treatments-xml/data/03/A3/7F/03A37F0CFFEAFFF95BDC07CCA51EFC34.xml

<document id="762A9AF9350A0BE3EFCDD50ECEE751E9" ID-CLB-Dataset="20555" ID-DOI="10.11646/zootaxa.5100.3.7" ID-GBIF-Dataset="52af6630-501b-4db9-8033-197ac1c3d6b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="6201845" ID-ZooBank="C20EFA46-D72C-4C23-8CAE-AC5DD0A27C08" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645430864307" checkinUser="plazi" docAuthor="Cruz, Paulo Vilela, Molineri, Carlos &amp; Hamada, Neusa" docDate="2022" docId="03A37F0CFFEAFFF95BDC07CCA51EFC34" docLanguage="en" docName="zootaxa.5100.3.7.pdf" docOrigin="Zootaxa 5100 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Campsurus fortuitus Cruz &amp; Molineri &amp; Hamada 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="440" masterDocId="FF9A0774FFE9FFFC5B4B020EA47FFF91" masterDocTitle="Two new species and new records of Campsurus Eaton (Ephemeroptera: Polymitarcyidae) in Amazon" masterLastPageNumber="444" masterPageNumber="435" pageNumber="438" updateTime="1699314969245" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0251BB74A6669130D54D9FF6C714CB44">Two new species and new records of Campsurus Eaton (Ephemeroptera: Polymitarcyidae) in Amazon</mods:title>
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<mods:namePart id="B75E00244EF8B9B6A97CFBC5A1DC6978">Cruz, Paulo Vilela</mods:namePart>
<mods:affiliation id="A70416003A5226A5B6D41A8A5A3B59F8">Universidade Federal de Rondônia - UNIR, Programa de Pós-Graduação em Ciências Ambientais - PPGCA, Programa de Pós- Graduação em Conservação e Uso de Recursos Naturais - PPGReN, Laboratório de Biodiversidade e Conservação - LABICON, CEP 76940 - 000, Rolim de Moura, Rondônia, Brazil &amp; Instituto Nacional de Pesquisas da Amazônia - INPA, Coordenação de Pesquisas em Biodiversidade, Laboratório de Citotaxonomia e Insetos Aquáticos - LACIA, CEP 69067 - 375, Manaus, Amazonas, Brazil.</mods:affiliation>
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<mods:namePart id="3534AE7ACF37560B06B69DA1161E742A">Molineri, Carlos</mods:namePart>
<mods:affiliation id="BDAAB02854446DC5B5160A9DF5FEDD8F">Instituto de Biodiversidad Neotropical, CONICET-UNT (Consejo Nacional de Investigaciones Científicas y Técnicas - Universidad Nacional de Tucumán), Facultad de Ciencias Naturales e Instituto M. Lillo, Tucumán, Argentina.</mods:affiliation>
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<mods:namePart id="6EDDBA36B974D4F228DC9A8BFF2F27A1">Hamada, Neusa</mods:namePart>
<mods:nameIdentifier id="4FFDED884564A087A3B5EE183819BC2D" type="ORCID">0000-0002-3526-5426</mods:nameIdentifier>
<mods:affiliation id="D985F666EBAF0B2E172094E466A7B7EF">Instituto Nacional de Pesquisas da Amazônia - INPA, Coordenação de Pesquisas em Biodiversidade, Laboratório de Citotaxonomia e Insetos Aquáticos - LACIA, CEP 69067 - 375, Manaus, Amazonas, Brazil. &amp; neusaha @ gmail. com; https: // orcid. org / 0000 - 0002 - 3526 - 5426</mods:affiliation>
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<treatment id="03A37F0CFFEAFFF95BDC07CCA51EFC34" ID-DOI="http://doi.org/10.5281/zenodo.6314276" ID-GBIF-Taxon="193422462" ID-Zenodo-Dep="6314276" LSID="urn:lsid:plazi:treatment:03A37F0CFFEAFFF95BDC07CCA51EFC34" httpUri="http://treatment.plazi.org/id/03A37F0CFFEAFFF95BDC07CCA51EFC34" lastPageId="5" lastPageNumber="440" pageId="3" pageNumber="438">
<subSubSection id="C3109D91FFEAFFFF5BDC07CCA589FA4C" box="[151,502,1474,1501]" pageId="3" pageNumber="438" type="nomenclature">
<paragraph id="8BB5CE1AFFEAFFFF5BDC07CCA589FA4C" blockId="3.[151,502,1474,1537]" box="[151,502,1474,1501]" pageId="3" pageNumber="438">
<heading id="D0FD7976FFEAFFFF5BDC07CCA589FA4C" bold="true" box="[151,502,1474,1501]" fontSize="11" level="1" pageId="3" pageNumber="438" reason="1">
<emphasis id="B97E1208FFEAFFFF5BDC07CCA589FA4C" bold="true" box="[151,502,1474,1501]" pageId="3" pageNumber="438">
<taxonomicName id="4C0AB599FFEAFFFF5BDC07CCA5EEFA4D" authority="Cruz &amp; Molineri &amp; Hamada, 2022" authorityName="Cruz &amp; Molineri &amp; Hamada" authorityYear="2022" box="[151,401,1474,1500]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="438" phylum="Arthropoda" rank="species" species="fortuitus" status="sp. nov.">
<emphasis id="B97E1208FFEAFFFF5BDC07CCA5EEFA4D" bold="true" box="[151,401,1474,1500]" italics="true" pageId="3" pageNumber="438">Campsurus fortuitus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24DAF73FFEAFFFF5AD207CDA589FA4C" box="[409,502,1475,1501]" pageId="3" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C3109D91FFEAFFFF5BDC07E8A53BF990" box="[151,324,1510,1537]" pageId="3" pageNumber="438" type="description">
<paragraph id="8BB5CE1AFFEAFFFF5BDC07E8A53BF990" blockId="3.[151,502,1474,1537]" box="[151,324,1510,1537]" pageId="3" pageNumber="438">
(
<figureCitation id="1331D29FFFEAFFFF5BD407E8A543F990" box="[159,316,1510,1537]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1968]" captionTargetBox="[235,1350,186,1912]" captionTargetId="figure-18@4.[230,1358,181,1920]" captionTargetPageId="4" captionText="FIGURE 2. Campsurus fortuitus sp. nov. male imago.A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards)." figureDoi="http://doi.org/10.5281/zenodo.6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="3" pageNumber="438">Figures 2 A–I</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3109D91FFEAFFFF5BDC0420A52AF901" pageId="3" pageNumber="438" type="materials_examined">
<paragraph id="8BB5CE1AFFEAFFFF5BDC0420A52AF901" blockId="3.[151,1437,1582,2041]" pageId="3" pageNumber="438">
<emphasis id="B97E1208FFEAFFFF5BDC0420A5FFF9D9" bold="true" box="[151,384,1582,1608]" pageId="3" pageNumber="438">Material examined.</emphasis>
<materialsCitation id="3B62C447FFEAFFFF5ACD0420A7CBF9FD" ID-GBIF-Occurrence="3470039308" collectingDate="2011-08-03" collectionCode="INPA, V" collectorName="da Estrada &amp; A. Pes &amp; Cruz, A" country="Brazil" latitude="0.62747" location="Igarape da Estrada" longLatPrecision="1" longitude="-51.63756" municipality="Pedra Branca" pageId="3" pageNumber="438" specimenCode="BR 210" specimenCount="1" stateProvince="Amapa" typeStatus="holotype">
<typeStatus id="54B170B8FFEAFFFF5ACD0420A58CF9D9" box="[390,499,1582,1608]" pageId="3" pageNumber="438" type="holotype">Holotype</typeStatus>
: male imago (
<collectionCode id="ED1B56DFFFEAFFFF59C70421A6B0F9D8" box="[652,719,1583,1609]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="3" pageNumber="438">INPA</collectionCode>
) from 
<collectingCountry id="F31D8E8AFFEAFFFF58530420A71FF9D9" box="[792,864,1582,1608]" name="Brazil" pageId="3" pageNumber="438">Brazil</collectingCountry>
; 
<collectingRegion id="49CE00F8FFEAFFFF58220421A7C1F9D8" box="[873,958,1583,1609]" country="Brazil" name="Amapa" pageId="3" pageNumber="438">Amapá</collectingRegion>
; 
<collectingMunicipality id="6BD15460FFEAFFFF58830420A01FF9D8" box="[968,1120,1582,1609]" pageId="3" pageNumber="438">Pedra Branca</collectingMunicipality>
; 
<location id="8ED598C1FFEAFFFF5F210421A146F9D9" LSID="urn:lsid:plazi:treatment:03A37F0CFFEAFFF95BDC07CCA51EFC34:8ED598C1FFEAFFFF5F210421A146F9D9" box="[1130,1337,1582,1609]" country="Brazil" latitude="0.62747" longLatPrecision="1" longitude="-51.63756" municipality="Pedra Branca" name="Igarape da Estrada" pageId="3" pageNumber="438" stateProvince="Amapa">
Igarapé 
<collectorName id="26FFABCCFFEAFFFF5F8F0420A146F9D9" box="[1220,1337,1582,1608]" pageId="3" pageNumber="438">da Estrada</collectorName>
</location>
<specimenCode id="DBAC6661FFEAFFFF5E740421A1E8F9D8" box="[1343,1431,1583,1609]" collectionCode="BR" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="3" pageNumber="438">BR 210</specimenCode>
; 
<geoCoordinate id="EE3EA8DDFFEAFFFF5BDC045CA48BF9FC" box="[151,244,1618,1645]" degrees="0.62747" direction="north" orientation="latitude" pageId="3" pageNumber="438" precision="1" value="0.62747">0.62747</geoCoordinate>
/ 
<geoCoordinate id="EE3EA8DDFFEAFFFF5BB6045CA50FF9FC" box="[253,368,1618,1645]" degrees="51.63756" direction="west" orientation="longitude" pageId="3" pageNumber="438" precision="1" value="-51.63756">-51.63756</geoCoordinate>
; 
<date id="FFB4E8DAFFEAFFFF5A36045CA679F9FD" box="[381,518,1618,1644]" pageId="3" pageNumber="438" value="2011-08-03">
<collectingDate id="EFF01132FFEAFFFF5A36045CA679F9FD" box="[381,518,1618,1644]" pageId="3" pageNumber="438" value="2011-08-03">03.viii.2011</collectingDate>
</date>
; 
<collectorName id="26FFABCCFFEAFFFF5959045DA620F9FC" box="[530,607,1619,1645]" pageId="3" pageNumber="438">A. Pes</collectorName>
, P. 
<collectionCode id="ED1B56DFFFEAFFFF5934045DA6EBF9FC" box="[639,660,1619,1645]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="438" type="Museum">V</collectionCode>
. 
<collectorName id="26FFABCCFFEAFFFF59D4045DA684F9FC" box="[671,763,1619,1645]" pageId="3" pageNumber="438">Cruz, A</collectorName>
. Fernandes cols
</materialsCitation>
. 
<materialsCitation id="3B62C447FFEAFFFF588B045DA52EF901" ID-GBIF-Occurrence="3470039313" collectingDate="2011-08-03" collectionCode="INPA" collectorName="da Estrada &amp; A. Pes &amp; Cruz, A" country="Brazil" latitude="0.62747" location="Igarape da Estrada" longLatPrecision="1" longitude="-51.63756" municipality="Pedra Branca" pageId="3" pageNumber="438" specimenCount="1" stateProvince="Amapa" typeStatus="paratype">
<typeStatus id="54B170B8FFEAFFFF588B045DA04EF9FC" box="[960,1073,1619,1645]" pageId="3" pageNumber="438" type="paratype">Paratypes</typeStatus>
: One male imago (
<collectionCode id="ED1B56DFFFEAFFFF5E42045DA131F9FC" box="[1289,1358,1619,1645]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="3" pageNumber="438">INPA</collectionCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3109D91FFEAFFFF5B8C0494A55DF8F8" pageId="3" pageNumber="438" type="diagnosis">
<paragraph id="8BB5CE1AFFEAFFFF5B8C0494A55DF8F8" blockId="3.[151,1437,1582,2041]" pageId="3" pageNumber="438">
<emphasis id="B97E1208FFEAFFFF5B8C0494A59EF925" bold="true" box="[199,481,1690,1717]" pageId="3" pageNumber="438">Diagnosis. Male imago.</emphasis>
1) sternum IX with posterior margin almost straight (
<figureCitation id="1331D29FFFEAFFFF5F040494A0D3F924" box="[1103,1196,1690,1717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1968]" captionTargetBox="[235,1350,186,1912]" captionTargetId="figure-18@4.[230,1358,181,1920]" captionTargetPageId="4" captionText="FIGURE 2. Campsurus fortuitus sp. nov. male imago.A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards)." figureDoi="http://doi.org/10.5281/zenodo.6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="3" pageNumber="438">Fig. 2H</figureCitation>
); 2) pedestal of each side touching basally, inner projection short, thick, almost rounded, and with outer margin straight, outer projection short, slender and with apex broadly pointed (
<figureCitation id="1331D29FFFEAFFFF59DF04ECA736F96C" box="[660,841,1762,1789]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1968]" captionTargetBox="[235,1350,186,1912]" captionTargetId="figure-18@4.[230,1358,181,1920]" captionTargetPageId="4" captionText="FIGURE 2. Campsurus fortuitus sp. nov. male imago.A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards)." figureDoi="http://doi.org/10.5281/zenodo.6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="3" pageNumber="438">Figs 2D and 2H</figureCitation>
); 3) base of penis wide proximally, narrowing toward apex (
<figureCitation id="1331D29FFFEAFFFF5B970508A5E8F8B0" box="[220,407,1798,1825]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1968]" captionTargetBox="[235,1350,186,1912]" captionTargetId="figure-18@4.[230,1358,181,1920]" captionTargetPageId="4" captionText="FIGURE 2. Campsurus fortuitus sp. nov. male imago.A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards)." figureDoi="http://doi.org/10.5281/zenodo.6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="3" pageNumber="438">Figs 2D and 2H</figureCitation>
); 4) main lobe of penis bladelike, slightly curved ventrally and with distoventral gonopore (
<figureCitation id="1331D29FFFEAFFFF5BD40524A566F8D4" box="[159,281,1834,1861]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1968]" captionTargetBox="[235,1350,186,1912]" captionTargetId="figure-18@4.[230,1358,181,1920]" captionTargetPageId="4" captionText="FIGURE 2. Campsurus fortuitus sp. nov. male imago.A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards)." figureDoi="http://doi.org/10.5281/zenodo.6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="3" pageNumber="438">Figs 2D–I</figureCitation>
); 5) penis apex with expansion curved outwards (
<figureCitation id="1331D29FFFEAFFFF58130524A7AEF8D4" box="[856,977,1834,1861]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1968]" captionTargetBox="[235,1350,186,1912]" captionTargetId="figure-18@4.[230,1358,181,1920]" captionTargetPageId="4" captionText="FIGURE 2. Campsurus fortuitus sp. nov. male imago.A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards)." figureDoi="http://doi.org/10.5281/zenodo.6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="3" pageNumber="438">Figs 2D–I</figureCitation>
); 6) secondary lobe completely reduced (
<figureCitation id="1331D29FFFEAFFFF5BD40540A569F8F8" box="[159,278,1870,1897]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1968]" captionTargetBox="[235,1350,186,1912]" captionTargetId="figure-18@4.[230,1358,181,1920]" captionTargetPageId="4" captionText="FIGURE 2. Campsurus fortuitus sp. nov. male imago.A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards)." figureDoi="http://doi.org/10.5281/zenodo.6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="3" pageNumber="438">Figs 2D–I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3109D91FFEAFFF95B8C057CA157FDF3" lastPageId="5" lastPageNumber="440" pageId="3" pageNumber="438" type="description">
<paragraph id="8BB5CE1AFFEAFFF95B8C057CA157FDF3" blockId="3.[151,1437,1582,2041]" lastBlockId="5.[151,1437,151,933]" lastPageId="5" lastPageNumber="440" pageId="3" pageNumber="438">
<emphasis id="B97E1208FFEAFFFF5B8C057CA587F81D" bold="true" box="[199,504,1906,1933]" pageId="3" pageNumber="438">Description. Male imago.</emphasis>
Length (n=1, mm): body, 7.8; fore wing, 5.7; hind wing, 2.4. General coloration yellowish grey (
<figureCitation id="1331D29FFFEAFFFF5A000598A5DBF820" box="[331,420,1942,1969]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1968]" captionTargetBox="[235,1350,186,1912]" captionTargetId="figure-18@4.[230,1358,181,1920]" captionTargetPageId="4" captionText="FIGURE 2. Campsurus fortuitus sp. nov. male imago.A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards)." figureDoi="http://doi.org/10.5281/zenodo.6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="3" pageNumber="438">Fig. 2A</figureCitation>
). Head completely shaded with black dorsally, ventrally whitish, slightly tinged with black. Antenna with scape and pedicel grey, flagellum lost. Thorax. Pronotum shaded with gray, with paler transversal line on posterior margin; prosternum whitish, propleura whitish, shaded with gray. Mesonotum yellowish white with blackish medial line, parapsidal suture and posteroscutal protuberance slightly shaded with gray; pleura and sterna whitish, shaded with black; and gray around coxa. Metanotum yellowish. Legs. Foreleg greyish, except distal tarsomere and claws slightly paler. Middle and hind legs yellowish. Wings (
<figureCitation id="1331D29FFFECFFF958E502D1A01EFF68" box="[942,1121,223,249]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1968]" captionTargetBox="[235,1350,186,1912]" captionTargetId="figure-18@4.[230,1358,181,1920]" captionTargetPageId="4" captionText="FIGURE 2. Campsurus fortuitus sp. nov. male imago.A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards)." figureDoi="http://doi.org/10.5281/zenodo.6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="5" pageNumber="440">Figs 2B and 2C</figureCitation>
). Fore wing with membrane hyaline, shaded with light grey on C and Sc areas; veins grey; cross veins hyaline; MP 
<subScript id="178ECC5FFFECFFF95F15031BA019FEB2" attach="left" box="[1118,1126,277,291]" fontSize="6" pageId="5" pageNumber="440">2</subScript>
not connected to IMP, ICu 
<subScript id="178ECC5FFFECFFF95EDF031BA1E3FEB2" attach="left" box="[1428,1436,277,291]" fontSize="6" pageId="5" pageNumber="440">1</subScript>
not connected to CuA, ICu 
<subScript id="178ECC5FFFECFFF95A8D0337A5B1FED6" attach="left" box="[454,462,313,327]" fontSize="6" pageId="5" pageNumber="440">2</subScript>
directly connected to CuA. Hind wing similar to fore wing, but paler; intercalary vein of radial sector very short. Abdomen whitish translucent, shaded with gray, except medial line, intersegmental membranes, and lateral transversal dash on terga II–VII (
<figureCitation id="1331D29FFFECFFF959AA0361A745FE18" box="[737,826,367,393]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1968]" captionTargetBox="[235,1350,186,1912]" captionTargetId="figure-18@4.[230,1358,181,1920]" captionTargetPageId="4" captionText="FIGURE 2. Campsurus fortuitus sp. nov. male imago.A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards)." figureDoi="http://doi.org/10.5281/zenodo.6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="5" pageNumber="440">Fig. 2A</figureCitation>
) Abdominal sterna whitish, shaded with gray on inner margin of gill sclerites and very slightly on meson of sterna VII–IX. Genitalia (
<figureCitation id="1331D29FFFECFFF95F41039DA0FDFE3C" box="[1034,1154,403,429]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1968]" captionTargetBox="[235,1350,186,1912]" captionTargetId="figure-18@4.[230,1358,181,1920]" captionTargetPageId="4" captionText="FIGURE 2. Campsurus fortuitus sp. nov. male imago.A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards)." figureDoi="http://doi.org/10.5281/zenodo.6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="5" pageNumber="440">Figs 2D–I</figureCitation>
): sternum IX with almost straight posterior margin; pedestal of each side touching basally, inner projection short, thick, almost rounded, and with outer margin straight, outer projection short, slender and with apex broadly pointed; base of penis wide proximally, narrowing toward apex; main lobe of penis slender, bladelike and sclerotized, slightly curved ventrally and with distoventral gonopore, inner margin of apex (
<figureCitation id="1331D29FFFECFFF9584B002DA7A8FDAC" box="[768,983,547,573]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1968]" captionTargetBox="[235,1350,186,1912]" captionTargetId="figure-18@4.[230,1358,181,1920]" captionTargetPageId="4" captionText="FIGURE 2. Campsurus fortuitus sp. nov. male imago.A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards)." figureDoi="http://doi.org/10.5281/zenodo.6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="5" pageNumber="440">Figures 2D and 2E</figureCitation>
) with expansion curved outwards (eco), secondary lobe completely reduced, apical half of penis orangish. (Caudal filaments broken off and lost)
</paragraph>
</subSubSection>
<caption id="DF759E92FFEDFFF85BDC0596A0D6F869" ID-DOI="http://doi.org/10.5281/zenodo.6201857" ID-Zenodo-Dep="6201857" httpUri="https://zenodo.org/record/6201857/files/figure.png" pageId="4" pageNumber="439" startId="4.[151,250,1944,1968]" targetBox="[235,1350,186,1912]" targetPageId="4">
<paragraph id="8BB5CE1AFFEDFFF85BDC0596A0D6F869" blockId="4.[151,1436,1944,2040]" pageId="4" pageNumber="439">
<emphasis id="B97E1208FFEDFFF85BDC0596A56DF821" bold="true" box="[151,274,1944,1968]" pageId="4" pageNumber="439">FIGURE 2.</emphasis>
<taxonomicName id="4C0AB599FFEDFFF85A530596A598F821" authority="Cruz &amp; Molineri &amp; Hamada, 2022" authorityName="Cruz &amp; Molineri &amp; Hamada" authorityYear="2022" box="[280,487,1944,1968]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="439" phylum="Arthropoda" rank="species" species="fortuitus" status="sp. nov.">
<emphasis id="B97E1208FFEDFFF85A530596A598F821" box="[280,487,1944,1968]" italics="true" pageId="4" pageNumber="439">Campsurus fortuitus</emphasis>
</taxonomicName>
<emphasis id="B97E1208FFEDFFF85AA60596A642F821" bold="true" box="[493,573,1944,1968]" pageId="4" pageNumber="439">
<taxonomicNameLabel id="A24DAF73FFEDFFF85AA60596A642F821" box="[493,573,1944,1968]" pageId="4" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male imago. A. dorsal view; B. forewing; C. hind wing; D. half of genitalia in ventral view; E. half of genitalia in ventrolateral view; F. half of genitalia in lateral view; G. penis ventral view - twisted in stereoscope (paratype); H. genitalia in ventral view; I. genitalia in lateral view. (eco = expansion curved outwards).
</paragraph>
</caption>
<subSubSection id="C3109D91FFECFFF95B8C0065A540FD38" pageId="5" pageNumber="440" type="etymology">
<paragraph id="8BB5CE1AFFECFFF95B8C0065A540FD38" blockId="5.[151,1437,151,933]" pageId="5" pageNumber="440">
<emphasis id="B97E1208FFECFFF95B8C0065A531FD14" bold="true" box="[199,334,619,645]" pageId="5" pageNumber="440">Etymology.</emphasis>
The word “fortuit” in Latin means “random”. The species was collected in a random stop in the road for snack.
</paragraph>
</subSubSection>
<subSubSection id="C3109D91FFECFFF95B8C00BCA51EFC34" pageId="5" pageNumber="440" type="discussion">
<paragraph id="8BB5CE1AFFECFFF95B8C00BCA51EFC34" blockId="5.[151,1437,151,933]" pageId="5" pageNumber="440">
<emphasis id="B97E1208FFECFFF95B8C00BCA533FD5D" bold="true" box="[199,332,690,716]" pageId="5" pageNumber="440">Discussion.</emphasis>
<taxonomicName id="4C0AB599FFECFFF95A1C00BDA63FFD5C" authority="Cruz &amp; Molineri &amp; Hamada, 2022" authorityName="Cruz &amp; Molineri &amp; Hamada" authorityYear="2022" box="[343,576,691,717]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="fortuitus" status="sp. nov.">
<emphasis id="B97E1208FFECFFF95A1C00BDA63FFD5C" box="[343,576,691,717]" italics="true" pageId="5" pageNumber="440">Campsurus fortuitus</emphasis>
</taxonomicName>
<emphasis id="B97E1208FFECFFF9590100BDA6D8FD5C" bold="true" box="[586,679,691,717]" pageId="5" pageNumber="440">
<taxonomicNameLabel id="A24DAF73FFECFFF9590100BDA6D8FD5C" box="[586,679,691,717]" pageId="5" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has unusual genital characteristics, similar to 
<taxonomicName id="4C0AB599FFECFFF95F8F00BDA51AFD60" authority="Molineri &amp; Salles" authorityName="Molineri &amp; Salles" authorityYear="2017" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="povilla">
<emphasis id="B97E1208FFECFFF95F8F00BDA1E3FD5C" box="[1220,1436,691,717]" italics="true" pageId="5" pageNumber="440">Campsurus povilla</emphasis>
Molineri &amp; Salles
</taxonomicName>
, but they can be distinguished by i) penis with wide base, narrowing distally in 
<taxonomicName id="4C0AB599FFECFFF95FB900D9A10CFD60" authority="Cruz &amp; Molineri &amp; Hamada, 2022" authorityName="Cruz &amp; Molineri &amp; Hamada" authorityYear="2022" box="[1266,1395,727,753]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="fortuitus" status="sp. nov.">
<emphasis id="B97E1208FFECFFF95FB900D9A10CFD60" box="[1266,1395,727,753]" italics="true" pageId="5" pageNumber="440">C. fortuitus</emphasis>
</taxonomicName>
<emphasis id="B97E1208FFECFFF95E3000D9A4AFFC84" bold="true" pageId="5" pageNumber="440">
<taxonomicNameLabel id="A24DAF73FFECFFF95E3000D9A4B5FC84" pageId="5" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
while abruptly thinning basal to penile lobes in 
<taxonomicName id="4C0AB599FFECFFF959B100F5A715FC84" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[762,874,763,789]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="povilla">
<emphasis id="B97E1208FFECFFF959B100F5A715FC84" box="[762,874,763,789]" italics="true" pageId="5" pageNumber="440">C. povilla</emphasis>
</taxonomicName>
; ii) main lobe of penis slender in 
<taxonomicName id="4C0AB599FFECFFF95FBA00F5A10DFC84" authority="Cruz &amp; Molineri &amp; Hamada, 2022" authorityName="Cruz &amp; Molineri &amp; Hamada" authorityYear="2022" box="[1265,1394,763,789]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="fortuitus" status="sp. nov.">
<emphasis id="B97E1208FFECFFF95FBA00F5A10DFC84" box="[1265,1394,763,789]" italics="true" pageId="5" pageNumber="440">C. fortuitus</emphasis>
</taxonomicName>
<emphasis id="B97E1208FFECFFF95E3000F5A4B7FCA8" bold="true" pageId="5" pageNumber="440">
<taxonomicNameLabel id="A24DAF73FFECFFF95E3000F5A4B7FCA8" pageId="5" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, while robust in 
<taxonomicName id="4C0AB599FFECFFF95ACC0111A588FCA8" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[391,503,799,825]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="povilla">
<emphasis id="B97E1208FFECFFF95ACC0111A588FCA8" box="[391,503,799,825]" italics="true" pageId="5" pageNumber="440">C. povilla</emphasis>
</taxonomicName>
; iii) main lobe of penis with inner margin apex curved outwards in 
<taxonomicName id="4C0AB599FFECFFF95FB80111A10CFCA8" authority="Cruz &amp; Molineri &amp; Hamada, 2022" authorityName="Cruz &amp; Molineri &amp; Hamada" authorityYear="2022" box="[1267,1395,799,825]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="fortuitus" status="sp. nov.">
<emphasis id="B97E1208FFECFFF95FB80111A10CFCA8" box="[1267,1395,799,825]" italics="true" pageId="5" pageNumber="440">C. fortuitus</emphasis>
</taxonomicName>
<emphasis id="B97E1208FFECFFF95E300111A4B7FCCC" bold="true" pageId="5" pageNumber="440">
<taxonomicNameLabel id="A24DAF73FFECFFF95E300111A4B7FCCC" pageId="5" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, while apex rounded in 
<taxonomicName id="4C0AB599FFECFFF95A9B014DA642FCCC" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[464,573,835,861]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="povilla">
<emphasis id="B97E1208FFECFFF95A9B014DA642FCCC" box="[464,573,835,861]" italics="true" pageId="5" pageNumber="440">C. povilla</emphasis>
</taxonomicName>
; iv) secondary lobe of penis completely reduced in 
<taxonomicName id="4C0AB599FFECFFF95F3E014DA08BFCCC" authority="Cruz &amp; Molineri &amp; Hamada, 2022" authorityName="Cruz &amp; Molineri &amp; Hamada" authorityYear="2022" box="[1141,1268,835,861]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="fortuitus" status="sp. nov.">
<emphasis id="B97E1208FFECFFF95F3E014DA08BFCCC" box="[1141,1268,835,861]" italics="true" pageId="5" pageNumber="440">C. fortuitus</emphasis>
</taxonomicName>
<emphasis id="B97E1208FFECFFF95FB2014DA12EFCCC" bold="true" box="[1273,1361,835,861]" pageId="5" pageNumber="440">
<taxonomicNameLabel id="A24DAF73FFECFFF95FB2014DA12EFCCC" box="[1273,1361,835,861]" pageId="5" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, while greatly reduced but clearly present in 
<taxonomicName id="4C0AB599FFECFFF959750169A6D2FC10" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[574,685,871,897]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="povilla">
<emphasis id="B97E1208FFECFFF959750169A6D2FC10" box="[574,685,871,897]" italics="true" pageId="5" pageNumber="440">C. povilla</emphasis>
</taxonomicName>
; v) pedestal in 
<taxonomicName id="4C0AB599FFECFFF958120169A7A6FC10" authority="Cruz &amp; Molineri &amp; Hamada, 2022" authorityName="Cruz &amp; Molineri &amp; Hamada" authorityYear="2022" box="[857,985,871,897]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="fortuitus" status="sp. nov.">
<emphasis id="B97E1208FFECFFF958120169A7A6FC10" box="[857,985,871,897]" italics="true" pageId="5" pageNumber="440">C. fortuitus</emphasis>
</taxonomicName>
<emphasis id="B97E1208FFECFFF958AB0169A045FC10" bold="true" box="[992,1082,871,897]" pageId="5" pageNumber="440">
<taxonomicNameLabel id="A24DAF73FFECFFF958AB0169A045FC10" box="[992,1082,871,897]" pageId="5" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with inner projection straighter than in 
<taxonomicName id="4C0AB599FFECFFF95BA00185A525FC34" authorityName="Molineri &amp; Salles" authorityYear="2017" box="[235,346,907,933]" class="Insecta" family="Polymitarcyidae" genus="Campsurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="povilla">
<emphasis id="B97E1208FFECFFF95BA00185A525FC34" box="[235,346,907,933]" italics="true" pageId="5" pageNumber="440">C. povilla</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>