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<document id="5F81205A20A2B25E6255B80B77E83329" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627005272297" checkinUser="felipe" docAuthor="Published, First" docDate="2008" docId="03DF87D2FF27ABB99EC8134245023B0F" docLanguage="en" docName="zt01804p456part1.pdf" docOrigin="Zootaxa 1804" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Corethrella (Corethrella) nudistyla Borkent and Szadziewski" docType="treatment" docVersion="2" lastPageNumber="205" masterDocId="FFE6FFAAFFECAB759E5F1025465D3823" masterDocTitle="The Frog-Biting Midges of the World (Corethrellidae: Diptera)" masterLastPageNumber="456" masterPageNumber="1" pageNumber="204" updateTime="1699014769975" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="59CC78FE46A41FA17536C665AAC05202">The Frog-Biting Midges of the World (Corethrellidae: Diptera)</mods:title>
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<subSubSection id="C36C654FFF27ABBE9EC8134245223BA2" box="[151,895,871,897]" pageId="203" pageNumber="204" type="nomenclature">
<paragraph id="8BC936C4FF27ABBE9EC8134245223BA2" blockId="203.[151,895,871,897]" box="[151,895,871,897]" pageId="203" pageNumber="204">
<heading id="D08181A8FF27ABBE9EC8134245223BA2" bold="true" box="[151,895,871,897]" fontSize="11" level="1" pageId="203" pageNumber="204" reason="1">
<taxonomicName id="4C764D47FF27ABBE9EC8134245223BA2" ID-CoL="8BL6R" authority="Borkent and Szadziewski" authorityName="Borkent and Szadziewski" box="[151,895,871,897]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="203" pageNumber="204" phylum="Arthropoda" rank="species" species="nudistyla" subGenus="Corethrella">
<emphasis id="B902EAD6FF27ABBE9EC8134245223BA2" bold="true" box="[151,895,871,897]" pageId="203" pageNumber="204">
<emphasis id="B902EAD6FF27ABBE9EC81342477C3BA2" bold="true" box="[151,289,871,897]" italics="true" pageId="203" pageNumber="204">Corethrella</emphasis>
(
<emphasis id="B902EAD6FF27ABBE9F6D134247E13BA2" bold="true" box="[306,444,871,897]" italics="true" pageId="203" pageNumber="204">Corethrella</emphasis>
)
<emphasis id="B902EAD6FF27ABBE9F91134244613BA2" bold="true" box="[462,572,871,897]" italics="true" pageId="203" pageNumber="204">nudistyla</emphasis>
Borkent and Szadziewski
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF27ABBE9EC8138D43DD3B9C" box="[151,1408,936,959]" pageId="203" pageNumber="204" type="reference_group">
<paragraph id="8BC936C4FF27ABBE9EC8138D43DD3B9C" blockId="203.[151,1408,936,959]" box="[151,1408,936,959]" pageId="203" pageNumber="204">
<treatmentCitationGroup id="AB6611EAFF27ABBE9EC8138D43DD3B9C" box="[151,1408,936,959]" pageId="203" pageNumber="204">
<treatmentCitation id="0AD710D5FF27ABBE9EC8138D44A33B9C" author="Borkent, A. &amp; Szadziewski, R." box="[151,766,936,959]" page="460" pageId="203" pageNumber="204" year="1992">
<taxonomicName id="4C764D47FF27ABBE9EC8138D44A33B9C" ID-CoL="8BL6R" authority="Borkent and Szadziewski 1992: 460" authorityName="Borkent and Szadziewski" authorityPageNumber="460" authorityYear="1992" box="[151,766,936,959]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="203" pageNumber="204" phylum="Arthropoda" rank="species" species="nudistyla">
<emphasis id="B902EAD6FF27ABBE9EC8138D47243B9C" box="[151,377,936,959]" italics="true" pageId="203" pageNumber="204">Corethrella nudistyla</emphasis>
<materialsCitation id="3B1E3C99FF27ABBE9FDF138D44A33B9C" box="[384,766,936,959]" collectionCode="AMNH" country="Dominican Republic" location="Dominican Republic" pageId="203" pageNumber="204" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<bibRefCitation id="EFE74B35FF27ABBE9FDF138D44A33B9C" author="Borkent, A. &amp; Szadziewski, R." box="[384,766,936,959]" pageId="203" pageNumber="204" pagination="457 - 463" refId="ref236849" refString="Borkent, A. &amp; Szadziewski, R. (1992) The first records of fossil Corethrellidae (Diptera). Entomologica Scandinavica, 22, 457 - 463." type="journal article" year="1992">Borkent and Szadziewski 1992:460</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
. Type-locality:
<collectingCountry id="F3617654FF27ABBE9DFB138D42D93B9C" box="[932,1156,936,959]" name="Dominican Republic" pageId="203" pageNumber="204">Dominican Republic</collectingCountry>
.
<typeStatus id="54CD8866FF27ABBE9ACA138D42A53B9C" box="[1173,1272,936,959]" pageId="203" pageNumber="204" type="holotype">Holotype</typeStatus>
<specimenCount id="9D70FD4DFF27ABBE9AA0138C43523B9A" box="[1279,1295,937,953]" pageId="203" pageNumber="204" type="male"></specimenCount>
(AMNH).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF27ABBE9EC813C045883C04" pageId="203" pageNumber="204" type="diagnosis">
<paragraph id="8BC936C4FF27ABBE9EC813C045883C04" blockId="203.[151,1436,997,2023]" pageId="203" pageNumber="204">
DIAGNOSIS:
<emphasis id="B902EAD6FF27ABBE9F1413C047973BDD" box="[331,458,997,1022]" italics="true" pageId="203" pageNumber="204">Male adult</emphasis>
: only fossil species of
<taxonomicName id="4C764D47FF27ABBE9C8B13C045073BDD" box="[724,858,997,1022]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="203" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="B902EAD6FF27ABBE9C8B13C045073BDD" box="[724,858,997,1022]" italics="true" pageId="203" pageNumber="204">Corethrella</emphasis>
</taxonomicName>
with slender scales on midfemur (as in Fig. 74A) and with elongate R
<subScript id="17F23481FF27ABBE9FDD143847D73C08" attach="left" box="[386,394,1053,1067]" fontSize="6" pageId="203" pageNumber="204">2</subScript>
and R
<subScript id="17F23481FF27ABBE9F89143847833C08" attach="left" box="[470,478,1053,1067]" fontSize="6" pageId="203" pageNumber="204">3</subScript>
(as in Fig. 62A).
<emphasis id="B902EAD6FF27ABBE9CEA1428450D3C05" box="[693,848,1037,1062]" italics="true" pageId="203" pageNumber="204">Female adult</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF27ABBE9E99141D44F53FB9" pageId="203" pageNumber="204" type="description">
<paragraph id="8BC936C4FF27ABBE9E99141D47043F51" blockId="203.[151,1436,997,2023]" pageId="203" pageNumber="204">
DESCRIPTION:
<emphasis id="B902EAD6FF27ABBE9FC6141D44443C72" box="[409,537,1080,1105]" italics="true" pageId="203" pageNumber="204">Male adult</emphasis>
. Habitus as in Fig. 113C. Descriptive statistics: see
<tableCitation id="C6F4037FFF27ABBE9A20141D435F3C71" box="[1151,1282,1080,1106]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStart-2="TABLE 4" captionStart-3="TABLE 5" captionStartId-0="269.[151,241,151,175]" captionStartId-1="271.[151,241,153,177]" captionStartId-2="273.[151,241,151,175]" captionStartId-3="275.[151,241,151,175]" captionText-0="TABLE 2. Length of wing of males of species of Corethrella. Values are in mm." captionText-1="TABLE 3. Ratio of foreleg tarsomeres 3/4 of males of species of Corethrella." captionText-2="TABLE 4. Ratio of distance of dorsomedial seta from the gonocoxite base/length of the gonocoxite of males of species" captionText-3="TABLE 5. Ratio of distance of subbasal seta from the gonostylus base/ length of the gonostylus of males of species of Corethrella." pageId="203" pageNumber="204">Tables 25</tableCitation>
.
<emphasis id="B902EAD6FF27ABBE9B4B141D430A3C71" bold="true" box="[1300,1367,1080,1106]" pageId="203" pageNumber="204">Head</emphasis>
: Outline in anterior view laterally elongate (as in
<figureCitation id="134D2A41FF27ABBE9CF7144545583C59" box="[680,773,1120,1146]" captionStart="FIGURE 7" captionStartId="313.[151,255,1982,2006]" captionTargetBox="[177,1408,249,1939]" captionTargetId="figure-23@313.[177,1411,232,1956]" captionTargetPageId="313" captionText="FIGURE 7. Head capsules of female Corethrella, anterior view." pageId="203" pageNumber="204">Fig. 7G</figureCitation>
). Number of large setae on frons between ventromedial area of ommatida uncertain. Antenna: pedicel pedicel with at least one distinctive, more elongate, stout, dorsal or dorsolateral seta; flagellomeres as in
<figureCitation id="134D2A41FF27ABBE9CD1149544A93CE9" box="[654,756,1200,1226]" captionStart="FIGURE 25" captionStartId="331.[151,255,1947,1971]" captionTargetBox="[187,1396,238,1921]" captionTargetId="figure-23@331.[177,1411,197,1921]" captionTargetPageId="331" captionText="FIGURE 25. Antennae of Corethrella. A-F. Male. G-L. Female." pageId="203" pageNumber="204">Fig. 25F</figureCitation>
; sensilla coeloconica on flagellomeres 1, 913, flagellomere 1 with two sensilla coeloconica, others with one each; flagellomere 13 with well-developed apical bifurcation. Palpus with segment 3 somewhat swollen at or near midlength.
<emphasis id="B902EAD6FF27ABBE9A64152542CA3D39" bold="true" box="[1083,1175,1280,1306]" pageId="203" pageNumber="204">Thorax</emphasis>
: Posterior portion of dorsocentral row somewhat uncertain but probably with 3 elongate setae situated somewhat lateral to one another. Prescutal suture uncertain. Shapes of pleurites uncertain.
<emphasis id="B902EAD6FF27ABBE9D95157542533D49" bold="true" box="[970,1038,1360,1386]" pageId="203" pageNumber="204">Wing</emphasis>
: Appressed against abdomen but generally similar to those of other
<taxonomicName id="4C764D47FF27ABBE9C73155D44EE3DB2" box="[556,691,1400,1425]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="203" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="B902EAD6FF27ABBE9C73155D44EE3DB2" box="[556,691,1400,1425]" italics="true" pageId="203" pageNumber="204">Corethrella</emphasis>
</taxonomicName>
; veins (other than costa and wing margin) with well-developed scales. Other details uncertain.
<emphasis id="B902EAD6FF27ABBE9C74158544383D99" bold="true" box="[555,613,1440,1466]" pageId="203" pageNumber="204">Legs</emphasis>
: With some thick setae on midfemur, slender on other legs (mostly denuded) (also scales in patch of whip-like setae on posterior portion of hind tibia) Anterior claws of foreleg without a basal prong. Midleg with thick, subapical setae on each of at least tarsomeres 13. Apex of foreleg fifth tarsomeres undivided, with claws slightly subapical to apical. Apex of midleg fifth tarsomere absent. Foreleg claws unequal. Mid-, hind leg claws absent. Foreleg third tarsomere longer than fourth tarsomere. Empodia not visible.
<emphasis id="B902EAD6FF27ABBE9FF8164D44463EA1" bold="true" box="[423,539,1640,1666]" pageId="203" pageNumber="204">Genitalia</emphasis>
(
<figureCitation id="134D2A41FF27ABBE9C70164D44D13EA1" box="[559,652,1640,1666]" captionStart="FIGURE 2" captionStartId="308.[151,255,1947,1971]" captionTargetBox="[177,1410,197,1921]" captionTargetId="figure-63@308.[177,1411,197,1921]" captionTargetPageId="308" captionText="FIGURE 2. A. Unidentified females of Corethrella hovering and landing on calling male of Physalaemus pustulosus in Panama. B. Female adult C. wirthi resting on bark of tree in Georgia, USA. (A, taken by Greg and Mary Beth Dimijian; B, taken by Sturgis McKeever)." pageId="203" pageNumber="204">Fig. 2C</figureCitation>
in
<bibRefCitation id="EFE74B35FF27ABBE9CE7164D42693EA1" author="Borkent, A. &amp; Szadziewski, R." box="[696,1076,1640,1666]" pageId="203" pageNumber="204" pagination="457 - 463" refId="ref236849" refString="Borkent, A. &amp; Szadziewski, R. (1992) The first records of fossil Corethrellidae (Diptera). Entomologica Scandinavica, 22, 457 - 463." type="journal article" year="1992">Borkent and Szadziewski 1992</bibRefCitation>
): Gonocoxite gently tapering; anteromedial area with spicules similar in length to those elsewhere on gonocoxite; with well-defined dorsal row of setae, with setae 2, 3, 4 thicker than others, with row restricted to dorsal portion of gonocoxite. With one dorsomedial stout seta, more or less of even thickness for most of length, tapering near apex. Gonostylus (Fig. 99B) (in retracted position) evenly curved, slender, slender at midlength, tapered apically; one elongate, slender subbasal seta, situated anteriorly; apical seta slender, elongate, simple. Aedeagus mostly not visible, pointed apically.
</paragraph>
<paragraph id="8BC936C4FF27ABBE9E9917A544F53FB9" blockId="203.[151,1436,997,2023]" box="[198,680,1920,1946]" pageId="203" pageNumber="204">
<emphasis id="B902EAD6FF27ABBE9E9917A5444A3FBA" box="[198,535,1920,1945]" italics="true" pageId="203" pageNumber="204">Female adult and immatures</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF27ABB99E99178D45E13A51" lastPageId="204" lastPageNumber="205" pageId="203" pageNumber="204" type="distribution">
<paragraph id="8BC936C4FF27ABBE9E99178D43C73FC4" blockId="203.[151,1436,997,2023]" pageId="203" pageNumber="204">
DISTRIBUTION AND BIONOMICS:
<taxonomicName id="4C764D47FF27ABBE9CC3178D45CB3FE2" box="[668,918,1960,1985]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="203" pageNumber="204" phylum="Arthropoda" rank="species" species="nudistyla">
<emphasis id="B902EAD6FF27ABBE9CC3178D45CB3FE2" box="[668,918,1960,1985]" italics="true" pageId="203" pageNumber="204">Corethrella nudistyla</emphasis>
</taxonomicName>
is known only from Dominican amber (Fig. 139), dated at 19 million years old. Part of a female
<taxonomicName id="4C764D47FF27ABBE9D5E17E845C83FC4" box="[769,917,1997,2023]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="203" pageNumber="204" phylum="Arthropoda" rank="family">Psychodidae</taxonomicName>
was also in the amber bearing the
<typeStatus id="54CD8866FF27ABBE9B7317E843CB3FC4" box="[1324,1430,1997,2023]" pageId="203" pageNumber="204" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BC936C4FF20ABB99E9910BD45263901" blockId="204.[151,1436,152,812]" pageId="204" pageNumber="205">
TAXONOMIC DISCUSSION: The state of the specimen was described by
<bibRefCitation id="EFE74B35FF20ABB99A3510BD46B538F4" author="Borkent, A. &amp; Szadziewski, R." pageId="204" pageNumber="205" pagination="457 - 463" refId="ref236849" refString="Borkent, A. &amp; Szadziewski, R. (1992) The first records of fossil Corethrellidae (Diptera). Entomologica Scandinavica, 22, 457 - 463." type="journal article" year="1992">Borkent and Szadziewski (1992)</bibRefCitation>
. The specimen was further ground to enhance visibility. Contrary to
<bibRefCitation id="EFE74B35FF20ABB99A49109843C538F4" author="Borkent, A. &amp; Szadziewski, R." box="[1046,1432,189,215]" pageId="204" pageNumber="205" pagination="457 - 463" refId="ref236849" refString="Borkent, A. &amp; Szadziewski, R. (1992) The first records of fossil Corethrellidae (Diptera). Entomologica Scandinavica, 22, 457 - 463." type="journal article" year="1992">Borkent and Szadziewski (1992)</bibRefCitation>
, the male does have a subbasal seta on the gonostylus (hence the species is poorly named). In addition, an elongate, slender seta was visible at the apex of the gonostylus.
</paragraph>
<paragraph id="8BC936C4FF20ABB99E99110845E13A51" blockId="204.[151,1436,152,812]" pageId="204" pageNumber="205">
There are
<specimenCount id="9D70FD4DFF20ABB99F611108478D3964" box="[318,464,301,327]" pageId="204" pageNumber="205" type="female">three female</specimenCount>
<taxonomicName id="4C764D47FF20ABB99F87110844023965" box="[472,607,301,326]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="B902EAD6FF20ABB99F87110844023965" box="[472,607,301,326]" italics="true" pageId="204" pageNumber="205">Corethrella</emphasis>
</taxonomicName>
in the AMNH, any of which may or may not be the female of
<taxonomicName id="4C764D47FF20ABB99B1B1108468A3948" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="nudistyla">
<emphasis id="B902EAD6FF20ABB99B1B1108468A3948" italics="true" pageId="204" pageNumber="205">C. nudistyla</emphasis>
</taxonomicName>
(DR-8-103, DR-14-720, DR-14-999). Each was in excellent condition, mostly opaque and difficult, therefore, to see some features of taxonomic importance (DR-8-103 is shown in Fig. 113D). The shape of their flagellomeres and palpal segments were similar. I did not consider them worth describing and naming because they provide so little information and could not be distinguished from numbers of extant species (especially considering that their original pigmentation is unknown). A fourth specimen (DR-13, AMNH), had a somewhat longer first antennal flagellomere and may represent another species but, again, such small differences provide little taxonomic information. All of these
<specimenCount id="9D70FD4DFF20ABB99DCF1217427A3A6F" box="[912,1063,562,588]" pageId="204" pageNumber="205" type="female">four females</specimenCount>
provide no more phylogenetic information than is available from the male
<typeStatus id="54CD8866FF20ABB99CC5127D455D3A51" box="[666,768,600,626]" pageId="204" pageNumber="205" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C764D47FF20ABB99D78127D45E83A52" box="[807,949,600,625]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="nudistyla">
<emphasis id="B902EAD6FF20ABB99D78127D45E83A52" box="[807,949,600,625]" italics="true" pageId="204" pageNumber="205">C. nudistyla</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36C654FFF20ABB99E99125845023B0F" pageId="204" pageNumber="205" type="materials_examined">
<paragraph id="8BC936C4FF20ABB99E99125844A23AC1" blockId="204.[151,1436,152,812]" pageId="204" pageNumber="205">
<materialsCitation id="3B1E3C99FF20ABB99E99125844A73AC1" collectingDate="1989-06" collectionCode="AMNH" country="Canada" location="Male" municipality="Balsam" pageId="204" pageNumber="205" specimenCount="1" typeStatus="holotype">
MATERIAL EXAMINED:
<location id="8EA9601FFF20ABB99C7C125844023AB4" LSID="urn:lsid:plazi:treatment:03DF87D2FF27ABB99EC8134245023B0F:8EA9601FFF20ABB99C7C125844023AB4" box="[547,607,637,663]" country="Canada" municipality="Balsam" name="Male" pageId="204" pageNumber="205">Male</location>
adult in amber mounted in
<collectingCountry id="F3617654FF20ABB99D9F125842453AB4" box="[960,1048,637,663]" name="Canada" pageId="204" pageNumber="205">Canada</collectingCountry>
<collectingMunicipality id="6BADACBEFF20ABB99A7B125842213AB4" box="[1060,1148,637,663]" pageId="204" pageNumber="205">Balsam</collectingMunicipality>
on a microscope slide, labeled &quot;
<typeStatus id="54CD8866FF20ABB99EA1128747C03A9F" box="[254,413,674,700]" pageId="204" pageNumber="205" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C764D47FF20ABB99FF9128745BE3A9F" authority="Borkent and Szadziewski" authorityName="Borkent and Szadziewski" box="[422,995,674,700]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="nudistyla">
<emphasis id="B902EAD6FF20ABB99FF9128744FF3A98" box="[422,674,674,699]" italics="true" pageId="204" pageNumber="205">Corethrella nudistyla</emphasis>
Borkent and Szadziewski
</taxonomicName>
&quot;, &quot;Dominican amber, Jorge Wunderlich,
<date id="FFC81004FF20ABB99E8E12ED47163AC1" box="[209,331,712,738]" pageId="204" pageNumber="205" value="1989-06">
<collectingDate id="EF8CE9ECFF20ABB99E8E12ED47163AC1" box="[209,331,712,738]" pageId="204" pageNumber="205" value="1989-06">June 1989</collectingDate>
</date>
, AMNH No. W-DR-46-b&quot; (
<collectionCode id="ED67AE01FF20ABB99CCD12ED44AF3AC1" box="[658,754,712,738]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="204" pageNumber="205">AMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC936C4FF20ABB99E9912C845023B0F" blockId="204.[151,1436,152,812]" pageId="204" pageNumber="205">
DERIVATION OF SPECIFIC EPITHET: The name
<taxonomicName id="4C764D47FF20ABB99D6612C845F93B25" box="[825,932,749,774]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="204" pageNumber="205" phylum="Arthropoda" rank="species" species="nudistyla">
<emphasis id="B902EAD6FF20ABB99D6612C845F93B25" box="[825,932,749,774]" italics="true" pageId="204" pageNumber="205">nudistyla</emphasis>
</taxonomicName>
(bare, stylus) refers to the bare gonostylus originally (but mistakenly) thought to present in this species.
</paragraph>
</subSubSection>
</treatment>
</document>