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<document id="590DB04C7AF33C713F5A9CC6A14D372F" ID-CLB-Dataset="2931" ID-DOI="10.11646/zootaxa.5079.1.1" ID-GBIF-Dataset="010695ef-a142-42ad-81d7-2e7ed1f7e89e" ID-ISSN="1175-5326" ID-Zenodo-Dep="5766402" ID-ZooBank="2B118B81-7353-4A2C-A892-5466DFC83230" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638952199435" checkinUser="plazi" docAuthor="Takano, Kohei Takenaka, Gao, Jian-Jun, Hu, Yao-Guang, Li, Nan-Nan, Yafuso, Masako, Suwito, Awit, Repin, Rimi, Pungga, Runi Anak Sylvester, Meleng, Paulus Ak, Kaliang, Clement Het, Chong, Lucy &amp; Toda, Masanori J." docDate="2021" docId="03D75C2D454C2E19FF526347B1D2B5E7" docLanguage="en" docName="zootaxa.5079.1.1.pdf" docOrigin="Zootaxa 5079 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Colocasiomyia sarawakana Toda &amp; Yafuso 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="39" masterDocId="FFEE2455456A2E3FFFC56005B34CB33C" masterDocTitle="Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species" masterLastPageNumber="70" masterPageNumber="1" pageNumber="39" updateTime="1699472124804" updateUser="ExternalLinkService">
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<mods:title id="07C4EA98A6B14D026607042045A12042">Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species</mods:title>
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<treatment id="03D75C2D454C2E19FF526347B1D2B5E7" ID-DOI="http://doi.org/10.5281/zenodo.5786185" ID-GBIF-Taxon="191651677" ID-Zenodo-Dep="5786185" LSID="urn:lsid:plazi:treatment:03D75C2D454C2E19FF526347B1D2B5E7" httpUri="http://treatment.plazi.org/id/03D75C2D454C2E19FF526347B1D2B5E7" lastPageNumber="39" pageId="38" pageNumber="39">
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<emphasis id="B90A3129454C2E19FF526347B072B061" bold="true" box="[151,830,834,861]" pageId="38" pageNumber="39">
10)
<taxonomicName id="4C7E96B8454C2E19FF036347B198B060" ID-CoL="9HS2H" authority="Toda &amp; Yafuso" authorityName="Toda &amp; Yafuso" authorityYear="2021" box="[198,724,834,861]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="sarawakana" status="sp. nov.">
<emphasis id="B90A3129454C2E19FF036347B15EB061" bold="true" box="[198,530,834,861]" italics="true" pageId="38" pageNumber="39">Colocasiomyia sarawakana</emphasis>
Toda &amp; Yafuso
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,
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<paragraph id="8BC1ED3B454C2E19FF526362B1CBB0BD" blockId="38.[151,830,834,897]" box="[151,647,870,897]" pageId="38" pageNumber="39">
(
<figureCitation id="1345F1BE454C2E19FF5A6362B3B7B0BD" box="[159,251,871,897]" captionStart="FIGURE 4" captionStartId="10.[151,250,798,822]" captionTargetBox="[151,1435,181,773]" captionTargetId="figure-22@10.[151,1435,181,774]" captionTargetPageId="10" captionText="FIGURE 4. Characters 5 and 6 on the cibarium (dorsal view) of Colocasiomyia xenalocasiae (Okada) (A), C. cristata de Meijere (B) and C. sarawakana sp. nov. (C). Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensillum; s c, sensilla campaniformia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766410" httpUri="https://zenodo.org/record/5766410/files/figure.png" pageId="38" pageNumber="39">Figs 4C</figureCitation>
,
<figureCitation id="1345F1BE454C2E19FEC36362B27CB0BD" box="[262,304,871,897]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson &amp; Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada &amp; Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson &amp; Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda &amp; Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="38" pageNumber="39">9M</figureCitation>
,
<figureCitation id="1345F1BE454C2E19FEFE6363B228B0BD" box="[315,356,870,897]" captionStart="FIGURE 15" captionStartId="21.[151,250,1936,1960]" captionTargetBox="[151,1435,860,1912]" captionTargetId="figure-242@21.[151,1435,860,1912]" captionTargetPageId="21" captionText="FIGURE 15. Habitus (upper, lateral view; lower, dorsal view) of species of the Colocasiomyia cristata subgroup.A, C. ecornuta sp. nov. ♂; B, C. grandis sp. nov. ♂; C, C. vieti sp. nov. ♂; D, C. cristata de Meijere ♂; E, C. sulawesiana Okada &amp; Yafuso ♂; F, C. kinabaluana sp. nov. ♂; G, C. kotana sp. nov. ♂; and H, C. matthewsi sp. nov. ♂. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766434" httpUri="https://zenodo.org/record/5766434/files/figure.png" pageId="38" pageNumber="39">15J</figureCitation>
,
<figureCitation id="1345F1BE454C2E19FEB56362B133B0BD" box="[368,639,871,897]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">24A,B,D,F,H–J,L,N,P,R</figureCitation>
)
</paragraph>
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<taxonomicName id="4C7E96B8454C2E19FF5263AEB1A1B0FF" authority=": Sultana et al., 2006: 694" authorityName="Sultana" authorityPageNumber="694" authorityYear="2006" box="[151,749,939,963]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="undefined-3">
<emphasis id="B90A3129454C2E19FF5263AEB263B0FF" box="[151,303,939,963]" italics="true" pageId="38" pageNumber="39">Colocasiomyia</emphasis>
sp.3
<treatmentCitation id="0ADFCB2A454C2E19FEA063AEB1A1B0FF" author="Sultana, F. &amp; Hu, Y. G. &amp; Toda, M. J. &amp; Takenaka, K. &amp; Yafuso, M." box="[357,749,939,963]" page="694" pageId="38" pageNumber="39" year="2006">
aff.
<emphasis id="B90A3129454C2E19FE4B63AEB2A9B0FF" box="[398,485,939,963]" italics="true" pageId="38" pageNumber="39">diconica</emphasis>
:
<bibRefCitation id="EFEF90CA454C2E19FE3763AEB1A1B0FF" author="Sultana, F. &amp; Hu, Y. G. &amp; Toda, M. J. &amp; Takenaka, K. &amp; Yafuso, M." box="[498,749,939,963]" pageId="38" pageNumber="39" pagination="684 - 702" refId="ref33499" refString="Sultana, F., Hu, Y. G., Toda, M. J., Takenaka, K. &amp; Yafuso, M. (2006) Phylogeny and classification of Colocasiomyia (Diptera, Drosophilidae), and its evolution of pollination mutualism with aroid plants. Systematic Entomology, 31, 684 - 702. https: // doi. org / 10.1111 / j. 1365 - 3113.2006.00344. x" type="journal article" year="2006">
Sultana
<emphasis id="B90A3129454C2E19FD8063A9B13DB0FF" box="[581,625,939,963]" italics="true" pageId="38" pageNumber="39">et al</emphasis>
., 2006: 694
</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0ADFCB2A454C2E19FD3263AEB74CB0FF" author="Toda, M. J. &amp; Lakim, M. B." box="[759,1024,939,963]" page="263" pageId="38" pageNumber="39" year="2011">
<bibRefCitation id="EFEF90CA454C2E19FD3263AEB74CB0FF" author="Toda, M. J. &amp; Lakim, M. B." box="[759,1024,939,963]" pageId="38" pageNumber="39" pagination="262 - 270" refId="ref33989" refString="Toda, M. J. &amp; Lakim, M. B. (2011) Genus Colocasiomyia (Drosophilidae: Diptera) in Sabah, Bornean Malaysia: high species diversity and use of host aroid inflorescences. Entomological Science, 14, 262 - 270. https: // doi. org / 10.1111 / j. 1479 - 8298.2011.00452. x" type="journal article" year="2011">Toda &amp; Lakim, 2011: 263</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ADFCB2A454C2E19FBCF63AEB7B9B0FF" author="Takano, T. K. &amp; Suwito, A. &amp; Gao, J. J. &amp; Yin, J. T." box="[1034,1269,939,963]" page="26" pageId="38" pageNumber="39" year="2011">
<bibRefCitation id="EFEF90CA454C2E19FBCF63AEB7B9B0FF" author="Takano, T. K. &amp; Suwito, A. &amp; Gao, J. J. &amp; Yin, J. T." box="[1034,1269,939,963]" pageId="38" pageNumber="39" pagination="19 - 28" refId="ref33707" refString="Takano, T. K., Suwito, A., Gao, J. J. &amp; Yin, J. T. (2011) Molecular phylogeny of the cristata species group of the genus Colocasiomyia (Diptera: Drosophilidae). Low Temperature Science, 69, 19 - 28." type="journal article" year="2011">
Takano
<emphasis id="B90A3129454C2E19FB9E63A9B7CBB0FF" box="[1115,1159,939,963]" italics="true" pageId="38" pageNumber="39">et al</emphasis>
., 2011: 26
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.
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</paragraph>
</subSubSection>
<subSubSection id="C364BEB0454C2E19FF5263F4B0EFB74B" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="8BC1ED3B454C2E19FF5263F4B0EFB74B" blockId="38.[151,1437,1009,1756]" pageId="38" pageNumber="39">
<emphasis id="B90A3129454C2E19FF5263F4B25EB737" bold="true" box="[151,274,1009,1035]" pageId="38" pageNumber="39">Diagnosis.</emphasis>
Additional dorsocentral setae on presutural area present (
<figureCitation id="1345F1BE454C2E19FC4663F4B0ABB737" box="[899,999,1009,1035]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">Fig. 24B</figureCitation>
).A pair of processes on male abdominal sternite VI nearly straight (
<figureCitation id="1345F1BE454C2E19FE796410B156B713" box="[444,538,1045,1071]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson &amp; Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada &amp; Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson &amp; Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda &amp; Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="38" pageNumber="39">Fig. 9M</figureCitation>
). Epandrium roundish on posteroventral margin; anteroventral elongation broader than cercus in lateral view (
<figureCitation id="1345F1BE454C2E19FE02643CB163B76F" box="[455,559,1081,1107]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">Fig. 24H</figureCitation>
). Female abdominal sternite VI longer than wide (
<figureCitation id="1345F1BE454C2E19FB9C643CB7F7B76F" box="[1113,1211,1081,1107]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">Fig. 24F</figureCitation>
). Oviscapt narrower than hypoproct, apically somewhat pointed in lateral view (
<figureCitation id="1345F1BE454C2E19FCF46458B0DBB74B" box="[817,919,1117,1143]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">Fig. 24L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C364BEB0454C2E19FF026484B607B538" pageId="38" pageNumber="39" type="description">
<paragraph id="8BC1ED3B454C2E19FF026484B713B7A7" blockId="38.[151,1437,1009,1756]" box="[199,1119,1153,1180]" pageId="38" pageNumber="39">
<emphasis id="B90A3129454C2E19FF026484B21DB7A7" bold="true" box="[199,337,1153,1179]" pageId="38" pageNumber="39">Description</emphasis>
(
<specimenCount id="9D7826B2454C2E19FEA46484B23AB7A7" box="[353,374,1153,1179]" pageId="38" pageNumber="39" type="male">♂</specimenCount>
and
<specimenCount id="9D7826B2454C2E19FE686487B28DB7A0" box="[429,449,1154,1180]" pageId="38" pageNumber="39" type="female">♀</specimenCount>
). Head. Supralateral setae outside prementum 3–4 per side.
</paragraph>
<paragraph id="8BC1ED3B454C2E19FF0264A0B735B7FC" blockId="38.[151,1437,1009,1756]" box="[199,1145,1189,1216]" pageId="38" pageNumber="39">Thorax. Basal scutellar setae long; posterior tip beyond half of apical scutellar setae.</paragraph>
<paragraph id="8BC1ED3B454C2E19FF0264CFB1FEB7D8" blockId="38.[151,1437,1009,1756]" box="[199,690,1225,1252]" pageId="38" pageNumber="39">Legs. Mid tibia with 2–3 apical, stout setae.</paragraph>
<paragraph id="8BC1ED3B454C2E19FF0264E8B006B63B" blockId="38.[151,1437,1009,1756]" box="[199,842,1261,1288]" pageId="38" pageNumber="39">
Abdomen. Female sternite IV wider than long (
<figureCitation id="1345F1BE454C2E19FD1F64E8B072B63B" box="[730,830,1261,1287]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">Fig. 24F</figureCitation>
).
</paragraph>
<paragraph id="8BC1ED3B454C2E19FF026514B021B64F" blockId="38.[151,1437,1009,1756]" pageId="38" pageNumber="39">
Male terminalia. Epandrium with 9–10 setae on posteroventral portion (
<figureCitation id="1345F1BE454C2E19FC186514B708B617" box="[989,1092,1297,1323]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">Fig. 24H</figureCitation>
). Median piece of subepandrial sclerite broad, somewhat quadrate plate; lateral pieces absent. Cercus with 37–38 setae, extended below, apically forming somewhat broad lobe with many small setae (
<figureCitation id="1345F1BE454C2E19FD3D655CB02DB64F" box="[760,865,1369,1395]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">Fig. 24H</figureCitation>
).
</paragraph>
<paragraph id="8BC1ED3B454C2E19FF026578B745B6AB" blockId="38.[151,1437,1009,1756]" box="[199,1033,1405,1431]" pageId="38" pageNumber="39">
Female terminalia. Oviscapt with approximately 12 ovisensilla (
<figureCitation id="1345F1BE454C2E19FC536578B0B0B6AB" box="[918,1020,1405,1431]" captionStart="FIGURE 24" captionStartId="37.[151,250,1798,1822]" captionTargetBox="[179,1407,478,1774]" captionTargetId="figure-110@37.[179,1407,478,1774]" captionTargetPageId="37" captionText="FIGURE 24. Colocasiomyia sabahana Toda, sp. nov. (♂ and ♀ paratypes and puparium from the Park Headquarters area, Mt. Kinabalu, Sabah, Malaysia; C,E,G,K,M,O,Q) and C. sarawakana Toda &amp; Yafuso, sp. nov. (♂ and ♀ paratypes and puparium from Kuching, Sarawak, Malaysia; A,B,D,F,H–J,L,N,P,R). A, Proboscis (caudal view); B, thorax (dorsal view); C, D, male abdominal sternites (ventral view); E, F, female abdominal sternites (ventral view); G, H, periphallic organs (lateral view); I, J, phallic organs (I, lateral view; J, ventral view); K, L, female terminalia (lateral view); M, N, puparium (ventral habitus); O, P, mouth hook; Q, R, cephalopharyngeal skeleton. Arrowheads indicate diagnostic characters. Scale bars: 0.1 mm in A, C–L, Q and R, 0.5 mm in B, 1.0 mm in M and N." figureDoi="http://doi.org/10.5281/zenodo.5766454" httpUri="https://zenodo.org/record/5766454/files/figure.png" pageId="38" pageNumber="39">Fig. 24L</figureCitation>
).
</paragraph>
<paragraph id="8BC1ED3B454C2E19FF0265A4B607B538" blockId="38.[151,1437,1009,1756]" pageId="38" pageNumber="39">
Indices (range of
<specimenCount id="9D7826B2454C2E19FE4C65A4B2F6B687" box="[393,442,1441,1467]" pageId="38" pageNumber="39" type="male">10♂</specimenCount>
and
<specimenCount id="9D7826B2454C2E19FE2A65A4B16EB687" box="[495,546,1441,1467]" pageId="38" pageNumber="39" type="female">10♀</specimenCount>
): FW/HW = 0.57–0.63, ch/o = 0.31–0.53, prorb = 0.81–1.07, rcorb = 0.20–0.60, vb = 0.20–0.36, dcl = 0.54–0.74, sctl = 0.63–0.89, sterno = 0.43–0.80, orbito = 0.36–0.69, dcp = 0.78–1.25, sctlp = 1.10–1.67, C = 2.12–2.53, 4c = 0.88–1.01, 4v = 1.44–2.11, 5x = 0.81–1.25, ac = 2.06–2.62, M = 0.31–0.50.
</paragraph>
</subSubSection>
<subSubSection id="C364BEB0454C2E19FF026608B663B5A8" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="8BC1ED3B454C2E19FF026608B279B577" blockId="38.[151,1437,1009,1756]" pageId="38" pageNumber="39">
<materialsCitation id="3B16E766454C2E19FF026608B279B577" ID-GBIF-Occurrence="3418896519" collectingDate="2004-11-27" collectionCode="RDID" country="Malaysia" location="Kuching" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B90A3129454C2E19FF026608B277B51B" bold="true" box="[199,315,1549,1575]" pageId="38" pageNumber="39">
<typeStatus id="54C55399454C2E19FF026608B27AB51B" box="[199,310,1549,1575]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D7826B2454C2E19FE80660BB216B514" box="[325,346,1550,1576]" pageId="38" pageNumber="39" type="male">♂</specimenCount>
(
<collectionCode id="ED6F75FE454C2E19FEAE6608B2F9B51B" box="[363,437,1549,1575]" pageId="38" pageNumber="39">RDID</collectionCode>
), “
<location id="8EA1BBE0454C2E19FE106608B177B51B" LSID="urn:lsid:plazi:treatment:03D75C2D454C2E19FF526347B1D2B5E7:8EA1BBE0454C2E19FE106608B177B51B" box="[469,571,1549,1575]" country="Malaysia" name="Kuching" pageId="38" pageNumber="39">Kuching</location>
,
<collectingCountry id="F369ADAB454C2E19FD8C6608B1E0B51B" box="[585,684,1549,1575]" name="Malaysia" pageId="38" pageNumber="39">Sarawak</collectingCountry>
,
<collectingCountry id="F369ADAB454C2E19FD7F6608B06FB51B" box="[698,803,1549,1575]" name="Malaysia" pageId="38" pageNumber="39">Malaysia</collectingCountry>
,
<date id="FFC0CBFB454C2E19FCF46608B0E7B51B" box="[817,939,1549,1575]" pageId="38" pageNumber="39" value="2004-11-27">
<collectingDate id="EF843213454C2E19FCF46608B0E7B51B" box="[817,939,1549,1575]" pageId="38" pageNumber="39" value="2004-11-27">27.xi.2004</collectingDate>
</date>
, ex
<taxonomicName id="4C7E96B8454C2E19FC186608B26BB577" authority=", Lucy Chong &amp; K. T. Takano" authorityName="Lucy Chong &amp; K. T. Takano" class="Liliopsida" family="Araceae" genus="Alocasia" kingdom="Plantae" order="Alismatales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="macrorrhizos">
<emphasis id="B90A3129454C2E19FC186608B792B51B" box="[989,1246,1549,1575]" italics="true" pageId="38" pageNumber="39">Alocasia macrorrhizos</emphasis>
, Lucy Chong &amp; K.T. Takano
</taxonomicName>
”.
</materialsCitation>
</paragraph>
<paragraph id="8BC1ED3B454C2E19FF026650B663B5A8" blockId="38.[151,1437,1009,1756]" pageId="38" pageNumber="39">
<materialsCitation id="3B16E766454C2E19FF026650B094B553" ID-GBIF-Occurrence="3418896544" box="[199,984,1621,1648]" collectingDate="2004-11-27" collectionCode="RDID" country="Malaysia" location="Kuching" pageId="38" pageNumber="39" specimenCount="23" specimenCount-female="12" specimenCount-male="11" typeStatus="paratype">
<emphasis id="B90A3129454C2E19FF026650B20AB553" bold="true" box="[199,326,1621,1647]" pageId="38" pageNumber="39">
<typeStatus id="54C55399454C2E19FF026650B20DB553" box="[199,321,1621,1647]" pageId="38" pageNumber="39" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="F369ADAB454C2E19FE956650B2F7B553" box="[336,443,1621,1647]" name="Malaysia" pageId="38" pageNumber="39">Malaysia</collectingCountry>
:
<specimenCount id="9D7826B2454C2E19FE0D6650B2B5B553" box="[456,505,1621,1647]" pageId="38" pageNumber="39" type="male">11♂</specimenCount>
,
<specimenCount id="9D7826B2454C2E19FDCD6650B177B553" box="[520,571,1621,1647]" pageId="38" pageNumber="39" type="female">12♀</specimenCount>
, same data as the holotype (
<collectionCode id="ED6F75FE454C2E19FC406650B083B553" box="[901,975,1621,1647]" pageId="38" pageNumber="39">RDID</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B16E766454C2E19FC236650B665B5A8" ID-GBIF-Occurrence="3418896936" collectingDate="2004-07-29" collectionCode="BORN, KPSP, SEHU" country="Malaysia" latitude="6.046111" location="Mt. Kinabalu" longLatPrecision="21" longitude="116.7025" municipality="Poring" pageId="38" pageNumber="39" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Sabah" typeStatus="paratype">
<specimenCount id="9D7826B2454C2E19FC236650B747B553" box="[998,1035,1621,1647]" pageId="38" pageNumber="39" type="male">3♂</specimenCount>
,
<specimenCount id="9D7826B2454C2E19FBDC6650B771B553" box="[1049,1085,1621,1647]" pageId="38" pageNumber="39" type="female">3♀</specimenCount>
,
<collectingMunicipality id="6BA57741454C2E19FB886650B7D7B553" box="[1101,1179,1621,1647]" pageId="38" pageNumber="39">Poring</collectingMunicipality>
,
<location id="8EA1BBE0454C2E19FB6D6653B608B553" LSID="urn:lsid:plazi:treatment:03D75C2D454C2E19FF526347B1D2B5E7:8EA1BBE0454C2E19FB6D6653B608B553" box="[1192,1348,1621,1648]" country="Malaysia" latitude="6.046111" longLatPrecision="21" longitude="116.7025" municipality="Poring" name="Mt. Kinabalu" pageId="38" pageNumber="39" stateProvince="Sabah">Mt. Kinabalu</location>
,
<collectingRegion id="49BA23D9454C2E19FA946650B6D4B553" box="[1361,1432,1621,1647]" country="Malaysia" name="Sabah" pageId="38" pageNumber="39">Sabah</collectingRegion>
,
<geoCoordinate id="EE4A8BFC454C2E19FF52667CB25FB5AF" box="[151,275,1657,1683]" degrees="6" direction="north" minutes="02" orientation="latitude" pageId="38" pageNumber="39" precision="15" seconds="46" value="6.046111">6°02’46”N</geoCoordinate>
<geoCoordinate id="EE4A8BFC454C2E19FEDC667CB2E3B5AF" box="[281,431,1657,1683]" degrees="116" direction="east" minutes="42" orientation="longitude" pageId="38" pageNumber="39" precision="15" seconds="09" value="116.7025">116°42’09”E</geoCoordinate>
,
<date id="FFC0CBFB454C2E19FE7F667CB171B5AF" box="[442,573,1657,1683]" pageId="38" pageNumber="39" value="2004-07-29">
<collectingDate id="EF843213454C2E19FE7F667CB171B5AF" box="[442,573,1657,1683]" pageId="38" pageNumber="39" value="2004-07-29">29.vii.2004</collectingDate>
</date>
, ex
<taxonomicName id="4C7E96B8454C2E19FDAC667CB0A7B5AF" authority=", K. T. Takano" authorityName="K. T. Takano" box="[617,1003,1657,1684]" class="Liliopsida" family="Araceae" genus="Alocasia" kingdom="Plantae" order="Alismatales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="scabriscula">
<emphasis id="B90A3129454C2E19FDAC667CB01FB5AF" box="[617,851,1657,1683]" italics="true" pageId="38" pageNumber="39">Alocasia scabriscula</emphasis>
, K.T. Takano
</taxonomicName>
leg. (
<collectionCode id="ED6F75FE454C2E19FBEF667CB730B5AF" box="[1066,1148,1657,1683]" country="Malaysia" httpUri="http://grbio.org/cool/y0pn-rrd0" name="Institute for Tropical Biology and Conservation, Borneensis" pageId="38" pageNumber="39">BORN</collectionCode>
,
<collectionCode id="ED6F75FE454C2E19FB4D667CB787B5AF" box="[1160,1227,1657,1683]" pageId="38" pageNumber="39">KPSP</collectionCode>
,
<collectionCode id="ED6F75FE454C2E19FB10667CB652B5AF" box="[1237,1310,1657,1683]" pageId="38" pageNumber="39">SEHU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C364BEB0454C2E19FF026698B010B584" box="[199,860,1693,1720]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8BC1ED3B454C2E19FF026698B010B584" blockId="38.[151,1437,1009,1756]" box="[199,860,1693,1720]" pageId="38" pageNumber="39">
<emphasis id="B90A3129454C2E19FF026698B22DB58B" bold="true" box="[199,353,1693,1719]" pageId="38" pageNumber="39">Distribution.</emphasis>
Borneo (
<collectingCountry id="F369ADAB454C2E19FE0C6698B162B58B" box="[457,558,1693,1719]" name="Malaysia" pageId="38" pageNumber="39">Sarawak</collectingCountry>
,
<collectingRegion id="49BA23D9454C2E19FDFC6698B1CCB58B" box="[569,640,1693,1719]" country="Malaysia" name="Sabah" pageId="38" pageNumber="39">Sabah</collectingRegion>
, West Kalimantan).
</paragraph>
</subSubSection>
<subSubSection id="C364BEB0454C2E19FF0266C4B1D2B5E7" box="[199,670,1729,1756]" pageId="38" pageNumber="39" type="etymology">
<paragraph id="8BC1ED3B454C2E19FF0266C4B1D2B5E7" blockId="38.[151,1437,1009,1756]" box="[199,670,1729,1756]" pageId="38" pageNumber="39">
<emphasis id="B90A3129454C2E19FF0266C4B202B5E7" bold="true" box="[199,334,1729,1755]" pageId="38" pageNumber="39">Etymology.</emphasis>
Referring to the
<typeStatus id="54C55399454C2E19FDC866C7B171B5E0" box="[525,573,1730,1756]" pageId="38" pageNumber="39">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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