum="Arthropoda" rank="genus">
<emphasis id="36E8EAE4FF88AF0563D5AEE2FEB0A0A1" box="[205,294,2012,2033]" italics="true" pageId="11" pageNumber="564">Ectopria</emphasis>
</taxonomicName>
is omitted from the key because it is probable that the Neotropical species belong in other genera.
</paragraph>
</footnote>
<paragraph id="042336F6FF8FAF0263DFA9A9FE59A64D" blockId="12.[151,1436,151,285]" pageId="12" pageNumber="565">
Ovipositor (
<figureCitation id="9CA72A73FF8FAF02624DA9A9FE3BA7E0" box="[341,429,151,176]" captionStart="FIGURE 26" captionStartId="13.[151,250,1721,1743]" captionTargetId="figure-201@13.[367,1220,289,1714]" captionTargetPageId="13" captionText="FIGURE 26. Neoeubria inbionis, ovipositor, A—dorsal view, B—left lateral view." figureDoi="http://doi.org/10.5281/zenodo.5116317" httpUri="https://zenodo.org/record/5116317/files/figure.png" pageId="12" pageNumber="565">Fig. 26</figureCitation>
) with bacula long, 1.4 times as long as coxites, thin, gently curved; only partially sclerotized. Coxites 0.7 times as long as bacula; joined medially in basal half, divergent medially in apical half; laterally gently sinuate. Styli short, one-segmented. Long, thin accessory sclerite dorsally in basal third of membrane between coxites.
</paragraph>
<caption id="50E3667EFF8FAF02638FAF4AFC49A1DA" ID-DOI="http://doi.org/10.5281/zenodo.4919047" ID-Zenodo-Dep="4919047" box="[151,991,1652,1674]" httpUri="https://zenodo.org/record/4919047/files/figure.png" pageId="12" pageNumber="565" startId="12.[151,250,1652,1674]" targetBox="[315,1279,349,1618]" targetPageId="12">
<paragraph id="042336F6FF8FAF02638FAF4AFC49A1DA" blockId="12.[151,991,1652,1674]" box="[151,991,1652,1674]" pageId="12" pageNumber="565">
<emphasis id="36E8EAE4FF8FAF02638FAF4AFE89A1DA" bold="true" box="[151,287,1652,1674]" pageId="12" pageNumber="565">FIGURE 25.</emphasis>
<taxonomicName id="C39C4D75FF8FAF02623DAF4AFE7FA1DA" box="[293,489,1652,1674]" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="565" phylum="Arthropoda" rank="species" species="inbionis">
<emphasis id="36E8EAE4FF8FAF02623DAF4AFE7FA1DA" box="[293,489,1652,1674]" italics="true" pageId="12" pageNumber="565">Neoeubria inbionis</emphasis>
</taxonomicName>
, aedeagus, A— dorsal view, B—left lateral view.
</paragraph>
</caption>
<paragraph id="042336F6FF8FAF0263DFAF88FE33A0BF" blockId="12.[151,1436,1718,2031]" pageId="12" pageNumber="565">
Immature specimens examined.
<materialsCitation id="B4F43CABFF8FAF026153AF88FD09A1A3" ID-GBIF-Occurrence="3325438301" collectingDate="2000-06-09" collectionCode="VI" collectorName="William D. Shepard" country="Costa Rica" location="Rio San Lorenzo" municipality="Alta Masis" pageId="12" pageNumber="565" specimenCount="1" specimenCount-larva="1" stateProvince="Alajuela">
<emphasis id="36E8EAE4FF8FAF026153AF88FC94A19F" bold="true" box="[587,770,1718,1743]" pageId="12" pageNumber="565">
<collectingCountry id="7C8B7666FF8FAF026153AF88FD6BA19F" box="[587,765,1718,1743]" name="Costa Rica" pageId="12" pageNumber="565">COSTA RICA</collectingCountry>
:
</emphasis>
<collectingRegion id="C658F814FF8FAF026012AF88FCFAA19F" box="[778,876,1718,1743]" country="Costa Rica" name="Alajuela" pageId="12" pageNumber="565">Alajuela</collectingRegion>
,
<collectingMunicipality id="E447AC8CFF8FAF026060AF88FC60A19F" box="[888,1014,1718,1743]" pageId="12" pageNumber="565">Alta Masis</collectingMunicipality>
,
<date id="70221036FF8FAF02671AAF88FBEFA19F" box="[1026,1145,1718,1743]" pageId="12" pageNumber="565" value="2000-06-09">
<collectingDate id="6066E9DEFF8FAF02671AAF88FBEFA19F" box="[1026,1145,1718,1743]" pageId="12" pageNumber="565" value="2000-06-09">
9
<collectionCode id="628DAE33FF8FAF026700AF89FBA0A19E" box="[1048,1078,1719,1742]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="565">VI</collectionCode>
2000
</collectingDate>
</date>
,
<location id="0143602DFF8FAF02679DAF89FAD5A19E" LSID="urn:lsid:plazi:treatment:8C3587E0FF88AF00638FAD8FFDC5A1AF:0143602DFF8FAF02679DAF89FAD5A19E" box="[1157,1347,1718,1743]" country="Costa Rica" municipality="Alta Masis" name="Rio San Lorenzo" pageId="12" pageNumber="565" stateProvince="Alajuela">Río San Lorenzo</location>
[WDS- A-1302] //
<collectorName id="A9695320FF8FAF02620EAFE4FE6AA1A3" box="[278,508,1754,1779]" pageId="12" pageNumber="565">William D. Shepard</collectorName>
, leg. (
<specimenCount id="129AFD7FFF8FAF02615EAFE4FD0EA1A3" box="[582,664,1754,1779]" pageId="12" pageNumber="565" type="larva">1 larva</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="B4F43CABFF8FAF0261B5AFE4FF67A06B" ID-GBIF-Occurrence="3325438310" collectingDate="2000-01-18" country="Costa Rica" location="Las Pailas Trail" pageId="12" pageNumber="565" specimenCount="25" specimenCount-larva="24" specimenCount-pupa="1" stateProvince="Guanacaste">
<collectingRegion id="C658F814FF8FAF0261B5AFE4FCE2A1A2" box="[685,884,1754,1779]" country="Costa Rica" name="Guanacaste" pageId="12" pageNumber="565">Guanacaste Prov.</collectingRegion>
,
<collectorName id="A9695320FF8FAF026098AFE5FA8AA1A3" box="[896,1308,1754,1779]" pageId="12" pageNumber="565">Parque Nacional Rincón de la Vieja</collectorName>
,
<location id="0143602DFF8FAF026630AFE5FF59A047" LSID="urn:lsid:plazi:treatment:8C3587E0FF88AF00638FAD8FFDC5A1AF:0143602DFF8FAF026630AFE5FF59A047" country="Costa Rica" name="Las Pailas Trail" pageId="12" pageNumber="565" stateProvince="Guanacaste">Las Pailas Trail</location>
,
<date id="70221036FF8FAF0263C2AFC0FEDBA047" box="[218,333,1790,1815]" pageId="12" pageNumber="565" value="2000-01-18">
<collectingDate id="6066E9DEFF8FAF0263C2AFC0FEDBA047" box="[218,333,1790,1815]" pageId="12" pageNumber="565" value="2000-01-18">18-I-2000</collectingDate>
</date>
,
<collectorName id="A9695320FF8FAF02624FAFC0FE24A047" box="[343,434,1790,1815]" pageId="12" pageNumber="565">William</collectorName>
D.
<collectorName id="A9695320FF8FAF0262C3AFC0FDA1A047" box="[475,567,1790,1815]" pageId="12" pageNumber="565">Shepard</collectorName>
&amp;
<collectorName id="A9695320FF8FAF026142AFC0FD30A047" box="[602,678,1790,1815]" pageId="12" pageNumber="565">Cheryl</collectorName>
B.
<collectorName id="A9695320FF8FAF0261D5AFC1FC97A046" box="[717,769,1791,1814]" pageId="12" pageNumber="565">Barr</collectorName>
, collected on wood in seep basin [WDS-A-1283](
<specimenCount id="129AFD7FFF8FAF026635AFC0FA0EA047" box="[1325,1432,1790,1815]" pageId="12" pageNumber="565" type="larva">24 larvae</specimenCount>
,
<specimenCount id="129AFD7FFF8FAF02638FAE1CFF7EA06B" box="[151,232,1826,1851]" pageId="12" pageNumber="565" type="pupa">1 pupa</specimenCount>
)
</materialsCitation>
; data same, except
<date id="70221036FF8FAF0262D7AE1CFDC0A06B" box="[463,598,1826,1851]" pageId="12" pageNumber="565" value="2001-06-14">14-VI-2001</date>
, William D. Shepard, leg. [WDS-A-1386] (
<specimenCount id="129AFD7FFF8FAF02675FAE1CFB33A06B" box="[1095,1189,1826,1851]" pageId="12" pageNumber="565" type="larva">3 larvae</specimenCount>
,
<specimenCount id="129AFD7FFF8FAF0267AAAE1CFA86A06B" box="[1202,1296,1826,1851]" pageId="12" pageNumber="565" type="pupa">3 pupae</specimenCount>
); data same, except
<date id="70221036FF8FAF0263FEAE78FEFBA00F" box="[230,365,1862,1887]" pageId="12" pageNumber="565" value="2003-06-15">15-VI-2003</date>
, William D. Shepard &amp; Cheryl B. Barr [WDS-A-1541] (
<specimenCount id="129AFD7FFF8FAF0260FDAE78FBC6A00F" box="[997,1104,1862,1887]" pageId="12" pageNumber="565" type="larva">11 larvae</specimenCount>
);
<materialsCitation id="B4F43CABFF8FAF02677BAE78FBE3A0D3" ID-GBIF-Occurrence="3325438311" collectingDate="2001-06-14" collectorName="William D. Shepard" country="Costa Rica" location="Catarata" municipality="Quebrada Pailas" pageId="12" pageNumber="565" specimenCount="1" specimenCount-larva="1" stateProvince="Guanacaste Prov.">
data same, except
<collectingMunicipality id="E447AC8CFF8FAF026637AE78FF4CA0D3" pageId="12" pageNumber="565">Quebrada Pailas</collectingMunicipality>
below
<location id="0143602DFF8FAF026228AE54FE06A0D3" LSID="urn:lsid:plazi:treatment:8C3587E0FF88AF00638FAD8FFDC5A1AF:0143602DFF8FAF026228AE54FE06A0D3" box="[304,400,1898,1923]" country="Costa Rica" municipality="Quebrada Pailas" name="Catarata" pageId="12" pageNumber="565" stateProvince="Guanacaste Prov.">Catarata</location>
,
<date id="70221036FF8FAF026286AE54FDB2A0D3" box="[414,548,1898,1923]" pageId="12" pageNumber="565" value="2001-06-14">
<collectingDate id="6066E9DEFF8FAF026286AE54FDB2A0D3" box="[414,548,1898,1923]" pageId="12" pageNumber="565" value="2001-06-14">14-VI-2001</collectingDate>
</date>
,
<collectorName id="A9695320FF8FAF026129AE54FC81A0D3" box="[561,791,1898,1923]" pageId="12" pageNumber="565">William D. Shepard</collectorName>
, leg. [WDS-A-1387] (
<specimenCount id="129AFD7FFF8FAF026703AE54FBFBA0D3" box="[1051,1133,1898,1923]" pageId="12" pageNumber="565" type="larva">1 larva</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="B4F43CABFF8FAF026799AE54FA0EA0F7" ID-GBIF-Occurrence="3325438309" collectingDate="2002-08-10" collectorName="William D. Shepard" country="Nicaragua" location="Rio Bartola" municipality="Refugio Bartola" pageId="12" pageNumber="565" specimenCount="1" specimenCount-larva="1" stateProvince="Rio San Juan">
<emphasis id="36E8EAE4FF8FAF026799AE54FAAFA0D3" bold="true" box="[1153,1337,1898,1923]" pageId="12" pageNumber="565">
<collectingCountry id="7C8B7666FF8FAF026799AE54FAA5A0D3" box="[1153,1331,1898,1923]" name="Nicaragua" pageId="12" pageNumber="565">NICARAGUA</collectingCountry>
:
</emphasis>
<collectingRegion id="C658F814FF8FAF02665AAE55FF5BA0F6" country="Nicaragua" name="Rio San Juan" pageId="12" pageNumber="565">Río San Juan</collectingRegion>
,
<collectingMunicipality id="E447AC8CFF8FAF0263C4AEB0FE00A0F7" box="[220,406,1934,1959]" pageId="12" pageNumber="565">Refugio Bartola</collectingMunicipality>
,
<date id="70221036FF8FAF0262BDAEB0FDD4A0F7" box="[421,578,1934,1959]" pageId="12" pageNumber="565" value="2002-08-10">
<collectingDate id="6066E9DEFF8FAF0262BDAEB0FDD4A0F7" box="[421,578,1934,1959]" pageId="12" pageNumber="565" value="2002-08-10">10 VIII 2002</collectingDate>
</date>
, riffle 3,
<location id="0143602DFF8FAF0261ABAEB1FCACA0F7" LSID="urn:lsid:plazi:treatment:8C3587E0FF88AF00638FAD8FFDC5A1AF:0143602DFF8FAF0261ABAEB1FCACA0F7" box="[691,826,1934,1959]" country="Nicaragua" municipality="Refugio Bartola" name="Rio Bartola" pageId="12" pageNumber="565" stateProvince="Rio San Juan">Río Bartola</location>
,
<collectorName id="A9695320FF8FAF026052AEB0FBA2A0F7" box="[842,1076,1934,1959]" pageId="12" pageNumber="565">William D. Shepard</collectorName>
, leg. [WDS-A-1492] (
<specimenCount id="129AFD7FFF8FAF026624AEB0FA07A0F7" box="[1340,1425,1934,1959]" pageId="12" pageNumber="565" type="larva">1 larva</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="B4F43CABFF8FAF02638FAE8CFB17A09B" ID-GBIF-Occurrence="3325438308" box="[151,1153,1969,1995]" collectingDate="2004-03" collectorName="Checo Colon-Gaud" country="Panama" location="Fortuna Forest Res." pageId="12" pageNumber="565" specimenCount="1" specimenCount-larva="1" stateProvince="Chiriqui">
<emphasis id="36E8EAE4FF8FAF02638FAE8CFE8CA09B" bold="true" box="[151,282,1970,1995]" pageId="12" pageNumber="565">
<collectingCountry id="7C8B7666FF8FAF02638FAE8CFE80A09B" box="[151,278,1970,1995]" name="Panama" pageId="12" pageNumber="565">PANAMA</collectingCountry>
:
</emphasis>
<collectingRegion id="C658F814FF8FAF026239AE8CFE17A09B" box="[289,385,1970,1995]" country="Panama" name="Chiriqui" pageId="12" pageNumber="565">Chiriquí</collectingRegion>
,
<location id="0143602DFF8FAF026293AE8DFDFFA09A" LSID="urn:lsid:plazi:treatment:8C3587E0FF88AF00638FAD8FFDC5A1AF:0143602DFF8FAF026293AE8DFDFFA09A" box="[395,617,1971,1994]" country="Panama" name="Fortuna Forest Res." pageId="12" pageNumber="565" stateProvince="Chiriqui">Fortuna Forest Res.</location>
,
<date id="70221036FF8FAF02616CAE8CFD68A09B" box="[628,766,1970,1995]" pageId="12" pageNumber="565" value="2004-03">
<collectingDate id="6066E9DEFF8FAF02616CAE8CFD68A09B" box="[628,766,1970,1995]" pageId="12" pageNumber="565" value="2004-03">March 2004</collectingDate>
</date>
,
<collectorName id="A9695320FF8FAF026010AE8CFC73A09B" box="[776,997,1969,1995]" pageId="12" pageNumber="565">Checo Colón-Gaud</collectorName>
, leg. (
<specimenCount id="129AFD7FFF8FAF026732AE8CFBEFA09B" box="[1066,1145,1970,1995]" pageId="12" pageNumber="565" type="larva">1 larva</specimenCount>
)
</materialsCitation>
. All immature specimens are deposited in
<collectionCode id="628DAE33FF8FAF026257AEE8FE36A0BF" box="[335,416,2006,2031]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="12" pageNumber="565" type="University or college">EMEC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="4C86657DFF8EAF0363DFA9A9FF6FA785" pageId="13" pageNumber="566" type="etymology">
<paragraph id="042336F6FF8EAF0363DFA9A9FF6FA785" blockId="13.[151,1436,151,248]" pageId="13" pageNumber="566">
<emphasis id="36E8EAE4FF8EAF0363DFA9A9FED8A7E0" bold="true" box="[199,334,151,176]" inLineHeading="true" pageId="13" pageNumber="566" reason="1">Etymology.</emphasis>
Named in honor of INBio, the Instituto Nacional de Biodiversidad in
<collectingCountry id="7C8B7666FF8EAF036791A9A9FA9DA7E0" box="[1161,1291,151,176]" name="Costa Rica" pageId="13" pageNumber="566">Costa Rica</collectingCountry>
. The case is genitive.
</paragraph>
</subSubSection>
<subSubSection id="4C86657DFF8EAF0063DFA9E1FDC5A1AF" lastPageId="14" lastPageNumber="567" pageId="13" pageNumber="566" type="distribution">
<paragraph id="042336F6FF8EAF0363DFA9E1FA93A7A8" blockId="13.[151,1436,151,248]" box="[199,1285,223,248]" pageId="13" pageNumber="566">
<emphasis id="36E8EAE4FF8EAF0363DFA9E1FEF6A7A8" bold="true" box="[199,352,223,248]" inLineHeading="true" pageId="13" pageNumber="566" reason="1">Distribution.</emphasis>
<collectingCountry id="7C8B7666FF8EAF03627FA9E1FE4BA7A8" box="[359,477,223,248]" name="Nicaragua" pageId="13" pageNumber="566">Nicaragua</collectingCountry>
,
<collectingCountry id="7C8B7666FF8EAF0362F0A9E1FDF3A7A8" box="[488,613,223,248]" name="Costa Rica" pageId="13" pageNumber="566">Costa Rica</collectingCountry>
,
<collectingCountry id="7C8B7666FF8EAF036168A9DEFD5FA7A7" box="[624,713,224,247]" name="Panama" pageId="13" pageNumber="566">Panama</collectingCountry>
and
<collectingCountry id="7C8B7666FF8EAF0361E7A9E1FCC8A7A8" box="[767,862,223,248]" name="Ecuador" pageId="13" pageNumber="566">Ecuador</collectingCountry>
, based on adult and larval specimens.
</paragraph>
<caption id="50E3667EFF8EAF03638FAF87FC74A19F" ID-DOI="http://doi.org/10.5281/zenodo.5116317" ID-Zenodo-Dep="5116317" box="[151,994,1721,1743]" httpUri="https://zenodo.org/record/5116317/files/figure.png" pageId="13" pageNumber="566" startId="13.[151,250,1721,1743]" targetBox="[375,1214,297,1699]" targetPageId="13">
<paragraph id="042336F6FF8EAF03638FAF87FC74A19F" blockId="13.[151,994,1721,1743]" box="[151,994,1721,1743]" pageId="13" pageNumber="566">
<emphasis id="36E8EAE4FF8EAF03638FAF87FEB6A19F" bold="true" box="[151,288,1721,1743]" pageId="13" pageNumber="566">FIGURE 26.</emphasis>
<taxonomicName id="C39C4D75FF8EAF03623DAF87FE7FA19F" box="[293,489,1721,1743]" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="566" phylum="Arthropoda" rank="species" species="inbionis">
<emphasis id="36E8EAE4FF8EAF03623DAF87FE7FA19F" box="[293,489,1721,1743]" italics="true" pageId="13" pageNumber="566">Neoeubria inbionis</emphasis>
</taxonomicName>
, ovipositor, A—dorsal view, B—left lateral view.
</paragraph>
</caption>
<paragraph id="042336F6FF8EAF0363DFAFC2FDB6A0BD" blockId="13.[151,1437,1788,2029]" pageId="13" pageNumber="566">
<emphasis id="36E8EAE4FF8EAF0363DFAFC2FEBCA045" bold="true" box="[199,298,1788,1813]" inLineHeading="true" pageId="13" pageNumber="566" reason="1">Habitat.</emphasis>
The
<typeStatus id="DB278854FF8EAF03627EAFC2FE00A044" box="[358,406,1788,1812]" pageId="13" pageNumber="566">type</typeStatus>
locality in Parque Nacional Rincón de la Vieja at an
<quantity id="C3649B13FF8EAF0360E4AFC2FBD3A043" box="[1020,1093,1788,1813]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="13" pageNumber="566" unit="m" value="780.0">780 m</quantity>
is a series of seeps in a small basin connected by a spring run to a narrow, slow-flowing forest stream which is a tributary of the Río
<collectingRegion id="C658F814FF8EAF036635AE21FA0FA068" box="[1325,1433,1823,1848]" country="United States of America" name="Colorado" pageId="13" pageNumber="566">Colorado</collectingRegion>
. The entire area around the seeps and both streams is heavily forested and generally heavily shaded. In the seep basin the water is only about
<quantity id="C3649B13FF8EAF0362FBAE59FDA1A02F" box="[483,567,1895,1920]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="13" pageNumber="566" unit="cm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 cm</quantity>
deep over a substrate composed of a thick deposit of silt and fine detritus on which lie sticks and larger pieces of rotting, waterlogged wood (
<figureCitation id="9CA72A73FF8EAF036072AEB2FC29A0F5" box="[874,959,1932,1957]" captionStart="FIGURE 27" captionStartId="14.[151,250,985,1007]" captionTargetBox="[281,1305,195,963]" captionTargetId="figure-438@14.[281,1305,195,963]" captionTargetPageId="14" captionText="FIGURE 27. Shallow, seep pool with waterlogged wood at Parque Nacional Rincón de la Vieja, type locality of Neoeubria inbionis." figureDoi="http://doi.org/10.5281/zenodo.4919049" httpUri="https://zenodo.org/record/4919049/files/figure.png" pageId="13" pageNumber="566">Fig. 27</figureCitation>
). In the basin the water is extremely slowmoving but in a couple of meters it begins to flow downhill in a narrow spring run which is crossed by the Las Pailas Trail between Stops 3 and 4.
</paragraph>
<caption id="50E3667EFF8DAF00638FAAE7FF78A35D" ID-DOI="http://doi.org/10.5281/zenodo.4919049" ID-Zenodo-Dep="4919049" httpUri="https://zenodo.org/record/4919049/files/figure.png" pageId="14" pageNumber="567" startId="14.[151,250,985,1007]" targetBox="[281,1305,195,963]" targetPageId="14">
<paragraph id="042336F6FF8DAF00638FAAE7FF78A35D" blockId="14.[151,1436,984,1038]" pageId="14" pageNumber="567">
<emphasis id="36E8EAE4FF8DAF00638FAAE7FEB4A4BF" bold="true" box="[151,290,985,1007]" pageId="14" pageNumber="567">FIGURE 27.</emphasis>
Shallow, seep pool with waterlogged wood at Parque Nacional Rincón de la Vieja, type locality of
<taxonomicName id="C39C4D75FF8DAF006629AAE7FF7FA35E" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="species" species="inbionis">
<emphasis id="36E8EAE4FF8DAF006629AAE7FF7FA35E" italics="true" pageId="14" pageNumber="567">Neoeubria inbionis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="042336F6FF8DAF0063DFAD05FB4EA223" blockId="14.[151,1437,1082,1791]" pageId="14" pageNumber="567">
Larvae and pupae of
<taxonomicName id="C39C4D75FF8DAF0062DBAD05FDD4A303" box="[451,578,1082,1107]" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="species" species="inbionis">
<emphasis id="36E8EAE4FF8DAF0062DBAD05FDD4A303" box="[451,578,1082,1107]" italics="true" pageId="14" pageNumber="567">N. inbionis</emphasis>
</taxonomicName>
were collected on pieces of decomposing wood found in the seep basin. Larvae were positioned below the water’s surface and pupae were above. The water is likely hypoxic because of the fine organic detritus substrate, coupled with the lack of sunlight for aquatic photosynthesizers due to the heavily-shaded nature of the site. Possession of a plastron facilitates larval survival in this water.
<taxonomicName id="C39C4D75FF8DAF0067E9AD98FAF1A3EF" authorityName="Shepard &amp; Barr" authorityYear="2014" box="[1265,1383,1190,1215]" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="36E8EAE4FF8DAF0067E9AD98FAF1A3EF" box="[1265,1383,1190,1215]" italics="true" pageId="14" pageNumber="567">Neoeubria</emphasis>
</taxonomicName>
was the only psephenid present in the seep area, and the only other co-occurring aquatic byrrhoid
<taxonomicName id="C39C4D75FF8DAF0067D1ADF4FAD2A3B3" authorityName="Linnaeus" authorityYear="1758" box="[1225,1348,1226,1251]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
was an unidentified ptilodactylid larva. Other arthropods present in the seep area included aquatic
<taxonomicName id="C39C4D75FF8DAF0067B9ADD0FA8CA257" box="[1185,1306,1262,1287]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
,
<taxonomicName id="C39C4D75FF8DAF00663EADD0FECEA27B" class="Insecta" family="Belostomatidae" genus="Belostoma" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="36E8EAE4FF8DAF00663EADD0FA0AA257" box="[1318,1436,1262,1287]" italics="true" pageId="14" pageNumber="567">Belostoma</emphasis>
(Belostomatidae)
</taxonomicName>
and
<taxonomicName id="C39C4D75FF8DAF006280AC2CFD3CA27B" box="[408,682,1298,1323]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="36E8EAE4FF8DAF006280AC2CFD90A27B" box="[408,518,1298,1323]" italics="true" pageId="14" pageNumber="567">Ambrysus</emphasis>
(Naucoridae)
</taxonomicName>
, and the crustacean
<taxonomicName id="C39C4D75FF8DAF006087AC2CFB31A27B" authority="(Amphipoda)" baseAuthorityName="Amphipoda" box="[927,1191,1298,1323]" class="Malacostraca" family="Hyalellidae" genus="Hyallela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="14" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="36E8EAE4FF8DAF006087AC2CFB96A27B" box="[927,1024,1298,1323]" italics="true" pageId="14" pageNumber="567">Hyallela</emphasis>
(Amphipoda)
</taxonomicName>
. No specimens were collected from the spring run formed by the seeps. A single larva was collected in a second, larger stream, Quebrada Pailas, a tributary of the Río
<collectingRegion id="C658F814FF8DAF006154AC64FD21A223" box="[588,695,1370,1395]" country="United States of America" name="Colorado" pageId="14" pageNumber="567">Colorado</collectingRegion>
, which is also located along the Las Pailas Trail.
</paragraph>
<paragraph id="042336F6FF8DAF0063DFAC40FC95A153" blockId="14.[151,1437,1082,1791]" pageId="14" pageNumber="567">
The other sites at which larvae were collected by the senior author are also forest streams, although with rocky substrates and faster flow. Although the particular microhabitat of the larvae at these sites is unknown, at all of them submerged wood was common.
<taxonomicName id="C39C4D75FF8DAF00615EACF8FC88A28F" box="[582,798,1478,1503]" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="species" species="inbionis">
<emphasis id="36E8EAE4FF8DAF00615EACF8FC88A28F" box="[582,798,1478,1503]" italics="true" pageId="14" pageNumber="567">Neoeubria inbionis</emphasis>
</taxonomicName>
has been collected at elevations ranging from as low as 40+ m in
<collectingCountry id="7C8B7666FF8DAF00621BACD4FEEFA153" box="[259,377,1514,1539]" name="Nicaragua" pageId="14" pageNumber="567">Nicaragua</collectingCountry>
, to as high as
<quantity id="C3649B13FF8DAF00610FACD4FDCBA152" box="[535,605,1514,1539]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="14" pageNumber="567" unit="m" value="780.0">780 m</quantity>
in
<collectingCountry id="7C8B7666FF8DAF006199ACD4FD69A153" box="[641,767,1514,1539]" name="Costa Rica" pageId="14" pageNumber="567">Costa Rica</collectingCountry>
.
</paragraph>
<paragraph id="042336F6FF8DAF0063DFAF30FAAFA13E" blockId="14.[151,1437,1082,1791]" pageId="14" pageNumber="567">
The Ecuadorian adults were all taken in Malaise traps which were set in a forested area to catch flies. Although we could obtain no further information beyond the label data, collection of adults via Malaise traps indicates that
<taxonomicName id="C39C4D75FF8DAF00669AAF0DFF67A13F" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="species" species="inbionis">
<emphasis id="36E8EAE4FF8DAF00669AAF0DFF67A13F" italics="true" pageId="14" pageNumber="567">N. inbionis</emphasis>
</taxonomicName>
adults behave like other eubriine adults and fly near the aquatic habitat in which the larvae occur.
</paragraph>
<paragraph id="042336F6FF8DAF0063DFAF44FDC5A1AF" blockId="14.[151,1437,1082,1791]" pageId="14" pageNumber="567">
<emphasis id="36E8EAE4FF8DAF0063DFAF44FEDFA1C3" bold="true" box="[199,329,1658,1683]" inLineHeading="true" pageId="14" pageNumber="567" reason="1">Phylogeny.</emphasis>
In the recent phylogeny of the
<taxonomicName id="C39C4D75FF8DAF0061B2AF44FCA0A1C3" box="[682,822,1658,1683]" class="Insecta" family="Psephenidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="family">Psephenidae</taxonomicName>
by
<bibRefCitation id="600D4B07FF8DAF006079AF45FB88A1C3" author="Lee, C. - F. &amp; Sato, M. &amp; Shepard, W. D. &amp; Jach, M. A." box="[865,1054,1658,1683]" pageId="14" pageNumber="567" pagination="502 - 538" refId="ref6628" refString="Lee, C. - F., Sato, M., Shepard, W. D. &amp; Jach, M. A. (2007) Phylogeny of the Psephenidae (Coleoptera: Byrrhoidea) based on larval, pupal and adult characters. Systematic Entomology, 32 (3), 502 - 538. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2006.00374. x" type="journal article" year="2007">Lee et al. (2007)</bibRefCitation>
,
<taxonomicName id="C39C4D75FF8DAF006732AF44FB36A1C3" authorityName="Shepard &amp; Barr" authorityYear="2014" box="[1066,1184,1658,1683]" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="36E8EAE4FF8DAF006732AF44FB36A1C3" box="[1066,1184,1658,1683]" italics="true" pageId="14" pageNumber="567">Neoeubria</emphasis>
</taxonomicName>
is included as “Genus A.” In the most parsimonious tree
<taxonomicName id="C39C4D75FF8DAF006138AFA0FD00A1E7" authorityName="Shepard &amp; Barr" authorityYear="2014" box="[544,662,1694,1719]" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="36E8EAE4FF8DAF006138AFA0FD00A1E7" box="[544,662,1694,1719]" italics="true" pageId="14" pageNumber="567">Neoeubria</emphasis>
</taxonomicName>
is placed in a basal trichotomy within the subfamily
<taxonomicName id="C39C4D75FF8DAF0067EDAFA0FAF0A1E7" authorityName="Lacordaire" authorityYear="1857" box="[1269,1382,1694,1719]" class="Insecta" family="Psephenidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="subFamily" subFamily="Eubriinae">Eubriinae</taxonomicName>
. The trichotomy positions
<taxonomicName id="C39C4D75FF8DAF00629DAFFCFE6DA18B" authorityName="Shepard &amp; Barr" authorityYear="2014" box="[389,507,1730,1755]" class="Insecta" family="Psephenidae" genus="Neoeubria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="36E8EAE4FF8DAF00629DAFFCFE6DA18B" box="[389,507,1730,1755]" italics="true" pageId="14" pageNumber="567">Neoeubria</emphasis>
</taxonomicName>
in one branch,
<taxonomicName id="C39C4D75FF8DAF0061B0AFFCFCA9A18B" authorityName="Blackburn" authorityYear="1892" box="[680,831,1730,1755]" class="Insecta" family="Psephenidae" genus="Sclerocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="36E8EAE4FF8DAF0061B0AFFCFCA9A18B" box="[680,831,1730,1755]" italics="true" pageId="14" pageNumber="567">Sclerocyphon</emphasis>
</taxonomicName>
+
<taxonomicName id="C39C4D75FF8DAF006046AFFCFC6FA18B" authorityName="Waterhouse" authorityYear="1876" box="[862,1017,1730,1755]" class="Insecta" family="Psephenidae" genus="Tychepsephus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="36E8EAE4FF8DAF006046AFFCFC6FA18B" box="[862,1017,1730,1755]" italics="true" pageId="14" pageNumber="567">Tychepsephus</emphasis>
</taxonomicName>
in another branch, and the remainder of the eubriine genera in a third branch.
</paragraph>
</subSubSection>
</treatment>
</document>
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