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<document id="73FD229DD313EB41D6EC995D66151B48" ID-DOI="10.11646/zootaxa.5536.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14023827" ID-ZooBank="C47EE4D9-D5B5-4B67-8547-856FBAEFE670" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1730455362568" checkinUser="plazi" docAuthor="Wang, Shuxia, Liu, Chen &amp; David, Lees" docDate="2024" docId="03E41827FFA10A45FD8EFAA1FDF4FD67" docLanguage="en" docName="zootaxa.5536.1.1.pdf" docOrigin="Zootaxa 5536 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Promalactis apiciserrulata Wang &amp; Liu &amp; David 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="21" masterDocId="FFDD605FFFB20A51FD19FF9BFFEEFFCE" masterDocTitle="Partial revision of Promalactis Meyrick (Lepidoptera: Oecophoridae), based on specimens deposited in the Natural History Museum, London" masterLastPageNumber="58" masterPageNumber="1" pageNumber="20" updateTime="1730824655839" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="301BBB6B65EE7E99AAC1FC6A956AEA18">Partial revision of Promalactis Meyrick (Lepidoptera: Oecophoridae), based on specimens deposited in the Natural History Museum, London</mods:title>
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<mods:namePart id="759AFB7E83F6113446279670DAA0A165">Wang, Shuxia</mods:namePart>
<mods:affiliation id="D5AE0434B8E311A7943D093F855E387D">College of Life Sciences, Nankai University, Tianjin 300071, China</mods:affiliation>
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<mods:namePart id="E030B18A77C3BA6E1D4F2D133726D2D5">Liu, Chen</mods:namePart>
<mods:nameIdentifier id="65CA081E8AF32C24525A55EBC1F7902D" type="ORCID">0000-0002-9111-6644</mods:nameIdentifier>
<mods:affiliation id="519065443EA05E53F6778D6B5E97C6B3">College of Life Sciences, Nankai University, Tianjin 300071, China</mods:affiliation>
<mods:nameIdentifier id="C71452C812E6EA3D13BA3D507E380268" type="email">348143302@qq.com</mods:nameIdentifier>
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<mods:namePart id="299E366690F42AFD4392BC5C635306F4">David, Lees</mods:namePart>
<mods:nameIdentifier id="B06BE73BB3A451AA2F8B894194B3DFF2" type="ORCID">0000-0002-7580-8560</mods:nameIdentifier>
<mods:affiliation id="3789470F9AB3B3B148EE1941C85C51DF">Department of Science, Natural History Museum, London, United Kingdom, SW 7 5 BD</mods:affiliation>
<mods:nameIdentifier id="BC7518884E0C15C8F9463E93A8D9D94D" type="email">david.lees@nhm.ac.uk</mods:nameIdentifier>
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<mods:title id="4D847BF005BAE71BA8E5FD7114E58015">Zootaxa</mods:title>
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<mods:date id="AA4CA06CF108E02A33BE38861748A3C3">2024</mods:date>
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<mods:number id="0B446497EF3B1A2C522A166CB081724D">2024-11-01</mods:number>
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<mods:number id="1E612B981F64E27C7D61A59C913F234A">5536</mods:number>
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<treatment id="03E41827FFA10A45FD8EFAA1FDF4FD67" ID-DOI="http://doi.org/10.5281/zenodo.14023873" ID-Zenodo-Dep="14023873" LSID="urn:lsid:plazi:treatment:03E41827FFA10A45FD8EFAA1FDF4FD67" httpUri="http://treatment.plazi.org/id/03E41827FFA10A45FD8EFAA1FDF4FD67" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="C357FABAFFA10A42FD8EFAA1FD7AFA9B" box="[151,660,1338,1365]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8BF2A931FFA10A42FD8EFAA1FD7AFA9B" blockId="19.[151,660,1338,1401]" box="[151,660,1338,1365]" pageId="19" pageNumber="20">
<heading id="D0BA1E5DFFA10A42FD8EFAA1FD7AFA9B" bold="true" box="[151,660,1338,1365]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<emphasis id="B9397523FFA10A42FD8EFAA1FD7AFA9B" bold="true" box="[151,660,1338,1365]" pageId="19" pageNumber="20">
<taxonomicName id="4C4DD2B2FFA10A42FD8EFAA1FDC7FA9B" authority="Wang" authorityName="Wang &amp; Liu &amp; David" authorityYear="2024" box="[151,553,1338,1365]" class="Insecta" family="Oecophoridae" genus="Promalactis" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="apiciserrulata" status="sp. nov.">
<emphasis id="B9397523FFA10A42FD8EFAA1FE36FA9B" bold="true" box="[151,472,1338,1365]" italics="true" pageId="19" pageNumber="20">Promalactis apiciserrulata</emphasis>
Wang
</taxonomicName>
,
<taxonomicNameLabel id="A20AC858FFA10A42FF2FFAA0FD7AFA9B" box="[566,660,1339,1365]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFA10A42FD8EFAC4FEC9FAB7" box="[151,295,1374,1401]" pageId="19" pageNumber="20" type="description">
<paragraph id="8BF2A931FFA10A42FD8EFAC4FEC9FAB7" blockId="19.[151,660,1338,1401]" box="[151,295,1374,1401]" pageId="19" pageNumber="20">
(
<figureCitation id="1376B5B4FFA10A42FD86FAC4FF18FAB7" box="[159,246,1375,1401]" captionStart="FIGURES 17–23" captionStartId="18.[151,264,1842,1866]" captionTargetBox="[184,1404,193,1806]" captionTargetId="figure-23@18.[172,1416,181,1819]" captionTargetPageId="18" captionText="FIGURES 17–23. Adults of Promalactis spp. 17. P. vulgaris sp. nov.; 18. P. apicibilobata sp. nov.; 19. P. apiciserrulata sp. nov.; 20. P. bacilliforma sp. nov.; 21. campanimorpha sp. nov.; 22. P. ciliavalva sp. nov.; 23. P. abortiva sp. nov. All males and holotypes. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023837" httpUri="https://zenodo.org/record/14023837/files/figure.png" pageId="19" pageNumber="20">Figs 19</figureCitation>
,
<figureCitation id="1376B5B4FFA10A42FC1BFAC5FEF1FAB7" box="[258,287,1374,1401]" captionStart="FIGURES 52–57" captionStartId="35.[151,264,1927,1951]" captionTargetBox="[186,1393,208,1895]" captionTargetId="figure-20@35.[159,1428,181,1903]" captionTargetPageId="35" captionText="FIGURES 52–57. Male genitalia of Promalactis spp. 52. P. apiciserrulata sp. nov.; 53. P. bacilliforma sp. nov.; 54. P. campanimorpha sp. nov.; 55. P. ciliavalva sp. nov.; 56. P. abortiva sp. nov.; 57. P. magnisaccata sp. nov. All holotypes. Scale bar: 52–53, 55–57 = 0.2 mm; 54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14023853" httpUri="https://zenodo.org/record/14023853/files/figure.png" pageId="19" pageNumber="20">52</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFA10A42FD8EFA3CFD9BFA2B" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BF2A931FFA10A42FD8EFA3CFD9BFA2B" blockId="19.[151,1437,1446,2013]" pageId="19" pageNumber="20">
<emphasis id="B9397523FFA10A42FD8EFA3CFEABFA0E" bold="true" box="[151,325,1446,1473]" pageId="19" pageNumber="20">Type material.</emphasis>
<materialsCitation id="3B25A36CFFA10A42FC55FA3DFD9BFA2B" collectingDate="1989-04-30" collectionCode="NHMUK" collectorName="M. G. Allen &amp; K. R. Tuck" country="Brunei" elevation="365" location="Ulu Temburong" pageId="19" pageNumber="20" specimenCode="NHMUK010316992" specimenCount="1" specimenCount-male="1" stateProvince="Temburong" typeStatus="holotype">
<emphasis id="B9397523FFA10A42FC55FA3DFE46FA0E" bold="true" box="[332,424,1446,1472]" pageId="19" pageNumber="20">
<collectingCountry id="F35AE9A1FFA10A42FC55FA3DFE4DFA0E" box="[332,419,1446,1472]" name="Brunei" pageId="19" pageNumber="20">Brunei</collectingCountry>
:
</emphasis>
<typeStatus id="54F61793FFA10A42FCB6FA3CFDF9FA0F" box="[431,535,1447,1473]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4B62B8FFA10A42FF06FA3CFDD8FA0F" box="[543,566,1447,1473]" count="1" pageId="19" pageNumber="20" type="male">♂</specimenCount>
,
<location id="8E92FFEAFFA10A42FF5BFA3CFD18FA0F" LSID="urn:lsid:plazi:treatment:03E41827FFA10A45FD8EFAA1FDF4FD67:8E92FFEAFFA10A42FF5BFA3CFD18FA0F" box="[578,758,1447,1473]" country="Brunei" name="Ulu Temburong" pageId="19" pageNumber="20" stateProvince="Temburong">Ulu Temburong</location>
,
<quantity id="4CB504D4FFA10A42FE18FA3DFCA5FA0F" box="[769,843,1446,1473]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="19" pageNumber="20" unit="m" value="365.0">
<elevation id="00604E02FFA10A42FE18FA3DFCA5FA0F" box="[769,843,1446,1473]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="19" pageNumber="20" unit="m" value="365.0">365 m</elevation>
</quantity>
,
<date id="FFF38FF1FFA10A42FE4FFA3CFC3BFA0F" box="[854,981,1447,1473]" pageId="19" pageNumber="20" value="1989-04-30">
<collectingDate id="EFB77619FFA10A42FE4FFA3CFC3BFA0F" box="[854,981,1447,1473]" pageId="19" pageNumber="20" value="1989-04-30">30.IV.1989</collectingDate>
</date>
, LP 283, GR 843887, leg.
<collectorName id="26B8CCE7FFA10A42F819FA3CFA91FA0F" box="[1280,1407,1447,1473]" pageId="19" pageNumber="20">M.G. Allen</collectorName>
&amp;
<collectorName id="26B8CCE7FFA10A42FD8EFA50FEE0FA2B" box="[151,270,1483,1509]" pageId="19" pageNumber="20">K.R. Tuck</collectorName>
, slide No.
<specimenCode id="DBEB014AFFA10A42FC9CFA50FD9FFA2B" box="[389,625,1483,1509]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="20" type="Museum">NHMUK010316992</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFA10A42FDDEFA75FDA9F99F" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BF2A931FFA10A42FDDEFA75FDA9F99F" blockId="19.[151,1437,1446,2013]" pageId="19" pageNumber="20">
<emphasis id="B9397523FFA10A42FDDEFA75FEACF9C6" bold="true" box="[199,322,1518,1544]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
The new species is distinguishable in the male genitalia by the costa with a spine-shaped process extending outward parallelly with the costal margin, the valva serrate at rounded apex, and the phallus with a sub-rectangular, apex-serrate plate distally.
</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFA10A45FDDEF9C1FE7FFDF3" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
<paragraph id="8BF2A931FFA10A42FDDEF9C1FCDFF9BB" blockId="19.[151,1437,1446,2013]" box="[199,817,1626,1653]" pageId="19" pageNumber="20">
<emphasis id="B9397523FFA10A42FDDEF9C1FEBFF9BA" bold="true" box="[199,337,1626,1652]" pageId="19" pageNumber="20">Description</emphasis>
. Adult (
<figureCitation id="1376B5B4FFA10A42FCB4F9C0FDEFF9BB" box="[429,513,1627,1653]" captionStart="FIGURES 17–23" captionStartId="18.[151,264,1842,1866]" captionTargetBox="[184,1404,193,1806]" captionTargetId="figure-23@18.[172,1416,181,1819]" captionTargetPageId="18" captionText="FIGURES 17–23. Adults of Promalactis spp. 17. P. vulgaris sp. nov.; 18. P. apicibilobata sp. nov.; 19. P. apiciserrulata sp. nov.; 20. P. bacilliforma sp. nov.; 21. campanimorpha sp. nov.; 22. P. ciliavalva sp. nov.; 23. P. abortiva sp. nov. All males and holotypes. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14023837" httpUri="https://zenodo.org/record/14023837/files/figure.png" pageId="19" pageNumber="20">Fig. 19</figureCitation>
). Forewing length
<quantity id="4CB504D4FFA10A42FFCDF9C1FCC3F9BB" box="[724,813,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="19" pageNumber="20" unit="mm" value="4.5">4.5 mm</quantity>
.
</paragraph>
<paragraph id="8BF2A931FFA10A42FDDEF9E4FE37F8CB" blockId="19.[151,1437,1446,2013]" pageId="19" pageNumber="20">
<emphasis id="B9397523FFA10A42FDDEF9E4FEEAF957" box="[199,260,1663,1689]" italics="true" pageId="19" pageNumber="20">Head</emphasis>
. Vertex snowy white, frons grey, occiput rust brown. Labial palpus with second segment dark brown on outer surface, pale yellow on inner surface; third segment black except white at base and apex. Antennal scape white dorsally, pale grey ventrally; flagellum black, alternating with white on dorsal surface except basal several flagellomeres entirely white.
</paragraph>
<paragraph id="8BF2A931FFA10A45FDDEF894FB6EFF37" blockId="19.[151,1437,1446,2013]" lastBlockId="20.[151,1437,151,681]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="B9397523FFA10A42FDDEF894FEF6F8E7" box="[199,280,1807,1833]" italics="true" pageId="19" pageNumber="20">Thorax</emphasis>
. Mesonotum and tegula rust brown, mesonotum white apically. Forewing rust brown; markings white edged with black scales: costal spot sub-ovate, from distal 1/3 extending to above posterior angle of cell; basal streak slender, reaching base of costal margin, beyond it placed a dot below costal margin; antemedian streak from middle of dorsum to basal 1/3 of costal margin, slightly curved outward, widened anteriorly; dorsal streak from before tornus oblique outward to before posterior angle of cell; apical spot ovate; fringe greyish black except orange yellow on extension of apical spot. Hindwing and fringe grey. Foreleg black except femur pale grey ventrally, tibia with white spot at base and middle, with a tuft of white scales at apex, tarsus white at apex of basal tarsomere and on entire second tarsomere; midleg grey ventrally, black dorsally, tibia with a white spot at base, with a tuft of white scales at apex, tarsus white at apices of basal two and apical tarsomeres; hindleg missing.
</paragraph>
<paragraph id="8BF2A931FFA60A45FDDEFE98FB8AFDD7" blockId="20.[151,1437,151,681]" pageId="20" pageNumber="21">
<emphasis id="B9397523FFA60A45FDDEFE98FE87FED3" box="[199,361,259,285]" italics="true" pageId="20" pageNumber="21">Male genitalia</emphasis>
(
<figureCitation id="1376B5B4FFA60A45FC6FFE98FE2BFED3" box="[374,453,258,285]" captionStart="FIGURES 52–57" captionStartId="35.[151,264,1927,1951]" captionTargetBox="[186,1393,208,1895]" captionTargetId="figure-20@35.[159,1428,181,1903]" captionTargetPageId="35" captionText="FIGURES 52–57. Male genitalia of Promalactis spp. 52. P. apiciserrulata sp. nov.; 53. P. bacilliforma sp. nov.; 54. P. campanimorpha sp. nov.; 55. P. ciliavalva sp. nov.; 56. P. abortiva sp. nov.; 57. P. magnisaccata sp. nov. All holotypes. Scale bar: 52–53, 55–57 = 0.2 mm; 54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14023853" httpUri="https://zenodo.org/record/14023853/files/figure.png" pageId="20" pageNumber="21">Fig. 52</figureCitation>
). Uncus elongately slender, slightly wide at base. Gnathos separate from base; lateral lobe enlarged basally, slender and widely apart distally, pointed apically. Tegumen uniformly wide, inverted V-shaped. Valva sub-rectangular, with a fold running from below base to before basal 2/3 of costal margin, apex obtusely rounded, serrate; costa with a fine spine-like process arising from beyond middle and extending outward parallelly with costal margin; sacculus wide at base, narrowed and ending at distal 1/3 of ventral margin of valva. Saccus elongate triangular, 1/2 length of valva, wide at base, narrowed to narrowly rounded apex. Juxta sub-rectangular, weakly sclerotized; basal lobe short, digitate. Phallus about same length as valva, with a large, sub-rectangular, apex-serrate plate distally; cornutus long and curved, longer than 1/2 length of phallus.
</paragraph>
<paragraph id="8BF2A931FFA60A45FDDEFDB8FE7FFDF3" blockId="20.[151,1437,151,681]" box="[199,401,547,573]" pageId="20" pageNumber="21">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFA60A45FDDEFDDDFE55FDAF" box="[199,443,582,609]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BF2A931FFA60A45FDDEFDDDFE55FDAF" blockId="20.[151,1437,151,681]" box="[199,443,582,609]" pageId="20" pageNumber="21">
<emphasis id="B9397523FFA60A45FDDEFDDDFE8FFDAE" bold="true" box="[199,353,582,608]" pageId="20" pageNumber="21">Distribution.</emphasis>
<collectingCountry id="F35AE9A1FFA60A45FC7EFDDCFE59FDAF" box="[359,439,583,609]" name="Brunei" pageId="20" pageNumber="21">Brunei</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C357FABAFFA60A45FDDEFDF0FDF4FD67" pageId="20" pageNumber="21" type="etymology">
<paragraph id="8BF2A931FFA60A45FDDEFDF0FDF4FD67" blockId="20.[151,1437,151,681]" pageId="20" pageNumber="21">
<emphasis id="B9397523FFA60A45FDDEFDF0FEA0FD4B" bold="true" box="[199,334,619,645]" pageId="20" pageNumber="21">Etymology.</emphasis>
The specific epithet is compounded from the Latin
<emphasis id="B9397523FFA60A45FEB8FDF0FC38FD4A" box="[929,982,619,644]" italics="true" pageId="20" pageNumber="21">apex</emphasis>
and
<emphasis id="B9397523FFA60A45F916FDF0FB6FFD4B" box="[1039,1153,619,645]" italics="true" pageId="20" pageNumber="21">serrulatus</emphasis>
, referring to the apically serrate valva of the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>
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