treatments-xml/data/03/D2/40/03D24013FD5EFFD1FF36FBB1C6FBF94A.xml
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<document id="DCB09E942C1E44DB4E94CACFABCBE2E8" ID-DOI="10.11646/zootaxa.3941.1.2" ID-GBIF-Dataset="9a54c25b-e43f-4e60-881a-fd262e98310f" ID-ISSN="1175-5326" ID-Zenodo-Dep="288211" ID-ZooBank="ECDCBC06-96AC-4D91-9C24-7A0A30A3E375" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461023909168" checkinUser="plazi" docAuthor="Hibino, Yusuke" docDate="2015" docId="03D24013FD5EFFD1FF36FBB1C6FBF94A" docLanguage="en" docName="zt03941p078.pdf" docOrigin="Zootaxa 3941 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Apterichtus jeffwilliamsi Hibino, 2015, n. sp." docType="treatment" docUuid="6310D486-11EF-4A43-8FB3-E3883A457F71" docUuidSource="ZooBank" docVersion="8" lastPageNumber="65" masterDocId="FFEB386BFD53FFC1FFA1FFE3C366FFAF" masterDocTitle="A review of the finless snake eels of the genus Apterichtus (Anguilliformes: Ophichthidae), with the description of five new species" masterLastPageNumber="78" masterPageNumber="49" pageNumber="62" updateTime="1698571358071" updateUser="plazi">
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<mods:title id="3FB6A4A74909777EFD7AE5D5AD1AC6F5">A review of the finless snake eels of the genus Apterichtus (Anguilliformes: Ophichthidae), with the description of five new species</mods:title>
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<mods:namePart id="5ACF7855B23DAF76F1598DDA669322C1">Hibino, Yusuke</mods:namePart>
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<mods:date id="4D42691D7A2C02F7FE676BD52A29439B">2015</mods:date>
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<treatment id="03D24013FD5EFFD1FF36FBB1C6FBF94A" ID-DOI="http://doi.org/10.5281/zenodo.6116405" ID-GBIF-Taxon="127687763" ID-Zenodo-Dep="6116405" LSID="urn:lsid:zoobank.org:act:6310D486-11EF-4A43-8FB3-E3883A457F71" httpUri="http://treatment.plazi.org/id/03D24013FD5EFFD1FF36FBB1C6FBF94A" lastPageId="16" lastPageNumber="65" pageId="13" pageNumber="62">
<subSubSection id="C361A28EFD5EFFCCFF36FBB1C2CDFB21" pageId="13" pageNumber="62" type="nomenclature">
<paragraph id="8BC4F105FD5EFFCCFF36FBB1C16FFBC2" blockId="13.[151,521,1106,1166]" box="[151,521,1106,1133]" pageId="13" pageNumber="62">
<heading id="D08C4669FD5EFFCCFF36FBB1C16FFBC2" bold="true" box="[151,521,1106,1133]" fontSize="11" level="1" pageId="13" pageNumber="62" reason="1">
<emphasis id="B90F2D17FD5EFFCCFF36FBB1C16FFBC2" bold="true" box="[151,521,1106,1133]" pageId="13" pageNumber="62">
<taxonomicName id="4C7B8A86FD5EFFCCFF36FBB1C2D9FBC3" ID-CoL="682C5" box="[151,447,1106,1132]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="13" pageNumber="62" phylum="Chordata" rank="species" species="jeffwilliamsi" status="sp. nov.">
<emphasis id="B90F2D17FD5EFFCCFF36FBB1C2D9FBC3" bold="true" box="[151,447,1106,1132]" italics="true" pageId="13" pageNumber="62">Apterichtus jeffwilliamsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C906CFD5EFFCCFE66FBB0C16FFBC2" box="[455,521,1107,1133]" pageId="13" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC4F105FD5EFFCCFF36FB96C2CDFB21" blockId="13.[151,521,1106,1166]" box="[151,427,1141,1166]" pageId="13" pageNumber="62">
<figureCitation id="1340ED80FD5EFFCCFF36FB96C24AFB21" box="[151,300,1141,1166]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStartId-0="14.[151,250,1525,1547]" captionStartId-1="14.[151,250,2003,2025]" captionStartId-2="15.[151,250,1953,1975]" captionTargetBox-0="[340,1246,481,1503]" captionTargetBox-1="[195,1398,1637,1967]" captionTargetBox-2="[497,1106,201,1924]" captionTargetId-0="figure@14.[340,1247,481,1504]" captionTargetId-1="figure@14.[189,1398,1628,1982]" captionTargetId-2="figure@15.[480,1106,193,1932]" captionTargetPageId-0="14" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIGURE 8. Holotype of Apterichtus jeffwilliamsi, USNM 363869, 305 mm, showing fresh coloration. Photograph by J. Williams." captionText-1="FIGURE 9. Head of holotype of Apterichtus jeffwilliamsi, USNM 363869, 305 mm. Drawn by Y. Hibino." captionText-2="FIGURE 10. Dentition of holotype of Apterichtus jeffwilliamsi, USNM 363869, 305 mm. Drawn by Y. Hibino." httpUri-0="https://zenodo.org/record/288219/files/figure.png" httpUri-1="https://zenodo.org/record/288220/files/figure.png" httpUri-2="https://zenodo.org/record/288221/files/figure.png" pageId="13" pageNumber="62">Figures 810</figureCitation>
,
<tableCitation id="C6F9C4BEFD5EFFCCFE99FB96C2F5FB21" box="[312,403,1141,1166]" captionStart="TABLE 1" captionStartId="5.[151,239,400,423]" captionTargetBox="[151,1159,593,1203]" captionTargetPageId="5" captionText="TABLE 1. Characteristics of the species of Apterichtus. Abbreviations are: MVF = mean vertebral formula; POP = preopercular pore; STP = supratemporal pore. Preanal vertebral counts of A. gracilis and A. monodi are estimates based on preanal lateral-line pores. Sources are: 1, this study; 2, Blache &amp; Bauchot 1972; 3, McCosker et al. 1989; 4, McCosker &amp; Randall 2005; 5, Pereira &amp; Aguiar 1979; 6, Hibino et al. 2014; 7, Machida &amp; Ohta 1994." httpUri="http://table.plazi.org/id/DF04A18DFD56FFC4FF36FE73C701FDA8" pageId="13" pageNumber="62" tableUuid="DF04A18DFD56FFC4FF36FE73C701FDA8">Tables 1</tableCitation>
,
<tableCitation id="C6F9C4BEFD5EFFCCFE3BFB95C2CDFB21" box="[410,427,1142,1166]" captionStart="TABLE 3" captionStartId="16.[151,239,152,175]" captionTargetBox="[151,954,248,731]" captionTargetPageId="16" captionText="TABLE 3. Counts and proportions (in thousandths) of the holotype and 4 paratypes of Apterichtus jeffwilliamsi n. sp. TL = total length. HL = head length." httpUri="http://table.plazi.org/id/DF04A18DFD43FFD1FF36FF7BC125FF60" pageId="13" pageNumber="62" tableUuid="DF04A18DFD43FFD1FF36FF7BC125FF60">3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C361A28EFD5EFFCCFF36FB5EC666FA25" pageId="13" pageNumber="62" type="materials_examined">
<paragraph id="8BC4F105FD5EFFCCFF36FB5EC2EAFAB1" blockId="13.[151,1437,1213,2030]" pageId="13" pageNumber="62">
<emphasis id="B90F2D17FD5EFFCCFF36FB5EC26DFB79" bold="true" box="[151,267,1213,1238]" pageId="13" pageNumber="62">
<typeStatus id="54C04FA7FD5EFFCCFF36FB5EC260FB79" box="[151,262,1213,1238]" pageId="13" pageNumber="62" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED6A69C0FD5EFFCCFEBBFB5DC20DFB79" LSID="urn:lsid:biocol.org:col:34871" box="[282,363,1214,1238]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="62">USNM</collectionCode>
<quantity id="4C835CE0FD5EFFCCFEDBFB5EC12FFB7A" box="[378,585,1213,1238]" metricMagnitude="5" metricUnit="m" metricValue="3.63869305" pageId="13" pageNumber="62" unit="mm" value="3.63869305E8">363869, 305 mm</quantity>
TL, female, Vanua Lava
<collectingCountry id="F36CB195FD5EFFCCFC2CFB5EC0B2FB79" box="[909,980,1213,1238]" name="Iceland" pageId="13" pageNumber="62">Island</collectingCountry>
, Banks Islands,
<collectingCountry id="F36CB195FD5EFFCCFB03FB5DC798FB7A" box="[1186,1278,1214,1237]" name="Vanuatu" pageId="13" pageNumber="62">Vanuatu</collectingCountry>
(13°.7461S, 167°.4061E), depth
<quantity id="4C835CE0FD5EFFCCFE26FB02C287FB56" box="[391,481,1249,1274]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.6" metricValueMin="0.9" pageId="13" pageNumber="62" unit="m" value="12.5" valueMax="16.0" valueMin="9.0">916 m</quantity>
, rippled sand bottom, collected by J.T. Williams, D.G. Smith, R.D. Mooi, and D.J. Bray,
<date id="FFC5D7C5FD5EFFCCFF78FAE5C2EEFAB1" box="[217,392,1285,1310]" pageId="13" pageNumber="62" value="1997-03-23">23 March 1997</date>
.
</paragraph>
<paragraph id="8BC4F105FD5EFFCCFF66FACAC666FA25" blockId="13.[151,1437,1213,2030]" pageId="13" pageNumber="62">
<emphasis id="B90F2D17FD5EFFCCFF66FACAC220FAED" bold="true" box="[199,326,1321,1346]" pageId="13" pageNumber="62">
<typeStatus id="54C04FA7FD5EFFCCFF66FACAC227FAED" box="[199,321,1321,1346]" pageId="13" pageNumber="62" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Four specimens,
<quantity id="4C835CE0FD5EFFCCFDB6FAC9C1D6FAEE" box="[535,688,1322,1346]" metricMagnitude="-1" metricUnit="m" metricValue="1.975" metricValueMax="2.83" metricValueMin="1.1199999999999999" pageId="13" pageNumber="62" unit="mm" value="197.5" valueMax="283.0" valueMin="112.0">112283 mm</quantity>
TL.
<collectionCode id="ED6A69C0FD5EFFCCFD4EFAC9C026FAED" LSID="urn:lsid:biocol.org:col:34871" box="[751,832,1322,1346]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="62">USNM</collectionCode>
427494 (formerly
<collectionCode id="ED6A69C0FD5EFFCCFB81FAC9C717FAED" LSID="urn:lsid:biocol.org:col:34871" box="[1056,1137,1322,1346]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="62">USNM</collectionCode>
363869), two specimens,
<quantity id="4C835CE0FD5EFFCCFF36FAADC24DFACA" box="[151,299,1358,1382]" metricMagnitude="-1" metricUnit="m" metricValue="1.975" metricValueMax="2.83" metricValueMin="1.1199999999999999" pageId="13" pageNumber="62" unit="mm" value="197.5" valueMax="283.0" valueMin="112.0">112283 mm</quantity>
TL, collected with
<typeStatus id="54C04FA7FD5EFFCCFDA7FAAEC10AFAC9" box="[518,620,1357,1382]" pageId="13" pageNumber="62" type="holotype">holotype</typeStatus>
.
<collectionCode id="ED6A69C0FD5EFFCCFDD7FAADC1A1FAC9" LSID="urn:lsid:biocol.org:col:34871" box="[630,711,1358,1382]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="62">USNM</collectionCode>
350117, two specimens,
<quantity id="4C835CE0FD5EFFCCFC43FAADC711FACA" box="[994,1143,1358,1382]" metricMagnitude="-1" metricUnit="m" metricValue="1.635" metricValueMax="1.78" metricValueMin="1.49" pageId="13" pageNumber="62" unit="mm" value="163.5" valueMax="178.0" valueMin="149.0">149178 mm</quantity>
TL, Ranon Bay, Ambrym
<collectingCountry id="F36CB195FD5EFFCCFF36FA92C3B8FA25" box="[151,222,1393,1418]" name="Iceland" pageId="13" pageNumber="62">Island</collectingCountry>
,
<collectingCountry id="F36CB195FD5EFFCCFF48FA91C223FA26" box="[233,325,1394,1417]" name="Vanuatu" pageId="13" pageNumber="62">Vanuatu</collectingCountry>
(16°.1406S, 168°.1167E), depth
<quantity id="4C835CE0FD5EFFCCFD6CFA91C073FA26" box="[717,789,1394,1418]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="13" pageNumber="62" unit="m" value="5.5" valueMax="7.0" valueMin="4.0">47 m</quantity>
, collected by J.T. Williams and D.G. Smith.
</paragraph>
</subSubSection>
<subSubSection id="C361A28EFD5EFFCCFF66FA76C744F95D" pageId="13" pageNumber="62" type="diagnosis">
<paragraph id="8BC4F105FD5EFFCCFF66FA76C2CEF991" blockId="13.[151,1437,1213,2030]" pageId="13" pageNumber="62">
<emphasis id="B90F2D17FD5EFFCCFF66FA76C227FA01" bold="true" box="[199,321,1429,1454]" pageId="13" pageNumber="62">Diagnosis.</emphasis>
An elongate species with: tail 1.71.8, head 1316, and body depth
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total length; 4 preopercular pores and 59 pores in supratemporal canal; teeth conical, uniserial on jaws and vomer; a single vomerine tooth; anterior nostril non-tubular, a hole; snout sharp, slope approximately 30° relative to underside; body coloration pinkish white, with numerous brown to orange spots less than eye diameter; and MVF 58140, total vertebrae 138142.
</paragraph>
<paragraph id="8BC4F105FD5EFFCCFF66F9AAC744F95D" blockId="13.[151,1437,1213,2030]" pageId="13" pageNumber="62">
<emphasis id="B90F2D17FD5EFFCCFF66F9AAC05DF9CD" bold="true" box="[199,827,1609,1634]" pageId="13" pageNumber="62">
Counts and measurements (in mm) of the
<typeStatus id="54C04FA7FD5EFFCCFD6CF9AAC050F9CD" box="[717,822,1609,1634]" pageId="13" pageNumber="62" type="holotype">holotype</typeStatus>
.
</emphasis>
Total length 305; head 18.8; trunk 106.9; tail 179.3; body depth at gill openings 4.5; body width at gill openings 3.9; body depth at anus 3.9; body width at anus 4.1; head depth at branchial basket 5.8; head width at branchial basket 3.9; snout length 3.3; tip of snout to rictus 9.2; tip of snout to tip of lower jaw 4.0; eye diameter 1.4; interorbital distance 1.1; gill-opening length 2.1; isthmus width 0.8. Vertebral formula 58141. Lateral-line pores 59, 7 pores in branchial region.
</paragraph>
</subSubSection>
<subSubSection id="C361A28EFD5EFFD1FF66F91EC602FB2F" lastPageId="16" lastPageNumber="65" pageId="13" pageNumber="62" type="description">
<paragraph id="8BC4F105FD5EFFCFFF66F91EC13CFE02" blockId="13.[151,1437,1213,2030]" lastBlockId="14.[151,1437,151,429]" lastPageId="14" lastPageNumber="63" pageId="13" pageNumber="62">
<emphasis id="B90F2D17FD5EFFCCFF66F91EC23FF8B9" bold="true" box="[199,345,1789,1814]" pageId="13" pageNumber="62">Description.</emphasis>
Body elongate (
<figureCitation id="1340ED80FD5EFFCCFDB1F91EC132F8B9" box="[528,596,1789,1814]" captionStart="FIGURE 8" captionStartId="14.[151,250,1525,1547]" captionTargetBox="[340,1246,481,1503]" captionTargetId="figure@14.[340,1247,481,1504]" captionTargetPageId="14" captionText="FIGURE 8. Holotype of Apterichtus jeffwilliamsi, USNM 363869, 305 mm, showing fresh coloration. Photograph by J. Williams." httpUri="https://zenodo.org/record/288219/files/figure.png" pageId="13" pageNumber="62">Fig. 8</figureCitation>
), subcylindrical throughout, snout and tail tip sharply pointed; depth at gill openings
<quantity id="4C835CE0FD5EFFCCFEA5F8C2C201F895" box="[260,359,1825,1850]" metricMagnitude="0" metricUnit="m" metricValue="1.7526" metricValueMax="1.8542" metricValueMin="1.6509999999999998" pageId="13" pageNumber="62" unit="in" value="69.0" valueMax="73.0" valueMin="65.0">6573 in</quantity>
TL. Branchial basket slightly wider and deeper than body. Head and trunk
<quantity id="4C835CE0FD5EFFCCFB1DF8C1C648F895" box="[1212,1326,1825,1850]" metricMagnitude="-2" metricUnit="m" metricValue="5.968999999999999" metricValueMax="6.096" metricValueMin="5.842" pageId="13" pageNumber="62" unit="in" value="2.3499999999999996" valueMax="2.4" valueMin="2.3">2.32.4 in</quantity>
TL; head
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TL,
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trunk. Snout pointed, long, almost equal to twice eye diameter, weak ridges on one-third of mid-lateral snout; short median groove ventrally on snout, beyond anterior edge of posterior nostril. Slope of dorsal surface of snout approximately 30° relative to underside of snout. Lower jaw included, its tip located between verticals through anterior margin and center of eye; snout extends beyond tip of lower jaw, distance from tip of snout to anterior tip of lower jaw almost equal to twice eye diameter; upper and lower lips meet when mouth is closed. Mouth moderately elongate. Rictus well behind rear margin of eye. Edge of upper lip smooth, a distinct crease extending from a vertical through rear margin of eye to end of rictus; a crease on lower lip same as on upper in adult specimens (
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TL), extending from a vertical through rear margin of eye to just before preopercle. Eye moderately developed,
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upper jaw and
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head, its anterior margin slightly beyond tip of lower jaw, its center in advance of middle of upper jaw. Anterior nostril non-tubular, small, hole-shaped with a pair of small flaps, located in anterior third of snout, invisible from above. Posterior nostril opens in outer lip beneath anterior margin of eye. Branchial openings low, ventral; branchial region modestly expanded, creating a bulbous region in posterior half of head.
</paragraph>
<caption id="DF04A18DFD5DFFCFFF36FA16C39CF985" httpUri="https://zenodo.org/record/288219/files/figure.png" pageId="14" pageNumber="63" targetBox="[340,1246,481,1503]" targetPageId="14">
<paragraph id="8BC4F105FD5DFFCFFF36FA16C39CF985" blockId="14.[151,1436,1525,1578]" pageId="14" pageNumber="63">
<emphasis id="B90F2D17FD5DFFCFFF36FA16C271F9A5" bold="true" box="[151,279,1525,1547]" pageId="14" pageNumber="63">FIGURE 8.</emphasis>
Holotype of
<taxonomicName id="4C7B8A86FD5DFFCFFE0AFA16C1C1F9A4" box="[427,679,1525,1547]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="14" pageNumber="63" phylum="Chordata" rank="species" species="jeffwilliamsi">
<emphasis id="B90F2D17FD5DFFCFFE0AFA16C1C1F9A4" box="[427,679,1525,1547]" italics="true" pageId="14" pageNumber="63">Apterichtus jeffwilliamsi</emphasis>
</taxonomicName>
, USNM 363869, 305 mm, showing fresh coloration. Photograph by J. Williams.
</paragraph>
</caption>
<caption id="DF04A18DFD5DFFCFFF36F830C7AAF846" box="[151,1228,2003,2025]" httpUri="https://zenodo.org/record/288220/files/figure.png" pageId="14" pageNumber="63" targetBox="[195,1398,1637,1967]" targetPageId="14">
<paragraph id="8BC4F105FD5DFFCFFF36F830C7AAF846" blockId="14.[151,1228,2003,2025]" box="[151,1228,2003,2025]" pageId="14" pageNumber="63">
<emphasis id="B90F2D17FD5DFFCFFF36F830C274F847" bold="true" box="[151,274,2003,2025]" pageId="14" pageNumber="63">FIGURE 9.</emphasis>
Head of holotype of
<taxonomicName id="4C7B8A86FD5DFFCFFE4BF830C184F846" box="[490,738,2003,2025]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="14" pageNumber="63" phylum="Chordata" rank="species" species="jeffwilliamsi">
<emphasis id="B90F2D17FD5DFFCFFE4BF830C184F846" box="[490,738,2003,2025]" italics="true" pageId="14" pageNumber="63">Apterichtus jeffwilliamsi</emphasis>
</taxonomicName>
, USNM 363869, 305 mm. Drawn by Y. Hibino.
</paragraph>
</caption>
<caption id="DF04A18DFD5CFFCEFF36F842C664F818" box="[151,1282,1953,1975]" httpUri="https://zenodo.org/record/288221/files/figure.png" pageId="15" pageNumber="64" targetBox="[497,1106,201,1924]" targetPageId="15">
<paragraph id="8BC4F105FD5CFFCEFF36F842C664F818" blockId="15.[151,1282,1953,1975]" box="[151,1282,1953,1975]" pageId="15" pageNumber="64">
<emphasis id="B90F2D17FD5CFFCEFF36F842C279F819" bold="true" box="[151,287,1953,1975]" pageId="15" pageNumber="64">FIGURE 10.</emphasis>
Dentition of holotype of
<taxonomicName id="4C7B8A86FD5CFFCEFD80F842C07FF818" box="[545,793,1953,1975]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="15" pageNumber="64" phylum="Chordata" rank="species" species="jeffwilliamsi">
<emphasis id="B90F2D17FD5CFFCEFD80F842C07FF818" box="[545,793,1953,1975]" italics="true" pageId="15" pageNumber="64">Apterichtus jeffwilliamsi</emphasis>
</taxonomicName>
, USNM 363869, 305 mm. Drawn by Y. Hibino.
</paragraph>
</caption>
<caption id="DF04A18DFD43FFD1FF36FF7BC125FF60" ID-Table-UUID="DF04A18DFD43FFD1FF36FF7BC125FF60" httpUri="http://table.plazi.org/id/DF04A18DFD43FFD1FF36FF7BC125FF60" pageId="16" pageNumber="65" targetBox="[151,954,248,731]" targetIsTable="true" targetPageId="16">
<paragraph id="8BC4F105FD43FFD1FF36FF7BC125FF60" blockId="16.[151,1436,151,207]" pageId="16" pageNumber="65">
<emphasis id="B90F2D17FD43FFD1FF36FF7BC3D1FF60" bold="true" pageId="16" pageNumber="65">
TABLE 3. Counts and proportions (in thousandths) of the holotype and 4 paratypes of
<taxonomicName id="4C7B8A86FD43FFD1FBD3FF74C6E6FF00" box="[1138,1408,151,175]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="16" pageNumber="65" phylum="Chordata" rank="species" species="jeffwilliamsi" status="sp. nov.">
<emphasis id="B90F2D17FD43FFD1FBD3FF74C6E6FF00" bold="true" box="[1138,1408,151,175]" italics="true" pageId="16" pageNumber="65">Apterichtus jeffwilliamsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C906CFD43FFD1FA27FF7BC3D1FF60" pageId="16" pageNumber="65" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
TL = total length. HL = head length.
</paragraph>
</caption>
<paragraph id="8BC4F105FD43FFD1FD86FF1BC0D2FD74" pageId="16" pageNumber="65">
<table id="F97B03A5FD43003EFF36FF1BC0DCFD74" box="[151,954,248,731]" gridcols="4" gridrows="15" pageId="16" pageNumber="65">
<tr id="354BF347FD43003EFF36FF1BC0DCFEA1" box="[151,954,248,270]" gridrow="0" pageId="16" pageNumber="65" rowspan-0="1">
<th id="769A9A3BFD43003EFD80FF1BC1E6FEA1" box="[545,640,248,270]" gridcol="1" gridrow="0" pageId="16" pageNumber="65">holotype</th>
<th id="769A9A3BFD43003EFD68FF1BC064FEA1" box="[713,770,248,270]" gridcol="2" gridrow="0" pageId="16" pageNumber="65">mean</th>
<th id="769A9A3BFD43003EFCFAFF1BC0DCFEA1" box="[859,954,248,270]" gridcol="3" gridrow="0" pageId="16" pageNumber="65">range</th>
</tr>
<tr id="354BF347FD43003EFF36FED5C0DCFEE3" box="[151,954,310,332]" gridrow="1" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FED5C2CBFEE3" box="[151,429,310,332]" gridcol="0" gridrow="1" pageId="16" pageNumber="65">TL (mm)</th>
<td id="769A9A3BFD43003EFD80FED5C1E6FEE3" box="[545,640,310,332]" gridcol="1" gridrow="1" pageId="16" pageNumber="65">305</td>
<td id="769A9A3BFD43003EFD68FED5C064FEE3" box="[713,770,310,332]" gridcol="2" gridrow="1" pageId="16" pageNumber="65">---</td>
<td id="769A9A3BFD43003EFCFAFED5C0DCFEE3" box="[859,954,310,332]" gridcol="3" gridrow="1" pageId="16" pageNumber="65">112305</td>
</tr>
<tr id="354BF347FD43003EFF36FEB6C0DCFEC4" box="[151,954,341,363]" gridrow="2" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FEB6C2CBFEC4" box="[151,429,341,363]" gridcol="0" gridrow="2" pageId="16" pageNumber="65">HL/TL</th>
<td id="769A9A3BFD43003EFD80FEB6C1E6FEC4" box="[545,640,341,363]" gridcol="1" gridrow="2" pageId="16" pageNumber="65">62</td>
<td id="769A9A3BFD43003EFD68FEB6C064FEC4" box="[713,770,341,363]" gridcol="2" gridrow="2" pageId="16" pageNumber="65">70</td>
<td id="769A9A3BFD43003EFCFAFEB6C0DCFEC4" box="[859,954,341,363]" gridcol="3" gridrow="2" pageId="16" pageNumber="65">6280</td>
</tr>
<tr id="354BF347FD43003EFF36FE97C0DCFE25" box="[151,954,372,394]" gridrow="3" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FE97C2CBFE25" box="[151,429,372,394]" gridcol="0" gridrow="3" pageId="16" pageNumber="65">Head and trunk/TL</th>
<td id="769A9A3BFD43003EFD80FE97C1E6FE25" box="[545,640,372,394]" gridcol="1" gridrow="3" pageId="16" pageNumber="65">412</td>
<td id="769A9A3BFD43003EFD68FE97C064FE25" box="[713,770,372,394]" gridcol="2" gridrow="3" pageId="16" pageNumber="65">425</td>
<td id="769A9A3BFD43003EFCFAFE97C0DCFE25" box="[859,954,372,394]" gridcol="3" gridrow="3" pageId="16" pageNumber="65">412441</td>
</tr>
<tr id="354BF347FD43003EFF36FE71C0DCFE07" box="[151,954,402,424]" gridrow="4" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FE71C2CBFE07" box="[151,429,402,424]" gridcol="0" gridrow="4" pageId="16" pageNumber="65">Tail/TL</th>
<td id="769A9A3BFD43003EFD80FE71C1E6FE07" box="[545,640,402,424]" gridcol="1" gridrow="4" pageId="16" pageNumber="65">588</td>
<td id="769A9A3BFD43003EFD68FE71C064FE07" box="[713,770,402,424]" gridcol="2" gridrow="4" pageId="16" pageNumber="65">574</td>
<td id="769A9A3BFD43003EFCFAFE71C0DCFE07" box="[859,954,402,424]" gridcol="3" gridrow="4" pageId="16" pageNumber="65">558588</td>
</tr>
<tr id="354BF347FD43003EFF36FE52C0DCFE68" box="[151,954,433,455]" gridrow="5" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FE52C2CBFE68" box="[151,429,433,455]" gridcol="0" gridrow="5" pageId="16" pageNumber="65">Upper jaw/HL</th>
<td id="769A9A3BFD43003EFD80FE52C1E6FE68" box="[545,640,433,455]" gridcol="1" gridrow="5" pageId="16" pageNumber="65">491</td>
<td id="769A9A3BFD43003EFD68FE52C064FE68" box="[713,770,433,455]" gridcol="2" gridrow="5" pageId="16" pageNumber="65">422</td>
<td id="769A9A3BFD43003EFCFAFE52C0DCFE68" box="[859,954,433,455]" gridcol="3" gridrow="5" pageId="16" pageNumber="65">345491</td>
</tr>
<tr id="354BF347FD43003EFF36FE33C0DCFE49" box="[151,954,464,486]" gridrow="6" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FE33C2CBFE49" box="[151,429,464,486]" gridcol="0" gridrow="6" pageId="16" pageNumber="65">Snout/HL</th>
<td id="769A9A3BFD43003EFD80FE33C1E6FE49" box="[545,640,464,486]" gridcol="1" gridrow="6" pageId="16" pageNumber="65">178</td>
<td id="769A9A3BFD43003EFD68FE33C064FE49" box="[713,770,464,486]" gridcol="2" gridrow="6" pageId="16" pageNumber="65">151</td>
<td id="769A9A3BFD43003EFCFAFE33C0DCFE49" box="[859,954,464,486]" gridcol="3" gridrow="6" pageId="16" pageNumber="65">134178</td>
</tr>
<tr id="354BF347FD43003EFF36FE0DC0DCFDAB" box="[151,954,494,516]" gridrow="7" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FE0DC2CBFDAB" box="[151,429,494,516]" gridcol="0" gridrow="7" pageId="16" pageNumber="65">Eye/HL</th>
<td id="769A9A3BFD43003EFD80FE0DC1E6FDAB" box="[545,640,494,516]" gridcol="1" gridrow="7" pageId="16" pageNumber="65">75</td>
<td id="769A9A3BFD43003EFD68FE0DC064FDAB" box="[713,770,494,516]" gridcol="2" gridrow="7" pageId="16" pageNumber="65">74</td>
<td id="769A9A3BFD43003EFCFAFE0DC0DCFDAB" box="[859,954,494,516]" gridcol="3" gridrow="7" pageId="16" pageNumber="65">6887</td>
</tr>
<tr id="354BF347FD43003EFF36FDEEC0DCFD8C" box="[151,954,525,547]" gridrow="8" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FDEEC2CBFD8C" box="[151,429,525,547]" gridcol="0" gridrow="8" pageId="16" pageNumber="65">Interorbital/HL</th>
<td id="769A9A3BFD43003EFD80FDEEC1E6FD8C" box="[545,640,525,547]" gridcol="1" gridrow="8" pageId="16" pageNumber="65">56</td>
<td id="769A9A3BFD43003EFD68FDEEC064FD8C" box="[713,770,525,547]" gridcol="2" gridrow="8" pageId="16" pageNumber="65">45</td>
<td id="769A9A3BFD43003EFCFAFDEEC0DCFD8C" box="[859,954,525,547]" gridcol="3" gridrow="8" pageId="16" pageNumber="65">2557</td>
</tr>
<tr id="354BF347FD43003EFF36FDCFC0DCFDED" box="[151,954,556,578]" gridrow="9" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FDCFC2CBFDED" box="[151,429,556,578]" gridcol="0" gridrow="9" pageId="16" pageNumber="65">Gill opening/HL</th>
<td id="769A9A3BFD43003EFD80FDCFC1E6FDED" box="[545,640,556,578]" gridcol="1" gridrow="9" pageId="16" pageNumber="65">112</td>
<td id="769A9A3BFD43003EFD68FDCFC064FDED" box="[713,770,556,578]" gridcol="2" gridrow="9" pageId="16" pageNumber="65">125</td>
<td id="769A9A3BFD43003EFCFAFDCFC0DCFDED" box="[859,954,556,578]" gridcol="3" gridrow="9" pageId="16" pageNumber="65">112138</td>
</tr>
<tr id="354BF347FD43003EFF36FDA9C0DCFDCF" box="[151,954,586,608]" gridrow="10" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FDA9C2CBFDCF" box="[151,429,586,608]" gridcol="0" gridrow="10" pageId="16" pageNumber="65">Isthmus/HL</th>
<td id="769A9A3BFD43003EFD80FDA9C1E6FDCF" box="[545,640,586,608]" gridcol="1" gridrow="10" pageId="16" pageNumber="65">40</td>
<td id="769A9A3BFD43003EFD68FDA9C064FDCF" box="[713,770,586,608]" gridcol="2" gridrow="10" pageId="16" pageNumber="65">35</td>
<td id="769A9A3BFD43003EFCFAFDA9C0DCFDCF" box="[859,954,586,608]" gridcol="3" gridrow="10" pageId="16" pageNumber="65">2243</td>
</tr>
<tr id="354BF347FD43003EFF36FD8AC0DCFDD0" box="[151,954,617,639]" gridrow="11" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FD8AC2CBFDD0" box="[151,429,617,639]" gridcol="0" gridrow="11" pageId="16" pageNumber="65">Depth at gill opening/TL</th>
<td id="769A9A3BFD43003EFD80FD8AC1E6FDD0" box="[545,640,617,639]" gridcol="1" gridrow="11" pageId="16" pageNumber="65">15</td>
<td id="769A9A3BFD43003EFD68FD8AC064FDD0" box="[713,770,617,639]" gridcol="2" gridrow="11" pageId="16" pageNumber="65">15</td>
<td id="769A9A3BFD43003EFCFAFD8AC0DCFDD0" box="[859,954,617,639]" gridcol="3" gridrow="11" pageId="16" pageNumber="65">1415</td>
</tr>
<tr id="354BF347FD43003EFF36FD6BC0DCFD31" box="[151,954,648,670]" gridrow="12" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FD6BC2CBFD31" box="[151,429,648,670]" gridcol="0" gridrow="12" pageId="16" pageNumber="65">Width/depth at gill opening</th>
<td id="769A9A3BFD43003EFD80FD6BC1E6FD31" box="[545,640,648,670]" gridcol="1" gridrow="12" pageId="16" pageNumber="65">870</td>
<td id="769A9A3BFD43003EFD68FD6BC064FD31" box="[713,770,648,670]" gridcol="2" gridrow="12" pageId="16" pageNumber="65">801</td>
<td id="769A9A3BFD43003EFCFAFD6BC0DCFD31" box="[859,954,648,670]" gridcol="3" gridrow="12" pageId="16" pageNumber="65">709870</td>
</tr>
<tr id="354BF347FD43003EFF36FD45C0DCFD13" box="[151,954,678,700]" gridrow="13" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FD45C2CBFD13" box="[151,429,678,700]" gridcol="0" gridrow="13" pageId="16" pageNumber="65">Preanal vertebrae</th>
<td id="769A9A3BFD43003EFD80FD45C1E6FD13" box="[545,640,678,700]" gridcol="1" gridrow="13" pageId="16" pageNumber="65">58</td>
<td id="769A9A3BFD43003EFD68FD45C064FD13" box="[713,770,678,700]" gridcol="2" gridrow="13" pageId="16" pageNumber="65">57.8</td>
<td id="769A9A3BFD43003EFCFAFD45C0DCFD13" box="[859,954,678,700]" gridcol="3" gridrow="13" pageId="16" pageNumber="65">5758</td>
</tr>
<tr id="354BF347FD43003EFF36FD26C0DCFD74" box="[151,954,709,731]" gridrow="14" pageId="16" pageNumber="65">
<th id="769A9A3BFD43003EFF36FD26C2CBFD74" box="[151,429,709,731]" gridcol="0" gridrow="14" pageId="16" pageNumber="65">Total vertebrae</th>
<td id="769A9A3BFD43003EFD80FD26C1E6FD74" box="[545,640,709,731]" gridcol="1" gridrow="14" pageId="16" pageNumber="65">141</td>
<td id="769A9A3BFD43003EFD68FD26C064FD74" box="[713,770,709,731]" gridcol="2" gridrow="14" pageId="16" pageNumber="65">139.6</td>
<td id="769A9A3BFD43003EFCFAFD26C0DCFD74" box="[859,954,709,731]" gridcol="3" gridrow="14" pageId="16" pageNumber="65">138142</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC4F105FD43FFD1FF66FCC7C6F0FC2A" blockId="16.[151,1437,804,1765]" pageId="16" pageNumber="65">
Head pores (
<figureCitation id="1340ED80FD43FFD1FEF4FCC7C2FEFC93" box="[341,408,804,829]" captionStart="FIGURE 9" captionStartId="14.[151,250,2003,2025]" captionTargetBox="[195,1398,1637,1967]" captionTargetId="figure@14.[189,1398,1628,1982]" captionTargetPageId="14" captionText="FIGURE 9. Head of holotype of Apterichtus jeffwilliamsi, USNM 363869, 305 mm. Drawn by Y. Hibino." httpUri="https://zenodo.org/record/288220/files/figure.png" pageId="16" pageNumber="65">Fig. 9</figureCitation>
) small but apparent. Single median interorbital and temporal pores. Supraorbital pores 1+6, infraorbital pores 5+5 (one
<typeStatus id="54C04FA7FD43FFD1FE71FCABC156FCCF" box="[464,560,840,864]" pageId="16" pageNumber="65" type="paratype">paratype</typeStatus>
has 5+6 left), supratemporal pores 5 to 9 (
<quantity id="4C835CE0FD43FFD1FBBBFCABC720FCCF" box="[1050,1094,839,864]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="16" pageNumber="65" unit="in" value="7.0">7 in</quantity>
<typeStatus id="54C04FA7FD43FFD1FBEEFCA4C7D3FCCF" box="[1103,1205,839,864]" pageId="16" pageNumber="65" type="holotype">holotype</typeStatus>
), lower jaw pores 5, preopercular pores 4. Lateral-line pores before anus 58-59, 7 pores in branchial region, almost complete to tail tip.
</paragraph>
<paragraph id="8BC4F105FD43FFD1FF66FC6CC7AFFC5F" blockId="16.[151,1437,804,1765]" pageId="16" pageNumber="65">
Teeth (
<figureCitation id="1340ED80FD43FFD1FEB5FC6CC200FC07" box="[276,358,911,936]" captionStart="FIGURE 10" captionStartId="15.[151,250,1953,1975]" captionTargetBox="[497,1106,201,1924]" captionTargetId="figure@15.[480,1106,193,1932]" captionTargetPageId="15" captionText="FIGURE 10. Dentition of holotype of Apterichtus jeffwilliamsi, USNM 363869, 305 mm. Drawn by Y. Hibino." httpUri="https://zenodo.org/record/288221/files/figure.png" pageId="16" pageNumber="65">Fig. 10</figureCitation>
) uniserial, small, conical and slightly recurved posteriorly. Intermaxillary with a chevron of 3-7 teeth (the largest in the jaw), followed by a short gap and a single vomerine tooth, same size as jaw teeth. Jaw teeth uniserial, small, 17-26 teeth in each side of upper jaw and 16-24 teeth in each side of lower jaw.
</paragraph>
<paragraph id="8BC4F105FD43FFD1FF66FC1FC151FBF2" blockId="16.[151,1437,804,1765]" pageId="16" pageNumber="65">
Ground color (
<figureCitation id="1340ED80FD43FFD1FECFFC1FC2D4FBBB" box="[366,434,1020,1045]" captionStart="FIGURE 8" captionStartId="14.[151,250,1525,1547]" captionTargetBox="[340,1246,481,1503]" captionTargetId="figure@14.[340,1247,481,1504]" captionTargetPageId="14" captionText="FIGURE 8. Holotype of Apterichtus jeffwilliamsi, USNM 363869, 305 mm, showing fresh coloration. Photograph by J. Williams." httpUri="https://zenodo.org/record/288219/files/figure.png" pageId="16" pageNumber="65">Fig. 8</figureCitation>
) of head and body pinkish white, with numerous spots, reddish brown to orange toward tip of tail. In alcohol preservative head dark green, body yellowish brown. Body spots nearly fade out but remain as dark brown spots on head and breast.
</paragraph>
<paragraph id="8BC4F105FD43FFD1FF66FB84C602FB2F" blockId="16.[151,1437,804,1765]" box="[199,1380,1127,1152]" pageId="16" pageNumber="65">
<emphasis id="B90F2D17FD43FFD1FF66FB84C398FB2F" bold="true" box="[199,254,1127,1152]" pageId="16" pageNumber="65">Size.</emphasis>
The largest specimen examined is
<quantity id="4C835CE0FD43FFD1FD26FB84C182FBD0" box="[647,740,1127,1152]" metricMagnitude="-1" metricUnit="m" metricValue="3.05" pageId="16" pageNumber="65" unit="mm" value="305.0">305 mm</quantity>
TL, a female with ripened eggs,
<quantity id="4C835CE0FD43FFD1FBF7FB8BC7BFFBD0" box="[1110,1241,1128,1152]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="16" pageNumber="65" unit="mm" value="0.8" valueMax="0.9" valueMin="0.7">0.7-0.9 mm</quantity>
in diameter.
</paragraph>
</subSubSection>
<subSubSection id="C361A28EFD43FFD1FF66FB6FC399FB67" pageId="16" pageNumber="65" type="etymology">
<paragraph id="8BC4F105FD43FFD1FF66FB6FC399FB67" blockId="16.[151,1437,804,1765]" pageId="16" pageNumber="65">
<emphasis id="B90F2D17FD43FFD1FF66FB6FC228FB0A" bold="true" box="[199,334,1164,1189]" pageId="16" pageNumber="65">Etymology.</emphasis>
Named in honor of Jeffrey T. Williams of the National Museum of Natural History (USNM), the collector.
</paragraph>
</subSubSection>
<subSubSection id="C361A28EFD43FFD1FF66FB37C24EFABF" pageId="16" pageNumber="65" type="distribution">
<paragraph id="8BC4F105FD43FFD1FF66FB37C24EFABF" blockId="16.[151,1437,804,1765]" pageId="16" pageNumber="65">
<emphasis id="B90F2D17FD43FFD1FF66FB37C206FB42" bold="true" box="[199,352,1236,1261]" pageId="16" pageNumber="65">Distribution.</emphasis>
Known only from the
<typeStatus id="54C04FA7FD43FFD1FDC0FB36C1F7FB42" box="[609,657,1237,1261]" pageId="16" pageNumber="65">type</typeStatus>
specimens which were captured at
<collectingCountry id="F36CB195FD43FFD1FB80FB36C7E6FB43" box="[1057,1152,1237,1260]" name="Vanuatu" pageId="16" pageNumber="65">Vanuatu</collectingCountry>
, in
<quantity id="4C835CE0FD43FFD1FB08FB37C79FFB43" box="[1193,1273,1236,1261]" metricMagnitude="1" metricUnit="m" metricValue="1.0" metricValueMax="1.6" metricValueMin="0.4" pageId="16" pageNumber="65" unit="m" value="10.0" valueMax="16.0" valueMin="4.0">4-16 m</quantity>
over a rippled sand bottom.
</paragraph>
</subSubSection>
<subSubSection id="C361A28EFD43FFD1FF66FAFFC6FBF94A" pageId="16" pageNumber="65" type="discussion">
<paragraph id="8BC4F105FD43FFD1FF66FAFFC6FBF94A" blockId="16.[151,1437,804,1765]" pageId="16" pageNumber="65">
<emphasis id="B90F2D17FD43FFD1FF66FAFFC25DFA9A" bold="true" box="[199,315,1308,1333]" pageId="16" pageNumber="65">Remarks.</emphasis>
<taxonomicName id="4C7B8A86FD43FFD1FEE5FAFFC13EFA9B" box="[324,600,1308,1332]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="16" pageNumber="65" phylum="Chordata" rank="species" species="jeffwilliamsi">
<emphasis id="B90F2D17FD43FFD1FEE5FAFFC13EFA9B" box="[324,600,1308,1332]" italics="true" pageId="16" pageNumber="65">Apterichtus jeffwilliamsi</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C7B8A86FD43FFD1FC95FAFFC0D9FA9B" box="[820,959,1308,1332]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="16" pageNumber="65" phylum="Chordata" rank="species" species="nariculus">
<emphasis id="B90F2D17FD43FFD1FC95FAFFC0D9FA9B" box="[820,959,1308,1332]" italics="true" pageId="16" pageNumber="65">A. nariculus</emphasis>
</taxonomicName>
in the shape and condition of its anterior nostril.
<collectingCountry id="F36CB195FD43FFD1FF4CFAA3C26AFAF7" box="[237,268,1344,1368]" name="American Samoa" pageId="16" pageNumber="65">As</collectingCountry>
mentioned in the generic remarks section of this paper, these are the only species within the genus that lack tubular anterior nostrils. Their other characteristics are clearly shared by their congeners.
<taxonomicName id="4C7B8A86FD43FFD1FABAFA87C244FA0F" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="16" pageNumber="65" phylum="Chordata" rank="species" species="jeffwilliamsi">
<emphasis id="B90F2D17FD43FFD1FABAFA87C244FA0F" italics="true" pageId="16" pageNumber="65">Apterichtus jeffwilliamsi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C7B8A86FD43FFD1FE16FA6BC126FA0F" box="[439,576,1416,1440]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="16" pageNumber="65" phylum="Chordata" rank="species" species="nariculus">
<emphasis id="B90F2D17FD43FFD1FE16FA6BC126FA0F" box="[439,576,1416,1440]" italics="true" pageId="16" pageNumber="65">A. nariculus</emphasis>
</taxonomicName>
in having a less obtuse and longer snout (profile is 30° vs. ca. 40°; length is equal to or more than twice eye diameter vs. equal to or less than twice eye diameter), a shorter trunk (35.0-36.2% vs. 40.0%. of TL), and more preopercular pores (4 vs. 3).
<taxonomicName id="4C7B8A86FD43FFD1FC8BFA33C759FA47" box="[810,1087,1488,1512]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="16" pageNumber="65" phylum="Chordata" rank="species" species="jeffwilliamsi">
<emphasis id="B90F2D17FD43FFD1FC8BFA33C759FA47" box="[810,1087,1488,1512]" italics="true" pageId="16" pageNumber="65">Apterichtus jeffwilliamsi</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C7B8A86FD43FFD1FB75FA33C63AFA47" box="[1236,1372,1488,1512]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="16" pageNumber="65" phylum="Chordata" rank="species" species="klazingai">
<emphasis id="B90F2D17FD43FFD1FB75FA33C63AFA47" box="[1236,1372,1488,1512]" italics="true" pageId="16" pageNumber="65">A. klazingai</emphasis>
</taxonomicName>
in its body coloration and in some proportional characters; the new species differs from the latter in having 4 preopercular pores (vs. 3), 5 to 9 supratemporal pores (vs. 5), a more acute and longer snout (its profile ca. 30° vs. ca. 4045°; its length equal to or more than twice eye diameter vs. equal to or less than twice eye diameter), and the shape of its anterior nostril (a hole vs. a tube). The supratemporal pore condition of
<taxonomicName id="4C7B8A86FD43FFD1FBC8F983C673F9D7" box="[1129,1301,1632,1656]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="16" pageNumber="65" phylum="Chordata" rank="species" species="jeffwilliamsi">
<emphasis id="B90F2D17FD43FFD1FBC8F983C673F9D7" box="[1129,1301,1632,1656]" italics="true" pageId="16" pageNumber="65">A. jeffwilliamsi</emphasis>
</taxonomicName>
is variable, ranging from 5 to 9 (5, n=1; 7, n=2; 9, n=2). Most species of
<taxonomicName id="4C7B8A86FD43FFD1FCD9F967C09FF933" box="[888,1017,1668,1692]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="16" pageNumber="65" phylum="Chordata" rank="genus">
<emphasis id="B90F2D17FD43FFD1FCD9F967C09FF933" box="[888,1017,1668,1692]" italics="true" pageId="16" pageNumber="65">Apterichtus</emphasis>
</taxonomicName>
have a singular supratemporal pore pattern (except for
<taxonomicName id="4C7B8A86FD43FFD1FED1F94BC2BBF910" box="[368,477,1704,1727]" class="Actinopterygii" family="Ophichthidae" genus="Apterichtus" kingdom="Animalia" order="Anguilliformes" pageId="16" pageNumber="65" phylum="Chordata" rank="species" species="moseri">
<emphasis id="B90F2D17FD43FFD1FED1F94BC2BBF910" box="[368,477,1704,1727]" italics="true" pageId="16" pageNumber="65">A. moseri</emphasis>
</taxonomicName>
, which has 5 or 6 pores,
<tableCitation id="C6F9C4BEFD43FFD1FD5EF944C034F96F" box="[767,850,1703,1728]" captionStart="TABLE 1" captionStartId="5.[151,239,400,423]" captionTargetBox="[151,1159,593,1203]" captionTargetPageId="5" captionText="TABLE 1. Characteristics of the species of Apterichtus. Abbreviations are: MVF = mean vertebral formula; POP = preopercular pore; STP = supratemporal pore. Preanal vertebral counts of A. gracilis and A. monodi are estimates based on preanal lateral-line pores. Sources are: 1, this study; 2, Blache &amp; Bauchot 1972; 3, McCosker et al. 1989; 4, McCosker &amp; Randall 2005; 5, Pereira &amp; Aguiar 1979; 6, Hibino et al. 2014; 7, Machida &amp; Ohta 1994." httpUri="http://table.plazi.org/id/DF04A18DFD56FFC4FF36FE73C701FDA8" pageId="16" pageNumber="65" tableUuid="DF04A18DFD56FFC4FF36FE73C701FDA8">Table 1</tableCitation>
). One of the nine-pored specimens has an irregular row; the standard number of this species can be inferred to be 5 or 7, but it is unclear based on our limited material.
</paragraph>
</subSubSection>
</treatment>
</document>