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<document id="9C8902E3D62572A3528343F16552A746" ID-DOI="10.5281/zenodo.12753216" ID-GBIF-Dataset="b05b7a19-692a-402e-a30e-eae2072a494a" ID-ISSN="2713-8615" ID-Zenodo-Dep="12753216" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1721182221815" checkinUser="felipe" docAuthor="Blakemore, R. J." docDate="2012" docId="03E487FD3F5E36536DD9FE74FEA2FEDB" docLanguage="en" docName="JSpeciesRes.1.1.35-43.pdf" docOrigin="Journal of Species Research 1 (1)" docStyle="DocumentStyle:4D398BFF45FF8CA9E08F546BB06AE372.2:JSpeciesRes.2012-.journal_article" docStyleId="4D398BFF45FF8CA9E08F546BB06AE372" docStyleName="JSpeciesRes.2012-.journal_article" docStyleVersion="2" docTitle="Amynthas carnosus NSMT An" docType="treatment" docVersion="5" lastPageNumber="38" masterDocId="FFDDFF853F5F36566D7BFF9EFFD4FFBB" masterDocTitle="Amynthas carnosus (Goto &amp; Hatai, 1899) redescribed on its neotype (Oligochaeta: Megadrilacea: Megascolecidae)" masterLastPageNumber="43" masterPageNumber="35" pageNumber="36" updateTime="1721394291871" updateUser="guilherme" zenodo-license-document="CC-BY-4.0">
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<mods:title id="266D00C1059F31178EA13D089EF6B80C">Amynthas carnosus (Goto &amp; Hatai, 1899) redescribed on its neotype (Oligochaeta: Megadrilacea: Megascolecidae)</mods:title>
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<mods:namePart id="0AA2FCFAE554CFCD7BEB2BD8A2149206">Blakemore, R. J.</mods:namePart>
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<mods:date id="D74E56BFE0002FC7B4C25B52DB94C7B0">2012</mods:date>
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<treatment id="03E487FD3F5E36536DD9FE74FEA2FEDB" ID-DOI="http://doi.org/10.5281/zenodo.12784057" ID-Zenodo-Dep="12784057" LSID="urn:lsid:plazi:treatment:03E487FD3F5E36536DD9FE74FEA2FEDB" httpUri="http://treatment.plazi.org/id/03E487FD3F5E36536DD9FE74FEA2FEDB" lastPageId="5" lastPageNumber="38" pageId="1" pageNumber="36">
<subSubSection id="C35765603F5E36576DD9FE74FDBBFDB9" box="[162,623,490,514]" pageId="1" pageNumber="36" type="nomenclature">
<paragraph id="8BF236EB3F5E36576DD9FE74FDBBFDB9" blockId="1.[162,768,490,1991]" box="[162,623,490,514]" pageId="1" pageNumber="36">
<heading id="D0BA81873F5E36576DD9FE74FDBBFDB9" bold="true" box="[162,623,490,514]" fontSize="10" level="2" pageId="1" pageNumber="36" reason="4">
<taxonomicName id="4C4D4D683F5E36576DD9FE74FDBBFDB9" authority="(Goto &amp; Hatai, 1899)" authorityName="NSMT An" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" box="[162,623,490,514]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B939EAF93F5E36576DD9FE74FDBBFDB9" bold="true" box="[162,623,490,514]" pageId="1" pageNumber="36">
<emphasis id="B939EAF93F5E36576DD9FE74FEAFFDB9" bold="true" box="[162,379,490,514]" italics="true" pageId="1" pageNumber="36">Amynthas carnosus</emphasis>
(
<bibRefCitation id="EFDC4B1A3F5E36576CFDFE74FDBCFDB9" author="Goto, S. &amp; S. Hatai" box="[390,616,490,514]" pageId="1" pageNumber="36" pagination="13 - 24" refId="ref6050" refString="Goto, S. and S. Hatai. 1899. New or imperfectly known species of earthworms. No. 2. Annotations Zoologicae Japonensis 3 (1): 13 - 24." type="journal article" year="1899">Goto &amp; Hatai, 1899</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35765603F5E36576DD9FD94FBFBF91D" pageId="1" pageNumber="36" type="reference_group">
<paragraph id="8BF236EB3F5E36576DD9FD94FEA7FD19" blockId="1.[162,768,490,1991]" pageId="1" pageNumber="36">
<treatmentCitationGroup id="AB5D11C53F5E36576DD9FD94FEA7FD19" pageId="1" pageNumber="36">
<treatmentCitation id="0AEC10FA3F5E36576DD9FD94FD54FD9A" author="Goto, S. &amp; S. Hatai" box="[162,640,522,545]" page="15" pageId="1" pageNumber="36" year="1899">
<taxonomicName id="4C4D4D683F5E36576DD9FD94FD54FD9A" authority="Goto &amp; Hatai, 1899: 15" authorityName="Goto &amp; Hatai" authorityPageNumber="15" authorityYear="1899" box="[162,640,522,545]" class="Clitellata" family="Megascolecidae" genus="Perichaeta" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="carnosa">
<emphasis id="B939EAF93F5E36576DD9FD94FEA1FD9A" box="[162,373,522,545]" italics="true" pageId="1" pageNumber="36">Perichaeta carnosa</emphasis>
<bibRefCitation id="EFDC4B1A3F5E36576C06FD94FD54FD9A" author="Goto, S. &amp; S. Hatai" box="[381,640,522,545]" pageId="1" pageNumber="36" pagination="13 - 24" refId="ref6050" refString="Goto, S. and S. Hatai. 1899. New or imperfectly known species of earthworms. No. 2. Annotations Zoologicae Japonensis 3 (1): 13 - 24." type="journal article" year="1899">Goto &amp; Hatai, 1899: 15</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 24, figs. 4, 5. [From
<collectingRegion id="4989F8093F5E36576C63FDB4FEB4FDFA" box="[280,352,554,577]" country="Japan" name="Tokyo" pageId="1" pageNumber="36">Tokyo</collectingRegion>
. Described on
<specimenCount id="9D4BFD623F5E36576C81FDB4FD41FDFA" box="[506,661,554,577]" count="2" pageId="1" pageNumber="36" type="generic">two specimens</specimenCount>
- see Note below.
<typeStatus id="54F688493F5E36576C7CFDD4FE9CFDDA" box="[263,328,586,609]" pageId="1" pageNumber="36">Types</typeStatus>
not known despite extensive searches by current author for 10 yrs in
<collectingCountry id="F35A767B3F5E36576C83FDF4FDE1FD3A" box="[504,565,618,641]" name="Japan" pageId="1" pageNumber="36">Japan</collectingCountry>
(cf.
<bibRefCitation id="EFDC4B1A3F5E36576F11FDF4FEB4FD19" author="Blakemore, R. J. &amp; R. Ueshima" pageId="1" pageNumber="36" pagination="59 - 83" refId="ref5419" refString="Blakemore, R. J. and R. Ueshima. 2011. Catalogue of Annelida: Oligochaeta in University of Tokyo Zoology Museum (ZMUT). In: R. Ushima (ed.), Catalogue of Invertebrate Collection Deposited in the Department of Zoology, the University Museum, the University of Tokyo (3), Phylum Annelida (Class Polychaeta, Oligochaeta, and Hirudinida). Material Reports, No. 90, 59 - 83 [Available from: http: // umdb. um. u-tokyo. ac. jp / DDoubutu / invertebrate _ en / index. html]." type="book chapter" year="2011">Blakemore &amp; Ueshima, 2011</bibRefCitation>
)].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF236EB3F5E36576DD9FD35FEF8FD59" blockId="1.[162,768,490,1991]" pageId="1" pageNumber="36">
<taxonomicName id="4C4D4D683F5E36576DD9FD35FEABFD79" baseAuthorityName="Rosa" baseAuthorityYear="1891" box="[162,383,683,706]" class="Clitellata" family="Megascolecidae" genus="Amyntas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="hawayanus">
<emphasis id="B939EAF93F5E36576DD9FD35FEABFD79" box="[162,383,683,706]" italics="true" pageId="1" pageNumber="36">Amyntas hawayanus</emphasis>
</taxonomicName>
(part.): Beddard, 1900: 645; Gates, 1932: 433.
</paragraph>
<paragraph id="8BF236EB3F5E36576DD9FD75FEFDFB98" blockId="1.[162,768,490,1991]" pageId="1" pageNumber="36">
<taxonomicName id="4C4D4D683F5E36576DD9FD75FEB9FCB9" authority="(Goto &amp; Hatai, 1899)" box="[162,365,747,770]" class="Clitellata" family="Megascolecidae" genus="Pheretima" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="carnosa">
<emphasis id="B939EAF93F5E36576DD9FD75FEB9FCB9" box="[162,365,747,770]" italics="true" pageId="1" pageNumber="36">Pheretima carnosa</emphasis>
</taxonomicName>
: Michaelsen, 1900: 260;
<treatmentCitation id="0AEC10FA3F5E36576F06FD75FF34FC99" author="Chen, Y." page="274" pageId="1" pageNumber="36" year="1936">
<bibRefCitation id="EFDC4B1A3F5E36576F06FD75FF34FC99" author="Chen, Y." pageId="1" pageNumber="36" pagination="269 - 306" refId="ref5800" refString="Chen, Y. 1936. On the terrestrial Oligochaeta from Szechuan II with the notes on Gates' types. Contribibutions of Biological Lababoratory of Science Society of China. Zoology 11: 269 - 306." type="journal article" year="1936">Chen, 1936: 274</bibRefCitation>
</treatmentCitation>
(when describing
<taxonomicName id="4C4D4D683F5E36576CE5FC95FD4CFC99" baseAuthorityName="Chen" baseAuthorityYear="1936" box="[414,664,779,802]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="subSpecies" species="pingi" subSpecies="chungkingensis">
<emphasis id="B939EAF93F5E36576CE5FC95FD4CFC99" box="[414,664,779,802]" italics="true" pageId="1" pageNumber="36">P. pingi chungkingensis</emphasis>
</taxonomicName>
, that possibly merits elevation to species level, remarked that
<taxonomicName id="4C4D4D683F5E36576DC2FCD2FE05FCD9" authority="Stephenson, 1925" authorityName="Stephenson" authorityYear="1925" box="[185,465,843,867]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="pingi">
<emphasis id="B939EAF93F5E36576DC2FCD2FED9FCD8" box="[185,269,844,867]" italics="true" pageId="1" pageNumber="36">P. pingi</emphasis>
Stephenson, 1925
</taxonomicName>
was probably a synonym of
<taxonomicName id="4C4D4D683F5E36576DC2FCF2FEDBFC38" authority="(Goto &amp; Hatai, 1899)" box="[185,271,876,899]" class="Clitellata" family="Megascolecidae" genus="Pheretima" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="carnosa">
<emphasis id="B939EAF93F5E36576DC2FCF2FEDBFC38" box="[185,271,876,899]" italics="true" pageId="1" pageNumber="36">carnosa</emphasis>
</taxonomicName>
);
<treatmentCitation id="0AEC10FA3F5E36576C5CFCF2FDF4FC38" author="Kobayashi, S." box="[295,544,876,899]" page="115" pageId="1" pageNumber="36" year="1936">
<bibRefCitation id="EFDC4B1A3F5E36576C5CFCF2FDF4FC38" author="Kobayashi, S." box="[295,544,876,899]" pageId="1" pageNumber="36" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi, 1936a: 115</bibRefCitation>
</treatmentCitation>
, tab. 1, text figs. 1-3 (syns.
<emphasis id="B939EAF93F5E36576D86FC12FE4AFC18" box="[253,414,908,931]" italics="true" pageId="1" pageNumber="36">
<taxonomicName id="4C4D4D683F5E36576D86FC12FEE1FC18" authorityName="Stephenson" authorityYear="1925" box="[253,309,908,931]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="pingi">pingi</taxonomicName>
,
<taxonomicName id="4C4D4D683F5E36576C3BFC12FE4AFC18" authorityName="Kobayashi" authorityYear="1934" box="[320,414,908,931]" class="Clitellata" family="Megascolecidae" genus="Pheretima" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="kyamikia">kyamikia</taxonomicName>
</emphasis>
), 1938: 161;
<bibRefCitation id="EFDC4B1A3F5E36576F57FC12FD12FC18" author="Ohfuchi, S." box="[556,710,908,931]" pageId="1" pageNumber="36" pagination="31 - 136" refId="ref6660" refString="Ohfuchi, S. 1937. On the species possessing four pairs of spermathecae in the genus Pheretima, together with the variability of some external and internal characteristics. Saito Ho-On Kai Museum Research Bulletin 12: 31 - 136." type="journal article" year="1937">Ohfuchi, 1937</bibRefCitation>
b: 56, fig. 9, Pl. 1, 4;
<treatmentCitation id="0AEC10FA3F5E36576C20FC32FD9EFC78" author="Kobayashi, S." box="[347,586,940,963]" page="161" pageId="1" pageNumber="36" year="1938">
<bibRefCitation id="EFDC4B1A3F5E36576C20FC32FD9EFC78" author="Kobayashi, S." box="[347,586,940,963]" pageId="1" pageNumber="36" pagination="89 - 170" refId="ref6515" refString="Kobayashi, S. 1938. Earthworms of Korea I. Science Report of the Tohoku Imperial University 13 (2): 89 - 170." type="journal article" year="1938">Kobayashi, 1938: 161</bibRefCitation>
</treatmentCitation>
;?
<treatmentCitation id="0AEC10FA3F5E36576F24FC32FD2EFC78" author="Chen, Y." box="[607,762,940,963]" page="9" pageId="1" pageNumber="36" year="1959">
<bibRefCitation id="EFDC4B1A3F5E36576F24FC32FD2EFC78" author="Chen, Y." box="[607,762,940,963]" pageId="1" pageNumber="36" refId="ref5837" refString="Chen, Y. 1959. Chinese Animal Atlas. Annelida. Science Press, Beijing [In Chinese]." type="book" year="1959">Chen, 1959: 9</bibRefCitation>
</treatmentCitation>
, fig. 9 (part. syn.
<taxonomicName id="4C4D4D683F5E36576C0CFC52FE7BFC58" authorityName="Stephenson" authorityYear="1925" box="[375,431,972,995]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="pingi">
<emphasis id="B939EAF93F5E36576C0CFC52FE7BFC58" box="[375,431,972,995]" italics="true" pageId="1" pageNumber="36">pingi</emphasis>
</taxonomicName>
);
<treatmentCitation id="0AEC10FA3F5E36576CB1FC52FD59FC58" author="Gates, G. E." box="[458,653,972,995]" page="149" pageId="1" pageNumber="36" year="1972">
<bibRefCitation id="EFDC4B1A3F5E36576CB1FC52FD59FC58" author="Gates, G. E." box="[458,653,972,995]" pageId="1" pageNumber="36" pagination="1 - 326" refId="ref5988" refString="Gates, G. E. 1972. Burmese Earthworms, an introduction to the systematics and biology of Megadrile oligochaetes with special reference to South-East Asia. Transactions of the American Philosophical Society 62 (7): 1 - 326 [Available from: http: // www. jstor. org / stable / 1006214 in 2011]." type="journal article" year="1972">Gates, 1972: 149</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AEC10FA3F5E36576FE7FC52FECEFBB8" author="Ishizuka, K." page="75" pageId="1" pageNumber="36" year="2001">
<bibRefCitation id="EFDC4B1A3F5E36576FE7FC52FECEFBB8" author="Ishizuka, K." pageId="1" pageNumber="36" pagination="1 - 125" refId="ref6222" refString="Ishizuka, K. 2001. Taxonomic study of the genus Pheretima s. lat (Oligochaeta, Megascolecidae) from Japan. Bulletin of Seikei University 33 (3): 1 - 125." type="journal article" year="2001">Ishizuka, 2001: 75</bibRefCitation>
</treatmentCitation>
, fig. 32;
<treatmentCitation id="0AEC10FA3F5E36576C07FC72FD8CFBB8" author="Nakamura, Y." box="[380,600,1004,1027]" page="27" pageId="1" pageNumber="36" year="1999">
<bibRefCitation id="EFDC4B1A3F5E36576C07FC72FD8CFBB8" author="Nakamura, Y." box="[380,600,1004,1027]" pageId="1" pageNumber="36" pagination="1 - 78" refId="ref6631" refString="Nakamura, Y. 1999. Checklist of Earthworms of Pheretima Genus Group (Megascolecidae: Oligochaeta) of the World. Edaphologia 64: 1 - 78." type="journal article" year="1999">Nakamura, 1999: 27</bibRefCitation>
</treatmentCitation>
(name misspelt “canosa”).
</paragraph>
<paragraph id="8BF236EB3F5E36576DD9FBB3FE75FA9F" blockId="1.[162,768,490,1991]" pageId="1" pageNumber="36">
<treatmentCitation id="0AEC10FA3F5E36576DD9FBB3FD98FBF8" author="Kobayashi, S." box="[162,588,1068,1092]" page="1" pageId="1" pageNumber="36" year="1934">
<taxonomicName id="4C4D4D683F5E36576DD9FBB3FD98FBF8" authority="Kobayashi, 1934: 1" authorityName="Kobayashi" authorityPageNumber="1" authorityYear="1934" box="[162,588,1068,1092]" class="Clitellata" family="Megascolecidae" genus="Pheretima" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="kyamikia">
<emphasis id="B939EAF93F5E36576DD9FBB3FEA2FBFF" box="[162,374,1069,1092]" italics="true" pageId="1" pageNumber="36">Pheretima kyamikia</emphasis>
<bibRefCitation id="EFDC4B1A3F5E36576C05FBB2FD98FBF8" author="Kobayashi, S." box="[382,588,1068,1091]" pageId="1" pageNumber="36" pagination="1 - 11" refId="ref6366" refString="Kobayashi, S. 1934. Three new Korean earthworms belonging to the genus Pheretima, together with the wider range of the distribution of Pheretima hilgendorfi (Michaelsen). Journal of Chosen Natural History Society 19: 1 - 11." type="journal article" year="1934">Kobayashi, 1934: 1</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 1-3. [From
<collectingCountry id="F35A767B3F5E36576DC2FBD2FE94FBD8" box="[185,320,1100,1123]" name="North Korea" pageId="1" pageNumber="36">North Korea</collectingCountry>
.
<typeStatus id="54F688493F5E36576C31FBD2FE5EFBD8" box="[330,394,1100,1123]" pageId="1" pageNumber="36">Types</typeStatus>
unknown, but a non-type specimen from “(Tetsugen) Kogen-do” sent by S. Kobayashi was added in synonymy of
<taxonomicName id="4C4D4D683F5E36576CCCFB13FDDAFB1F" authorityName="Stephenson" authorityYear="1925" box="[439,526,1165,1188]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="pingi">
<emphasis id="B939EAF93F5E36576CCCFB13FDDAFB1F" box="[439,526,1165,1188]" italics="true" pageId="1" pageNumber="36">P. pingi</emphasis>
</taxonomicName>
by
<treatmentCitation id="0AEC10FA3F5E36576F43FB13FD28FB1F" author="Gates, G. E." box="[568,764,1165,1188]" page="465" pageId="1" pageNumber="36" year="1939">
<bibRefCitation id="EFDC4B1A3F5E36576F43FB13FD28FB1F" author="Gates, G. E." box="[568,764,1165,1188]" pageId="1" pageNumber="36" pagination="405 - 507" refId="ref5892" refString="Gates, G. E. 1939. On some species of Chinese earthworms with special reference to specimens collected in Szechuan by Dr. D. C. Graham. Proceedings of the United States National Museum 85: 405 - 507 [Available from: si-dppr. si. edu / dspace / bitstream / 10088 / 16290 / USNMP- 85 _ 3040 _ 1939. pdf]." type="journal article" year="1939">Gates (1939: 465)</bibRefCitation>
</treatmentCitation>
.
<bibRefCitation id="EFDC4B1A3F5E36576DC2FB33FE56FB7F" author="Kobayashi, S." box="[185,386,1197,1220]" pageId="1" pageNumber="36" pagination="1 - 11" refId="ref6366" refString="Kobayashi, S. 1934. Three new Korean earthworms belonging to the genus Pheretima, together with the wider range of the distribution of Pheretima hilgendorfi (Michaelsen). Journal of Chosen Natural History Society 19: 1 - 11." type="journal article" year="1934">Kobayashis (1934</bibRefCitation>
: fig. 1) is the same as
<bibRefCitation id="EFDC4B1A3F5E36576F17FB33FD28FB7F" author="Chen, Y." box="[620,764,1197,1220]" pageId="1" pageNumber="36" refId="ref5837" refString="Chen, Y. 1959. Chinese Animal Atlas. Annelida. Science Press, Beijing [In Chinese]." type="book" year="1959">Chens (1959</bibRefCitation>
: fig. 9i variation) for
<taxonomicName id="4C4D4D683F5E36576CDBFB53FE22FB5F" authority="(Goto &amp; Hatai, 1899)" box="[416,502,1229,1252]" class="Clitellata" family="Megascolecidae" genus="Pheretima" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="carnosa">
<emphasis id="B939EAF93F5E36576CDBFB53FE22FB5F" box="[416,502,1229,1252]" italics="true" pageId="1" pageNumber="36">carnosa</emphasis>
</taxonomicName>
(syn.
<taxonomicName id="4C4D4D683F5E36576F42FB53FDA4FB5F" authorityName="Stephenson" authorityYear="1925" box="[569,624,1229,1252]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="pingi">
<emphasis id="B939EAF93F5E36576F42FB53FDA4FB5F" box="[569,624,1229,1252]" italics="true" pageId="1" pageNumber="36">pingi</emphasis>
</taxonomicName>
). Named for Korean vernacular “kyamiki” meaning “
<emphasis id="B939EAF93F5E36576F0CFB73FD07FABF" box="[631,723,1261,1284]" italics="true" pageId="1" pageNumber="36">dog bait</emphasis>
” or “
<emphasis id="B939EAF93F5E36576DBEFA90FE51FA9E" box="[197,389,1294,1317]" italics="true" pageId="1" pageNumber="36">useless as fishbait</emphasis>
”].
</paragraph>
<paragraph id="8BF236EB3F5E36576DD9FAB0FF3DFA1E" blockId="1.[162,768,490,1991]" pageId="1" pageNumber="36">
<treatmentCitationGroup id="AB5D11C53F5E36576DD9FAB0FF3DFA1E" pageId="1" pageNumber="36">
<treatmentCitation id="0AEC10FA3F5E36576DD9FAB0FD75FAFF" author="Kobayashi, S." box="[162,673,1325,1349]" page="168" pageId="1" pageNumber="36" year="1936">
<taxonomicName id="4C4D4D683F5E36576DD9FAB0FD75FAFF" authority="Kobayashi, 1936 b: 168" authorityName="Kobayashi" authorityPageNumber="168" authorityYear="1936" box="[162,673,1325,1349]" class="Clitellata" family="Megascolecidae" genus="Pheretima" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="monstrifera">
<emphasis id="B939EAF93F5E36576DD9FAB0FE43FAFE" box="[162,407,1326,1349]" italics="true" pageId="1" pageNumber="36">Pheretima monstrifera</emphasis>
<bibRefCitation id="EFDC4B1A3F5E36576CDAFAB3FD75FAFF" author="Kobayashi, S." box="[417,673,1325,1348]" pageId="1" pageNumber="36" pagination="139 - 184" refId="ref6452" refString="Kobayashi, S. 1936 b. Earthworms from Koryo, Korea. Science Report of the Tohoku Imperial University 11 (1): 139 - 184." type="journal article" year="1936">Kobayashi, 1936b: 168</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 11. [From
<collectingRegion id="4989F8093F5E36576C79FAD0FEEAFADE" box="[258,318,1358,1381]" country="South Korea" name="Seoul" pageId="1" pageNumber="36">Seoul</collectingRegion>
and other localities. Agreeing with
<bibRefCitation id="EFDC4B1A3F5E36576FC6FAD0FE91FA3E" author="Kobayashi, S." pageId="1" pageNumber="36" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi (1936a</bibRefCitation>
: tab. 1, and text-figs. 1
<collectionCode id="ED5CAE2E3F5E36576F38FAF0FDB9FA3E" box="[579,621,1390,1413]" pageId="1" pageNumber="36">XII</collectionCode>
&amp; 2I)].
<emphasis id="B939EAF93F5E36576FB6FAF3FF3DFA1E" bold="true" pageId="1" pageNumber="36">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF236EB3F5E36576DD9FA30FD73F93D" blockId="1.[162,768,490,1991]" pageId="1" pageNumber="36">
<taxonomicName id="4C4D4D683F5E36576DD9FA30FEA4FA7E" authorityName="NSMT An" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" box="[162,368,1454,1477]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B939EAF93F5E36576DD9FA30FEA4FA7E" box="[162,368,1454,1477]" italics="true" pageId="1" pageNumber="36">Amynthas carnosus</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AEC10FA3F5E36576C04FA30FD42FA7E" author="Sims, R. W. &amp; E. G. Easton" box="[383,662,1454,1477]" page="235" pageId="1" pageNumber="36" year="1972">
<bibRefCitation id="EFDC4B1A3F5E36576C04FA30FD42FA7E" author="Sims, R. W. &amp; E. G. Easton" box="[383,662,1454,1477]" pageId="1" pageNumber="36" pagination="1 - 169" refId="ref6864" refString="Sims, R. W. and E. G. Easton. 1972. A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnaean Society, London 4: 1 - 169." type="journal article" year="1972">Sims &amp; Easton, 1972: 235</bibRefCitation>
</treatmentCitation>
[lapsus in
<taxonomicName id="4C4D4D683F5E36576DC2FA50FE99FA5E" baseAuthorityName="Rosa" baseAuthorityYear="1891" box="[185,333,1486,1509]" class="Clitellata" family="Megascolecidae" genus="Amyntas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="hawayanus">
<emphasis id="B939EAF93F5E36576DC2FA50FE99FA5E" box="[185,333,1486,1509]" italics="true" pageId="1" pageNumber="36">A. hawayanus</emphasis>
</taxonomicName>
(
<taxonomicName id="4C4D4D683F5E36576C22FA4EFE69FA5E" ID-CoL="84WCW" baseAuthorityName="Kinberg" baseAuthorityYear="1867" box="[345,445,1486,1509]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="gracilis">
=
<emphasis id="B939EAF93F5E36576C15FA50FE69FA5E" box="[366,445,1486,1509]" italics="true" pageId="1" pageNumber="36">gracilis</emphasis>
</taxonomicName>
) group];
<treatmentCitation id="0AEC10FA3F5E36576F59FA50FD2BFA5E" author="Blakemore, R. J." box="[546,767,1486,1509]" page="13" pageId="1" pageNumber="36" year="2003">
<bibRefCitation id="EFDC4B1A3F5E36576F59FA50FD2BFA5E" author="Blakemore, R. J." box="[546,767,1486,1509]" pageId="1" pageNumber="36" refId="ref4978" refString="Blakemore, R. J. 2003. Japanese Earthworms (Annelida: Oligochaeta): a review and checklist of species. Organisms, Diversity and Evolution 3 (3): 241 - 244 [Available from: 2003 - 11 http: // www. senckenberg. de / odes / 03 - 11. htm]." type="url" year="2003">Blakemore, 2003: 13</bibRefCitation>
</treatmentCitation>
+43 addendum (syns.?
<emphasis id="B939EAF93F5E36576CCEFA70FD5DF9BE" box="[437,649,1518,1541]" italics="true" pageId="1" pageNumber="36">
distichus,
<taxonomicName id="4C4D4D683F5E36576F59FA70FD51F9BE" authorityName="Kobayashi" authorityYear="1934" box="[546,645,1518,1541]" class="Clitellata" family="Megascolecidae" genus="Pheretima" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="kyamikia">kyamikia</taxonomicName>
,
</emphasis>
<taxonomicName id="4C4D4D683F5E36576FF4FA70FD2FF9BE" authorityName="Y.Hong &amp; James" authorityYear="2001" box="[655,763,1518,1541]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="youngtai">
?
<emphasis id="B939EAF93F5E36576FE0FA70FD2FF9BE" box="[667,763,1518,1541]" italics="true" pageId="1" pageNumber="36">youngtai</emphasis>
</taxonomicName>
<emphasis id="B939EAF93F5E36576F80FA70FECBF99D" italics="true" pageId="1" pageNumber="36">
,
<taxonomicName id="4C4D4D683F5E36576DC2F991FECBF99D" authorityName="Y.Hong &amp; James" authorityYear="2001" box="[185,287,1551,1574]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="sangyeoli">sangyeoli</taxonomicName>
</emphasis>
); 2004; 2007; 2008 (syns.
<taxonomicName id="4C4D4D683F5E36576F3DF990FD5FF99D" authorityName="Stephenson" authorityYear="1925" box="[582,651,1550,1574]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="pingi">
?
<emphasis id="B939EAF93F5E36576F29F991FD5FF99D" box="[594,651,1551,1574]" italics="true" pageId="1" pageNumber="36">pingi</emphasis>
</taxonomicName>
<emphasis id="B939EAF93F5E36576FF0F991FD2BF99D" box="[651,767,1551,1574]" italics="true" pageId="1" pageNumber="36">
,
<taxonomicName id="4C4D4D683F5E36576FE3F991FD2FF99D" authorityName="Kobayashi" authorityYear="1934" box="[664,763,1551,1574]" class="Clitellata" family="Megascolecidae" genus="Pheretima" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="kyamikia">kyamikia</taxonomicName>
,
</emphasis>
<taxonomicName id="4C4D4D683F5E36576DC2F9B0FEF0F9FD" authorityName="Y.Hong &amp; James" authorityYear="2001" box="[185,292,1582,1606]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="youngtai">
?
<emphasis id="B939EAF93F5E36576DBEF9B1FEF0F9FD" box="[197,292,1583,1606]" italics="true" pageId="1" pageNumber="36">youngtai</emphasis>
</taxonomicName>
<emphasis id="B939EAF93F5E36576C5FF9B1FE52F9FD" box="[292,390,1583,1606]" italics="true" pageId="1" pageNumber="36">, sanyeoli</emphasis>
);
<treatmentCitation id="0AEC10FA3F5E36576CE6F9B0FD52F9FE" author="Shen, H. - P. &amp; C. - F. Tsai &amp; S. - C. Tsai" box="[413,646,1582,1606]" page="484" pageId="1" pageNumber="36" year="2003">
<bibRefCitation id="EFDC4B1A3F5E36576CE6F9B0FD52F9FE" author="Shen, H. - P. &amp; C. - F. Tsai &amp; S. - C. Tsai" box="[413,646,1582,1606]" pageId="1" pageNumber="36" pagination="479 - 490" refId="ref6702" refString="Shen, H. - P., C. - F. Tsai and S. - C. Tsai. 2003. Six new earthworms of the genus Amynthas (Oligochaeta: Megascolecidae) from Central Taiwan. Zoolological Studies 42 (4): 479 - 490 [Available from: https: // zoolstud. sinica. edu. tw / Journals / 42.4 / 279. pdf]." type="journal article" year="2003">
Shen
<emphasis id="B939EAF93F5E36576CA2F9B1FDDAF9FD" box="[473,526,1583,1606]" italics="true" pageId="1" pageNumber="36">et al.</emphasis>
, 2003: 484
</bibRefCitation>
</treatmentCitation>
(syn.
<taxonomicName id="4C4D4D683F5E36576FBEF9B1FF3AF9DD" authorityName="Y.Hong &amp; James" authorityYear="2001" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="sangyeoli">
<emphasis id="B939EAF93F5E36576FBEF9B1FF3AF9DD" italics="true" pageId="1" pageNumber="36">sangyeoli</emphasis>
</taxonomicName>
); [?non
<treatmentCitation id="0AEC10FA3F5E36576C2BF9D1FD95F9DD" author="Shen, H. - P. &amp; S. - C. Tsai &amp; C. - F. Tsai &amp; J. - H. Chen" box="[336,577,1615,1638]" page="95" pageId="1" pageNumber="36" year="2005">
<bibRefCitation id="EFDC4B1A3F5E36576C2BF9D1FD95F9DD" author="Shen, H. - P. &amp; S. - C. Tsai &amp; C. - F. Tsai &amp; J. - H. Chen" box="[336,577,1615,1638]" pageId="1" pageNumber="36" pagination="95 - 100" refId="ref6776" refString="Shen, H. - P., S. - C. Tsai, C. - F. Tsai and J. - H. Chen. 2005. Occurrence of the Earthworm Amynthas carnosus (Goto and Hatai, 1899) in the Northern Taiwan. Endemic Species Research 7 (1): 95 - 100 [Available from: https: // tesri. coa. gov. tw / files / tesri _ protect / 124 / 070108. pdf]." type="journal article" year="2005">
Shen
<emphasis id="B939EAF93F5E36576CEBF9D1FE1EF9DD" box="[400,458,1615,1638]" italics="true" pageId="1" pageNumber="36">et al.</emphasis>
, 2005: 95
</bibRefCitation>
</treatmentCitation>
; nec.
<treatmentCitation id="0AEC10FA3F5E36576FFDF9D1FECEF93D" author="Tsai, C. - F. &amp; H. - P. Shen &amp; S. - C. Tsai &amp; K. - J. Lin &amp; H. - L. Hsieh &amp; S. - P. Yo" page="38" pageId="1" pageNumber="36" year="2009">
<bibRefCitation id="EFDC4B1A3F5E36576FFDF9D1FECEF93D" author="Tsai, C. - F. &amp; H. - P. Shen &amp; S. - C. Tsai &amp; K. - J. Lin &amp; H. - L. Hsieh &amp; S. - P. Yo" pageId="1" pageNumber="36" pagination="33 - 48" refId="ref7004" refString="Tsai, C. - F., H. - P. Shen, S. - C. Tsai, K. - J. Lin, H. - L. Hsieh and S. - P. Yo. 2009. A checklist of oligochaetes (Annelida) from Taiwan and adjacent islands. Zootaxa 2133: 33 - 48." type="journal article" year="2009">
Tsai
<emphasis id="B939EAF93F5E36576FC4F9D1FD2DF9DD" box="[703,761,1615,1638]" italics="true" pageId="1" pageNumber="36">et al.</emphasis>
, 2009: 38
</bibRefCitation>
</treatmentCitation>
; nec.
<treatmentCitation id="0AEC10FA3F5E36576C2CF9F1FD93F93D" author="Chang, C. - H. &amp; H. - P. Shen &amp; J. - H. Chen" box="[343,583,1647,1670]" page="32" pageId="1" pageNumber="36" year="2009">
<bibRefCitation id="EFDC4B1A3F5E36576C2CF9F1FD93F93D" author="Chang, C. - H. &amp; H. - P. Shen &amp; J. - H. Chen" box="[343,583,1647,1670]" pageId="1" pageNumber="36" refId="ref5762" refString="Chang, C. - H., H. - P. Shen and J. - H. Chen. 2009. Earthworm fauna of Taiwan. National University Press, Taipei, 167 p." type="book" year="2009">
Chang
<emphasis id="B939EAF93F5E36576CDFF9F1FE0EF93D" box="[420,474,1647,1670]" italics="true" pageId="1" pageNumber="36">et al.</emphasis>
, 2009: 32
</bibRefCitation>
</treatmentCitation>
, fig. 12].
</paragraph>
<paragraph id="8BF236EB3F5E36576DD9F911FD48F91D" blockId="1.[162,768,490,1991]" box="[162,668,1679,1702]" pageId="1" pageNumber="36">
<treatmentCitationGroup id="AB5D11C53F5E36576DD9F911FD48F91D" box="[162,668,1679,1702]" pageId="1" pageNumber="36">
<taxonomicName id="4C4D4D683F5E36576DD9F911FE58F91D" baseAuthorityName="Baird" baseAuthorityYear="1869" box="[162,396,1679,1702]" class="Clitellata" family="Megascolecidae" genus="Pheretima" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="diffringens">
<emphasis id="B939EAF93F5E36576DD9F911FE58F91D" box="[162,396,1679,1702]" italics="true" pageId="1" pageNumber="36">Pheretima diffringens</emphasis>
</taxonomicName>
(part.):
<treatmentCitation id="0AEC10FA3F5E36576CA4F911FD4CF91D" author="Gates, G. E." box="[479,664,1679,1702]" page="149" pageId="1" pageNumber="36" year="1972">
<bibRefCitation id="EFDC4B1A3F5E36576CA4F911FD4CF91D" author="Gates, G. E." box="[479,664,1679,1702]" pageId="1" pageNumber="36" pagination="1 - 326" refId="ref5988" refString="Gates, G. E. 1972. Burmese Earthworms, an introduction to the systematics and biology of Megadrile oligochaetes with special reference to South-East Asia. Transactions of the American Philosophical Society 62 (7): 1 - 326 [Available from: http: // www. jstor. org / stable / 1006214 in 2011]." type="journal article" year="1972">Gates, 1972: 149</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF236EB3F5E36576DD9F931FDA6F97D" blockId="1.[162,768,490,1991]" box="[162,626,1711,1734]" pageId="1" pageNumber="36">
<treatmentCitationGroup id="AB5D11C53F5E36576DD9F931FDA6F97D" box="[162,626,1711,1734]" pageId="1" pageNumber="36">
<taxonomicName id="4C4D4D683F5E36576DD9F931FEB7F97D" ID-CoL="84WCW" baseAuthorityName="Kinberg" baseAuthorityYear="1867" box="[162,355,1711,1734]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="gracilis">
<emphasis id="B939EAF93F5E36576DD9F931FEB7F97D" box="[162,355,1711,1734]" italics="true" pageId="1" pageNumber="36">Amynthas gracilis</emphasis>
</taxonomicName>
(part.):
<treatmentCitation id="0AEC10FA3F5E36576CCCF931FDBBF97D" author="Easton, E. G." box="[439,623,1711,1734]" page="50" pageId="1" pageNumber="36" year="1981">
<bibRefCitation id="EFDC4B1A3F5E36576CCCF931FDBBF97D" author="Easton, E. G." box="[439,623,1711,1734]" pageId="1" pageNumber="36" pagination="33 - 65" refId="ref5858" refString="Easton, E. G. 1981. Japanese earthworms: a synopsis of the Megadrile species. Bulletin of the British Museum (Natural History) Zoology 40: 33 - 65." type="journal article" year="1981">Easton, 1981: 50</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF236EB3F5E36576DD9F951FA57FEFB" blockId="1.[162,768,490,1991]" lastBlockId="1.[828,1433,265,1349]" pageId="1" pageNumber="36">
<taxonomicName id="4C4D4D683F5E36576DD9F951FEBBF95D" authorityName="Y.Hong &amp; James" authorityYear="2001" box="[162,367,1743,1766]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="youngtai">
<emphasis id="B939EAF93F5E36576DD9F951FEBBF95D" box="[162,367,1743,1766]" italics="true" pageId="1" pageNumber="36">Amynthas youngtai</emphasis>
</taxonomicName>
<bibRefCitation id="EFDC4B1A3F5E36576C0CF951FD89F95D" author="Hong, Y. &amp; S. W. James" box="[375,605,1743,1766]" pageId="1" pageNumber="36" pagination="269 - 275" refId="ref6085" refString="Hong, Y. and S. W. James. 2001. Five new earthworms of the genus Amynthas Kinberg, 1867 (Megascolecidae) with four pairs of spermathecae. Zoolological Studies 40 (4): 269 - 275." type="journal article" year="2001">Hong &amp; James, 2001</bibRefCitation>
b: 269, fig. 1A-C. [From
<collectingRegion id="4989F8093F5E36576C57F971FE8DF8BD" box="[300,345,1775,1798]" country="South Korea" name="Jeju" pageId="1" pageNumber="36">Jeju</collectingRegion>
(=Quelpart),
<collectingCountry id="F35A767B3F5E36576C87F971FD94F8BD" box="[508,576,1775,1798]" name="South Korea" pageId="1" pageNumber="36">Korea</collectingCountry>
- the same place whence
<bibRefCitation id="EFDC4B1A3F5E36576C6BF891FE2BF89D" author="Kobayashi, S." box="[272,511,1807,1830]" pageId="1" pageNumber="36" pagination="89 - 170" refId="ref6515" refString="Kobayashi, S. 1938. Earthworms of Korea I. Science Report of the Tohoku Imperial University 13 (2): 89 - 170." type="journal article" year="1938">Kobayashi (1938: 161)</bibRefCitation>
had already recorded
<taxonomicName id="4C4D4D683F5E36576F92F88EFECEF8FC" authorityName="NSMT An" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B939EAF93F5E36576F92F88EFECEF8FC" italics="true" pageId="1" pageNumber="36">A. carnosus</emphasis>
</taxonomicName>
- apparently from some other locations too.
<typeStatus id="54F688493F5E36576DC2F8CEFF2EF8DC" box="[185,250,1872,1895]" pageId="1" pageNumber="36">Types</typeStatus>
put in Korean Institute of Biodiversity Research (
<collectionCode id="ED5CAE2E3F5E36576DBAF8EEFEC4F83C" box="[193,272,1904,1927]" pageId="1" pageNumber="36">KIBIO</collectionCode>
),
<collectingRegion id="4989F8093F5E36576C58F8EEFEA8F83C" box="[291,380,1904,1927]" country="South Korea" name="Jeonbuk" pageId="1" pageNumber="36">Jeonbuk</collectingRegion>
National University, but details are unclear. Uncertainty due to segmental miscounts between authors fig. 1 and somewhat confused description but, nevertheless, it mostly agrees, at least with
<bibRefCitation id="EFDC4B1A3F5E36576E29FEB7FBC2FEFB" author="Kobayashi, S." box="[850,1046,297,320]" pageId="1" pageNumber="36" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi (1936a</bibRefCitation>
: tab. 1 and text-figs. 1
<collectionCode id="ED5CAE2E3F5E36576985FEB7FAFCFEFB" box="[1278,1320,297,320]" pageId="1" pageNumber="36">VII</collectionCode>
&amp; 2X)].
</paragraph>
<paragraph id="8BF236EB3F5E36576E47FED4FB27FE59" blockId="1.[828,1433,265,1349]" pageId="1" pageNumber="36">
<taxonomicName id="4C4D4D683F5E36576E47FED4FBE2FEDA" authorityName="Y.Hong &amp; James" authorityYear="2001" box="[828,1078,330,353]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="kimhaeiensis">
<emphasis id="B939EAF93F5E36576E47FED4FBE2FEDA" box="[828,1078,330,353]" italics="true" pageId="1" pageNumber="36">Amynthas kimhaeiensis</emphasis>
</taxonomicName>
<bibRefCitation id="EFDC4B1A3F5E36576945FED7FAF1FEDB" author="Hong, Y. &amp; S. W. James" box="[1086,1317,329,352]" pageId="1" pageNumber="36" pagination="269 - 275" refId="ref6085" refString="Hong, Y. and S. W. James. 2001. Five new earthworms of the genus Amynthas Kinberg, 1867 (Megascolecidae) with four pairs of spermathecae. Zoolological Studies 40 (4): 269 - 275." type="journal article" year="2001">Hong &amp; James, 2001</bibRefCitation>
b: 270, fig. 2A-C. [From
<collectingCountry id="F35A767B3F5E36576E90FEF4FBE6FE3A" box="[1003,1074,362,385]" name="South Korea" pageId="1" pageNumber="36">Korea</collectingCountry>
.
<typeStatus id="54F688493F5E36576944FEF4FBA3FE3A" box="[1087,1143,362,385]" pageId="1" pageNumber="36">Type</typeStatus>
in
<collectingCountry id="F35A767B3F5E365769D9FEF4FB3DFE3A" box="[1186,1257,362,385]" name="South Korea" pageId="1" pageNumber="36">Korea</collectingCountry>
. Description of spermathecal pores and figures do not quite correspond, yet apparently agreeing at least with
<bibRefCitation id="EFDC4B1A3F5E365769AEFE34FA41FE7A" author="Kobayashi, S." box="[1237,1429,426,449]" pageId="1" pageNumber="36" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi (1936a</bibRefCitation>
: text-figs. 1XI or
<collectionCode id="ED5CAE2E3F5E3657697EFE54FBFEFE5A" box="[1029,1066,458,481]" pageId="1" pageNumber="36">XII</collectionCode>
&amp; 2II)].
<emphasis id="B939EAF93F5E365769F0FE54FB27FE59" bold="true" box="[1163,1267,458,482]" pageId="1" pageNumber="36">Syn. nov.</emphasis>
</paragraph>
<paragraph id="8BF236EB3F5E36576E47FE74FC23FD59" blockId="1.[828,1433,265,1349]" pageId="1" pageNumber="36">
<taxonomicName id="4C4D4D683F5E36576E47FE74FBDBFDBA" authorityName="Y.Hong &amp; James" authorityYear="2001" box="[828,1039,490,513]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="sangyeoli">
<emphasis id="B939EAF93F5E36576E47FE74FBDBFDBA" box="[828,1039,490,513]" italics="true" pageId="1" pageNumber="36">Amynthas sangyeoli</emphasis>
</taxonomicName>
<bibRefCitation id="EFDC4B1A3F5E3657696DFE74FB2CFDBA" author="Hong, Y. &amp; S. W. James" box="[1046,1272,490,513]" pageId="1" pageNumber="36" pagination="269 - 275" refId="ref6085" refString="Hong, Y. and S. W. James. 2001. Five new earthworms of the genus Amynthas Kinberg, 1867 (Megascolecidae) with four pairs of spermathecae. Zoolological Studies 40 (4): 269 - 275." type="journal article" year="2001">Hong &amp; James, 2001</bibRefCitation>
b: 271, fig. 3A-C. [Korean
<typeStatus id="54F688493F5E36576EB4FD94FBDDFD9A" box="[975,1033,522,545]" pageId="1" pageNumber="36">types</typeStatus>
, materials and type-material locations seem rather confused. Markings near male and spermathecal pores, and the spermatheca exactly the same as Goto &amp; Hatais corrected figures from 102 years earlier, as well as those of
<bibRefCitation id="EFDC4B1A3F5E365769FAFD15FAEDFD19" author="Kobayashi, S." box="[1153,1337,651,674]" pageId="1" pageNumber="36" pagination="1 - 11" refId="ref6366" refString="Kobayashi, S. 1934. Three new Korean earthworms belonging to the genus Pheretima, together with the wider range of the distribution of Pheretima hilgendorfi (Michaelsen). Journal of Chosen Natural History Society 19: 1 - 11." type="journal article" year="1934">Kobayashi (1934</bibRefCitation>
;
<bibRefCitation id="EFDC4B1A3F5E36576833FD15FA58FD19" author="Kobayashi, S." box="[1352,1420,651,674]" pageId="1" pageNumber="36" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">1936a</bibRefCitation>
),
<bibRefCitation id="EFDC4B1A3F5E36576E29FD35FC2EFD79" author="Ohfuchi, S." box="[850,1018,683,706]" pageId="1" pageNumber="36" pagination="31 - 136" refId="ref6660" refString="Ohfuchi, S. 1937. On the species possessing four pairs of spermathecae in the genus Pheretima, together with the variability of some external and internal characteristics. Saito Ho-On Kai Museum Research Bulletin 12: 31 - 136." type="journal article" year="1937">Ohfuchi (1937)</bibRefCitation>
,
<bibRefCitation id="EFDC4B1A3F5E36576973FD35FB5DFD79" author="Chen, Y." box="[1032,1161,683,706]" pageId="1" pageNumber="36" pagination="269 - 306" refId="ref5800" refString="Chen, Y. 1936. On the terrestrial Oligochaeta from Szechuan II with the notes on Gates' types. Contribibutions of Biological Lababoratory of Science Society of China. Zoology 11: 269 - 306." type="journal article" year="1936">Chen (1936</bibRefCitation>
;
<bibRefCitation id="EFDC4B1A3F5E365769ECFD35FB04FD79" author="Chen, Y." box="[1175,1232,683,706]" pageId="1" pageNumber="36" refId="ref5837" refString="Chen, Y. 1959. Chinese Animal Atlas. Annelida. Science Press, Beijing [In Chinese]." type="book" year="1959">1959</bibRefCitation>
), etc. dating from ~69 years ago].
</paragraph>
<paragraph id="8BF236EB3F5E36576E47FD75FBD8FCD8" blockId="1.[828,1433,265,1349]" pageId="1" pageNumber="36">
<taxonomicName id="4C4D4D683F5E36576E47FD75FBDBFCB9" authorityName="Y.Hong &amp; James" authorityYear="2001" box="[828,1039,747,770]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="sinsiensis">
<emphasis id="B939EAF93F5E36576E47FD75FBDBFCB9" box="[828,1039,747,770]" italics="true" pageId="1" pageNumber="36">Amynthas sinsiensis</emphasis>
</taxonomicName>
<bibRefCitation id="EFDC4B1A3F5E3657696DFD75FB2CFCB9" author="Hong, Y. &amp; S. W. James" box="[1046,1272,747,770]" pageId="1" pageNumber="36" pagination="269 - 275" refId="ref6085" refString="Hong, Y. and S. W. James. 2001. Five new earthworms of the genus Amynthas Kinberg, 1867 (Megascolecidae) with four pairs of spermathecae. Zoolological Studies 40 (4): 269 - 275." type="journal article" year="2001">Hong &amp; James, 2001</bibRefCitation>
b: 272, fig. 4A-C. [From
<collectingCountry id="F35A767B3F5E36576EC1FC95FC2BFC99" box="[954,1023,779,802]" name="South Korea" pageId="1" pageNumber="36">Korea</collectingCountry>
.
<typeStatus id="54F688493F5E36576972FC95FB98FC99" box="[1033,1100,779,802]" pageId="1" pageNumber="36">Types</typeStatus>
? Agreeing both with
<taxonomicName id="4C4D4D683F5E36576842FC92FC75FCF8" baseAuthorityName="Kobayashi" baseAuthorityYear="1936" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="monstriferus">
<emphasis id="B939EAF93F5E36576842FC92FC75FCF8" italics="true" pageId="1" pageNumber="36">A. monstriferus</emphasis>
</taxonomicName>
and
<bibRefCitation id="EFDC4B1A3F5E36576EADFCB5FB4EFCF9" author="Kobayashi, S." box="[982,1178,811,834]" pageId="1" pageNumber="36" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi (1936a</bibRefCitation>
: text-figs. 1I or II or
<collectionCode id="ED5CAE2E3F5E36576807FCB5FA4CFCF9" box="[1404,1432,811,834]" pageId="1" pageNumber="36">IV</collectionCode>
&amp; 2I)].
<emphasis id="B939EAF93F5E36576ED8FCD5FBD8FCD8" bold="true" box="[931,1036,843,867]" pageId="1" pageNumber="36">Syn. nov.</emphasis>
</paragraph>
<paragraph id="8BF236EB3F5E36576E47FCF2FC6FFC5F" blockId="1.[828,1433,265,1349]" pageId="1" pageNumber="36">
<taxonomicName id="4C4D4D683F5E36576E47FCF2FB83FC38" authorityName="Y.Hong &amp; James" authorityYear="2001" box="[828,1111,876,899]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="baemsagolensis">
<emphasis id="B939EAF93F5E36576E47FCF2FB83FC38" box="[828,1111,876,899]" italics="true" pageId="1" pageNumber="36">Amynthas baemsagolensis</emphasis>
</taxonomicName>
<bibRefCitation id="EFDC4B1A3F5E3657691AFCF2FA9BFC38" author="Hong, Y. &amp; S. W. James" box="[1121,1359,876,899]" pageId="1" pageNumber="36" pagination="269 - 275" refId="ref6085" refString="Hong, Y. and S. W. James. 2001. Five new earthworms of the genus Amynthas Kinberg, 1867 (Megascolecidae) with four pairs of spermathecae. Zoolological Studies 40 (4): 269 - 275." type="journal article" year="2001">Hong &amp; James, 2001</bibRefCitation>
b: 274, fig. 5A-C. [From
<collectingCountry id="F35A767B3F5E36576976FC12FB85FC18" box="[1037,1105,908,931]" name="South Korea" pageId="1" pageNumber="36">Korea</collectingCountry>
.
<typeStatus id="54F688493F5E36576920FC12FB4AFC18" box="[1115,1182,908,931]" pageId="1" pageNumber="36">Types</typeStatus>
? Agreeing both with
<taxonomicName id="4C4D4D683F5E365768FAFC12FC6CFC78" authorityName="Y.Hong &amp; James" authorityYear="2001" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="sinsiensis">
<emphasis id="B939EAF93F5E365768FAFC12FC6CFC78" italics="true" pageId="1" pageNumber="36">A. sinsiensis</emphasis>
</taxonomicName>
and
<bibRefCitation id="EFDC4B1A3F5E36576E8AFC32FB63FC78" author="Kobayashi, S." box="[1009,1207,940,963]" pageId="1" pageNumber="36" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi (1936a</bibRefCitation>
: text-figs. 1I &amp; 2I)].
<emphasis id="B939EAF93F5E36576E29FC52FC6FFC5F" bold="true" box="[850,955,972,996]" pageId="1" pageNumber="36">Syn. nov.</emphasis>
</paragraph>
<paragraph id="8BF236EB3F5E36576E47FC72FBB4FAFE" blockId="1.[828,1433,265,1349]" pageId="1" pageNumber="36">
?
<treatmentCitation id="0AEC10FA3F5E36576E3CFC72FA9DFBB8" author="James, S. W. &amp; H. - T. Shih &amp; H. - W. Chang" box="[839,1353,1004,1027]" httpUri="http://treatment.plazi.org/id/03E61D4ED63BFF8D57C4CBCFF957FEFB" page="1012" pageId="1" pageNumber="36" year="2005">
<taxonomicName id="4C4D4D683F5E36576E3CFC72FA9DFBB8" authority="James et al. 2005: 1012" authorityName="James et al. 2005:" authorityPageNumber="1012" authorityYear="2005" box="[839,1353,1004,1027]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="monsoonus">
<emphasis id="B939EAF93F5E36576E3CFC72FBE6FBB8" box="[839,1074,1004,1027]" italics="true" pageId="1" pageNumber="36">Amynthas monsoonus</emphasis>
<bibRefCitation id="EFDC4B1A3F5E36576947FC72FA9DFBB8" author="James, S. W. &amp; H. - T. Shih &amp; H. - W. Chang" box="[1084,1353,1004,1027]" pageId="1" pageNumber="36" pagination="1007 - 1028" refId="ref6258" refString="James, S. W., H. - T. Shih and H. - W. Chang. 2005. Seven new species of Amynthas (Clitellata: Megascolecidae) and new earthworm records from Taiwan. Journal of Natural History 39 (14): 1007 - 1028." type="journal article" year="2005">
James
<emphasis id="B939EAF93F5E365769FCFC72FB14FBB8" box="[1159,1216,1004,1027]" italics="true" pageId="1" pageNumber="36">et al.</emphasis>
2005: 1012
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. [From
<collectingCountry id="F35A767B3F5E36576E29FB92FC71FB98" box="[850,933,1036,1059]" name="Taiwan" pageId="1" pageNumber="36">Taiwan</collectingCountry>
. Cf. regarded as a junior synonym of
<taxonomicName id="4C4D4D683F5E36576839FB93FB89FBF8" authority="Tsai et al., 1999" authorityName="Tsai" authorityYear="1999" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="tungpuensis">
<emphasis id="B939EAF93F5E36576839FB93FC76FBFF" italics="true" pageId="1" pageNumber="36">A. tungpuensis</emphasis>
Tsai
<emphasis id="B939EAF93F5E36576E9AFBB3FBCDFBFF" box="[993,1049,1069,1092]" italics="true" pageId="1" pageNumber="36">et al.</emphasis>
, 1999
</taxonomicName>
by
<bibRefCitation id="EFDC4B1A3F5E365769F2FBB2FA91FBF8" author="Tsai, C. - F. &amp; H. - P. Shen &amp; S. - C. Tsai &amp; K. - J. Lin &amp; H. - L. Hsieh &amp; S. - P. Yo" box="[1161,1349,1068,1092]" pageId="1" pageNumber="36" pagination="33 - 48" refId="ref7004" refString="Tsai, C. - F., H. - P. Shen, S. - C. Tsai, K. - J. Lin, H. - L. Hsieh and S. - P. Yo. 2009. A checklist of oligochaetes (Annelida) from Taiwan and adjacent islands. Zootaxa 2133: 33 - 48." type="journal article" year="2009">
Tsai
<emphasis id="B939EAF93F5E365769BAFBB3FB2DFBFF" box="[1217,1273,1069,1092]" italics="true" pageId="1" pageNumber="36">et al.</emphasis>
(2009)
</bibRefCitation>
. Probable instant synonymy of all nine “new” species was already suggested to the authors in manuscript review, prior to publication in a referees report sent to journal editor
<date id="FFF3102B3F5E36576EE2FB33FC26FB7F" box="[921,1010,1197,1220]" pageId="1" pageNumber="36" value="2004-01-08">8.I.2004</date>
(as online in
<bibRefCitation id="EFDC4B1A3F5E365769FEFB33FA9FFB7F" author="Blakemore, R. J." box="[1157,1355,1197,1220]" pageId="1" pageNumber="36" refId="ref5323" refString="Blakemore, R. J. 2010 b. Unravelling some Kinki worms supplement: [Available from: http: // opuscula. elte. hu / Tomus 41 _ 2 / Blakemore _ App 2. pdf]." type="url" year="2010">Blakemore, 2010b</bibRefCitation>
and as attached in Appendix), but this advice was manifestly ignored. Their named specimen agrees somewhat with
<bibRefCitation id="EFDC4B1A3F5E36576E29FA93FBF1FA9F" author="Kobayashi, S." box="[850,1061,1293,1316]" pageId="1" pageNumber="36" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashis (1936a</bibRefCitation>
: text-figs. 1
<collectionCode id="ED5CAE2E3F5E365769E7FA93FB0CFA9F" box="[1180,1240,1293,1316]" pageId="1" pageNumber="36">XVII</collectionCode>
&amp; 2I) and
<bibRefCitation id="EFDC4B1A3F5E36576835FA93FC40FAFE" author="Chen, Y." pageId="1" pageNumber="36" refId="ref5837" refString="Chen, Y. 1959. Chinese Animal Atlas. Annelida. Science Press, Beijing [In Chinese]." type="book" year="1959">Chens (1959</bibRefCitation>
: fig. 9 variations)].
</paragraph>
<paragraph id="8BF236EB3F5E36576E47FAF3FBFBF91D" blockId="1.[828,1433,1389,1991]" pageId="1" pageNumber="36">
<emphasis id="B939EAF93F5E36576E47FAF3FCA3FA3E" bold="true" box="[828,887,1389,1413]" pageId="1" pageNumber="36">Note.</emphasis>
<bibRefCitation id="EFDC4B1A3F5E36576E06FAF0FBA9FA3E" author="Goto, S. &amp; S. Hatai" box="[893,1149,1390,1413]" pageId="1" pageNumber="36" pagination="13 - 24" refId="ref6050" refString="Goto, S. and S. Hatai. 1899. New or imperfectly known species of earthworms. No. 2. Annotations Zoologicae Japonensis 3 (1): 13 - 24." type="journal article" year="1899">Goto &amp; Hatai (1899: 15)</bibRefCitation>
said: “
<emphasis id="B939EAF93F5E365769BCFAF0FB99F9BD" italics="true" pageId="1" pageNumber="36">
<specimenCount id="9D4BFD623F5E365769BCFAF0FAB2FA3E" box="[1223,1382,1390,1413]" count="2" pageId="1" pageNumber="36" type="generic">Two specimens</specimenCount>
presenting a difference of some importance in the genital papillae around the male pores. We shall base our description on the larger specimen, which is also provided with more genital papillae
</emphasis>
.” They then proceeded to poorly describe both specimens that they said were otherwise identical and thereby commenced introduction of 100+ years of confusion and uncertainty. These two original
<typeStatus id="54F688493F5E36576E47F9F1FC4CF93D" box="[828,920,1647,1670]" pageId="1" pageNumber="36" type="syntype">syntypes</typeStatus>
are not traceable in any known Japanese collection (as already noted).
</paragraph>
</subSubSection>
<subSubSection id="C35765603F5E36556E47F931FEC1FE9B" lastPageId="3" lastPageNumber="38" pageId="1" pageNumber="36" type="materials_examined">
<paragraph id="8BF236EB3F5E36556E47F931FEC1FE9B" blockId="1.[828,1433,1389,1991]" lastBlockId="3.[162,770,265,1991]" lastPageId="3" lastPageNumber="38" pageId="1" pageNumber="36">
<emphasis id="B939EAF93F5E36576E47F931FBCCF97C" bold="true" box="[828,1048,1711,1735]" pageId="1" pageNumber="36">Material inspected.</emphasis>
<materialsCitation id="3B253CB63F5E3655695BF931FEC1FE9B" collectionCode="NMST" lastPageId="3" lastPageNumber="38" location="Tokyo" pageId="1" pageNumber="36" specimenCount="2" stateProvince="Tokyo" typeStatus="neotype">
<collectingRegion id="4989F8093F5E3657695BF931FBBCF97D" box="[1056,1128,1711,1734]" country="Japan" name="Tokyo" pageId="1" pageNumber="36">Tokyo</collectingRegion>
,
<collectionCode id="ED5CAE2E3F5E3657690FF931FB6AF97D" box="[1140,1214,1711,1734]" pageId="1" pageNumber="36">NMST</collectionCode>
An435, formol preserved
<typeStatus id="54F688493F5E36576EF1F951FC30F95D" box="[906,996,1743,1766]" pageId="1" pageNumber="36" type="neotype">Neotype</typeStatus>
(
<figureCitation id="13762A6E3F5E36576E89F951FBE6F95D" box="[1010,1074,1743,1766]" captionStart="Fig" captionStartId="2.[160,193,1913,1933]" captionTargetBox="[429,1135,270,1852]" captionTargetId="figure-23@2.[419,1191,250,1939]" captionTargetPageId="2" captionText="Fig. 1. Amynthas carnosus (Goto &amp; Hatai, 1899) NSMT An435 Neotype showing prostomium, ventral view of body with spermathecae and prostates in situ (GM=Genital Marking with internal glands); also septal glands on dorsal blood vessel and intestinal caecum (Ihs=left side)." figureDoi="http://doi.org/10.5281/zenodo.12753218" httpUri="https://zenodo.org/record/12753218/files/figure.png" pageId="1" pageNumber="36">Fig. 1</figureCitation>
) labeled “
<taxonomicName id="4C4D4D683F5E365769D3F94EFCAAF8BD" authority="(Goto and Hatai)" baseAuthorityName="Goto and Hatai" class="Clitellata" family="Megascolecidae" genus="Pheretima" kingdom="Animalia" order="Crassiclitellata" pageId="1" pageNumber="36" phylum="Annelida" rank="species" species="carnosa">
<emphasis id="B939EAF93F5E365769D3F94EFAF1F95C" box="[1192,1317,1744,1767]" italics="true" pageId="1" pageNumber="36">Ph carnosa</emphasis>
(Goto and Hatai)
</taxonomicName>
[Kanji for Sendai-city] 1923-1925” part of Saito Ho-on Kai Museum Collection, possibly inspected and labeled by Dr Hatai who was original author, or one of his students e.g. Shinryo Ohfuchi. The date precludes it being a syntype. One mature specimen with tip of tail missing, previously undissected and here figured and dissected with small tissue samples taken to attempt DNA
<quantity id="4CB59B0E3F5D36546E71F8DCFCEFF8ED" box="[778,827,1858,1878]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="37" unit="mm" value="1.0">1 mm</quantity>
barcoding.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF3266633F5D36546DDBF8E7FF07F878" ID-DOI="http://doi.org/10.5281/zenodo.12753218" ID-Zenodo-Dep="12753218" httpUri="https://zenodo.org/record/12753218/files/figure.png" pageId="2" pageNumber="37" startId="2.[160,193,1913,1933]" targetBox="[429,1135,270,1852]" targetPageId="2" targetType="figure">
<paragraph id="8BF236EB3F5D36546DDBF8E7FF07F878" blockId="2.[160,1431,1913,1987]" pageId="2" pageNumber="37">
<emphasis id="B939EAF93F5D36546DDBF8E7FF08F836" bold="true" box="[160,220,1913,1933]" pageId="2" pageNumber="37">Fig. 1.</emphasis>
<taxonomicName id="4C4D4D683F5D36546D99F8E7FD6BF836" authority="(Goto &amp; Hatai, 1899) NSMT An" authorityName="NSMT An" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" box="[226,703,1913,1933]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="2" pageNumber="37" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B939EAF93F5D36546D99F8E7FE47F836" box="[226,403,1913,1933]" italics="true" pageId="2" pageNumber="37">Amynthas carnosus</emphasis>
(
<bibRefCitation id="EFDC4B1A3F5D36546CE7F8E7FD86F836" author="Goto, S. &amp; S. Hatai" box="[412,594,1913,1933]" pageId="2" pageNumber="37" pagination="13 - 24" refId="ref6050" refString="Goto, S. and S. Hatai. 1899. New or imperfectly known species of earthworms. No. 2. Annotations Zoologicae Japonensis 3 (1): 13 - 24." type="journal article" year="1899">Goto &amp; Hatai, 1899</bibRefCitation>
) NSMT An
</taxonomicName>
435 Neotype showing prostomium, ventral view of body with spermathecae and prostates
<emphasis id="B939EAF93F5D36546D81F80AFEE1F813" box="[250,309,1940,1960]" italics="true" pageId="2" pageNumber="37">in situ</emphasis>
(GM=Genital Marking with internal glands); also septal glands on dorsal blood vessel and intestinal caecum (Ihs=left side).
</paragraph>
</caption>
<subSubSection id="C35765603F5C36556DD9FEB7FD3AFD9A" pageId="3" pageNumber="38" type="diagnosis">
<paragraph id="8BF236EB3F5C36556DD9FEB7FD3AFD9A" blockId="3.[162,770,265,1991]" pageId="3" pageNumber="38">
<emphasis id="B939EAF93F5C36556DD9FEB7FEC2FEFA" bold="true" box="[162,278,297,321]" pageId="3" pageNumber="38">Diagnosis.</emphasis>
Size
<quantity id="4CB59B0E3F5C36556C2AFEB7FE0EFEFB" box="[337,474,297,320]" metricMagnitude="-1" metricUnit="m" metricValue="1.785" metricValueMax="2.4699999999999998" metricValueMin="1.1" pageId="3" pageNumber="38" unit="mm" value="178.5" valueMax="247.0" valueMin="110.0">110-247 mm</quantity>
. Spermathecal pores in 5/6/ 7/8/9 or rarely in 6/7/8/9 (Goto &amp; Hatai confused them in 5/6/7/8). Dorsal pores typically from 12/13. Genital markings typically closely paired mid-ventral and presetal in 8-9 and often also in 18-19 with other pairs just posterior (and sometimes anteriorly) median to the male pores; some variation apparently acceptable, including complete absence of markings. Intestinal caeca simple.
</paragraph>
</subSubSection>
<subSubSection id="C35765603F5C36556DD9FDB4FEF3FB98" pageId="3" pageNumber="38" type="distribution">
<paragraph id="8BF236EB3F5C36556DD9FDB4FEF3FB98" blockId="3.[162,770,265,1991]" pageId="3" pageNumber="38">
<emphasis id="B939EAF93F5C36556DD9FDB4FEECFDF9" bold="true" box="[162,312,554,578]" pageId="3" pageNumber="38">Distribution.</emphasis>
<collectingCountry id="F35A767B3F5C36556C39FDB4FE54FDFA" box="[322,384,554,577]" name="Japan" pageId="3" pageNumber="38">Japan</collectingCountry>
(
<bibRefCitation id="EFDC4B1A3F5C36556CF5FDB4FDA1FDFA" author="Goto, S. &amp; S. Hatai" box="[398,629,554,577]" pageId="3" pageNumber="38" pagination="13 - 24" refId="ref6050" refString="Goto, S. and S. Hatai. 1899. New or imperfectly known species of earthworms. No. 2. Annotations Zoologicae Japonensis 3 (1): 13 - 24." type="journal article" year="1899">Goto &amp; Hatai, 1899</bibRefCitation>
;
<bibRefCitation id="EFDC4B1A3F5C36556FFFFDB4FF33FDDA" author="Kobayashi, S." pageId="3" pageNumber="38" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi, 1936a</bibRefCitation>
;
<bibRefCitation id="EFDC4B1A3F5C36556D8DFDD4FE4DFDDA" author="Ohfuchi, S." box="[246,409,586,609]" pageId="3" pageNumber="38" pagination="31 - 136" refId="ref6660" refString="Ohfuchi, S. 1937. On the species possessing four pairs of spermathecae in the genus Pheretima, together with the variability of some external and internal characteristics. Saito Ho-On Kai Museum Research Bulletin 12: 31 - 136." type="journal article" year="1937">Ohfuchi, 1937</bibRefCitation>
;
<bibRefCitation id="EFDC4B1A3F5C36556CD3FDD4FDECFDDA" author="Easton, E. G." box="[424,568,586,609]" pageId="3" pageNumber="38" pagination="33 - 65" refId="ref5858" refString="Easton, E. G. 1981. Japanese earthworms: a synopsis of the Megadrile species. Bulletin of the British Museum (Natural History) Zoology 40: 33 - 65." type="journal article" year="1981">Easton, 1981</bibRefCitation>
- from Kyushu to Tohoku and
<collectingRegion id="4989F8093F5C36556C57FDF5FE41FD39" box="[300,405,619,642]" country="Japan" name="Hokkaido" pageId="3" pageNumber="38">Hokkaido</collectingRegion>
from reports he quotes by Kobayashi, 1936, 1941 and
<collectingRegion id="4989F8093F5C36556CD2FD15FDFEFD19" box="[425,554,651,674]" country="Japan" name="Yamaguchi" pageId="3" pageNumber="38">Yamaguchi</collectingRegion>
, 1962);
<collectingCountry id="F35A767B3F5C36556FF0FD15FD1AFD19" box="[651,718,651,674]" name="South Korea" pageId="3" pageNumber="38">Korea</collectingCountry>
and
<collectingRegion id="4989F8093F5C36556DD9FD35FF25FD78" box="[162,241,683,707]" country="South Korea" name="Jeju" pageId="3" pageNumber="38">Jeju-do</collectingRegion>
(=Quelpart)
<collectingCountry id="F35A767B3F5C36556CF6FD35FE1BFD79" box="[397,463,683,706]" name="Iceland" pageId="3" pageNumber="38">Island</collectingCountry>
(
<bibRefCitation id="EFDC4B1A3F5C36556CA0FD35FD72FD79" author="Kobayashi, S." box="[475,678,683,706]" pageId="3" pageNumber="38" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi, 1936a</bibRefCitation>
;
<bibRefCitation id="EFDC4B1A3F5C36556FCFFD35FD2FFD79" author="Kobayashi, S." box="[692,763,683,706]" pageId="3" pageNumber="38" pagination="139 - 184" refId="ref6452" refString="Kobayashi, S. 1936 b. Earthworms from Koryo, Korea. Science Report of the Tohoku Imperial University 11 (1): 139 - 184." type="journal article" year="1936">1936b</bibRefCitation>
;
<bibRefCitation id="EFDC4B1A3F5C36556DD9FD55FF0EFD59" author="Kobayashi, S." box="[162,218,715,738]" pageId="3" pageNumber="38" pagination="333 - 351" refId="ref6481" refString="Kobayashi, S. 1937. Preliminary survey of the earthworms of Quelpart Island. Science Report of the Tohoku Imperial University (B) 11 (3): 333 - 351." type="journal article" year="1937">1937</bibRefCitation>
;
<bibRefCitation id="EFDC4B1A3F5C36556D9DFD55FECBFD59" author="Kobayashi, S." box="[230,287,715,738]" pageId="3" pageNumber="38" pagination="89 - 170" refId="ref6515" refString="Kobayashi, S. 1938. Earthworms of Korea I. Science Report of the Tohoku Imperial University 13 (2): 89 - 170." type="journal article" year="1938">1938</bibRefCitation>
);
<collectingCountry id="F35A767B3F5C36556C48FD55FEA7FD59" box="[307,371,715,738]" name="China" pageId="3" pageNumber="38">China</collectingCountry>
(
<bibRefCitation id="EFDC4B1A3F5C36556C05FD55FE2AFD59" author="Chen, Y." box="[382,510,715,738]" pageId="3" pageNumber="38" refId="ref5837" refString="Chen, Y. 1959. Chinese Animal Atlas. Annelida. Science Press, Beijing [In Chinese]." type="book" year="1959">Chen, 1959</bibRefCitation>
); claimed from
<collectingRegion id="4989F8093F5C36556FD3FD55FD2FFD59" box="[680,763,715,738]" country="China" name="Jiangsu" pageId="3" pageNumber="38">Jiangsu</collectingRegion>
,
<collectingRegion id="4989F8093F5C36556DD9FD75FED0FCB9" box="[162,260,747,770]" country="China" name="Zhejiang" pageId="3" pageNumber="38">Zhejiang</collectingRegion>
,
<collectingRegion id="4989F8093F5C36556C69FD75FE83FCB9" box="[274,343,747,770]" country="China" name="Anhui" pageId="3" pageNumber="38">Anhui</collectingRegion>
,
<collectingRegion id="4989F8093F5C36556C1EFD75FE06FCB9" box="[357,466,747,770]" country="China" name="Shandong" pageId="3" pageNumber="38">Shandong</collectingRegion>
,
<collectingCountry id="F35A767B3F5C36556C9BFD75FDB0FCB9" box="[480,612,747,770]" name="Hong Kong" pageId="3" pageNumber="38">Hong Kong</collectingCountry>
, Sichuan, and Beijing [from Chinese Agricultural Academy of Science website: www.agrionline.net.cn/zhuanti/index.htm (2005), whence
<taxonomicName id="4C4D4D683F5C36556D86FCD2FE18FCD8" box="[253,460,844,867]" class="Clitellata" family="Megascolecidae" genus="Pheretima" kingdom="Animalia" order="Crassiclitellata" pageId="3" pageNumber="38" phylum="Annelida" rank="species" species="carnosa">
<emphasis id="B939EAF93F5C36556D86FCD2FE18FCD8" box="[253,460,844,867]" italics="true" pageId="3" pageNumber="38">Pheretima carnosa</emphasis>
</taxonomicName>
(sic) is dubiously described with either three or four pairs of spermathecae in 5/6/7/8, 8/9, possibly including
<taxonomicName id="4C4D4D683F5C36556CD6FC12FDD3FC18" authorityName="Stephenson" authorityYear="1925" box="[429,519,908,931]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="3" pageNumber="38" phylum="Annelida" rank="species" species="pingi">
<emphasis id="B939EAF93F5C36556CD6FC12FDD3FC18" box="[429,519,908,931]" italics="true" pageId="3" pageNumber="38">P. pingi</emphasis>
</taxonomicName>
subspecies];
<collectingCountry id="F35A767B3F5C36556FDAFC12FD2BFC18" box="[673,767,908,931]" name="Vietnam" pageId="3" pageNumber="38">Vietnam</collectingCountry>
(unconfirmed as
<taxonomicName id="4C4D4D683F5C36556C28FC32FE70FC78" authorityName="Stephenson" authorityYear="1925" box="[339,420,940,963]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="3" pageNumber="38" phylum="Annelida" rank="species" species="pingi">
<emphasis id="B939EAF93F5C36556C28FC32FE70FC78" box="[339,420,940,963]" italics="true" pageId="3" pageNumber="38">P. pingi</emphasis>
</taxonomicName>
). Relatively recently
<taxonomicName id="4C4D4D683F5C36556FFEFC32FD2BFC78" authorityName="NSMT An" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" box="[645,767,940,963]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="3" pageNumber="38" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="B939EAF93F5C36556FFEFC32FD2BFC78" box="[645,767,940,963]" italics="true" pageId="3" pageNumber="38">A. carnosus</emphasis>
</taxonomicName>
was (?mis-)described from
<collectingCountry id="F35A767B3F5C36556CB8FC52FDC6FC58" box="[451,530,972,995]" name="Taiwan" pageId="3" pageNumber="38">Taiwan</collectingCountry>
by
<bibRefCitation id="EFDC4B1A3F5C36556F40FC52FD2FFC58" author="Shen, H. - P. &amp; S. - C. Tsai &amp; C. - F. Tsai &amp; J. - H. Chen" box="[571,763,972,995]" pageId="3" pageNumber="38" pagination="95 - 100" refId="ref6776" refString="Shen, H. - P., S. - C. Tsai, C. - F. Tsai and J. - H. Chen. 2005. Occurrence of the Earthworm Amynthas carnosus (Goto and Hatai, 1899) in the Northern Taiwan. Endemic Species Research 7 (1): 95 - 100 [Available from: https: // tesri. coa. gov. tw / files / tesri _ protect / 124 / 070108. pdf]." type="journal article" year="2005">
Shen
<emphasis id="B939EAF93F5C36556F02FC52FD7BFC58" box="[633,687,972,995]" italics="true" pageId="3" pageNumber="38">et al.</emphasis>
(2005)
</bibRefCitation>
; this requires confirmation as noted by
<bibRefCitation id="EFDC4B1A3F5C36556F31FC72FEF7FB98" author="Blakemore, R. J. &amp; C. - H. Chang &amp; S. - C. Chuang &amp; M. T. Ito &amp; S. James &amp; J. - H. Chen" pageId="3" pageNumber="38" pagination="226 - 236" refId="ref5530" refString="Blakemore, R. J., C. - H. Chang, S. - C. Chuang, M. T. Ito, S. James and J. - H. Chen. 2006. Biodiversity of Earthworms in Taiwan: a species checklist with the confirmation and new records of the exotic lumbricids Eisenia fetida and Eiseniella tetraedra. Taiwania 51 (3): 226 - 236 [Available from: http: // tai 2. ntu. edu. tw / udth / bin / taiwania. exe / dl? id = 507 or http: // tai 2. ntu. edu. tw / taiwania / pdf / tai. 2006.51.3. 226. pdf or http: // www. press. ntu. edu. tw / ejournal / Files / taiwan / 200609 / 10. pdf]." type="journal article" year="2006">
Blakemore
<emphasis id="B939EAF93F5C36556FB1FC72FCD6FBB8" box="[714,770,1004,1027]" italics="true" pageId="3" pageNumber="38">et al.</emphasis>
(2006: 228)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35765603F5C36536DD9FBB2FEA2FEDB" lastPageId="5" lastPageNumber="40" pageId="3" pageNumber="38" type="description">
<paragraph id="8BF236EB3F5C36536DD9FBB2FEA2FEDB" blockId="3.[162,770,265,1991]" lastBlockId="5.[162,767,265,352]" lastPageId="5" lastPageNumber="40" pageId="3" pageNumber="38">
<emphasis id="B939EAF93F5C36556DD9FBB2FEF2FBFF" bold="true" box="[162,294,1068,1092]" pageId="3" pageNumber="38">Description</emphasis>
(
<typeStatus id="54F688493F5C36556C49FBB2FE5AFBF8" box="[306,398,1068,1091]" pageId="3" pageNumber="38" type="neotype">Neotype</typeStatus>
compared to the original descriptions and synonymy above, excluding typical
<taxonomicName id="4C4D4D683F5C36556FDDFBD3FD2BFBDF" authorityName="Stephenson" authorityYear="1925" box="[678,767,1101,1124]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="3" pageNumber="38" phylum="Annelida" rank="species" species="pingi">
<emphasis id="B939EAF93F5C36556FDDFBD3FD2BFBDF" box="[678,767,1101,1124]" italics="true" pageId="3" pageNumber="38">A. pingi</emphasis>
</taxonomicName>
characteristics). Body length 180+ mm (cf. 143-153 by Goto &amp; Hatai or 110-247 by
<bibRefCitation id="EFDC4B1A3F5C36556CB4FB13FD59FB1F" author="Kobayashi, S." box="[463,653,1165,1188]" pageId="3" pageNumber="38" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi, 1936a</bibRefCitation>
), segments 111+ (cf. 106-126 by Goto &amp; Hatai or 110-179 by Kobayashi). Dark brown dorsum with darker clitellum. First dorsal pore 12/13 (cf. 13/14 Goto &amp; Hatai lapsus, or 11/ 12/
<quantity id="4CB59B0E3F5C36556DBDFA93FF28FA9F" box="[198,252,1293,1316]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="3" pageNumber="38" unit="in" value="13.0">13 in</quantity>
some other accounts). Setae 24-69. Spermathecal pores wide in 5/6/7/8/9 (initially confused by Goto &amp; Hatai as 5/6/7/8). Genital markings closely paired presetally in 8 and 9 (7 and 8 according to Goto &amp; Hatai, lapsus -
<figureCitation id="13762A6E3F5C36556DACFA10FECEFA1E" box="[215,282,1422,1445]" captionStart="Fig" captionStartId="3.[828,861,1635,1655]" captionTargetBox="[833,1428,268,1594]" captionTargetId="figure-622@3.[825,1434,262,1630]" captionTargetPageId="3" captionText="Fig. 2. Amynthas carnosus: upper Goto &amp; Hatais original fig. 4; lower Ph. kyamikia Kobayashi, 1934, figs. 1-3 (as exactly similar in syns. youngtai and sangyeoli)." figureDoi="http://doi.org/10.5281/zenodo.12753220" httpUri="https://zenodo.org/record/12753220/files/figure.png" pageId="3" pageNumber="38">Fig. 2</figureCitation>
), in 18 and often in 19; other markings just anterioventral to superficial male pores in 18 postsetally (
<typeStatus id="54F688493F5C36556DD2FA50FF2BFA5E" box="[169,255,1486,1509]" pageId="3" pageNumber="38" type="neotype">neotype</typeStatus>
in agreement except that 19rhs presetal is unilateral -
<figureCitation id="13762A6E3F5C36556D8CFA70FEE1F9BE" box="[247,309,1518,1541]" captionStart="Fig" captionStartId="2.[160,193,1913,1933]" captionTargetBox="[429,1135,270,1852]" captionTargetId="figure-23@2.[419,1191,250,1939]" captionTargetPageId="2" captionText="Fig. 1. Amynthas carnosus (Goto &amp; Hatai, 1899) NSMT An435 Neotype showing prostomium, ventral view of body with spermathecae and prostates in situ (GM=Genital Marking with internal glands); also septal glands on dorsal blood vessel and intestinal caecum (Ihs=left side)." figureDoi="http://doi.org/10.5281/zenodo.12753218" httpUri="https://zenodo.org/record/12753218/files/figure.png" pageId="3" pageNumber="38">Fig. 1</figureCitation>
); in Goto &amp; Hatais second specimen and 50% of Kobayashis material both these mid-ventral pairs in 18 and 19 were absent and sometimes no markings were near male pores either, while a few other specimens lacked markings entirely or had extra markings in 18. Considerable variation was permitted in both preclitellar and postclitellar marking locations by
<bibRefCitation id="EFDC4B1A3F5C36556F41F931FCD4F97D" author="Kobayashi, S." box="[570,768,1711,1734]" pageId="3" pageNumber="38" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi (1936a)</bibRefCitation>
-
<figureCitation id="13762A6E3F5C36556DD4F951FF25F95D" box="[175,241,1743,1766]" captionStart="Fig" captionStartId="4.[160,193,1949,1969]" captionTargetBox="[160,1410,247,1923]" captionTargetId="figure-23@4.[96,1411,211,1992]" captionText="Fig. 3. Variation in genital markings and spermathecal pores from Kobayashi (1936: text-figs. 1-2) showing permissible variations as determined by his detailed study." figureDoi="http://doi.org/10.5281/zenodo.12784055" httpUri="https://zenodo.org/record/12784055/files/figure.png" pageId="3" pageNumber="38">Fig. 3</figureCitation>
. Sessile glands correspond to the markings internally. No accessory pore glands noted, neither near to spermathecal nor to male pores (cf.
<bibRefCitation id="EFDC4B1A3F5C36556F4AF88EFD20F89C" author="Shen, H. - P. &amp; S. - C. Tsai &amp; C. - F. Tsai &amp; J. - H. Chen" box="[561,756,1808,1831]" pageId="3" pageNumber="38" pagination="95 - 100" refId="ref6776" refString="Shen, H. - P., S. - C. Tsai, C. - F. Tsai and J. - H. Chen. 2005. Occurrence of the Earthworm Amynthas carnosus (Goto and Hatai, 1899) in the Northern Taiwan. Endemic Species Research 7 (1): 95 - 100 [Available from: https: // tesri. coa. gov. tw / files / tesri _ protect / 124 / 070108. pdf]." type="journal article" year="2005">
Shen
<emphasis id="B939EAF93F5C36556F0AF88EFD7EF89C" box="[625,682,1808,1831]" italics="true" pageId="3" pageNumber="38">et al.</emphasis>
, 2005
</bibRefCitation>
). Internally the pharyngeal mass extends to 4 and tufted meroic nephridia are in 5 and 6.
<collectingRegion id="4989F8093F5C36556C86F8CEFDECF8DC" box="[509,568,1872,1895]" country="Spain" name="Ceuta" pageId="3" pageNumber="38">Septa</collectingRegion>
8/9/10 are aborted or 8/9 sometimes retained and displaced by gizzard (
<bibRefCitation id="EFDC4B1A3F5C36556FAFF8EEFEA8F81C" author="Kobayashi, S." pageId="3" pageNumber="38" pagination="115 - 138" refId="ref6409" refString="Kobayashi, S. 1936 a. Distribution and some external characteristics of Pheretima (Ph.) carnosa (Goto et Hatai) from Korea. Science Report of the Tohoku Imperial University 11 (1): 115 - 138." type="journal article" year="1936">Kobayashi, 1936a: 116</bibRefCitation>
); 10/11/12 and sometimes 12/13 are strong; thereafter membranous. Spermathecae in 6-9, the first pair often smaller (and sometimes absent) the last two pairs after septum 7/8; with diverticula about half the length of duct plus ampulla. Last hearts in 13. From ca. 15/16 distinct paired septal glands on dorsal blood vessel occur (
<figureCitation id="13762A6E3F5C36556EF8F8EEFC10F83C" box="[899,964,1904,1927]" captionStart="Fig" captionStartId="2.[160,193,1913,1933]" captionTargetBox="[429,1135,270,1852]" captionTargetId="figure-23@2.[419,1191,250,1939]" captionTargetPageId="2" captionText="Fig. 1. Amynthas carnosus (Goto &amp; Hatai, 1899) NSMT An435 Neotype showing prostomium, ventral view of body with spermathecae and prostates in situ (GM=Genital Marking with internal glands); also septal glands on dorsal blood vessel and intestinal caecum (Ihs=left side)." figureDoi="http://doi.org/10.5281/zenodo.12753218" httpUri="https://zenodo.org/record/12753218/files/figure.png" pageId="3" pageNumber="38">Fig. 1</figureCitation>
). Seminal vesicles with dorsal appendages in 11 and 12. Ovaries in 13 with vestigial ovisacs on posterior of 13/14. Intestine origin in 15 with simple caeca from 27 extnding forward to 24 or 23. Gut contains yellow soil in
<typeStatus id="54F688493F5A36536C60FEB7FEACFEFB" box="[283,376,297,320]" pageId="5" pageNumber="40" type="neotype">Neotype</typeStatus>
, i.e., low organic content suggesting a geophageous diet.
</paragraph>
</subSubSection>
</treatment>
</document>