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<document id="8B41CBBA359A4223071B940FBB11461D" ID-CLB-Dataset="2931" ID-DOI="10.11646/zootaxa.5079.1.1" ID-GBIF-Dataset="010695ef-a142-42ad-81d7-2e7ed1f7e89e" ID-ISSN="1175-5326" ID-Zenodo-Dep="5766402" ID-ZooBank="2B118B81-7353-4A2C-A892-5466DFC83230" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638952199435" checkinUser="plazi" docAuthor="Takano, Kohei Takenaka, Gao, Jian-Jun, Hu, Yao-Guang, Li, Nan-Nan, Yafuso, Masako, Suwito, Awit, Repin, Rimi, Pungga, Runi Anak Sylvester, Meleng, Paulus Ak, Kaliang, Clement Het, Chong, Lucy &amp; Toda, Masanori J." docDate="2021" docId="03D75C2D454F2E19FF526092B1D2B1CD" docLanguage="en" docName="zootaxa.5079.1.1.pdf" docOrigin="Zootaxa 5079 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Colocasiomyia sabahana Takano &amp; Gao &amp; Hu &amp; Li &amp; Yafuso &amp; Suwito &amp; Repin &amp; Pungga &amp; Meleng &amp; Kaliang &amp; Chong &amp; Toda 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="39" masterDocId="FFEE2455456A2E3FFFC56005B34CB33C" masterDocTitle="Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species" masterLastPageNumber="70" masterPageNumber="1" pageNumber="38" updateTime="1699472124804" updateUser="ExternalLinkService">
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<mods:title id="5E156E50A22D2B7684B13BE352326AEF">Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species</mods:title>
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<mods:namePart id="29AB92B096BB1C6354467177814FDB4E">Takano, Kohei Takenaka</mods:namePart>
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<mods:namePart id="43501670EB249B9037AD061E917F28A1">Hu, Yao-Guang</mods:namePart>
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<mods:namePart id="809D0B90D32E1A6A5E10A752FA824245">Li, Nan-Nan</mods:namePart>
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<mods:namePart id="9A6032377FA8C0DF7C3E04E197B480F2">Yafuso, Masako</mods:namePart>
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<mods:namePart id="8E6E641421C7E527346340548B9FC42E">Suwito, Awit</mods:namePart>
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<mods:namePart id="F73727FDA38FEE7FAAF81AC6DE3F5C8B">Repin, Rimi</mods:namePart>
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<mods:namePart id="E846614B2D2BC5126962FFD6C56C5030">Pungga, Runi Anak Sylvester</mods:namePart>
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<mods:namePart id="33C5B430BB3B7EE0399615A40E53AC62">Meleng, Paulus Ak</mods:namePart>
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<mods:namePart id="98FB1F184627C9AD7ABBD1CB27E56410">Kaliang, Clement Het</mods:namePart>
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<mods:namePart id="5BD148FAFA7291E76ECD935B4E493B5D">Chong, Lucy</mods:namePart>
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<mods:namePart id="C55604BA02096695013AC1D3574E4026">Toda, Masanori J.</mods:namePart>
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<mods:title id="0D3A897C5E9A1A39868BF11A9FDE4F92">Zootaxa</mods:title>
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<mods:date id="BEB01C28F1637E53E2D419346665D829">2021</mods:date>
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<mods:number id="2C58E00554DF20C3F5018A7B9A6E2896">2021-12-08</mods:number>
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<treatment id="03D75C2D454F2E19FF526092B1D2B1CD" ID-DOI="http://doi.org/10.5281/zenodo.5786181" ID-GBIF-Taxon="191651673" ID-Zenodo-Dep="5786181" LSID="urn:lsid:plazi:treatment:03D75C2D454F2E19FF526092B1D2B1CD" httpUri="http://treatment.plazi.org/id/03D75C2D454F2E19FF526092B1D2B1CD" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<subSubSection id="C364BEB0454F2E1AFF526092B1DBB38D" box="[151,663,151,178]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8BC1ED3B454F2E1AFF526092B1DBB38D" blockId="37.[151,663,151,214]" box="[151,663,151,178]" pageId="37" pageNumber="38">
<emphasis id="B90A3129454F2E1AFF526092B1DBB38D" bold="true" box="[151,663,151,178]" pageId="37" pageNumber="38">
9)
<taxonomicName id="4C7E96B8454F2E1AFF726092B160B38D" ID-CoL="9HY7W" authority="Toda" authorityName="Takano &amp; Gao &amp; Hu &amp; Li &amp; Yafuso &amp; Suwito &amp; Repin &amp; Pungga &amp; Meleng &amp; Kaliang &amp; Chong &amp; Toda" authorityYear="2021" box="[183,556,151,178]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="sabahana" status="sp. nov.">
<emphasis id="B90A3129454F2E1AFF726092B2A9B38E" bold="true" box="[183,485,151,178]" italics="true" pageId="37" pageNumber="38">Colocasiomyia sabahana</emphasis>
Toda
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,
<taxonomicNameLabel id="A2398C52454F2E1AFDFC6092B1DBB38D" box="[569,663,151,177]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C364BEB0454F2E1AFF5260BEB12AB3E9" box="[151,614,187,214]" pageId="37" pageNumber="38" type="description">
<paragraph id="8BC1ED3B454F2E1AFF5260BEB12AB3E9" blockId="37.[151,663,151,214]" box="[151,614,187,214]" pageId="37" pageNumber="38">
(
<figureCitation id="1345F1BE454F2E1AFF5A60BEB3B5B3E9" box="[159,249,187,213]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson &amp; Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada &amp; Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson &amp; Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda &amp; Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="37" pageNumber="38">Figs 9L</figureCitation>
,
<figureCitation id="1345F1BE454F2E1AFEC060BEB207B3E9" box="[261,331,187,213]" captionStart="FIGURE 11" captionStartId="16.[151,250,1315,1339]" captionTargetBox="[151,1435,237,1292]" captionTargetId="figure-43@16.[151,1435,237,1292]" captionTargetPageId="16" captionText="FIGURE 11. Characters 40‒47 on the phallic organs of Colocasiomyia colocasiae (Duda) (A, lateral view; B, ventral view; C, hypandrium and gonocoxites, oblique lateral view), C. xenalocasiae (Okada) (D, lateral view; E, ventral view), C. sabahana sp. nov. (F, lateral view; G, ventral view) and C. cristata de Meijere (H, lateral view; I, ventral view).Abbreviations: aed, aedeagus; goncx, gonocoxite; hypd, hypandrium; phap, phallapodeme; ph sh, phallal sheath. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766424" httpUri="https://zenodo.org/record/5766424/files/figure.png" pageId="37" pageNumber="38">11F,G</figureCitation>
,
<figureCitation id="1345F1BE454F2E1AFE9360BEB231B3EA" box="[342,381,187,214]" captionStart="FIGURE 15" captionStartId="21.[151,250,1936,1960]" captionTargetBox="[151,1435,860,1912]" captionTargetId="figure-242@21.[151,1435,860,1912]" captionTargetPageId="21" captionText="FIGURE 15. Habitus (upper, lateral view; lower, dorsal view) of species of the Colocasiomyia cristata subgroup.A, C. ecornuta sp. nov. ♂; B, C. grandis sp. nov. ♂; C, C. vieti sp. nov. ♂; D, C. cristata de Meijere ♂; E, C. sulawesiana Okada &amp; Yafuso ♂; F, C. kinabaluana sp. nov. ♂; G, C. kotana sp. nov. ♂; and H, C. matthewsi sp. nov. ♂. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766434" httpUri="https://zenodo.org/record/5766434/files/figure.png" pageId="37" pageNumber="38">15I</figureCitation>
,
<figureCitation id="1345F1BE454F2E1AFE4F60BEB112B3E9" box="[394,606,187,213]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="37" pageNumber="38">24C,E,G,K,M,O,Q</figureCitation>
)
</paragraph>
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<paragraph id="8BC1ED3B454F2E1AFF5260FFB7B5B22E" blockId="37.[151,1273,250,274]" box="[151,1273,250,274]" pageId="37" pageNumber="38">
<treatmentCitationGroup id="AB6ECA15454F2E1AFF5260FFB7B5B22E" box="[151,1273,250,274]" pageId="37" pageNumber="38">
<taxonomicName id="4C7E96B8454F2E1AFF5260FFB1A0B22E" authority=": Sultana et al., 2006: 694" authorityName="Sultana" authorityPageNumber="694" authorityYear="2006" box="[151,748,250,274]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="sp.4">
<emphasis id="B90A3129454F2E1AFF5260FFB263B22E" box="[151,303,250,274]" italics="true" pageId="37" pageNumber="38">Colocasiomyia</emphasis>
sp.4
<treatmentCitation id="0ADFCB2A454F2E1AFEA060FFB1A0B22E" author="Sultana, F. &amp; Hu, Y. G. &amp; Toda, M. J. &amp; Takenaka, K. &amp; Yafuso, M." box="[357,748,250,274]" page="694" pageId="37" pageNumber="38" year="2006">
aff.
<emphasis id="B90A3129454F2E1AFE4B60FFB2A9B22E" box="[398,485,250,274]" italics="true" pageId="37" pageNumber="38">diconica</emphasis>
:
<bibRefCitation id="EFEF90CA454F2E1AFE3760FFB1A0B22E" author="Sultana, F. &amp; Hu, Y. G. &amp; Toda, M. J. &amp; Takenaka, K. &amp; Yafuso, M." box="[498,748,250,274]" pageId="37" pageNumber="38" pagination="684 - 702" refId="ref33499" refString="Sultana, F., Hu, Y. G., Toda, M. J., Takenaka, K. &amp; Yafuso, M. (2006) Phylogeny and classification of Colocasiomyia (Diptera, Drosophilidae), and its evolution of pollination mutualism with aroid plants. Systematic Entomology, 31, 684 - 702. https: // doi. org / 10.1111 / j. 1365 - 3113.2006.00344. x" type="journal article" year="2006">
Sultana
<emphasis id="B90A3129454F2E1AFD8060FEB13DB22E" box="[581,625,250,274]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
., 2006: 694
</bibRefCitation>
</treatmentCitation>
</taxonomicName>
,
<treatmentCitation id="0ADFCB2A454F2E1AFD3360FFB74CB22E" author="Toda, M. J. &amp; Lakim, M. B." box="[758,1024,250,274]" page="264" pageId="37" pageNumber="38" year="2011">
<bibRefCitation id="EFEF90CA454F2E1AFD3360FFB74CB22E" author="Toda, M. J. &amp; Lakim, M. B." box="[758,1024,250,274]" pageId="37" pageNumber="38" pagination="262 - 270" refId="ref33989" refString="Toda, M. J. &amp; Lakim, M. B. (2011) Genus Colocasiomyia (Drosophilidae: Diptera) in Sabah, Bornean Malaysia: high species diversity and use of host aroid inflorescences. Entomological Science, 14, 262 - 270. https: // doi. org / 10.1111 / j. 1479 - 8298.2011.00452. x" type="journal article" year="2011">Toda &amp; Lakim, 2011: 264</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADFCB2A454F2E1AFBCF60FFB7B9B22E" author="Takano, T. K. &amp; Suwito, A. &amp; Gao, J. J. &amp; Yin, J. T." box="[1034,1269,250,274]" page="26" pageId="37" pageNumber="38" year="2011">
<bibRefCitation id="EFEF90CA454F2E1AFBCF60FFB7B9B22E" author="Takano, T. K. &amp; Suwito, A. &amp; Gao, J. J. &amp; Yin, J. T." box="[1034,1269,250,274]" pageId="37" pageNumber="38" pagination="19 - 28" refId="ref33707" refString="Takano, T. K., Suwito, A., Gao, J. J. &amp; Yin, J. T. (2011) Molecular phylogeny of the cristata species group of the genus Colocasiomyia (Diptera: Drosophilidae). Low Temperature Science, 69, 19 - 28." type="journal article" year="2011">
Takano
<emphasis id="B90A3129454F2E1AFB9F60FEB7CBB22E" box="[1114,1159,250,274]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
., 2011: 26
</bibRefCitation>
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.
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</paragraph>
</subSubSection>
<subSubSection id="C364BEB0454F2E1AFF526145B0D4B2FB" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="8BC1ED3B454F2E1AFF526145B0D4B2FB" blockId="37.[151,1437,320,455]" pageId="37" pageNumber="38">
<emphasis id="B90A3129454F2E1AFF526145B25EB266" bold="true" box="[151,274,320,346]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
Additional dorsocentral setae on presutural area absent.A pair of processes on male abdominal sternite VI slightly curved inward (
<figureCitation id="1345F1BE454F2E1AFE616161B2B0B243" box="[420,508,356,383]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson &amp; Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada &amp; Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson &amp; Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda &amp; Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="37" pageNumber="38">Fig. 9L</figureCitation>
). Epandrium triangularly angled at posteroventral corner; anteroventral elongation as broad as cercus in lateral view (
<figureCitation id="1345F1BE454F2E1AFDE7618DB1C0B29F" box="[546,652,392,419]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="37" pageNumber="38">Fig. 24G</figureCitation>
). Female abdominal sternite VI wider than long (
<figureCitation id="1345F1BE454F2E1AFB78618DB668B29F" box="[1213,1316,392,419]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="37" pageNumber="38">Fig. 24E</figureCitation>
). Oviscapt broader than hypoproct, apically roundish in lateral view (
<figureCitation id="1345F1BE454F2E1AFCE661A9B0C7B2FB" box="[803,907,428,455]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="37" pageNumber="38">Fig. 24K</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF01BDB3454F2E1AFF526703B176B4CA" ID-DOI="http://doi.org/10.5281/zenodo.5766454" ID-Zenodo-Dep="5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,250,1798,1822]" targetBox="[179,1407,478,1774]" targetPageId="37">
<paragraph id="8BC1ED3B454F2E1AFF526703B176B4CA" blockId="37.[151,1437,1798,2039]" pageId="37" pageNumber="38">
<emphasis id="B90A3129454F2E1AFF526703B26CB422" bold="true" box="[151,288,1798,1822]" pageId="37" pageNumber="38">FIGURE 24.</emphasis>
<taxonomicName id="4C7E96B8454F2E1AFEE36702B12EB423" authority="Toda" authorityName="Takano &amp; Gao &amp; Hu &amp; Li &amp; Yafuso &amp; Suwito &amp; Repin &amp; Pungga &amp; Meleng &amp; Kaliang &amp; Chong &amp; Toda" authorityYear="2021" box="[294,610,1799,1823]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="sabahana" status="sp. nov.">
<emphasis id="B90A3129454F2E1AFEE36702B16BB423" box="[294,551,1799,1823]" italics="true" pageId="37" pageNumber="38">Colocasiomyia sabahana</emphasis>
Toda
</taxonomicName>
,
<emphasis id="B90A3129454F2E1AFDA96702B1F0B423" bold="true" box="[620,700,1799,1823]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="A2398C52454F2E1AFDA96702B1F0B423" box="[620,700,1799,1823]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and
<taxonomicName id="4C7E96B8454F2E1AFD5F672EB095B47F" authority="Toda &amp; Yafuso" authorityName="Toda &amp; Yafuso" authorityYear="2021" box="[666,985,1835,1859]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="sarawakana" status="sp. nov.">
<emphasis id="B90A3129454F2E1AFD5F672EB1E7B47E" box="[666,683,1835,1858]" italics="true" pageId="37" pageNumber="38">C</emphasis>
.
<emphasis id="B90A3129454F2E1AFD7C672EB079B47F" box="[697,821,1835,1859]" italics="true" pageId="37" pageNumber="38">sarawakana</emphasis>
Toda &amp; Yafuso
</taxonomicName>
,
<emphasis id="B90A3129454F2E1AFC21672EB77AB47F" bold="true" box="[996,1078,1835,1859]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="A2398C52454F2E1AFC21672EB77AB47F" box="[996,1078,1835,1859]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N.
</paragraph>
</caption>
<subSubSection id="C364BEB0454C2E19FF026093B600B101" pageId="38" pageNumber="39" type="description">
<paragraph id="8BC1ED3B454C2E19FF026093B7A0B38D" blockId="38.[151,1436,150,753]" box="[199,1260,150,177]" pageId="38" pageNumber="39">
<emphasis id="B90A3129454C2E19FF026093B21DB38C" bold="true" box="[199,337,150,176]" pageId="38" pageNumber="39">Description</emphasis>
(
<specimenCount id="9D7826B2454C2E19FEA46092B23AB38D" box="[353,374,151,177]" pageId="38" pageNumber="39" type="male">♂</specimenCount>
and
<specimenCount id="9D7826B2454C2E19FE686092B28DB38D" box="[429,449,151,177]" pageId="38" pageNumber="39" type="female">♀</specimenCount>
). Head. Supralateral setae outside prementum approximately 2 per side.
</paragraph>
<paragraph id="8BC1ED3B454C2E19FF0260BEB709B3E9" blockId="38.[151,1436,150,753]" box="[199,1093,187,213]" pageId="38" pageNumber="39">Thorax. Basal scutellar setae short, not reaching to half of apical scutellar setae.</paragraph>
<paragraph id="8BC1ED3B454C2E19FF0260DAB1DAB3C5" blockId="38.[151,1436,150,753]" box="[199,662,223,249]" pageId="38" pageNumber="39">Legs. Mid tibia with 2 apical, stout setae.</paragraph>
<paragraph id="8BC1ED3B454C2E19FF026106B004B221" blockId="38.[151,1436,150,753]" box="[199,840,259,285]" pageId="38" pageNumber="39">
Abdomen. Female sternite IV as long as wide (
<figureCitation id="1345F1BE454C2E19FD136106B070B221" box="[726,828,259,285]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">Fig. 24E</figureCitation>
).
</paragraph>
<paragraph id="8BC1ED3B454C2E19FF026122B259B291" blockId="38.[151,1436,150,753]" pageId="38" pageNumber="39">
Male terminalia. Epandrium with approximately 1 seta on lateral portion and 14–16 setae on posteroventral portion (
<figureCitation id="1345F1BE454C2E19FF3C614EB228B259" box="[249,356,331,357]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">Fig. 24G</figureCitation>
). Median piece of subepandrial sclerite medially and anteriorly bilobed into long lobes; lateral pieces long, narrow. Cercus with 45–48 setae, extended below, apically forming small lobe with many small setae (
<figureCitation id="1345F1BE454C2E19FF656196B244B291" box="[160,264,403,429]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">Fig. 24G</figureCitation>
).
</paragraph>
<paragraph id="8BC1ED3B454C2E19FF0261B2B0C1B2ED" blockId="38.[151,1436,150,753]" box="[199,909,438,465]" pageId="38" pageNumber="39">
Female terminalia. Oviscapt with 14–15 ovisensilla (
<figureCitation id="1345F1BE454C2E19FCDD61B2B0CCB2ED" box="[792,896,439,465]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">Fig. 24K</figureCitation>
).
</paragraph>
<paragraph id="8BC1ED3B454C2E19FF0261DEB600B101" blockId="38.[151,1436,150,753]" pageId="38" pageNumber="39">
Indices (range of
<specimenCount id="9D7826B2454C2E19FE4C61DEB2F6B2C9" box="[393,442,475,501]" pageId="38" pageNumber="39" type="male">10♂</specimenCount>
and
<specimenCount id="9D7826B2454C2E19FE2A61DEB16EB2C9" box="[495,546,475,501]" pageId="38" pageNumber="39" type="female">10♀</specimenCount>
): FW/HW = 0.55–0.65, ch/o = 0.34–0.45, prorb = 0.83–1.04, rcorb = 0.22–0.47, vb = 0.25–0.39, dcl = (n/a), sctl = 0.51–0.77, sterno = 0.56–0.83, orbito = 0.48–0.77, dcp = 0.82–1.21, sctlp = 1.08–1.64, C = 1.62–2.56, 4c = 0.83–1.38, 4v = 1.31–1.94, 5x = 0.74–1.25, ac = 1.98–3.74, M = 0.25–0.56.
</paragraph>
</subSubSection>
<subSubSection id="C364BEB0454C2E19FF026242B75AB195" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="8BC1ED3B454C2E19FF026242B2F4B1B9" blockId="38.[151,1436,150,753]" pageId="38" pageNumber="39">
<materialsCitation id="3B16E766454C2E19FF026242B2F4B1B9" ID-GBIF-Occurrence="3418896836" collectingDate="2003-08-10" collectionCode="KPSP" country="Malaysia" location="Park Headquarters" municipality="Mt. Kinabalu" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B90A3129454C2E19FF026242B277B15D" bold="true" box="[199,315,583,609]" pageId="38" pageNumber="39">
<typeStatus id="54C55399454C2E19FF026242B27AB15D" box="[199,310,583,609]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D7826B2454C2E19FE896242B22DB15D" box="[332,353,583,609]" pageId="38" pageNumber="39" type="male">♂</specimenCount>
(
<collectionCode id="ED6F75FE454C2E19FEBF6242B28EB15D" box="[378,450,583,609]" pageId="38" pageNumber="39">KPSP</collectionCode>
), “
<location id="8EA1BBE0454C2E19FE2C6242B184B15D" LSID="urn:lsid:plazi:treatment:03D75C2D454F2E19FF526092B1D2B1CD:8EA1BBE0454C2E19FE2C6242B184B15D" box="[489,712,583,609]" country="Malaysia" municipality="Mt. Kinabalu" name="Park Headquarters" pageId="38" pageNumber="39" stateProvince="Sabah">Park Headquarters</location>
,
<collectingMunicipality id="6BA57741454C2E19FD186242B0CCB15D" box="[733,896,583,609]" pageId="38" pageNumber="39">Mt. Kinabalu</collectingMunicipality>
,
<collectingRegion id="49BA23D9454C2E19FC506242B090B15D" box="[917,988,583,609]" country="Malaysia" name="Sabah" pageId="38" pageNumber="39">Sabah</collectingRegion>
,
<collectingCountry id="F369ADAB454C2E19FC346242B716B15D" box="[1009,1114,583,609]" name="Malaysia" pageId="38" pageNumber="39">Malaysia</collectingCountry>
,
<date id="FFC0CBFB454C2E19FBB56242B7B6B15D" box="[1136,1274,583,609]" pageId="38" pageNumber="39" value="2003-08-10">
<collectingDate id="EF843213454C2E19FBB56242B7B6B15D" box="[1136,1274,583,609]" pageId="38" pageNumber="39" value="2003-08-10">10.viii.2003</collectingDate>
</date>
, ex
<taxonomicName id="4C7E96B8454C2E19FAFF6242B2E7B1B9" authority=", M. J. Toda" authorityName="M. J. Toda" class="Liliopsida" family="Araceae" genus="Alocasia" kingdom="Plantae" order="Alismatales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="scabriuscula">
<emphasis id="B90A3129454C2E19FAFF6242B26AB1B9" italics="true" pageId="38" pageNumber="39">Alocasia scabriuscula</emphasis>
, M.J. Toda
</taxonomicName>
”.
</materialsCitation>
</paragraph>
<paragraph id="8BC1ED3B454C2E19FF02628AB75AB195" blockId="38.[151,1436,150,753]" box="[199,1046,655,681]" pageId="38" pageNumber="39">
<materialsCitation id="3B16E766454C2E19FF02628AB75EB195" ID-GBIF-Occurrence="3418896473" box="[199,1042,655,681]" collectingDate="2003-08-10" collectionCode="KPSP, SEHU" country="Malaysia" location="Park Headquarters" municipality="Mt. Kinabalu" pageId="38" pageNumber="39" specimenCount="22" specimenCount-female="11" specimenCount-male="11" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="B90A3129454C2E19FF02628AB20AB195" bold="true" box="[199,326,655,681]" pageId="38" pageNumber="39">
<typeStatus id="54C55399454C2E19FF02628AB20DB195" box="[199,321,655,681]" pageId="38" pageNumber="39" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="F369ADAB454C2E19FE88628AB2F4B195" box="[333,440,655,681]" name="Malaysia" pageId="38" pageNumber="39">Malaysia</collectingCountry>
:
<specimenCount id="9D7826B2454C2E19FE07628AB2B8B195" box="[450,500,655,681]" pageId="38" pageNumber="39" type="male">11♂</specimenCount>
,
<specimenCount id="9D7826B2454C2E19FDC5628AB17EB195" box="[512,562,655,681]" pageId="38" pageNumber="39" type="female">11♀</specimenCount>
, same data as the holotype (
<collectionCode id="ED6F75FE454C2E19FCA9628AB0FDB195" box="[876,945,655,681]" pageId="38" pageNumber="39">KPSP</collectionCode>
,
<collectionCode id="ED6F75FE454C2E19FC79628AB746B195" box="[956,1034,655,681]" pageId="38" pageNumber="39">SEHU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C364BEB0454C2E19FF0262B7B153B1F2" box="[199,543,690,718]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8BC1ED3B454C2E19FF0262B7B153B1F2" blockId="38.[151,1436,150,753]" box="[199,543,690,718]" pageId="38" pageNumber="39">
<emphasis id="B90A3129454C2E19FF0262B7B22DB1F0" bold="true" box="[199,353,690,716]" pageId="38" pageNumber="39">Distribution.</emphasis>
Borneo (
<collectingRegion id="49BA23D9454C2E19FE0C62B6B15FB1F2" box="[457,531,691,718]" country="Malaysia" name="Sabah" pageId="38" pageNumber="39">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C364BEB0454C2E19FF0262D2B1D2B1CD" box="[199,670,727,753]" pageId="38" pageNumber="39" type="etymology">
<paragraph id="8BC1ED3B454C2E19FF0262D2B1D2B1CD" blockId="38.[151,1436,150,753]" box="[199,670,727,753]" pageId="38" pageNumber="39">
<emphasis id="B90A3129454C2E19FF0262D2B202B1CD" bold="true" box="[199,334,727,753]" pageId="38" pageNumber="39">Etymology.</emphasis>
Referring to the
<typeStatus id="54C55399454C2E19FDC862D2B171B1CD" box="[525,573,727,753]" pageId="38" pageNumber="39">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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