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<mods:title id="F6B4F09BD2FA9A56F9FC02AFA8EE7E4B">Chitons (Mollusca, Polyplacophora) from the Maldive islands</mods:title>
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<mods:namePart id="8CAADBE80E77AF02AD3C4155D0A0E9FE">Gori, Sandro</mods:namePart>
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<mods:namePart id="2CE3614E5C4F086C87C31C998DCD9D90">Bonfitto, Antonio</mods:namePart>
<mods:affiliation id="98CD8FFE8030126B20F3A333D9F33ABC">Dipartimento di Biologia evoluzionistica e sperimentale, Via Selmi 3, 40126 Bologna, Italy</mods:affiliation>
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<subSubSection id="C3640CE5FFDAFF84FF7FFBA6FDCAB16E" box="[151,520,1098,1124]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BC15F6EFFDAFF84FF7FFBA6FDCAB16E" blockId="14.[151,520,1098,1159]" box="[151,520,1098,1124]" pageId="14" pageNumber="15">
<heading id="D089E802FFDAFF84FF7FFBA6FDCAB16E" bold="true" box="[151,520,1098,1124]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="4C7E24EDFFDAFF84FF7FFBA6FDCAB16E" ID-CoL="3WC4J" authority="(Kaas, 1979)" baseAuthorityName="Kaas" baseAuthorityYear="1979" box="[151,520,1098,1124]" class="Polyplacophora" family="Chitonidae" genus="Lucilina" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="carnosa">
<emphasis id="B90A837CFFDAFF84FF7FFBA6FDCAB16E" bold="true" box="[151,520,1098,1124]" pageId="14" pageNumber="15">
<emphasis id="B90A837CFFDAFF84FF7FFBA6FEA6B16E" bold="true" box="[151,356,1098,1124]" italics="true" pageId="14" pageNumber="15">Lucilina carnosa</emphasis>
(
<bibRefCitation id="EFEF229FFFDAFF84FE9CFBA6FDC2B16E" author="Kaas, P." box="[372,512,1098,1124]" pageId="14" pageNumber="15" pagination="855 - 879" refId="ref21829" refString="Kaas, P. (1979) The Chitons (Mollusca: Polyplacophora) of Mozambique. Annals of the Natal Museum, 23, 855 - 879." type="journal article" year="1979">Kaas, 1979</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3640CE5FFDAFF84FF7FFB81FE6DB18D" box="[151,431,1133,1159]" pageId="14" pageNumber="15" type="description">
<paragraph id="8BC15F6EFFDAFF84FF7FFB81FE6DB18D" blockId="14.[151,520,1098,1159]" box="[151,431,1133,1159]" pageId="14" pageNumber="15">
(
<figureCitation id="134543EBFFDAFF84FF77FB81FE87B18D" box="[159,325,1133,1159]" captionStart="FIGURE 5" captionStartId="15.[151,257,1381,1405]" captionTargetBox="[151,1436,645,1360]" captionTargetId="figure-400@15.[151,1436,645,1360]" captionTargetPageId="15" captionText="FIGURE 5AB, Lucilina perligera Thiele, 1909, Maldive, M13, intermediate valve. A, dorsal view. B, tegmentum granules of the central area. CL, Lucilina carnosa (Kaas, 1979), Maldive. CD, head valve, MLD02A. C, dorsal view. D, detail of the sculpture. EF, intermediate valve, MLD02A. E, dorsal view. F, detail of the sculpture. GI, intermediate valve, M23. G, dorsal view. H, frontal view. I, detail of the sculpture. JL, tail valve, MLD02A. J. dorsal view. K, lateral view. L, detail of the sculpture of the antemucronal area. Scale bars 1 mm: A, C, E, G, H, J, K; 500 µm: I; 100 µm: B, D, F, L." figureDoi="http://doi.org/10.5281/zenodo.5304561" httpUri="https://zenodo.org/record/5304561/files/figure.png" pageId="14" pageNumber="15">Figures 5CL</figureCitation>
,
<figureCitation id="134543EBFFDAFF84FEB9FB81FE65B18D" box="[337,423,1133,1159]" captionStart="FIGURE 13" captionStartId="28.[151,256,1692,1716]" captionTargetBox="[243,1339,194,1667]" captionTargetId="figure-187@28.[243,1339,194,1667]" captionTargetPageId="28" captionText="FIGURE 13AB, Callochiton sp., S. Nilandu Atoll, Maldive, NHM 1903.4.11.71 (living specimen collected by J.S. Gardiner), close-up of the head and tail valves. CH, Lucilina carnosa (Kaas, 1979), valves, Maldive. CD, head valves, M13, showing differences in the number of ocelli rows, widths 3 and 3.1 mm, respectively. E, intermediate valve, M13, width 3.5 mm. F, tail valve, M13, width 3.3 mm. GH, anomalous tail valve, dorsal and posterior views, respectively, M20, width 3.2 mm. IK, Lucilina sp., valves, Maldive. I, head valve, M13, width 3.1 mm. J, intermediate valve, M20, width 4.7 mm. K, tail valve, M65, width 3.4 mm. LM, Craspedochiton cf. laqueatus (Sowerby, 1842), head and intermediate valves, Maldive, M65, widths 5.7 and 5.2 mm, respectively. N, Craspedochiton sp., tail valve, Maldive, M65, width 5.8 mm. OR, Cryptoplax burrowi E.A. Smith, 1884, NHM 1881.11.1.100, syntype, detached valves, coll. Coppinger, Port Molle, Queensland. OP, intermediate valve, length 4 mm, dorsal and right lateral views, respectively. Q, intermediate valve, length 4.2 mm. R, head valve, length 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5330696" httpUri="https://zenodo.org/record/5330696/files/figure.png" pageId="14" pageNumber="15">13CH</figureCitation>
)
</paragraph>
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<paragraph id="8BC15F6EFFDAFF84FF7FFB54FA59B1C5" blockId="14.[151,1436,1208,1391]" box="[151,1435,1208,1231]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB6E7840FFDAFF84FF7FFB54FA59B1C5" box="[151,1435,1208,1231]" pageId="14" pageNumber="15">
<treatmentCitation id="0ADF797FFFDAFF84FF7FFB54FDA1B1C5" author="Kaas, P." box="[151,611,1208,1231]" page="869" pageId="14" pageNumber="15" year="1979">
<taxonomicName id="4C7E24EDFFDAFF84FF7FFB54FDA1B1C5" ID-CoL="7CGJL" authority="Kaas 1979: 869" authorityName="Kaas" authorityPageNumber="869" authorityYear="1979" box="[151,611,1208,1231]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="carnosa" subGenus="Lucilina">
<emphasis id="B90A837CFFDAFF84FF7FFB54FE71B1C5" box="[151,435,1208,1231]" italics="true" pageId="14" pageNumber="15">Tonicia (Lucilina) carnosa</emphasis>
<bibRefCitation id="EFEF229FFFDAFF84FE52FB54FDA1B1C5" author="Kaas, P." box="[442,611,1208,1231]" pageId="14" pageNumber="15" pagination="855 - 879" refId="ref21829" refString="Kaas, P. (1979) The Chitons (Mollusca: Polyplacophora) of Mozambique. Annals of the Natal Museum, 23, 855 - 879." type="journal article" year="1979">Kaas 1979: 869</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 3, figs 110; 1986: 18, figs 4648; DellAngelo
<emphasis id="B90A837CFFDAFF84FB79FB54FB7DB1C5" box="[1169,1215,1208,1231]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 2004: 54, pl. 5, figs
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC15F6EFFDAFF84FF2FFB34FA5EB045" blockId="14.[151,1436,1208,1391]" pageId="14" pageNumber="15">
49;
<bibRefCitation id="EFEF229FFFDAFF84FF15FB34FE20B1E5" author="Kaas, P. &amp; Van Belle, R. A. &amp; Strack, H. L." box="[253,482,1240,1263]" pageId="14" pageNumber="15" refId="ref22247" refString="Kaas, P., Van Belle, R. A. &amp; Strack, H. L. (2006) Monograph of Living Chitons (Mollusca: Polyplacophora). Volume 6. Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae. Additions to Volumes 1 - 5. Brill, Leiden- Boston, 463 pp." type="book" year="2006">
Kaas
<emphasis id="B90A837CFFDAFF84FED0FB34FEA5B1E5" box="[312,359,1240,1263]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 2006: 323
</bibRefCitation>
, fig. 133.
<taxonomicName id="4C7E24EDFFDAFF84FF7FFB14FE81B005" ID-CoL="7CGJL" authorityName="Kaas" authorityYear="1979" box="[151,323,1272,1295]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="carnosa">
<emphasis id="B90A837CFFDAFF84FF7FFB14FE81B005" box="[151,323,1272,1295]" italics="true" pageId="14" pageNumber="15">Tonicia carnosa</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFEF229FFFDAFF84FEB8FB14FDDBB005" author="Ferreira, A. J." box="[336,537,1272,1295]" pageId="14" pageNumber="15" pagination="249 - 297" refId="ref21388" refString="Ferreira, A. J. (1983) Researches on the coast of Somalia. The chiton fauna (Mollusca Polyplacophora). Monitore Zoologico italiano, 9, 249 - 297." type="journal article" year="1983">Ferreira 1983: 272</bibRefCitation>
, 274.
<taxonomicName id="4C7E24EDFFDAFF84FF7FFAF4FE8EB025" ID-CoL="3WC4J" baseAuthorityName="Kaas" baseAuthorityYear="1979" box="[151,332,1304,1327]" class="Polyplacophora" family="Chitonidae" genus="Lucilina" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="carnosa">
<emphasis id="B90A837CFFDAFF84FF7FFAF4FE8EB025" box="[151,332,1304,1327]" italics="true" pageId="14" pageNumber="15">Lucilina carnosa</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFEF229FFFDAFF84FEB2FAF4FDC7B025" author="Kaas, P." box="[346,517,1304,1327]" pageId="14" pageNumber="15" pagination="367 - 375" refId="ref21942" refString="Kaas, P. (1996) Chitons (Mollusca: Polyplacophora) from the Seychelles with description of a new species. Zoologische Mededelingen, 70, 367 - 375." type="journal article" year="1996">Kaas 1996: 372</bibRefCitation>
;
<bibRefCitation id="EFEF229FFFDAFF84FDF8FAF4FD31B025" author="Schwabe, E." box="[528,755,1304,1327]" pageId="14" pageNumber="15" pagination="3143 - 3173" refId="ref23247" refString="Schwabe, E. (2004) The Polyplacophora (Mollusca) collected during the First International Marine Biodiversity Workshop for Rodrigues (western Indian Ocean), with the description of a new species. Journal of Natural History, 38, 3143 - 3173." type="journal article" year="2004">Schwabe 2004: 3156</bibRefCitation>
. Non
<taxonomicName id="4C7E24EDFFDAFF84FF38FAD4FE34B045" ID-CoL="7CGJL" authorityName="Kaas" authorityYear="1979" box="[208,502,1336,1359]" class="Polyplacophora" family="Chitonidae" genus="Tonicia" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="carnosa" subGenus="Lucilina">
<emphasis id="B90A837CFFDAFF84FF38FAD4FE34B045" box="[208,502,1336,1359]" italics="true" pageId="14" pageNumber="15">Tonicia (Lucilina) carnosa</emphasis>
</taxonomicName>
; Kaas 1985: 331;
<bibRefCitation id="EFEF229FFFDAFF84FD24FAD4FC11B045" author="Drivas, J. &amp; Jay, M." box="[716,979,1336,1359]" pageId="14" pageNumber="15" pagination="32 - 34" refId="ref21357" refString="Drivas, J. &amp; Jay, M. (1998) I Chitoni della Reunion. La Conchiglia, 30 (289 suppl.), 32 - 34." type="journal article" year="1998">Drivas &amp; Jay 1998: 32</bibRefCitation>
, fig. 3 [=
<taxonomicName id="4C7E24EDFFDAFF84FBA6FAD4FAA6B045" ID-CoL="3WC4S" authority="(Leloup, 1981)" baseAuthorityName="Leloup" baseAuthorityYear="1981" box="[1102,1380,1336,1359]" class="Polyplacophora" family="Chitonidae" genus="Lucilina" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="indica">
<emphasis id="B90A837CFFDAFF84FBA6FAD4FB70B045" box="[1102,1202,1336,1359]" italics="true" pageId="14" pageNumber="15">L. indica</emphasis>
(
<bibRefCitation id="EFEF229FFFDAFF84FB2DFAD4FA9EB045" author="Leloup, E." box="[1221,1372,1336,1359]" pageId="14" pageNumber="15" pagination="1 - 46" refId="ref22550" refString="Leloup, E. (1981) Chitons de Tulear, Reunion, Maurice et Tahiti. Bulletin Institut Royale des Sciences Naturelles de Belgique, 53 (3), 1 - 46, pls 1 - 4." type="journal article" year="1981">Leloup, 1981</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B90A837CFFDAFF84FA9CFAD4FA5EB045" box="[1396,1436,1336,1359]" italics="true" pageId="14" pageNumber="15">fide</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3640CE5FFDAFF84FF2FFAB4FE75B065" box="[199,439,1368,1391]" pageId="14" pageNumber="15" type="description">
<paragraph id="8BC15F6EFFDAFF84FF2FFAB4FE75B065" blockId="14.[151,1436,1208,1391]" box="[199,439,1368,1391]" pageId="14" pageNumber="15">
<bibRefCitation id="EFEF229FFFDAFF84FF2FFAB4FE6EB065" author="Schwabe, E." box="[199,428,1368,1391]" pageId="14" pageNumber="15" pagination="3143 - 3173" refId="ref23247" refString="Schwabe, E. (2004) The Polyplacophora (Mollusca) collected during the First International Marine Biodiversity Workshop for Rodrigues (western Indian Ocean), with the description of a new species. Journal of Natural History, 38, 3143 - 3173." type="journal article" year="2004">Schwabe 2004: 3153</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C3640CE5FFDAFF84FF7FFA71FD49B3E8" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BC15F6EFFDAFF84FF7FFA71FAA5B0BD" blockId="14.[151,1436,1437,2023]" box="[151,1383,1437,1463]" pageId="14" pageNumber="15">
<emphasis id="B90A837CFFDAFF84FF7FFA71FF1FB0BD" bold="true" box="[151,221,1437,1463]" pageId="14" pageNumber="15">Type:</emphasis>
<typeStatus id="54C5E1CCFFDAFF84FF0DFA71FE90B0BD" box="[229,338,1437,1463]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
NM H2576 (21.7 x
<quantity id="4C86F28BFFDAFF84FDAAFA71FD6CB0BD" box="[578,686,1437,1463]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="14" pageNumber="15" unit="mm" value="11.1">11.1 mm</quantity>
);
<typeStatus id="54C5E1CCFFDAFF84FD2AFA71FCE4B0BD" box="[706,806,1437,1463]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
NM H2575 (20.5 x
<quantity id="4C86F28BFFDAFF84FBFEFA71FBB0B0BD" box="[1046,1138,1437,1463]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="14" pageNumber="15" unit="mm" value="9.5">9.5 mm</quantity>
, now disarticulated).
</paragraph>
<paragraph id="8BC15F6EFFDAFF84FF2DFA2EFD09B0D6" blockId="14.[151,1436,1437,2023]" box="[197,715,1474,1500]" pageId="14" pageNumber="15">
<materialsCitation id="3B165533FFDAFF84FF2DFA2EFD0AB0D6" box="[197,712,1474,1500]" country="Mozambique" location="Conducia Bay" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<emphasis id="B90A837CFFDAFF84FF2DFA2EFEACB0D6" bold="true" box="[197,366,1474,1500]" pageId="14" pageNumber="15">
<typeStatus id="54C5E1CCFFDAFF84FF2DFA2EFEC0B0D6" box="[197,258,1474,1500]" pageId="14" pageNumber="15">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F3691FFEFFDAFF84FE9EFA2EFDD1B0D6" box="[374,531,1474,1500]" name="Mozambique" pageId="14" pageNumber="15">Mozambique</collectingCountry>
,
<location id="8EA109B5FFDAFF84FDF7FA2EFD0AB0D6" LSID="urn:lsid:plazi:treatment:03D7EE78FFDAFF85FF7FFBA6FE75B746:8EA109B5FFDAFF84FDF7FA2EFD0AB0D6" box="[543,712,1474,1500]" country="Mozambique" name="Conducia Bay" pageId="14" pageNumber="15">Conducia Bay</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC15F6EFFDAFF84FF2DFA0BFD49B3E8" blockId="14.[151,1436,1437,2023]" pageId="14" pageNumber="15">
<emphasis id="B90A837CFFDAFF84FF2DFA0BFE79B30B" bold="true" box="[197,443,1511,1537]" pageId="14" pageNumber="15">Material examined:</emphasis>
M13: 65 valves (11 head, 51 intermediate and 3 tail); M17: 5 valves (3 intermediate and 2 tail); M19: 3 valves (1 intermediate and 2 tail); M20: 18 valves (4 head, 12 intermediate and 2 tail); M23: 6 valves (1 head, 4 intermediate and 1 tail); M26: 10 valves (1 head, 8 intermediate and 1 tail); M65: 11 valves (10 intermediate and 1 tail); MLD01A: 1 tail valve; MLD02A: 25 valves (4 head, 16 intermediate and 5 tail); MLD03A: 9 valves (8 intermediate and 1 tail); MLD04B: 2 head valves; MLD05A: 3 valves (2 head and 1 intermediate); MLD05B: 8 valves (1 head and 7 intermediate); LB04: 1 intermediate valve; LB05: 31 valves (4 head, 23 intermediate and 4 tail).
</paragraph>
</subSubSection>
<subSubSection id="C3640CE5FFDAFF85FF2DF901FC64B708" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BC15F6EFFDAFF84FF2DF901FAEDB20D" blockId="14.[151,1436,1437,2023]" box="[197,1327,1773,1799]" pageId="14" pageNumber="15">
<emphasis id="B90A837CFFDAFF84FF2DF901FE82B20D" bold="true" box="[197,320,1773,1799]" pageId="14" pageNumber="15">Remarks:</emphasis>
Detailed descriptions of this species are given by
<bibRefCitation id="EFEF229FFFDAFF84FC64F901FBDEB20D" author="Kaas, P." box="[908,1052,1773,1799]" pageId="14" pageNumber="15" pagination="855 - 879" refId="ref21829" refString="Kaas, P. (1979) The Chitons (Mollusca: Polyplacophora) of Mozambique. Annals of the Natal Museum, 23, 855 - 879." type="journal article" year="1979">Kaas (1979)</bibRefCitation>
and
<bibRefCitation id="EFEF229FFFDAFF84FBBDF901FAEEB20D" author="Kaas, P. &amp; Van Belle, R. A. &amp; Strack, H. L." box="[1109,1324,1773,1799]" pageId="14" pageNumber="15" refId="ref22247" refString="Kaas, P., Van Belle, R. A. &amp; Strack, H. L. (2006) Monograph of Living Chitons (Mollusca: Polyplacophora). Volume 6. Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae. Additions to Volumes 1 - 5. Brill, Leiden- Boston, 463 pp." type="book" year="2006">
Kaas
<emphasis id="B90A837CFFDAFF84FB70F901FB09B20C" box="[1176,1227,1773,1798]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (2006)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BC15F6EFFDAFF85FF2DF8FEFC1AB466" blockId="14.[151,1436,1437,2023]" lastBlockId="15.[151,1436,152,588]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
The valves present a certain degree of variability in some characters. Particularly, the sculpture of the greater part of head valves agrees with the original description (
<bibRefCitation id="EFEF229FFFDAFF84FC92F8D4FBF1B258" author="Kaas, P." box="[890,1075,1848,1874]" pageId="14" pageNumber="15" pagination="855 - 879" refId="ref21829" refString="Kaas, P. (1979) The Chitons (Mollusca: Polyplacophora) of Mozambique. Annals of the Natal Museum, 23, 855 - 879." type="journal article" year="1979">Kaas 1979: 869</bibRefCitation>
<emphasis id="B90A837CFFDAFF84FBAEF8D4FC05B296" italics="true" pageId="14" pageNumber="15">
about 10 low, wide, radiating riblets, decorated with flat, inverted V-shaped warts directed towards umbo.
<taxonomicName id="4C7E24EDFFDAFF84FBFDF8B1FACCB27C" box="[1045,1294,1885,1910]" class="Polyplacophora" family="Ischnochitonidae" genus="Ischnochiton" kingdom="Animalia" order="Chitonida" pageId="14" pageNumber="15" phylum="Mollusca" rank="subSpecies" species="shallow" subSpecies="interstices">In shallow interstices</taxonomicName>
eight single rows of ocelli, each splitting up into two or three rows towards margin
</emphasis>
”) (
<figureCitation id="134543EBFFDAFF84FC06F86EFB8FB296" box="[1006,1101,1922,1948]" captionStart="FIGURE 5" captionStartId="15.[151,257,1381,1405]" captionTargetBox="[151,1436,645,1360]" captionTargetId="figure-400@15.[151,1436,645,1360]" captionTargetPageId="15" captionText="FIGURE 5AB, Lucilina perligera Thiele, 1909, Maldive, M13, intermediate valve. A, dorsal view. B, tegmentum granules of the central area. CL, Lucilina carnosa (Kaas, 1979), Maldive. CD, head valve, MLD02A. C, dorsal view. D, detail of the sculpture. EF, intermediate valve, MLD02A. E, dorsal view. F, detail of the sculpture. GI, intermediate valve, M23. G, dorsal view. H, frontal view. I, detail of the sculpture. JL, tail valve, MLD02A. J. dorsal view. K, lateral view. L, detail of the sculpture of the antemucronal area. Scale bars 1 mm: A, C, E, G, H, J, K; 500 µm: I; 100 µm: B, D, F, L." figureDoi="http://doi.org/10.5281/zenodo.5304561" httpUri="https://zenodo.org/record/5304561/files/figure.png" pageId="14" pageNumber="15">Figs 5C</figureCitation>
,
<figureCitation id="134543EBFFDAFF84FBB0F86EFB4CB296" box="[1112,1166,1922,1948]" captionStart="FIGURE 13" captionStartId="28.[151,256,1692,1716]" captionTargetBox="[243,1339,194,1667]" captionTargetId="figure-187@28.[243,1339,194,1667]" captionTargetPageId="28" captionText="FIGURE 13AB, Callochiton sp., S. Nilandu Atoll, Maldive, NHM 1903.4.11.71 (living specimen collected by J.S. Gardiner), close-up of the head and tail valves. CH, Lucilina carnosa (Kaas, 1979), valves, Maldive. CD, head valves, M13, showing differences in the number of ocelli rows, widths 3 and 3.1 mm, respectively. E, intermediate valve, M13, width 3.5 mm. F, tail valve, M13, width 3.3 mm. GH, anomalous tail valve, dorsal and posterior views, respectively, M20, width 3.2 mm. IK, Lucilina sp., valves, Maldive. I, head valve, M13, width 3.1 mm. J, intermediate valve, M20, width 4.7 mm. K, tail valve, M65, width 3.4 mm. LM, Craspedochiton cf. laqueatus (Sowerby, 1842), head and intermediate valves, Maldive, M65, widths 5.7 and 5.2 mm, respectively. N, Craspedochiton sp., tail valve, Maldive, M65, width 5.8 mm. OR, Cryptoplax burrowi E.A. Smith, 1884, NHM 1881.11.1.100, syntype, detached valves, coll. Coppinger, Port Molle, Queensland. OP, intermediate valve, length 4 mm, dorsal and right lateral views, respectively. Q, intermediate valve, length 4.2 mm. R, head valve, length 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5330696" httpUri="https://zenodo.org/record/5330696/files/figure.png" pageId="14" pageNumber="15">13D</figureCitation>
), but some head valves show one or two more ocelli rows, with a certain degree of variability (
<figureCitation id="134543EBFFDAFF84FC30F844FB81B2C8" box="[984,1091,1960,1986]" captionStart="FIGURE 13" captionStartId="28.[151,256,1692,1716]" captionTargetBox="[243,1339,194,1667]" captionTargetId="figure-187@28.[243,1339,194,1667]" captionTargetPageId="28" captionText="FIGURE 13AB, Callochiton sp., S. Nilandu Atoll, Maldive, NHM 1903.4.11.71 (living specimen collected by J.S. Gardiner), close-up of the head and tail valves. CH, Lucilina carnosa (Kaas, 1979), valves, Maldive. CD, head valves, M13, showing differences in the number of ocelli rows, widths 3 and 3.1 mm, respectively. E, intermediate valve, M13, width 3.5 mm. F, tail valve, M13, width 3.3 mm. GH, anomalous tail valve, dorsal and posterior views, respectively, M20, width 3.2 mm. IK, Lucilina sp., valves, Maldive. I, head valve, M13, width 3.1 mm. J, intermediate valve, M20, width 4.7 mm. K, tail valve, M65, width 3.4 mm. LM, Craspedochiton cf. laqueatus (Sowerby, 1842), head and intermediate valves, Maldive, M65, widths 5.7 and 5.2 mm, respectively. N, Craspedochiton sp., tail valve, Maldive, M65, width 5.8 mm. OR, Cryptoplax burrowi E.A. Smith, 1884, NHM 1881.11.1.100, syntype, detached valves, coll. Coppinger, Port Molle, Queensland. OP, intermediate valve, length 4 mm, dorsal and right lateral views, respectively. Q, intermediate valve, length 4.2 mm. R, head valve, length 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5330696" httpUri="https://zenodo.org/record/5330696/files/figure.png" pageId="14" pageNumber="15">Fig. 13C</figureCitation>
). The number of longitudinal grooves in the pleural areas of intermediate valves is variable, some valves show few and shorter grooves never reaching the front margin (
<figureCitation id="134543EBFFDBFF85FDFFFF74FD4EB5B8" box="[535,652,152,178]" captionStart="FIGURE 5" captionStartId="15.[151,257,1381,1405]" captionTargetBox="[151,1436,645,1360]" captionTargetId="figure-400@15.[151,1436,645,1360]" captionTargetPageId="15" captionText="FIGURE 5AB, Lucilina perligera Thiele, 1909, Maldive, M13, intermediate valve. A, dorsal view. B, tegmentum granules of the central area. CL, Lucilina carnosa (Kaas, 1979), Maldive. CD, head valve, MLD02A. C, dorsal view. D, detail of the sculpture. EF, intermediate valve, MLD02A. E, dorsal view. F, detail of the sculpture. GI, intermediate valve, M23. G, dorsal view. H, frontal view. I, detail of the sculpture. JL, tail valve, MLD02A. J. dorsal view. K, lateral view. L, detail of the sculpture of the antemucronal area. Scale bars 1 mm: A, C, E, G, H, J, K; 500 µm: I; 100 µm: B, D, F, L." figureDoi="http://doi.org/10.5281/zenodo.5304561" httpUri="https://zenodo.org/record/5304561/files/figure.png" pageId="15" pageNumber="16">Figs 5G, I</figureCitation>
), and so nearly the whole central area looks smooth (compare with
<figureCitation id="134543EBFFDBFF85FF7FFF51FED6B5DD" box="[151,276,189,215]" captionStart="FIGURE 5" captionStartId="15.[151,257,1381,1405]" captionTargetBox="[151,1436,645,1360]" captionTargetId="figure-400@15.[151,1436,645,1360]" captionTargetPageId="15" captionText="FIGURE 5AB, Lucilina perligera Thiele, 1909, Maldive, M13, intermediate valve. A, dorsal view. B, tegmentum granules of the central area. CL, Lucilina carnosa (Kaas, 1979), Maldive. CD, head valve, MLD02A. C, dorsal view. D, detail of the sculpture. EF, intermediate valve, MLD02A. E, dorsal view. F, detail of the sculpture. GI, intermediate valve, M23. G, dorsal view. H, frontal view. I, detail of the sculpture. JL, tail valve, MLD02A. J. dorsal view. K, lateral view. L, detail of the sculpture of the antemucronal area. Scale bars 1 mm: A, C, E, G, H, J, K; 500 µm: I; 100 µm: B, D, F, L." figureDoi="http://doi.org/10.5281/zenodo.5304561" httpUri="https://zenodo.org/record/5304561/files/figure.png" pageId="15" pageNumber="16">Figs 5EF</figureCitation>
,
<figureCitation id="134543EBFFDBFF85FEC9FF51FE96B5DD" box="[289,340,189,215]" captionStart="FIGURE 13" captionStartId="28.[151,256,1692,1716]" captionTargetBox="[243,1339,194,1667]" captionTargetId="figure-187@28.[243,1339,194,1667]" captionTargetPageId="28" captionText="FIGURE 13AB, Callochiton sp., S. Nilandu Atoll, Maldive, NHM 1903.4.11.71 (living specimen collected by J.S. Gardiner), close-up of the head and tail valves. CH, Lucilina carnosa (Kaas, 1979), valves, Maldive. CD, head valves, M13, showing differences in the number of ocelli rows, widths 3 and 3.1 mm, respectively. E, intermediate valve, M13, width 3.5 mm. F, tail valve, M13, width 3.3 mm. GH, anomalous tail valve, dorsal and posterior views, respectively, M20, width 3.2 mm. IK, Lucilina sp., valves, Maldive. I, head valve, M13, width 3.1 mm. J, intermediate valve, M20, width 4.7 mm. K, tail valve, M65, width 3.4 mm. LM, Craspedochiton cf. laqueatus (Sowerby, 1842), head and intermediate valves, Maldive, M65, widths 5.7 and 5.2 mm, respectively. N, Craspedochiton sp., tail valve, Maldive, M65, width 5.8 mm. OR, Cryptoplax burrowi E.A. Smith, 1884, NHM 1881.11.1.100, syntype, detached valves, coll. Coppinger, Port Molle, Queensland. OP, intermediate valve, length 4 mm, dorsal and right lateral views, respectively. Q, intermediate valve, length 4.2 mm. R, head valve, length 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5330696" httpUri="https://zenodo.org/record/5330696/files/figure.png" pageId="15" pageNumber="16">13E</figureCitation>
, illustrating a typical intermediate valve). The same occurs in the tail valve (a typical one, agreeing to the species description, is illustrated in
<figureCitation id="134543EBFFDBFF85FD12FF0EFCB0B5F6" box="[762,882,226,252]" captionStart="FIGURE 5" captionStartId="15.[151,257,1381,1405]" captionTargetBox="[151,1436,645,1360]" captionTargetId="figure-400@15.[151,1436,645,1360]" captionTargetPageId="15" captionText="FIGURE 5AB, Lucilina perligera Thiele, 1909, Maldive, M13, intermediate valve. A, dorsal view. B, tegmentum granules of the central area. CL, Lucilina carnosa (Kaas, 1979), Maldive. CD, head valve, MLD02A. C, dorsal view. D, detail of the sculpture. EF, intermediate valve, MLD02A. E, dorsal view. F, detail of the sculpture. GI, intermediate valve, M23. G, dorsal view. H, frontal view. I, detail of the sculpture. JL, tail valve, MLD02A. J. dorsal view. K, lateral view. L, detail of the sculpture of the antemucronal area. Scale bars 1 mm: A, C, E, G, H, J, K; 500 µm: I; 100 µm: B, D, F, L." figureDoi="http://doi.org/10.5281/zenodo.5304561" httpUri="https://zenodo.org/record/5304561/files/figure.png" pageId="15" pageNumber="16">Figs 5J, L</figureCitation>
,
<figureCitation id="134543EBFFDBFF85FC95FF0EFC6CB5F6" box="[893,942,226,252]" captionStart="FIGURE 13" captionStartId="28.[151,256,1692,1716]" captionTargetBox="[243,1339,194,1667]" captionTargetId="figure-187@28.[243,1339,194,1667]" captionTargetPageId="28" captionText="FIGURE 13AB, Callochiton sp., S. Nilandu Atoll, Maldive, NHM 1903.4.11.71 (living specimen collected by J.S. Gardiner), close-up of the head and tail valves. CH, Lucilina carnosa (Kaas, 1979), valves, Maldive. CD, head valves, M13, showing differences in the number of ocelli rows, widths 3 and 3.1 mm, respectively. E, intermediate valve, M13, width 3.5 mm. F, tail valve, M13, width 3.3 mm. GH, anomalous tail valve, dorsal and posterior views, respectively, M20, width 3.2 mm. IK, Lucilina sp., valves, Maldive. I, head valve, M13, width 3.1 mm. J, intermediate valve, M20, width 4.7 mm. K, tail valve, M65, width 3.4 mm. LM, Craspedochiton cf. laqueatus (Sowerby, 1842), head and intermediate valves, Maldive, M65, widths 5.7 and 5.2 mm, respectively. N, Craspedochiton sp., tail valve, Maldive, M65, width 5.8 mm. OR, Cryptoplax burrowi E.A. Smith, 1884, NHM 1881.11.1.100, syntype, detached valves, coll. Coppinger, Port Molle, Queensland. OP, intermediate valve, length 4 mm, dorsal and right lateral views, respectively. Q, intermediate valve, length 4.2 mm. R, head valve, length 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5330696" httpUri="https://zenodo.org/record/5330696/files/figure.png" pageId="15" pageNumber="16">13F</figureCitation>
). Some appreciable difference is recorded in the tail valve, where the front margin is straight or at most slightly concave, never convex, and 1013 irregularly and inequidistant slits are present (slit formula 8/1/
<quantity id="4C86F28BFFDBFF85FC7EFEC1FBC6B44D" box="[918,1028,301,327]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.302" metricValueMin="2.54" pageId="15" pageNumber="16" unit="in" value="11.5" valueMax="13.0" valueMin="10.0">1013 in</quantity>
contrast to the 8/1/810 formula reported by
<bibRefCitation id="EFEF229FFFDBFF85FECCFEBEFE20B466" author="Kaas, P. &amp; Van Belle, R. A. &amp; Strack, H. L." box="[292,482,338,364]" pageId="15" pageNumber="16" refId="ref22247" refString="Kaas, P., Van Belle, R. A. &amp; Strack, H. L. (2006) Monograph of Living Chitons (Mollusca: Polyplacophora). Volume 6. Suborder Ischnochitonina (concluded): Schizochitonidae; Chitonidae. Additions to Volumes 1 - 5. Brill, Leiden- Boston, 463 pp." type="book" year="2006">
Kaas
<emphasis id="B90A837CFFDBFF85FE8EFEBEFE63B461" box="[358,417,338,363]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2006
</bibRefCitation>
and 8/1/912 by DellAngelo
<emphasis id="B90A837CFFDBFF85FCA2FEBEFCBFB461" box="[842,893,338,363]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
. 2004).
</paragraph>
<paragraph id="8BC15F6EFFDBFF85FF2DFE94FB31B498" blockId="15.[151,1436,152,588]" box="[197,1267,376,402]" pageId="15" pageNumber="16">The differences described above seemingly fall within the limits of intraspecific variability.</paragraph>
<paragraph id="8BC15F6EFFDBFF85FF2DFE71FBA1B4D6" blockId="15.[151,1436,152,588]" pageId="15" pageNumber="16">
A tail valve from M20 shows a superficial incision in the postmucronal area, which caused the development of a new insertion plate under the one already existing (
<figureCitation id="134543EBFFDBFF85FC28FE2EFB94B4D6" box="[960,1110,450,476]" captionStart="FIGURE 13" captionStartId="28.[151,256,1692,1716]" captionTargetBox="[243,1339,194,1667]" captionTargetId="figure-187@28.[243,1339,194,1667]" captionTargetPageId="28" captionText="FIGURE 13AB, Callochiton sp., S. Nilandu Atoll, Maldive, NHM 1903.4.11.71 (living specimen collected by J.S. Gardiner), close-up of the head and tail valves. CH, Lucilina carnosa (Kaas, 1979), valves, Maldive. CD, head valves, M13, showing differences in the number of ocelli rows, widths 3 and 3.1 mm, respectively. E, intermediate valve, M13, width 3.5 mm. F, tail valve, M13, width 3.3 mm. GH, anomalous tail valve, dorsal and posterior views, respectively, M20, width 3.2 mm. IK, Lucilina sp., valves, Maldive. I, head valve, M13, width 3.1 mm. J, intermediate valve, M20, width 4.7 mm. K, tail valve, M65, width 3.4 mm. LM, Craspedochiton cf. laqueatus (Sowerby, 1842), head and intermediate valves, Maldive, M65, widths 5.7 and 5.2 mm, respectively. N, Craspedochiton sp., tail valve, Maldive, M65, width 5.8 mm. OR, Cryptoplax burrowi E.A. Smith, 1884, NHM 1881.11.1.100, syntype, detached valves, coll. Coppinger, Port Molle, Queensland. OP, intermediate valve, length 4 mm, dorsal and right lateral views, respectively. Q, intermediate valve, length 4.2 mm. R, head valve, length 3.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5330696" httpUri="https://zenodo.org/record/5330696/files/figure.png" pageId="15" pageNumber="16">Figs 13GH</figureCitation>
).
</paragraph>
<paragraph id="8BC15F6EFFDBFF85FF2DFE04FC64B708" blockId="15.[151,1436,152,588]" box="[197,934,488,514]" pageId="15" pageNumber="16">
This is the first report of this species from the
<collectingCountry id="F3691FFEFFDBFF85FD0BFE04FC61B708" box="[739,931,488,514]" name="Maldives" pageId="15" pageNumber="16">Maldive Islands</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3640CE5FFDBFF85FF2DFDE1FE75B746" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BC15F6EFFDBFF85FF2DFDE1FE75B746" blockId="15.[151,1436,152,588]" pageId="15" pageNumber="16">
<emphasis id="B90A837CFFDBFF85FF2DFDE1FEA5B72D" bold="true" box="[197,359,525,551]" pageId="15" pageNumber="16">Distribution:</emphasis>
<taxonomicName id="4C7E24EDFFDBFF85FE86FDE1FDF7B72C" baseAuthorityName="Kaas" baseAuthorityYear="1979" box="[366,565,525,550]" class="Polyplacophora" family="Chitonidae" genus="Lucilina" kingdom="Animalia" order="Chitonida" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="carnosa">
<emphasis id="B90A837CFFDBFF85FE86FDE1FDF7B72C" box="[366,565,525,550]" italics="true" pageId="15" pageNumber="16">Lucilina carnosa</emphasis>
</taxonomicName>
is known from
<collectingCountry id="F3691FFEFFDBFF85FD05FDE1FCBEB72D" box="[749,892,525,551]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
,
<collectingCountry id="F3691FFEFFDBFF85FC6FFDE1FBE7B72D" box="[903,1061,525,551]" name="Mozambique" pageId="15" pageNumber="16">Mozambique</collectingCountry>
,
<collectingCountry id="F3691FFEFFDBFF85FBD8FDE1FB54B72D" box="[1072,1174,525,551]" name="Réunion" pageId="15" pageNumber="16">Réunion</collectingCountry>
,
<collectingCountry id="F3691FFEFFDBFF85FB4AFDE1FAE0B72D" box="[1186,1314,525,551]" name="Seychelles" pageId="15" pageNumber="16">Seychelles</collectingCountry>
, Amirante and the
<collectingCountry id="F3691FFEFFDBFF85FF1CFDDEFE76B746" box="[244,436,562,588]" name="Maldives" pageId="15" pageNumber="16">Maldive Islands</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>