141 lines
13 KiB
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141 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.250.3715" ID-GBIF-Dataset="4f61fd14-c253-4e81-b31a-b04e06300223" ID-PMC="PMC3558971" ID-Pensoft-Pub="1313-2970-250-1" ID-PubMed="23378803" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-250-1" ModsDocOrigin="ZooKeys 250" ModsDocTitle="Introduction of the Exocelina ekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1451247866083" checkinUser="pensoft" docAuthor="Shaverdo, Helena V., Surbakti, Suriani, Hendrich, Lars & Balke, Michael" docDate="2012" docId="094D07C92BC41756DBCF114C25BB5C31" docLanguage="en" docName="ZooKeys 250: 1-76" docOrigin="ZooKeys 250" docSource="http://dx.doi.org/10.3897/zookeys.250.3715" docTitle="Exocelina kakapupu Shaverdo, Hendrich & Balke, sp. n." docType="treatment" docVersion="4" lastPageNumber="22" masterDocId="FFF3FFABFFA8FF93FFA65605CF65FFB4" masterDocTitle="Introduction of the Exocelina ekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="76" masterPageNumber="1" pageNumber="21" updateTime="1668155011676" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Introduction of the Exocelina ekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena V.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Surbakti, Suriani</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hendrich, Lars</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>250</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>76</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.250.3715</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.250.3715</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-250-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152039184" LSID="urn:lsid:zoobank.org:act:C54CD308-6991-4787-ACF7-5C5C2504B596" httpUri="http://treatment.plazi.org/id/094D07C92BC41756DBCF114C25BB5C31" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection pageId="20" pageNumber="21" type="nomenclature">
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<paragraph pageId="20" pageNumber="21">
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15.
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<taxonomicName LSID="urn:lsid:zoobank.org:act:C54CD308-6991-4787-ACF7-5C5C2504B596" authority="Shaverdo, Hendrich & Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina kakapupu" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="kakapupu">Exocelina kakapupu Shaverdo, Hendrich & Balke</taxonomicName>
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<taxonomicNameLabel pageId="20" pageNumber="21">sp. n.</taxonomicNameLabel>
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Figs 22
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<normalizedToken originalValue="A–E">A-E</normalizedToken>
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, 48
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</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="21" type="type locality">
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<paragraph pageId="20" pageNumber="21">Type locality.</paragraph>
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<paragraph pageId="20" pageNumber="21">
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Indonesia: Papua Province: Nabire/Paniai Regencies, road Nabire-Enarotali, 54th km,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-3.492">03°29.52'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="135.73183">135°43.91'E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="type material">
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<paragraph pageId="20" pageNumber="21">Type material.</paragraph>
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<paragraph lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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Holotype: male "IRIAN JAYA: Paniai Prov. road Nabire-Ilaga, km 54 26./27.8.1996, 750-800m leg. M. Balke (96 # 2)" (NHMW). Paratypes: 7 males
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<pageBreakToken pageId="21" pageNumber="22" start="start">with</pageBreakToken>
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the same label as the holotype (NHMW). 2 males "IR 24-W. New Guinea, track Nabire-Ilaga KM 54, basecamp 750m, 25.vii.1991 Balke &Hendrich leg." (NHMW, ZSM). 1 male "West New Guinea/Paniai Prov./IR 24 track Nabire-Ilaga km 54 basecamp, 750m, 25.7.1991 leg: Balke &Hendrich" (CLH). 1 male "West New Guinea/Paniai Prov./IR 19 track Nabire-Ilaga km 54 Basecamp, 750-800m, 16.-27.7.1991 leg: Balke & Hendrich" (CLH). 1 male "IRIAN JAYA: Paniai Prov. road Nabire-Ilaga, km 54 10.9.1996, 800m leg. M. Balke (96 # 20)" (NHMW), 1 male "IRIAN JAYA: Paniai Prov. road Nabire-Ilaga, km 54 10.9.1996, 900m leg. M. Balke (96 # 19)" (NHMW). 20 males "Indonesia: Papua, Road Nabire-Enarotali KM 60, 640m, 22.x.2011, 03 30.474S 135 42.611E, Uncen (PAP10)", one of them additionally with a label "DNA M. Balke 4912" (MZB, NHMW, ZSM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="additional material">
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<paragraph pageId="21" pageNumber="22">Additional material.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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12 females "Indonesia: Papua, Road Nabire-Enarotali KM 60, 640m, 22.x.2011, 03 30.474S 135 42.611E, Uncen (PAP10)", two of them additionally with labels "DNA M. Balke 4909", "DNA M. Balke 4913" (NHMW, ZSM). These females most likely belong to
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina kakapupu" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="kakapupu">Exocelina kakapupu</taxonomicName>
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sp. n. However they are not included in the type series, since four specimens of
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina weylandensis" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="weylandensis">Exocelina weylandensis</taxonomicName>
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sp. n. have been collected from this locality too, and, therefore, some of females might belong to this species. Also see in the paragraph of
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina irianensis" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="irianensis">Exocelina irianensis</taxonomicName>
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sp. n. and
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ekari" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ekari">Exocelina ekari</taxonomicName>
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sp. n.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="diagnosis">
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<paragraph pageId="21" pageNumber="22">Diagnosis.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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Beetle small, very similar to
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina soppi" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="soppi">Exocelina soppi</taxonomicName>
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sp. n. and
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina pseudosoppi" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pseudosoppi">Exocelina pseudosoppi</taxonomicName>
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sp. n. in size and coloration; pronotum without lateral bead; male antennomeres simple; male protarsomere 4 with large, thick, strongly curved anterolateral hook; median lobe with strong submedian constriction in ventral view and almost truncate apex in lateral view; paramere with notch on dorsal side and elongate subdistal part, with numerous, dense, long, thin setae; setae of proximal part longer, thicker, distinctly visible.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="description">
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<paragraph pageId="21" pageNumber="22">Description.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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Size and shape: Beetle small (TL-H 3.15-3.4 mm, TL 3.55-3.8 mm, MW 1.7-1.85 mm), with oblong-oval habitus, broadest at elytral middle. Coloration: as in
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina soppi" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="soppi">Exocelina soppi</taxonomicName>
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sp. n. and
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina pseudosoppi" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pseudosoppi">Exocelina pseudosoppi</taxonomicName>
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sp. n (Fig. 48).
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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Surface sculpture: Punctation as in
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina soppi" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="soppi">Exocelina soppi</taxonomicName>
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sp. n. and
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina pseudosoppi" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pseudosoppi">Exocelina pseudosoppi</taxonomicName>
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sp. n. but denser and coarser; microreticulation evidently stronger than in these species.
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</paragraph>
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<paragraph pageId="21" pageNumber="22">Structures: Pronotum without lateral bead or with weak traces of lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, anteriorly rounded, smooth, with very small anterolateral extensions. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct bead and few setae; neck and blade of prosternal process evenly jointed. Abdominal sternite 7 broadly truncate apically.</paragraph>
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<paragraph pageId="21" pageNumber="22">Male: Antenna simple (Fig. 22A). Protarsomere 4 with large, thick, strongly curved anterolateral hook. Protarsomere 5 ventrally with anterior row of 13 short setae and posterior row of 3 short setae (Fig. 22B). Abdominal sternite 7 with 2-7 lateral striae on each side. Median lobe with strong submedian constriction in ventral view and almost truncate apex in lateral view (Figs 22C, D). Paramere with notch on dorsal side and elongate subdistal part, with numerous, dense, long, thin setae; setae of proximal part longer, thicker, distinctly visible (Fig. 22E).</paragraph>
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<paragraph pageId="21" pageNumber="22">Female: Without evident differences in external morphology from male, except for abdominal sternite 7 without striae.</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="distribution">
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<paragraph pageId="21" pageNumber="22">Distribution.</paragraph>
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<paragraph pageId="21" pageNumber="22">Indonesia: Papua Province: Nabire and Paniai Regencies. This species is known only from the type locality area (Fig. 50).</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="etymology">
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<paragraph pageId="21" pageNumber="22">Etymology.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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The Ekari people sometimes called smaller water beetles
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<normalizedToken originalValue="“kakapupu”">"kakapupu"</normalizedToken>
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(
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<bibRefCitation author="Hendrich, L" journalOrPublisher="Latissimus - Newsletter of the Balfour-Browne Club" pageId="37" pageNumber="38" pagination="1 - 3" title="Kakapupu and others in fantastic New Guinea - An account of two hydroentomological expeditions to West New Guinea (Irian Jaya, Indonesia)." volume="1" year="1992">Hendrich and Balke 1992</bibRefCitation>
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). The species name is a noun in the nominative singular standing in apposition.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |