treatments-xml/data/03/FF/BF/03FFBF2FFFD6FFADFF2CFF45C59E06F9.xml
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<document id="3A2494321CAADE399C54866D38E39564" ID-DOI="10.11646/zootaxa.3972.4.3" ID-GBIF-Dataset="549c7fa4-ba3a-48b0-bc83-560ef7acfd1b" ID-ISSN="1175-5326" ID-Zenodo-Dep="243042" ID-ZooBank="CC1ED425-B897-4107-B39B-BA64D88A7980" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461189786377" checkinUser="plazi" docAuthor="Bilton, David T." docDate="2015" docId="03FFBF2FFFD6FFADFF2CFF45C59E06F9" docLanguage="en" docName="zt03972p520.pdf" docOrigin="Zootaxa 3972 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mesoceration foggoi Bilton, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="501" masterDocId="FFC6C757FFD3FFABFFBBFFD2C13F031C" masterDocTitle="New species and new records of Mesoceration Janssens, 1967 from South Africa (Coleoptera, Hydraenidae)" masterLastPageNumber="520" masterPageNumber="495" pageNumber="500" updateTime="1698600606930" updateUser="plazi">
<mods:mods id="265C6588E8AD40FCFE35490B4DA4514D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="49FB1625F11A51838DD9F7AC4D067AAE">
<mods:title id="83D26105193603C5D9553E7E15FA1AA8">New species and new records of Mesoceration Janssens, 1967 from South Africa (Coleoptera, Hydraenidae)</mods:title>
</mods:titleInfo>
<mods:name id="05CE47CBC5E7DD4CDFBFE3500E9BA883" type="personal">
<mods:role id="F4D823C8F14BDD64A0B91A38D477A2CA">
<mods:roleTerm id="F0991DD938D704EC934867E30433189C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B83DC4815CC8DD80C276187DBD23E33">Bilton, David T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2455129187C9BC89AB7B4DE89D12FE2A">text</mods:typeOfResource>
<mods:relatedItem id="264C4011BFE4D50DDAF01738BB3E10A0" type="host">
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<mods:title id="FFE7AB2545B9A912C6DD4ECF2B9358C7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C6FBA769E6761F699FA2D173B22EA2B9">
<mods:date id="282C2921FCA51DB7926BBB2BFE9B1A7A">2015</mods:date>
<mods:detail id="D3BEB95600A316C6A4431798388D907B" type="volume">
<mods:number id="D02641777128E40A28E6881F64F4E5E0">3972</mods:number>
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<mods:number id="93888DED8AB572B741A60393C26AF940">4</mods:number>
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<mods:classification id="17321298999D387006F1F852D32BAB75">journal article</mods:classification>
<mods:identifier id="70F514020C8A1AA912D6B8E214BE01BF" type="DOI">10.11646/zootaxa.3972.4.3</mods:identifier>
<mods:identifier id="3366090DC8F1EB9F79A42FB14E843FC6" type="GBIF-Dataset">549c7fa4-ba3a-48b0-bc83-560ef7acfd1b</mods:identifier>
<mods:identifier id="AD70BEC92370811CAF380FAF1D5FB8BA" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03FFBF2FFFD6FFADFF2CFF45C59E06F9" ID-DOI="http://doi.org/10.5281/zenodo.6107732" ID-GBIF-Taxon="119629315" ID-Zenodo-Dep="6107732" LSID="urn:lsid:plazi:treatment:03FFBF2FFFD6FFADFF2CFF45C59E06F9" httpUri="http://treatment.plazi.org/id/03FFBF2FFFD6FFADFF2CFF45C59E06F9" lastPageId="6" lastPageNumber="501" pageId="5" pageNumber="500">
<subSubSection id="C34C5DB2FFD6FFAEFF2CFF45C06603CF" pageId="5" pageNumber="500" type="nomenclature">
<paragraph id="8BE90E39FFD6FFAEFF2CFF45C0CB03AE" blockId="5.[151,500,151,211]" box="[151,500,151,178]" pageId="5" pageNumber="500">
<heading id="D0A1B955FFD6FFAEFF2CFF45C0CB03AE" bold="true" box="[151,500,151,178]" fontSize="11" level="1" pageId="5" pageNumber="500" reason="1">
<emphasis id="B922D22BFFD6FFAEFF2CFF45C0CB03AE" bold="true" box="[151,500,151,178]" pageId="5" pageNumber="500">
<taxonomicName id="4C5675BAFFD6FFAEFF2CFF45C0B003AD" ID-CoL="42236" box="[151,399,151,177]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="foggoi" status="sp. nov.">
<emphasis id="B922D22BFFD6FFAEFF2CFF45C0B003AD" bold="true" box="[151,399,151,177]" italics="true" pageId="5" pageNumber="500">Mesoceration foggoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2116F50FFD6FFAEFE2DFF4AC0CB03AE" box="[406,500,152,178]" pageId="5" pageNumber="500" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE90E39FFD6FFAEFF2CFF68C06603CF" blockId="5.[151,500,151,211]" box="[151,345,186,211]" pageId="5" pageNumber="500">
(
<figureCitation id="136D12BCFFD6FFAEFF24FF68C1DA03CF" box="[159,229,186,211]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[179,1408,203,1904]" captionTargetId="figure@3.[160,1426,195,1922]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Mesoceration holotypes. A) M. caniplenum sp. nov.; B) M. foggoi sp. nov.; C) M. helmei sp. nov.; D) M. hirsutum sp. nov." httpUri="https://zenodo.org/record/243043/files/figure.png" pageId="5" pageNumber="500">Figs 1</figureCitation>
,
<figureCitation id="136D12BCFFD6FFAEFF48FF69C03C03CF" box="[243,259,187,211]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[234,1357,212,1925]" captionTargetId="figure@4.[206,1380,193,1931]" captionTargetPageId="4" captionText="FIGURE 2. Aedeagi of Mesoceration species. A) M. caniplenum sp. nov., holotype; B) M. foggoi sp. nov., holotype; C) M. helmei sp. nov., holotype; D) M. hirsutum sp. nov., paratype. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/243044/files/figure.png" pageId="5" pageNumber="500">2</figureCitation>
,
<figureCitation id="136D12BCFFD6FFAEFEB4FF68C02203CF" box="[271,285,186,211]" captionStart="FIGURE 5" captionStartId="14.[151,250,1826,1848]" captionTargetBox="[236,1347,193,1806]" captionTargetId="figure@14.[236,1347,193,1807]" captionTargetPageId="14" captionText="FIGURE 5. Habitats of Mesoceration species in South Africa. A) KwaZulu-Natal, uKhahlamba Drakensberg National Park, Injisuthi, stream in Afromontane forest remnant below Van Heyningens Pass— M. caniplenum sp. nov., beetles collected from the wet rockface on the right hand side of the stream; B) Western Cape, Matsikammaberg, stream 1 km NW of Sewefontein farm— M. foggoi sp. nov., M. umbrosum; C) Western Cape, Cederberg, Breëwaterkloof stream, ca. 2 km SE of Uitkyk Pass, 1,050 m— M. hirsutum sp. nov., M. dissonum, M. granulovestum, M. jucundum, M. languidum, M. periscopum, M. piceum, M. repandum, M. rivulare; D) Western Cape, Franschhoek Pass, wet rock faces beside stream crossing R 45 road, 500 m— M. rugulosum sp. nov. Photos D. T. Bilton." httpUri="https://zenodo.org/record/243047/files/figure.png" pageId="5" pageNumber="500">5</figureCitation>
&amp;
<figureCitation id="136D12BCFFD6FFAEFEFAFF68C06F03CF" box="[321,336,186,211]" captionStart="FIGURE 6" captionStartId="15.[151,250,1860,1882]" captionTargetBox="[229,1347,193,1815]" captionTargetId="figure@15.[229,1347,193,1816]" captionTargetPageId="15" captionText="FIGURE 6. Habitats of Mesoceration species in South Africa. A) Western Cape, Matsikammaberg, permanent stream 1 km SE of Sewefontein farm— M. foggoi sp. nov., M. sewefonteinense sp. nov., M. umbrosum; B) Western Cape, False Bay, small stream on R 44 road below Boskloof Peak— M. sinclairi sp. nov., M. brevigranum; C) Western Cape, Groote Swartberg, permanent stream below Kangoberg on Gamkaskloof road— M. barriotum, M. littlekarroo, M. reticulatum, M. splendorum; D) Western Cape, Table Mountain National Park, Table Mountain, stream in Echo Valley— M. abstrictum, M. tabulare. Photos D. T. Bilton." httpUri="https://zenodo.org/record/243048/files/figure.png" pageId="5" pageNumber="500">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34C5DB2FFD6FFAEFF2CFED0C2CC019F" pageId="5" pageNumber="500" type="materials_examined">
<paragraph id="8BE90E39FFD6FFAEFF2CFED0C0390223" blockId="5.[151,1437,258,2011]" pageId="5" pageNumber="500">
<emphasis id="B922D22BFFD6FFAEFF2CFED0C00F0207" bold="true" box="[151,304,258,283]" pageId="5" pageNumber="500">
<typeStatus id="54EDB09BFFD6FFAEFF2CFED0C1ED0207" box="[151,210,258,283]" pageId="5" pageNumber="500">Type</typeStatus>
locality
</emphasis>
.
<emphasis id="B922D22BFFD6FFAEFEFBFED0C0E30207" bold="true" box="[320,476,258,283]" pageId="5" pageNumber="500">
<collectingCountry id="F3414EA9FFD6FFAEFEFBFED0C0E30207" box="[320,476,258,283]" name="South Africa" pageId="5" pageNumber="500">South Africa</collectingCountry>
</emphasis>
, Western Cape, Matsikammaberg, permanent stream
<quantity id="4CAEA3DCFFD6FFAEFBF9FED1C5410207" box="[1090,1150,258,283]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="500" unit="km" value="1.0">1 km</quantity>
SE of Sewefontein farm (
<figureCitation id="136D12BCFFD6FFAEFF24FEF4C1DC0223" box="[159,227,294,319]" captionStart="FIGURE 6" captionStartId="15.[151,250,1860,1882]" captionTargetBox="[229,1347,193,1815]" captionTargetId="figure@15.[229,1347,193,1816]" captionTargetPageId="15" captionText="FIGURE 6. Habitats of Mesoceration species in South Africa. A) Western Cape, Matsikammaberg, permanent stream 1 km SE of Sewefontein farm— M. foggoi sp. nov., M. sewefonteinense sp. nov., M. umbrosum; B) Western Cape, False Bay, small stream on R 44 road below Boskloof Peak— M. sinclairi sp. nov., M. brevigranum; C) Western Cape, Groote Swartberg, permanent stream below Kangoberg on Gamkaskloof road— M. barriotum, M. littlekarroo, M. reticulatum, M. splendorum; D) Western Cape, Table Mountain National Park, Table Mountain, stream in Echo Valley— M. abstrictum, M. tabulare. Photos D. T. Bilton." httpUri="https://zenodo.org/record/243048/files/figure.png" pageId="5" pageNumber="500">Fig. 6</figureCitation>
A).
</paragraph>
<paragraph id="8BE90E39FFD6FFAEFF7CFE98C01402B7" blockId="5.[151,1437,258,2011]" pageId="5" pageNumber="500">
<emphasis id="B922D22BFFD6FFAEFF7CFE98C04D027F" bold="true" box="[199,370,330,355]" pageId="5" pageNumber="500">
<typeStatus id="54EDB09BFFD6FFAEFF7CFE98C03D027F" box="[199,258,330,355]" pageId="5" pageNumber="500">Type</typeStatus>
material
</emphasis>
.
<emphasis id="B922D22BFFD6FFAEFE3DFE98C0CC027F" bold="true" box="[390,499,330,355]" pageId="5" pageNumber="500">
<typeStatus id="54EDB09BFFD6FFAEFE3DFE98C0CC027F" box="[390,499,330,355]" pageId="5" pageNumber="500" type="holotype">Holotype</typeStatus>
</emphasis>
(male): “
<date id="FFE828F9FFD6FFAEFDDDFE98C3DD027F" box="[614,738,330,355]" pageId="5" pageNumber="500" value="2014-09-21">21/ix/2014</date>
<collectingCountry id="F3414EA9FFD6FFAEFD55FE98C2BD027F" box="[750,898,330,355]" name="South Africa" pageId="5" pageNumber="500">South Africa</collectingCountry>
WC// Matsikammaberg stream//
<quantity id="4CAEA3DCFFD6FFAEFAB5FE99C473027F" box="[1294,1356,330,355]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="500" unit="km" value="1.0">1 km</quantity>
SE of Sewefontein farm// permanent D T Bilton leg.” (genitalia extracted and mounted on same card) and red
<typeStatus id="54EDB09BFFD6FFAEFA81FEBCC4A3029B" box="[1338,1436,366,391]" pageId="5" pageNumber="500" type="holotype">holotype</typeStatus>
label (
<collectionCode id="ED4796FCFFD6FFAEFF67FE41C02102B7" box="[220,286,403,427]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="5" pageNumber="500">SAM</collectionCode>
).
</paragraph>
<paragraph id="8BE90E39FFD6FFAEFF7CFE64C2CC019F" blockId="5.[151,1437,258,2011]" pageId="5" pageNumber="500">
<emphasis id="B922D22BFFD6FFAEFF7CFE64C00002D3" bold="true" box="[199,319,438,463]" pageId="5" pageNumber="500">
<typeStatus id="54EDB09BFFD6FFAEFF7CFE64C00002D3" box="[199,319,438,463]" pageId="5" pageNumber="500" type="paratype">Paratypes</typeStatus>
</emphasis>
(36):
<specimenCount id="9D50C5B0FFD6FFAEFE32FE65C0FB02D2" box="[393,452,439,463]" pageId="5" pageNumber="500" type="generic">13 ♂</specimenCount>
, 13 ♀ same data as
<typeStatus id="54EDB09BFFD6FFAEFD02FE64C22002D2" box="[697,799,438,462]" pageId="5" pageNumber="500" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D50C5B0FFD6FFAEFC96FE64C26502D2" box="[813,858,438,462]" pageId="5" pageNumber="500" type="generic">6 ♂</specimenCount>
<date id="FFE828F9FFD6FFAEFCD4FE64C2D502D3" box="[879,1002,438,463]" pageId="5" pageNumber="500" value="2014-09-21">21/ix/2014</date>
<collectingCountry id="F3414EA9FFD6FFAEFC4EFE64C5B802D3" box="[1013,1159,438,463]" name="South Africa" pageId="5" pageNumber="500">South Africa</collectingCountry>
WC// Matsikammaberg stream//
<quantity id="4CAEA3DCFFD6FFAEFEBAFE09C07C02EF" box="[257,323,474,499]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="500" unit="km" value="1.0">1 km</quantity>
NW of Sewefontein farm// D T Bilton leg.”;
<specimenCount id="9D50C5B0FFD6FFAEFC28FE09C2F702EE" box="[915,968,475,499]" pageId="5" pageNumber="500" type="generic">1 ♂</specimenCount>
, 1 ♀ “
<date id="FFE828F9FFD6FFAEFB90FE08C59902EF" box="[1067,1190,474,499]" pageId="5" pageNumber="500" value="2014-09-21">21/ix/2014</date>
<collectingCountry id="F3414EA9FFD6FFAEFB0DFE08C47002EF" box="[1206,1359,474,499]" name="South Africa" pageId="5" pageNumber="500">South Africa</collectingCountry>
WC// Matsikammaberg seepage stream// over rock at head of Elandskloof// D T Bilton leg.”;
<specimenCount id="9D50C5B0FFD6FFAEFB2FFE2DC5FE010A" box="[1172,1217,511,535]" pageId="5" pageNumber="500" type="generic">1 ♂</specimenCount>
<date id="FFE828F9FFD6FFAEFB6EFE2CC46E010B" box="[1237,1361,510,535]" pageId="5" pageNumber="500" value="2014-09-20">20/ix/2014</date>
<collectingCountry id="F3414EA9FFD6FFAEFAE0FE2CC1E00127" name="South Africa" pageId="5" pageNumber="500">South Africa</collectingCountry>
WC// Gifberg—stream in Gifberg Pass// above Vanrhynsdorp// rocky stream D T Bilton leg.”;
<specimenCount id="9D50C5B0FFD6FFAEFAA0FDF1C4790126" box="[1307,1350,547,571]" pageId="5" pageNumber="500" type="generic">1 ♂</specimenCount>
<date id="FFE828F9FFD6FFAEFAE3FDF0C1F00143" pageId="5" pageNumber="500" value="2014-09-20">20/ix/ 2014</date>
<collectingCountry id="F3414EA9FFD6FFAEFF6EFD94C05B0143" box="[213,356,582,607]" name="South Africa" pageId="5" pageNumber="500">South Africa</collectingCountry>
WC// Gifberg-vertical wet rock face// in Gifberg Pass nr. Vanrhynsdorp// A Foggo leg.” (
<collectionCode id="ED4796FCFFD6FFAEFAF2FD95C4A70143" box="[1353,1432,583,607]" pageId="5" pageNumber="500">CDTB</collectionCode>
,
<collectionCode id="ED4796FCFFD6FFAEFF2CFDB9C1E8019F" LSID="urn:lsid:biocol.org:col:33791" box="[151,215,619,643]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="500">MCZ</collectionCode>
,
<collectionCode id="ED4796FCFFD6FFAEFF59FDB9C015019E" box="[226,298,619,642]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="500">NMW</collectionCode>
,
<collectionCode id="ED4796FCFFD6FFAEFE8EFDB9C09D019F" box="[309,418,619,643]" pageId="5" pageNumber="500">OUMNH</collectionCode>
,
<collectionCode id="ED4796FCFFD6FFAEFE17FDB9C0D4019F" box="[428,491,619,643]" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="5" pageNumber="500">SAM</collectionCode>
,
<collectionCode id="ED4796FCFFD6FFAEFE4CFDB9C37B019F" LSID="urn:lsid:biocol.org:col:34248" box="[503,580,619,643]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="5" pageNumber="500">SANC</collectionCode>
,
<collectionCode id="ED4796FCFFD6FFAEFDF4FDB9C39D019F" box="[591,674,619,643]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="5" pageNumber="500">TMSA</collectionCode>
). All with red
<typeStatus id="54EDB09BFFD6FFAEFCF8FDB9C29C019F" box="[835,931,619,643]" pageId="5" pageNumber="500" type="paratype">paratype</typeStatus>
labels.
</paragraph>
</subSubSection>
<subSubSection id="C34C5DB2FFD6FFADFF7CFD5CC47D07BC" lastPageId="6" lastPageNumber="501" pageId="5" pageNumber="500" type="description">
<paragraph id="8BE90E39FFD6FFAEFF7CFD5CC59101D6" blockId="5.[151,1437,258,2011]" pageId="5" pageNumber="500">
<emphasis id="B922D22BFFD6FFAEFF7CFD5CC06E01BB" bold="true" box="[199,337,654,679]" pageId="5" pageNumber="500">Description</emphasis>
. Size:
<typeStatus id="54EDB09BFFD6FFAEFE18FD5CC32F01BB" box="[419,528,654,679]" pageId="5" pageNumber="500" type="holotype">Holotype</typeStatus>
: BL
<quantity id="4CAEA3DCFFD6FFAEFDF2FD5DC39B01BA" box="[585,676,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="500" unit="mm" value="1.7">1.7 mm</quantity>
; EL
<quantity id="4CAEA3DCFFD6FFAEFD66FD5CC27901BA" box="[733,838,654,679]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="500" unit="mm" value="1.05">1.05 mm</quantity>
; EW
<quantity id="4CAEA3DCFFD6FFAEFC33FD5CC2DB01BA" box="[904,996,654,679]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="500" unit="mm" value="0.6">0.6 mm</quantity>
.
<typeStatus id="54EDB09BFFD6FFAEFC4BFD5DC55D01BB" box="[1008,1122,655,679]" pageId="5" pageNumber="500" type="paratype">Paratypes</typeStatus>
: ♂s BL
<quantity id="4CAEA3DCFFD6FFAEFB7FFD5CC45301BA" box="[1220,1388,654,679]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.75" metricValueMin="1.65" pageId="5" pageNumber="500" unit="mm" value="1.7" valueMax="1.75" valueMin="1.65">1.651.75 mm</quantity>
; EL 1.0
<quantity id="4CAEA3DCFFD6FFAEFF73FD60C01001D6" box="[200,303,690,714]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="500" unit="mm" value="1.05">1.05 mm</quantity>
; EW
<quantity id="4CAEA3DCFFD6FFAEFED6FD60C33A01D6" box="[365,517,690,714]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="5" pageNumber="500" unit="mm" value="0.625" valueMax="0.65" valueMin="0.6">0.60.65 mm</quantity>
. ♀s BL
<quantity id="4CAEA3DCFFD6FFAEFDDAFD60C23801D6" box="[609,775,690,714]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.85" metricValueMin="1.65" pageId="5" pageNumber="500" unit="mm" value="1.75" valueMax="1.85" valueMin="1.65">1.651.85 mm</quantity>
; EL
<quantity id="4CAEA3DCFFD6FFAEFC87FD60C2EB01D6" box="[828,980,690,714]" metricMagnitude="-3" metricUnit="m" metricValue="1.075" metricValueMax="1.1" metricValueMin="1.05" pageId="5" pageNumber="500" unit="mm" value="1.075" valueMax="1.1" valueMin="1.05">1.051.1 mm</quantity>
; EW
<quantity id="4CAEA3DCFFD6FFAEFBAAFD60C59601D6" box="[1041,1193,690,714]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="5" pageNumber="500" unit="mm" value="0.625" valueMax="0.65" valueMin="0.6">0.60.65 mm</quantity>
.
</paragraph>
<paragraph id="8BE90E39FFD6FFAEFF7CFD04C4270047" blockId="5.[151,1437,258,2011]" pageId="5" pageNumber="500">
Colour: Dorsum (
<figureCitation id="136D12BCFFD6FFAEFE35FD04C0F101F3" box="[398,462,726,751]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[179,1408,203,1904]" captionTargetId="figure@3.[160,1426,195,1922]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Mesoceration holotypes. A) M. caniplenum sp. nov.; B) M. foggoi sp. nov.; C) M. helmei sp. nov.; D) M. hirsutum sp. nov." httpUri="https://zenodo.org/record/243043/files/figure.png" pageId="5" pageNumber="500">Fig. 1</figureCitation>
B) with head and pronotum dark piceous brown to black; ocelli slightly paler than rest of head. Elytra paler; orange-brown, suture pitchy, and explanate margins paler. Legs orange-brown, tarsi and femoro-tibial junctions infuscated; tarsal claws pale yellow. Maxillary palpi pitchy brown to black. Venter dark brown to black, with silvery hydrofuge vestiture; pronotal hypomeron and elytral pseudepipleurs paler.
</paragraph>
<paragraph id="8BE90E39FFD6FFAEFF7CFCB4C1E107DF" blockId="5.[151,1437,258,2011]" pageId="5" pageNumber="500">Head: Labrum slightly transverse, rounded apicolaterally with broad apicomedian emargination running approx. 0.4 of length. Sides of apicomedian emargination and anterior angles weakly raised. Surface shining, lacking microreticulation and with scattered fine punctures bearing long white decumbent setae. Frontoclypeal suture strong, arcuate. Clypeus, frons and vertex strongly shining. Clypeus with medium scattered punctures bearing decumbent setae; devoid of microreticulation over central 0.8, with weak longitudinal meshes at lateral margins. Central portion of frons sculptured as central clypeus; punctures slightly larger and sparser. Anteocellar sulci well-marked, broad, open laterally. Sulci and area inside compound eyes rugulosely microreticulated. Ocelli small, shining. Head behind ocelli with sharp declivity. Vertex and occiput rugulose, with transverse microreticulation. Compound eyes large, occupying approx. 0.5 lateral margin of head, 14 ommatidia in longest series.</paragraph>
<paragraph id="8BE90E39FFD6FFAEFF7CFB1DC2850536" blockId="5.[151,1437,258,2011]" pageId="5" pageNumber="500">Pronotum: Cordate, slightly transverse, broadest just before middle. Anterior margin arcuate over median 0.5, without hyaline border; posterior margin straight. Anterior angles obtusely rounded, posterior angles rectangular. Lateral margins sinuated from widest point to base; broadly marginated and crenulate. Surface shining, lacking microreticulation except for areas of transverse meshes in narrow strips in central 0.5 close to anterior and posterior margins. With 10 distinct foveae as follows: Anterior median fovea elongate; posterior median fovea transverse oval. Anterior admedian fovea circular; posterior admedian fovea oval, orientated anterolaterally. Anterior adlateral fovea circular, open laterally; posterior adlateral fovea oval, open laterally. Field of coarse punctures just behind anterior margin in central 0.5, punctures extending into anterior 0.5 of anterior median and anterior admedian foveae. Double row of coarse punctures along posterior margin in central 0.6; remainder of disc with scattered fine—medium punctures bearing long white decumbent setae.</paragraph>
<paragraph id="8BE90E39FFD6FFAEFF7CF9E4C07D0456" blockId="5.[151,1437,258,2011]" pageId="5" pageNumber="500">Elytra: Elongate oval, broadest at middle. Subparallel over middle 0.3, then attenuated to truncately rounded apex. Apical angles rounded, slightly emarginated around suture. Sides explanate; minutely serrate. Punctures of elytral series evident to apices; punctures relatively coarse; larger in anterior 0.5. Discal series striate impressed, especially in anterior 0.3. Series 1 and 2 confluent just behind middle, in front of posterior declivity. Series 5 and 6 confluent over anterior 0.3. Serial punctures bearing fine, white, decumbent setae, which reach anterior margin of next puncture in row. Elytral intervals minutely granulate; granules each with fine puncture posteriorly, which bears long, white, decumbent seta which overlaps next seta in row by approx. 0.5 of length. Interval 8 carinate almost to apex.</paragraph>
<paragraph id="8BE90E39FFD6FFAEFF7CF885C40E0472" blockId="5.[151,1437,258,2011]" box="[199,1329,1878,1903]" pageId="5" pageNumber="500">Legs: Apical protarsomere with two stout ventral setae; basal three protarsomeres with suction setae.</paragraph>
<paragraph id="8BE90E39FFD6FFADFF7CF8A9C4BD02E9" blockId="5.[151,1437,258,2011]" lastBlockId="6.[151,1437,151,1509]" lastPageId="6" lastPageNumber="501" pageId="5" pageNumber="500">
Venter: Mentum weakly shining, rugulosely microreticulate; meshes elongate along lateral margins and isodiametric in centre. Submentum shining, lacking microreticulation. Genae with transverse microreticulation anteriorly; hydrofuge vestiture posteriorly. Gula with transverse, open microreticulation. Prosternum with narrow, distinct, shining and glabrous central ridge. Rest of surface rugulose, with dense hydrofuge vestiture. Pronotal hypomeron broad, shining, lacking microreticulation. Mesoventrite and metaventrite with dense hydrofuge vestiture. Mesoventral plaques glabrous, forming inverted
<collectingCountry id="F3414EA9FFD5FFADFCF4FF32C25C03EB" box="[847,867,224,247]" name="Italy" pageId="6" pageNumber="501">Y</collectingCountry>
. Adlateral and admedian ridges also visible on mesoventrite. Metaventrite with broad, elongate median fovea over posterior 0.5. Metaventrite with small, shining glabrous patch apicomedially, close to metacoxae. Elytral pseudoepipleurs broad, shining, lacking microreticulation; epipleurs shining, ridge-like, with hydrofuge vestiture over anterior 0.2. Abdominal ventrites 1 4 with dense hydrofuge vestiture. Abdominal ventrite 5 glabrous over posterior
<quantity id="4CAEA3DCFFD5FFADFB90FEA2C5520294" box="[1067,1133,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="2.032" pageId="6" pageNumber="501" unit="in" value="0.8">0.8 in</quantity>
central 0.3, with scattered hydrofuge setae and dense, medium punctures bearing long, white, decumbent setae in posterior 0.5 of glabrous patch. Abdominal ventrite 6 shining, traces of transverse microreticulation and double row of irregular punctures along posterior margin, with long, decumbent setae in centre of row, and shorter, stouter, peg-like setae laterally.
</paragraph>
<paragraph id="8BE90E39FFD5FFADFF7CFE2DC39C0198" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="501">
Aedeagus: Elongate, with parameres attaching close to base and extending well beyond the apex of the main piece. Main piece relatively broad in ventral view, with strong, stout, curved apical tooth, and long, tube-like distal lobe, extending just beyond the paramere apices. Distal lobe bifurcated approx. 0.25 from apex—this structure being unique so far within the genus (
<figureCitation id="136D12BCFFD5FFADFD85FDBEC3BD0198" box="[574,642,620,645]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[234,1357,212,1925]" captionTargetId="figure@4.[206,1380,193,1931]" captionTargetPageId="4" captionText="FIGURE 2. Aedeagi of Mesoceration species. A) M. caniplenum sp. nov., holotype; B) M. foggoi sp. nov., holotype; C) M. helmei sp. nov., holotype; D) M. hirsutum sp. nov., paratype. Scale bar = 0.25 mm." httpUri="https://zenodo.org/record/243044/files/figure.png" pageId="6" pageNumber="501">Fig. 2</figureCitation>
B).
</paragraph>
<paragraph id="8BE90E39FFD5FFADFF7CFD5DC5D20009" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="501">Female: Elytra broader and flatter than in males, more strongly explanate along lateral margins. Apex more strongly truncated and emarginated around suture. Abdominal ventrite 5 with glabrous area occupying central 0.3, with weak transverse microreticulation and moderate, medium punctures over apical 0.2, each bearing a yellow decumbent seta. Setae longer centrally; shorter, stouter and peg-like laterally. Anterior tarsi simple.</paragraph>
<paragraph id="8BE90E39FFD5FFADFF7CFCCDC5430041" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="501">
Variation:
<typeStatus id="54EDB09BFFD5FFADFEF9FCF2C0900024" box="[322,431,800,824]" pageId="6" pageNumber="501" type="paratype">Paratypes</typeStatus>
vary somewhat in colouration, some specimens being paler than the
<typeStatus id="54EDB09BFFD5FFADFB4FFCCDC4660024" box="[1268,1369,799,824]" pageId="6" pageNumber="501" type="holotype">holotype</typeStatus>
, with yellowish brown elytra and brownish pronotum. This is not, apparently, due to tenerality.
</paragraph>
<paragraph id="8BE90E39FFD5FFADFF7CFCB5C47D07BC" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="501">
<emphasis id="B922D22BFFD5FFADFF7CFCB5C0EC009C" bold="true" box="[199,467,871,896]" pageId="6" pageNumber="501">Differential diagnosis.</emphasis>
A member of the
<taxonomicName id="4C5675BAFFD5FFADFD03FCBAC22E009C" box="[696,785,872,896]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="rivulare">
<emphasis id="B922D22BFFD5FFADFD03FCBAC22E009C" box="[696,785,872,896]" italics="true" pageId="6" pageNumber="501">rivulare</emphasis>
</taxonomicName>
group, which would key to
<emphasis id="B922D22BFFD5FFADFBD0FCBBC425009C" box="[1131,1306,872,896]" italics="true" pageId="6" pageNumber="501">
M.
<taxonomicName id="4C5675BAFFD5FFADFB2DFCBAC425009C" authority="Perkins &amp; Balfour-Browne, 1994" authorityName="Perkins &amp; Balfour-Browne" authorityYear="1994" box="[1174,1306,872,896]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="splendorum">splendorum</taxonomicName>
</emphasis>
<bibRefCitation id="EFC773C8FFD5FFADFA9DFCB5C0A600B8" author="Perkins" pageId="6" pageNumber="501" refString="Perkins, P. D. &amp; Balfour-Browne, J. (1994) A contribution to the taxonomy of aquatic and humicolous beetles of the family Hydraenidae in southern Africa. Fieldiana Zoology, 77, 1 - 159." type="journal article" year="1994">Perkins &amp; Balfour-Browne, 1994</bibRefCitation>
. Compared to
<emphasis id="B922D22BFFD5FFADFDF8FC5FC3D100B8" box="[579,750,908,932]" italics="true" pageId="6" pageNumber="501">
M.
<taxonomicName id="4C5675BAFFD5FFADFDD1FC5EC3D100B8" box="[618,750,908,932]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="splendorum">splendorum</taxonomicName>
</emphasis>
,
<emphasis id="B922D22BFFD5FFADFD44FC5FC25100B8" box="[767,878,908,932]" italics="true" pageId="6" pageNumber="501">
M.
<taxonomicName id="4C5675BAFFD5FFADFC9CFC5EC25100B8" box="[807,878,908,932]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="foggoi" status="sp. nov.">foggoi</taxonomicName>
</emphasis>
<emphasis id="B922D22BFFD5FFADFCCCFC5EC2EC00B9" bold="true" box="[887,979,908,933]" pageId="6" pageNumber="501">
<taxonomicNameLabel id="A2116F50FFD5FFADFCCCFC5EC2EC00B9" box="[887,979,908,933]" pageId="6" pageNumber="501" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is larger (BL
<quantity id="4CAEA3DCFFD5FFADFBC2FC5EC42100B8" box="[1145,1310,908,933]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.85" metricValueMin="1.65" pageId="6" pageNumber="501" unit="mm" value="1.75" valueMax="1.85" valueMin="1.65">1.651.85 mm</quantity>
vs. ca.
<quantity id="4CAEA3DCFFD5FFADFAC2FC5EC1F800D4" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="501" unit="mm" value="1.6">1.6 mm</quantity>
). Within the
<taxonomicName id="4C5675BAFFD5FFADFEE3FC62C08E00D4" box="[344,433,944,968]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="rivulare">
<emphasis id="B922D22BFFD5FFADFEE3FC62C08E00D4" box="[344,433,944,968]" italics="true" pageId="6" pageNumber="501">rivulare</emphasis>
</taxonomicName>
group, the new species can be further distinguished by the elytral series, which are wellmarked throughout, with the discal series striate impressed. The most similar species in size and colouration is
<emphasis id="B922D22BFFD5FFADFAC6FC07C1D4070C" italics="true" pageId="6" pageNumber="501">M. hantam</emphasis>
Bilton, 2014, which has less impressed elytral series, composed of smaller punctures. The aedeagus of
<emphasis id="B922D22BFFD5FFADFAC6FC2BC1E00728" italics="true" pageId="6" pageNumber="501">
M.
<taxonomicName id="4C5675BAFFD5FFADFF2CFBCEC1E00728" box="[151,223,1052,1076]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="foggoi" status="sp. nov.">foggoi</taxonomicName>
</emphasis>
<emphasis id="B922D22BFFD5FFADFF5DFBCEC07F0729" bold="true" box="[230,320,1052,1077]" pageId="6" pageNumber="501">
<taxonomicNameLabel id="A2116F50FFD5FFADFF5DFBCEC07F0729" box="[230,320,1052,1077]" pageId="6" pageNumber="501" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is closest to
<emphasis id="B922D22BFFD5FFADFE68FBCFC3580728" box="[467,615,1052,1076]" italics="true" pageId="6" pageNumber="501">M. rapidense</emphasis>
<bibRefCitation id="EFC773C8FFD5FFADFDD4FBCEC2350728" author="Perkins" box="[623,778,1052,1077]" pageId="6" pageNumber="501" refString="Perkins, P. D. (2008) New species and new collection records of Prosthetopine water beetles from southern Africa (Coleoptera: Hydraenidae). Zootaxa, 1864, 1 - 124." type="journal article" year="2008">Perkins, 2008</bibRefCitation>
[note—automatic emendation of
<taxonomicName id="4C5675BAFFD5FFADFB3DFBCFC1D10744" authority="Perkins" authorityName="Perkins" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="rapidensis">
<emphasis id="B922D22BFFD5FFADFB3DFBCFC4A40728" box="[1158,1435,1052,1076]" italics="true" pageId="6" pageNumber="501">Mesoceration rapidensis</emphasis>
Perkins
</taxonomicName>
] in overall structure, although these species differ in the detail of the apical tooth, and the bifurcated distal lobe of
<emphasis id="B922D22BFFD5FFADFF55FBB7C0630760" box="[238,348,1124,1148]" italics="true" pageId="6" pageNumber="501">
M.
<taxonomicName id="4C5675BAFFD5FFADFEAEFBB6C0630760" box="[277,348,1124,1148]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="foggoi" status="sp. nov.">foggoi</taxonomicName>
</emphasis>
<emphasis id="B922D22BFFD5FFADFEDEFBB6C0800761" bold="true" box="[357,447,1124,1149]" pageId="6" pageNumber="501">
<taxonomicNameLabel id="A2116F50FFD5FFADFEDEFBB6C0800761" box="[357,447,1124,1149]" pageId="6" pageNumber="501" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is so far unique within the genus.
<emphasis id="B922D22BFFD5FFADFCF2FBB7C2E10760" box="[841,990,1124,1148]" italics="true" pageId="6" pageNumber="501">M. rapidense</emphasis>
is also very different externally, with a much narrower forebody than
<emphasis id="B922D22BFFD5FFADFE50FB5BC36707BC" box="[491,600,1160,1184]" italics="true" pageId="6" pageNumber="501">
M.
<taxonomicName id="4C5675BAFFD5FFADFDAAFB5AC36707BC" box="[529,600,1160,1184]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="foggoi" status="sp. nov.">foggoi</taxonomicName>
</emphasis>
<emphasis id="B922D22BFFD5FFADFDE4FB55C38607BC" bold="true" box="[607,697,1159,1184]" pageId="6" pageNumber="501">
<taxonomicNameLabel id="A2116F50FFD5FFADFDE4FB55C38607BC" box="[607,697,1159,1184]" pageId="6" pageNumber="501" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, with weak elytral series, which are not striate impressed.
</paragraph>
</subSubSection>
<subSubSection id="C34C5DB2FFD5FFADFF7CFB7EC3960681" pageId="6" pageNumber="501" type="distribution">
<paragraph id="8BE90E39FFD5FFADFF7CFB7EC3960681" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="501">
<emphasis id="B922D22BFFD5FFADFF7CFB7EC06607D9" bold="true" box="[199,345,1196,1221]" pageId="6" pageNumber="501">Distribution</emphasis>
. Known to date only from a number of localities on the Gifberg-Matsikammaberg massif in the northern part of the fynbos biome of the Core Cape Floristic region (
<bibRefCitation id="EFC773C8FFD5FFADFC0FFB1DC5CC07F4" author="Manning" box="[948,1267,1231,1256]" pageId="6" pageNumber="501" refString="Manning, J. C. &amp; Goldblatt, P. (2012) The Greater Cape Floristic Region I: the Core Cape Flora. Strelitzia, 29, 1 - 853. [South African national Biodiversity Institute, Pretoria]" type="journal article" year="2012">Manning &amp; Goldblatt 2012</bibRefCitation>
). Apparently a relatively common and widespread species in these mountains, found abundantly in all running-water sites investigated in 2014. Taken together with two other apparent regional Gifberg-Matsikammaberg endemics,
<taxonomicName id="4C5675BAFFD5FFADFF2CFAEFC3690648" authority="Perkins, 2008" authorityName="Perkins" authorityYear="2008" box="[151,598,1340,1365]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="umbrosum">
<emphasis id="B922D22BFFD5FFADFF2CFAEFC08F0648" box="[151,432,1340,1364]" italics="true" pageId="6" pageNumber="501">Mesoceration umbrosum</emphasis>
<bibRefCitation id="EFC773C8FFD5FFADFE02FAEEC3690648" author="Perkins" box="[441,598,1340,1365]" pageId="6" pageNumber="501" refString="Perkins, P. D. (2008) New species and new collection records of Prosthetopine water beetles from southern Africa (Coleoptera: Hydraenidae). Zootaxa, 1864, 1 - 124." type="journal article" year="2008">Perkins, 2008</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C5675BAFFD5FFADFD29FAEFC2D40648" box="[658,1003,1340,1364]" class="Insecta" family="Hydraenidae" genus="Mesoceration" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="sewefonteinense" status="sp. nov.">
<emphasis id="B922D22BFFD5FFADFD29FAEFC2D40648" box="[658,1003,1340,1364]" italics="true" pageId="6" pageNumber="501">Mesoceration sewefonteinense</emphasis>
</taxonomicName>
<emphasis id="B922D22BFFD5FFADFC4EFAEEC56E0649" bold="true" box="[1013,1105,1340,1365]" pageId="6" pageNumber="501">
<taxonomicNameLabel id="A2116F50FFD5FFADFC4EFAEEC56E0649" box="[1013,1105,1340,1365]" pageId="6" pageNumber="501" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Particularly abundant where thin trickles of water flowed over algae-encrusted sheets of sandstone bedrock in stream margins (e.g.
<figureCitation id="136D12BCFFD5FFADFA8EFA8DC4450664" box="[1333,1402,1375,1400]" captionStart="FIGURE 5" captionStartId="14.[151,250,1826,1848]" captionTargetBox="[236,1347,193,1806]" captionTargetId="figure@14.[236,1347,193,1807]" captionTargetPageId="14" captionText="FIGURE 5. Habitats of Mesoceration species in South Africa. A) KwaZulu-Natal, uKhahlamba Drakensberg National Park, Injisuthi, stream in Afromontane forest remnant below Van Heyningens Pass— M. caniplenum sp. nov., beetles collected from the wet rockface on the right hand side of the stream; B) Western Cape, Matsikammaberg, stream 1 km NW of Sewefontein farm— M. foggoi sp. nov., M. umbrosum; C) Western Cape, Cederberg, Breëwaterkloof stream, ca. 2 km SE of Uitkyk Pass, 1,050 m— M. hirsutum sp. nov., M. dissonum, M. granulovestum, M. jucundum, M. languidum, M. periscopum, M. piceum, M. repandum, M. rivulare; D) Western Cape, Franschhoek Pass, wet rock faces beside stream crossing R 45 road, 500 m— M. rugulosum sp. nov. Photos D. T. Bilton." httpUri="https://zenodo.org/record/243047/files/figure.png" pageId="6" pageNumber="501">Fig. 5</figureCitation>
B). Some of the specimens collected were tenerals.
</paragraph>
</subSubSection>
<subSubSection id="C34C5DB2FFD5FFADFF7CFA75C59E06F9" pageId="6" pageNumber="501" type="etymology">
<paragraph id="8BE90E39FFD5FFADFF7CFA75C59E06F9" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="501">
<emphasis id="B922D22BFFD5FFADFF7CFA75C07906DC" bold="true" box="[199,326,1447,1472]" pageId="6" pageNumber="501">Etymology</emphasis>
. Named after my friend and colleague Dr Andy Foggo, who enthusiastically assisted with water beetle sampling on the Gifberg-Matsikammaberg, including the collection of the
<typeStatus id="54EDB09BFFD5FFADFB98FA1FC56C06F9" box="[1059,1107,1485,1509]" pageId="6" pageNumber="501">type</typeStatus>
series.
</paragraph>
</subSubSection>
</treatment>
</document>